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<document id="B20AD6200CA2DDE3EFE5E96D731B22A6" ID-CLB-Dataset="7631" ID-DOI="10.1206/0003-0090-423.1.1" ID-GBIF-Dataset="2e886aea-b59a-45a6-aeaa-2427d584a894" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615995240537" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2018" docId="03B187A8FFAEFFD4FCA93BDD7D3CFC99" docLanguage="en" docName="BulAmeMusNatHis.2018.423.1-97.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2018 (423)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-423.1.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Burmapeza radicis Grimaldi 2018, new species" docType="treatment" docVersion="7" lastPageNumber="97" masterDocId="FF88FFD0FF86FFFEFFAA39537F49FFF7" masterDocTitle="Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi" masterLastPageNumber="97" masterPageNumber="1" pageNumber="1" updateTime="1698933720305" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4AE9A3355E3B0CEDA2152313F0D37CA4">Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi</mods:title>
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<mods:namePart id="4901B0A55E9559320055F9A1C45E58AB">Grimaldi, David A.</mods:namePart>
<mods:affiliation id="809A4CFF07917BB7893D3C7AF74F9653">Division of Invertebrate Zoology American Museum of Natural History, New York</mods:affiliation>
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<mods:title id="62530804DB227D79E2F743F3C65BD6A6">Bulletin of the American Museum of Natural History</mods:title>
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<mods:date id="B681A63FAF5C3C909CB21AC9DC804A0A">2018</mods:date>
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<mods:number id="498C78C85DCA4914E7AEF4EE01B1B9E5">2018-10-24</mods:number>
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<treatment id="03B187A8FFAEFFD4FCA93BDD7D3CFC99" ID-DOI="http://doi.org/10.5281/zenodo.4631083" ID-GBIF-Taxon="180670805" ID-Zenodo-Dep="4631083" LSID="urn:lsid:plazi:treatment:03B187A8FFAEFFD4FCA93BDD7D3CFC99" httpUri="http://treatment.plazi.org/id/03B187A8FFAEFFD4FCA93BDD7D3CFC99" lastPageId="42" pageId="40">
<subSubSection id="C3026535FFAEFFD6FCA93BDD7B2CFD52" box="[771,1125,653,677]" pageId="40" type="nomenclature">
<paragraph id="8BA736BEFFAEFFD6FCA93BDD7B2CFD52" blockId="40.[771,1125,653,677]" box="[771,1125,653,677]" pageId="40">
<heading id="D0EF81D2FFAEFFD6FCA93BDD7B2CFD52" box="[771,1125,653,677]" centered="true" fontSize="9" level="2" pageId="40" reason="2">
<taxonomicName id="4C184D3DFFAEFFD6FCA93BDD7C9EFD52" ID-CoL="8B9Q6" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[771,983,653,677]" class="Insecta" family="Platypezidae" genus="Burmapeza" kingdom="Animalia" order="Diptera" pageId="40" phylum="Arthropoda" rank="species" species="radicis" status="sp. nov.">
<emphasis id="B96CEAACFFAEFFD6FCA93BDD7C9EFD52" bold="true" box="[771,983,653,677]" italics="true" pageId="40">Burmapeza radicis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFAEFFD6FC4E3BDD7B2CFD52" box="[996,1125,653,677]" pageId="40" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFAEFFD6FCC43BEC7CB3FD20" box="[878,1018,703,727]" pageId="40" type="description">
<paragraph id="8BA736BEFFAEFFD6FCC43BEC7CB3FD20" blockId="40.[878,1018,703,727]" box="[878,1018,703,727]" pageId="40">
<heading id="D0EF81D2FFAEFFD6FCC43BEC7CB3FD20" box="[878,1018,703,727]" centered="true" fontSize="9" level="2" pageId="40" reason="2">
<figureCitation id="13232A3BFFAEFFD6FCC43BEC7C9AFD20" box="[878,979,703,727]" captionStart="FIG" captionStartId="10.[109,151,1707,1728]" captionTargetBox="[121,1200,225,1680]" captionTargetId="figure-9@10.[108,1212,225,1680]" captionTargetPageId="10" captionText="FIG. 3. Burmapeza radicis, n. gen., n. sp. (Platypezidae), holotype female, JZC Bu178a, in Burmese amber. A. Dorsal habitus. B. Frontal view of head." figureDoi="http://doi.org/10.5281/zenodo.4613016" httpUri="https://zenodo.org/record/4613016/files/figure.png" pageId="40">Figures 3</figureCitation>
,
<figureCitation id="13232A3BFFAEFFD6FC4A3BEC7CB3FD20" box="[992,1018,703,727]" captionStart="FIG" captionStartId="41.[107,146,1378,1399]" captionTargetBox="[127,1183,237,1346]" captionTargetId="figure-145@41.[108,1212,224,1350]" captionTargetPageId="41" captionText="FIG.28. Burmapeza radicis, n. gen., n. sp.(Platypezidae), holotype female, JZC Bu178a, in Burmese amber.A. Wing. B. Thorax, dorsal, as preserved (slightly distorted by preservation). C. Female terminalia. D. Antenna, lateral." figureDoi="http://doi.org/10.5281/zenodo.4613080" httpUri="https://zenodo.org/record/4613080/files/figure.png" pageId="40">28</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFAEFFD6FD663BA27B37FCFE" box="[716,1150,752,777]" pageId="40" type="diagnosis">
<paragraph id="8BA736BEFFAEFFD6FD663BA27B37FCFE" blockId="40.[684,1213,752,1735]" box="[716,1150,752,777]" pageId="40">DIAGNOSIS: As for genus, by monotypy.</paragraph>
</subSubSection>
<subSubSection id="C3026535FFAEFFD4FD663A417E1AFCFC" lastPageId="42" pageId="40" type="description">
<paragraph id="8BA736BEFFAEFFD6FD663A417CEEFACC" blockId="40.[684,1213,752,1735]" pageId="40">
DESCRIPTION: (female only).
<emphasis id="B96CEAACFFAEFFD6FB8E3A417B21FCDD" bold="true" box="[1060,1128,786,810]" pageId="40">Head:</emphasis>
Somewhat flattened anteroposteriad, with broad frontal surface. Eyes bare, occupying all of lateral surface, well separated, no differentiation of facets. Median furrow of frons, interfrontal and fronto-orbital setae not observable (due to preservation); three pairs of small, stout ocellar setae present; row short, stout, postocular setae present near posterior margin of eye. Vibrissa, genal setae absent.
<emphasis id="B96CEAACFFAEFFD6FCEA3D687CE3FBA4" bold="true" box="[832,938,1083,1107]" pageId="40">Antenna:</emphasis>
Scape not visible, apical rim of pedicel with fine setae; arista dorsal; basal flagellomere short, laterall flattened, subcircular in lateral view; three aristomeres present, basal two minute, apical aristomere length 1.5× that of basal flagellomere, with minute micropubescence.
<emphasis id="B96CEAACFFAEFFD6FD5F3C517CCDFAED" bold="true" box="[757,900,1282,1306]" pageId="40">Mouthparts:</emphasis>
Palp well developed, clavate; labellum barely visible.
</paragraph>
<paragraph id="8BA736BEFFAEFFD4FD673C177D3EFECB" blockId="40.[684,1213,752,1735]" lastBlockId="42.[108,637,226,878]" lastPageId="42" pageId="40">
<emphasis id="B96CEAACFFAEFFD6FD673C177C6CFAAB" bold="true" box="[717,805,1348,1372]" pageId="40">Thorax:</emphasis>
Scutum relatively flat dorsally. Acrostichals numerous, very fine, not in rows; 3 pairs prescutellar setae (lateralmost pair the dorsocentrals), graded row of dorsocentrals absent; 1 supraalar, 34 notopleurals present. Prescutellum present; scutellum relatively large, setulose on dorsal surface, posterior rim with 4 setae and smaller setulae.
<emphasis id="B96CEAACFFAEFFD6FCCF3F787CE2F9B4" bold="true" box="[869,939,1579,1603]" pageId="40">Wing:</emphasis>
Relatively broad, short, width 0.5× the length; membrane entirely covered with dense microtrichia, even in cells. Two prehumeral costal setae present; vein C with 2 long rows of thick setulae that grade to fine setu- lae apicad; C terminates at apex of M
<subScript id="179C34FBFFAFFFD7FDC73C867D3FFA11" attach="right" box="[621,630,1493,1510]" fontSize="7" pageId="41">1</subScript>
. Pterostigma absent; slight tint between Sc and R
<subScript id="179C34FBFFAFFFD7FFD73F447FCFF9DF" attach="left" box="[125,134,1559,1576]" fontSize="7" pageId="41">1</subScript>
. Sc long, terminates near middle of wing; R
<subScript id="179C34FBFFAFFFD7FDD93F447D35F9DF" attach="left" box="[627,636,1559,1576]" fontSize="7" pageId="41">1</subScript>
long, runs very close to Sc at base. R
<subScript id="179C34FBFFAFFFD7FDBB3F6B7D65F9BE" attach="left" box="[529,556,1592,1609]" fontSize="7" pageId="41">4+5</subScript>
terminates at tip of wing, slightly deflexed in middle posteriad. Cells rm and bm complete; crossvein dm-cu 2.5× its length from wing margin; M
<subScript id="179C34FBFFAFFFD7FDE33FCF7D1BF95A" attach="left" box="[585,594,1692,1709]" fontSize="7" pageId="41">1</subScript>
-M
<subScript id="179C34FBFFAFFFD7FDD93FCF7D35F95A" attach="left" box="[627,636,1692,1709]" fontSize="7" pageId="41">2</subScript>
fork asymmetrical (M
<subScript id="179C34FBFFAFFFD7FEF33FEE7E2BF939" attach="left" box="[345,354,1725,1742]" fontSize="7" pageId="41">1</subScript>
branch longer, complete), apex of M
<subScript id="179C34FBFFAFFFD7FC883C867C62FA11" attach="left" box="[802,811,1493,1510]" fontSize="7" pageId="41">2</subScript>
abruptly incomplete; tips of CuA
<subScript id="179C34FBFFAFFFD7FB073C867BFFFA11" attach="left" box="[1197,1206,1493,1510]" fontSize="7" pageId="41">1</subScript>
, CuA
<subScript id="179C34FBFFAFFFD7FD753CA57DA1F9F0" attach="left" box="[735,744,1526,1543]" fontSize="7" pageId="41">2</subScript>
+A
<subScript id="179C34FBFFAFFFD7FCA03CA57C5AF9F0" attach="left" box="[778,787,1526,1543]" fontSize="7" pageId="41">1</subScript>
incomplete (gradually faded). Cell cup large, CuA
<subScript id="179C34FBFFAFFFD7FC823F447C78F9DF" attach="left" box="[808,817,1559,1576]" fontSize="7" pageId="41">2</subScript>
curved. Anal lobe and alula well developed.
<emphasis id="B96CEAACFFAFFFD7FC8D3F7E7C2BF9B2" bold="true" box="[807,866,1581,1605]" pageId="41">Legs:</emphasis>
Relatively unmodified, generalized. Femora stoutest and longest podomeres, especially metafemur; metatibia slightly curved femorad, with apical circlet of 5 stout, short setae. Legs without dark, bifid scales on tibiae or tarsi; with fine setulae only. Hind tarsomeres not expanded/flattened; tarsomere lengths 1&gt; 2&gt; 3&gt; 4 = 5. Pulvilli and empodium not observable.
</paragraph>
<caption id="DF676636FFAFFFD7FFC13C317BD9FA61" ID-DOI="http://doi.org/10.5281/zenodo.4613080" ID-Zenodo-Dep="4613080" httpUri="https://zenodo.org/record/4613080/files/figure.png" pageId="41" startId="41.[107,146,1378,1399]" targetBox="[127,1183,237,1346]" targetPageId="41">
<paragraph id="8BA736BEFFAFFFD7FFC13C317BD9FA61" blockId="41.[106,1214,1376,1430]" pageId="41">
FIG. 28.
<taxonomicName id="4C184D3DFFAFFFD7FF113C327E22FA8F" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[187,363,1376,1400]" class="Insecta" family="Platypezidae" genus="Burmapeza" kingdom="Animalia" order="Diptera" pageId="41" phylum="Arthropoda" rank="species" species="radicis" status="n. gen., n. sp.">
<emphasis id="B96CEAACFFAFFFD7FF113C327E22FA8F" box="[187,363,1376,1400]" italics="true" pageId="41">Burmapeza radicis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFAFFFD7FEDF3C327EBBFA8F" box="[373,498,1377,1401]" pageId="41" rank="species">n. gen., n. sp.</taxonomicNameLabel>
(
<taxonomicName id="4C184D3DFFAFFFD7FE563C337D32FA8F" box="[508,635,1376,1400]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="41" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
), holotype female, JZC Bu178a, in Burmese amber.
<emphasis id="B96CEAACFFAFFFD7FBCE3C337B3FFA8F" bold="true" box="[1124,1142,1376,1400]" pageId="41">A</emphasis>
. Wing.
<emphasis id="B96CEAACFFAFFFD7FFC03C2D7F33FA61" bold="true" box="[106,122,1406,1430]" pageId="41">B</emphasis>
. Thorax, dorsal, as preserved (slightly distorted by preservation).
<emphasis id="B96CEAACFFAFFFD7FD523C2E7C41FA62" bold="true" box="[760,776,1405,1429]" pageId="41">C</emphasis>
. Female terminalia.
<emphasis id="B96CEAACFFAFFFD7FC7A3C2D7CA1FA61" bold="true" box="[976,1000,1406,1430]" pageId="41">D.</emphasis>
Antenna, lateral.
</paragraph>
</caption>
<paragraph id="8BA736BEFFACFFD4FF2638167FB3FDF5" blockId="42.[108,637,226,878]" pageId="42">
<emphasis id="B96CEAACFFACFFD4FF2638167E4CFEAA" bold="true" box="[140,261,325,349]" pageId="42">Abdomen:</emphasis>
Relatively long, slightly flattened dorsoventrally; tergites well developed (8 visible), with fine setae; tergite 7 with 2 pairs of slightly longer setae; sternites well developed but not very visible. Cercus slender, elongate, ventroflexed.
</paragraph>
<paragraph id="8BA736BEFFACFFD4FF263B587E1AFCFC" blockId="42.[108,637,226,878]" pageId="42">
SPECIMENS EXAMINED:
<typeStatus id="54A3881CFFACFFD4FE263B587EBAFDD4" box="[396,499,523,547]" pageId="42" type="holotype">Holotype</typeStatus>
, female, JZC Bu178a, the better preserved of two specimens in the same piece of amber. The other specimen (178b) is partial (sex uncertain), and is designated as a
<typeStatus id="54A3881CFFACFFD4FF403BC37E04FD5F" box="[234,333,656,680]" pageId="42" type="paratype">paratype</typeStatus>
. The species description is taken mostly from the
<typeStatus id="54A3881CFFACFFD4FEC43BE37E99FD3F" box="[366,464,688,712]" pageId="42" type="holotype">holotype</typeStatus>
, but features of the head were derived from the
<typeStatus id="54A3881CFFACFFD4FE693B817D68FD1D" box="[451,545,722,746]" pageId="42" type="paratype">paratype</typeStatus>
because it is better preserved.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFACFFD4FF263A477D3CFC99" pageId="42" type="etymology">
<paragraph id="8BA736BEFFACFFD4FF263A477D3CFC99" blockId="42.[108,637,226,878]" pageId="42">
ETYMOLOGY: From Latin (“of the root”), in reference to the basal phylogenetic position of the genus and
<typeStatus id="54A3881CFFACFFD4FEA63A057E72FC99" box="[268,315,854,878]" pageId="42">type</typeStatus>
species within
<taxonomicName id="4C184D3DFFACFFD4FE4F3A057D38FC99" box="[485,625,854,878]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="42" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>