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<document id="D4D366D8E46FAFBFF03247A0BAAA8690" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFC99D2A4BA500E5DA3F21C4" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Benedictesmus aureus Shear &amp; Marek 2021, n. sp." docType="treatment" docVersion="6" lastPageNumber="116" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="113" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F0DDBF437DF6C2B61B35679BC8FE4B20" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="42E971E93ABA30548A74A65487A6E319">
<mods:title id="C051DA70BAB8AACB9F8022DC44CD08B5">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:name id="BF56340A5522B33C59F114FB4B715EB4" type="personal">
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<mods:roleTerm id="C8F26F3931007C3FE2DBA0EC968229CB">Author</mods:roleTerm>
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<mods:namePart id="5DC1D8EB706DB1A0743EBF28A63A0DC2">Shear, William A.</mods:namePart>
<mods:affiliation id="14FD2C78EB7785D8FE5F471EE8B5E7B8">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
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<mods:namePart id="CBDD3E137311B02A568D1176C8C842FE">Marek, Paul E.</mods:namePart>
<mods:nameIdentifier id="FC0BA37B4A4C240E22FD0CB8F9B7A7FF" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation id="B3208F2DA6B7EA7E3B9E3C48CEB5DF0A">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
<mods:nameIdentifier id="65455909BE95947FE2EF453C1AD2331D" type="email">pmarek@vt.edu</mods:nameIdentifier>
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<mods:typeOfResource id="E6C3430F7F53C988CA1BB9901D2DADAD">text</mods:typeOfResource>
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<mods:title id="6AD3488DA16141655DEECEF792D3A12D">Zootaxa</mods:title>
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<mods:part id="D6B885A4D0FA169E5F6354F6F7C93664">
<mods:date id="885621B982ABD1B03B1784CF6FE41760">2021</mods:date>
<mods:detail id="CBD092003532714AEC5600B02B936D8A" type="pubDate">
<mods:number id="70970A340B35B3F86595CB0E39339B5A">2021-05-24</mods:number>
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<mods:detail id="698E4CD490B7D8F8C8A52017552733E9" type="volume">
<mods:number id="5C3086AA0CF75450FFE73137E90D31AD">4975</mods:number>
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<mods:identifier id="F157008037976895157D31F85FCF606F" type="CLB-Dataset">6194</mods:identifier>
<mods:identifier id="9E0F2F4CE381E7819DA8654E1D0A3D71" type="DOI">10.11646/zootaxa.4975.1.3</mods:identifier>
<mods:identifier id="957730D1C0B67EE4D99772D9DE9FD836" type="GBIF-Dataset">529d48db-4299-4e2d-9aaf-6c4178c08b18</mods:identifier>
<mods:identifier id="C91D19F66E45FD6A5ACC05EB7685C436" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B18797FFC99D2A4BA500E5DA3F21C4" ID-DOI="http://doi.org/10.5281/zenodo.4805057" ID-GBIF-Taxon="182644919" ID-Zenodo-Dep="4805057" LSID="urn:lsid:plazi:treatment:03B18797FFC99D2A4BA500E5DA3F21C4" httpUri="http://treatment.plazi.org/id/03B18797FFC99D2A4BA500E5DA3F21C4" lastPageId="35" lastPageNumber="116" pageId="32" pageNumber="113">
<subSubSection id="C302650AFFC99D294BA500E5DB8025B4" box="[151,707,1503,1530]" pageId="32" pageNumber="113" type="nomenclature">
<paragraph id="8BA73681FFC99D294BA500E5DB8025B4" blockId="32.[151,707,1503,1565]" box="[151,707,1503,1530]" pageId="32" pageNumber="113">
<heading id="D0EF81EDFFC99D294BA500E5DB8025B4" bold="true" box="[151,707,1503,1530]" fontSize="11" level="1" pageId="32" pageNumber="113" reason="1">
<emphasis id="B96CEA93FFC99D294BA500E5DB8025B4" bold="true" box="[151,707,1503,1530]" pageId="32" pageNumber="113">
<taxonomicName id="4C184D02FFC99D294BA500E5DB3725B4" authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2021" box="[151,628,1503,1530]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="aureus" status="sp. nov.">
<emphasis id="B96CEA93FFC99D294BA500E5D8E525B4" bold="true" box="[151,422,1503,1530]" italics="true" pageId="32" pageNumber="113">Benedictesmus aureus</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFC99D2949B300E5DB8025B4" box="[641,707,1503,1529]" pageId="32" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFC99D294BA50339D99626E0" pageId="32" pageNumber="113" type="description">
<paragraph id="8BA73681FFC99D294BA50339D8192650" blockId="32.[151,707,1503,1565]" box="[151,346,1539,1565]" pageId="32" pageNumber="113">
<figureCitation id="13232A04FFC99D294BA50339D8662650" box="[151,293,1539,1565]" captionStart-0="FIGURES 96–100" captionStart-1="FIGURES 101–106" captionStartId-0="34.[151,264,1943,1967]" captionStartId-1="36.[151,264,1852,1876]" captionTargetBox-0="[180,1408,181,1918]" captionTargetBox-1="[151,1436,181,1828]" captionTargetId-0="figure-24@34.[179,1408,181,1919]" captionTargetId-1="figure-24@36.[151,1436,181,1828]" captionTargetPageId-0="34" captionTargetPageId-1="36" captionText-0="FIGURES 96–100. Benedictesmus aureus, n. gen., n. sp. 96. Male, dorsal view. 97. Anterior part of male, ventral view. 98. Head and anterior rings, dorsal view. 99. Midbody ring, dorsal view. 100. Pygidium, lateral view. SEM stub WS33-6." captionText-1="FIGURES 101–106. Species of Benedictesmus, n. gen. 101, Gonopods of Benedictesmus aureus, n. sp., ventral view. SEM stub WS33-6. 102–106. Benedictesmus ellenae, n. sp. 102. Head and anterior rings, dorsal view. 103. Midbody ring, dorsal view. 104. Pygidium, lateral view. 105. Gonopods in situ, lateroventral view. 106. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: p, pulvillus; pp, pulvillar process; ppf, prefemorite pore field; tz, terminal zone." figureDoi-0="http://doi.org/10.5281/zenodo.4805039" figureDoi-1="http://doi.org/10.5281/zenodo.4805047" httpUri-0="https://zenodo.org/record/4805039/files/figure.png" httpUri-1="https://zenodo.org/record/4805047/files/figure.png" pageId="32" pageNumber="113">Figs 96–101</figureCitation>
,
<figureCitation id="13232A04FFC99D294A020339D8192650" box="[304,346,1539,1565]" captionStart="FIGURES 133–137" captionStartId="42.[151,264,1252,1276]" captionTargetBox="[160,1429,181,1218]" captionTargetId="figure-24@42.[151,1436,181,1228]" captionTargetPageId="42" captionText="FIGURES 133–137. Drawings of gonopods of polydesmid species. 133. Right gonopod of Benedictesmus aureus, n. sp., mesal view. 134. Right gonopod of Benedictesmus ellenae,n. sp., lateral view. 135, Right gonopod of Benedictesmus yaquina, n. sp., lateral view. Fig. 136, 137, Benedictesmus timber.n. sp. 136, Right gonopod, lateral view. 137, Tip of right gonopod, mesal view. Abbreviation: p, pulvillus." figureDoi="http://doi.org/10.5281/zenodo.4805079" httpUri="https://zenodo.org/record/4805079/files/figure.png" pageId="32" pageNumber="113">133</figureCitation>
</paragraph>
<paragraph id="8BA73681FFC99D294BA50371D99626E0" blockId="32.[151,1437,1611,2034]" pageId="32" pageNumber="113">
<emphasis id="B96CEA93FFC99D294BA50371D9A72628" bold="true" box="[151,228,1611,1637]" pageId="32" pageNumber="113">Types.</emphasis>
<materialsCitation id="3B703CDCFFC99D294BDC0371D99226E0" ID-GBIF-Occurrence="3128519318" collectingDate="1972-03-10" collectionCode="CAS" collectorName="E. M. Benedict. All" county="Curry Co." location="Agness" municipality="Male" pageId="32" pageNumber="113" specimenCount="2" stateProvince="Oregon" typeStatus="holotype">
<collectingMunicipality id="6BC3ACFBFFC99D294BDC0371D86B2628" box="[238,296,1611,1637]" pageId="32" pageNumber="113">Male</collectingMunicipality>
<typeStatus id="54A38823FFC99D294A010371D8D62628" box="[307,405,1611,1637]" pageId="32" pageNumber="113" type="holotype">holotype</typeStatus>
and male and female
<typeStatus id="54A38823FFC99D2949AF0376DA4B262B" box="[669,776,1612,1638]" pageId="32" pageNumber="113" type="paratype">paratypes</typeStatus>
from
<locationDeviation id="9A152FA3FFC99D2948670371DDCA2628" box="[853,1161,1611,1638]" location="East of Gold Beach" pageId="32" pageNumber="113">
<quantity id="4CE09B64FFC99D2948670371DADA2628" box="[853,921,1611,1637]" metricMagnitude="4" metricUnit="m" metricValue="2.0921472" pageId="32" pageNumber="113" unit="mi" value="13.0">13 mi</quantity>
East of Gold Beach
</locationDeviation>
on the road to
<location id="8EC7605AFFC99D294E710371DCDB2628" LSID="urn:lsid:plazi:treatment:03B18797FFC99D2A4BA500E5DA3F21C4:8EC7605AFFC99D294E710371DCDB2628" box="[1347,1432,1611,1637]" county="Curry Co." municipality="Male" name="Agness" pageId="32" pageNumber="113" stateProvince="Oregon">Agness</location>
, 42.4932°, -124.2194°,
<collectingCounty id="62C64E0DFFC99D294AAC0355DB5726C4" box="[414,532,1647,1673]" pageId="32" pageNumber="113">Curry Co.</collectingCounty>
,
<collectingRegion id="49DCF863FFC99D2949110355DB3926C4" box="[547,634,1647,1673]" country="United States of America" name="Oregon" pageId="32" pageNumber="113">Oregon</collectingRegion>
, collected
<date id="FFA61041FFC99D2949CA0355DAE826C4" box="[760,939,1647,1673]" pageId="32" pageNumber="113" value="1972-03-10">
<collectingDate id="EFE2E9A9FFC99D2949CA0355DAE826C4" box="[760,939,1647,1673]" pageId="32" pageNumber="113" value="1972-03-10">10 March 1972</collectingDate>
</date>
by
<collectorName id="26ED5357FFC99D2948EE034ADD8226C4" box="[988,1217,1647,1674]" pageId="32" pageNumber="113">E. M. Benedict. All</collectorName>
types deposited in
<collectionCode id="ED09AE44FFC99D294BA503A9D99226E0" box="[151,209,1683,1709]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="113" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFC99D294BF5038DDC192754" pageId="32" pageNumber="113" type="diagnosis">
<paragraph id="8BA73681FFC99D294BF5038DDC192754" blockId="32.[151,1437,1611,2034]" pageId="32" pageNumber="113">
<emphasis id="B96CEA93FFC99D294BF5038DD801269C" bold="true" box="[199,322,1719,1745]" pageId="32" pageNumber="113">Diagnosis.</emphasis>
Distinct from
<emphasis id="B96CEA93FFC99D294AD60382DB18269C" box="[484,603,1719,1745]" italics="true" pageId="32" pageNumber="113">
<taxonomicName id="4C184D02FFC99D294AD60382DB14269C" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[484,599,1719,1745]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="ellenae" status="sp. nov.">B. ellenae</taxonomicName>
,
</emphasis>
<emphasis id="B96CEA93FFC99D294953038DDBE3269C" bold="true" box="[609,672,1719,1745]" pageId="32" pageNumber="113">
<taxonomicNameLabel id="A25F57E8FFC99D294953038DDBE3269C" box="[609,672,1719,1745]" pageId="32" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, which also has 18 rings in males, in the much shorter terminal zone of the gonopod and in having 20, rather than 18 setae in the anterior marginal row on the collum. The other two species of
<taxonomicName id="4C184D02FFC99D294BDA03C5D8CD2754" authorityName="Shear &amp; Marek" authorityYear="2021" box="[232,398,1791,1817]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEA93FFC99D294BDA03C5D8CD2754" box="[232,398,1791,1817]" italics="true" pageId="32" pageNumber="113">Benedictesmus</emphasis>
</taxonomicName>
have 19 rings in males. Females of all known species of
<taxonomicName id="4C184D02FFC99D294F3C03C5DDF72754" authorityName="Shear &amp; Marek" authorityYear="2021" box="[1038,1204,1791,1817]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEA93FFC99D294F3C03C5DDF72754" box="[1038,1204,1791,1817]" italics="true" pageId="32" pageNumber="113">Benedictesmus</emphasis>
</taxonomicName>
have 19 rings.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFC99D294BF50219DD922770" box="[199,1233,1827,1854]" pageId="32" pageNumber="113" type="etymology">
<paragraph id="8BA73681FFC99D294BF50219DD922770" blockId="32.[151,1437,1611,2034]" box="[199,1233,1827,1854]" pageId="32" pageNumber="113">
<emphasis id="B96CEA93FFC99D294BF50219D80D2770" bold="true" box="[199,334,1827,1853]" pageId="32" pageNumber="113">Etymology.</emphasis>
The species name, an adjective, refers to Gold Beach,
<collectingRegion id="49DCF863FFC99D2948870219DD4F2770" box="[949,1036,1827,1853]" country="United States of America" name="Oregon" pageId="32" pageNumber="113">Oregon</collectingRegion>
, the
<typeStatus id="54A38823FFC99D294F72021EDD332773" box="[1088,1136,1828,1854]" pageId="32" pageNumber="113">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFC99D2A4BF5027DDA9420B4" lastPageId="35" lastPageNumber="116" pageId="32" pageNumber="113" type="description">
<paragraph id="8BA73681FFC99D2A4BF5027DDA9420B4" blockId="32.[151,1437,1611,2034]" lastBlockId="35.[151,1436,150,393]" lastPageId="35" lastPageNumber="116" pageId="32" pageNumber="113">
<emphasis id="B96CEA93FFC99D294BF5027DD81A272C" bold="true" box="[199,345,1863,1889]" pageId="32" pageNumber="113">Description.</emphasis>
Male
<typeStatus id="54A38823FFC99D294A97027DDB48272C" box="[421,523,1863,1889]" pageId="32" pageNumber="113" type="holotype">holotype</typeStatus>
. Eighteen rings (
<figureCitation id="13232A04FFC99D2949FF027DDA67272F" box="[717,804,1863,1890]" captionStart="FIGURES 96–100" captionStartId="34.[151,264,1943,1967]" captionTargetBox="[180,1408,181,1918]" captionTargetId="figure-24@34.[179,1408,181,1919]" captionTargetPageId="34" captionText="FIGURES 96–100. Benedictesmus aureus, n. gen., n. sp. 96. Male, dorsal view. 97. Anterior part of male, ventral view. 98. Head and anterior rings, dorsal view. 99. Midbody ring, dorsal view. 100. Pygidium, lateral view. SEM stub WS33-6." figureDoi="http://doi.org/10.5281/zenodo.4805039" httpUri="https://zenodo.org/record/4805039/files/figure.png" pageId="32" pageNumber="113">Fig. 96</figureCitation>
). Length about 3.0 mm, greatest width
<quantity id="4CE09B64FFC99D294FD9027DDC17272F" box="[1259,1364,1863,1890]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="32" pageNumber="113" unit="mm" value="0.41">0.41 mm</quantity>
. Head densely setose, cuticle alveolate (
<figureCitation id="13232A04FFC99D2949360251DB1527C8" box="[516,598,1899,1925]" captionStart="FIGURES 96–100" captionStartId="34.[151,264,1943,1967]" captionTargetBox="[180,1408,181,1918]" captionTargetId="figure-24@34.[179,1408,181,1919]" captionTargetPageId="34" captionText="FIGURES 96–100. Benedictesmus aureus, n. gen., n. sp. 96. Male, dorsal view. 97. Anterior part of male, ventral view. 98. Head and anterior rings, dorsal view. 99. Midbody ring, dorsal view. 100. Pygidium, lateral view. SEM stub WS33-6." figureDoi="http://doi.org/10.5281/zenodo.4805039" httpUri="https://zenodo.org/record/4805039/files/figure.png" pageId="32" pageNumber="113">Fig. 98</figureCitation>
). Collum with anterior marginal row of 20 setae (
<figureCitation id="13232A04FFC99D294F5C0251DD8327C8" box="[1134,1216,1899,1925]" captionStart="FIGURES 96–100" captionStartId="34.[151,264,1943,1967]" captionTargetBox="[180,1408,181,1918]" captionTargetId="figure-24@34.[179,1408,181,1919]" captionTargetPageId="34" captionText="FIGURES 96–100. Benedictesmus aureus, n. gen., n. sp. 96. Male, dorsal view. 97. Anterior part of male, ventral view. 98. Head and anterior rings, dorsal view. 99. Midbody ring, dorsal view. 100. Pygidium, lateral view. SEM stub WS33-6." figureDoi="http://doi.org/10.5281/zenodo.4805039" httpUri="https://zenodo.org/record/4805039/files/figure.png" pageId="32" pageNumber="113">Fig. 98</figureCitation>
).Anterior metazonites with three rows of setae, transitioning to four rows about ring 5, setal tubercles becoming almost obsolete on midbody (
<figureCitation id="13232A04FFC99D294A390289D81F2780" box="[267,348,1971,1997]" captionStart="FIGURES 96–100" captionStartId="34.[151,264,1943,1967]" captionTargetBox="[180,1408,181,1918]" captionTargetId="figure-24@34.[179,1408,181,1919]" captionTargetPageId="34" captionText="FIGURES 96–100. Benedictesmus aureus, n. gen., n. sp. 96. Male, dorsal view. 97. Anterior part of male, ventral view. 98. Head and anterior rings, dorsal view. 99. Midbody ring, dorsal view. 100. Pygidium, lateral view. SEM stub WS33-6." figureDoi="http://doi.org/10.5281/zenodo.4805039" httpUri="https://zenodo.org/record/4805039/files/figure.png" pageId="32" pageNumber="113">Fig. 99</figureCitation>
) to posterior rings. Alveolate cuticle absent from metazonites posterior to collum. Epiproct (
<figureCitation id="13232A04FFC99D294E5D0289D98127BC" captionStart="FIGURES 96–100" captionStartId="34.[151,264,1943,1967]" captionTargetBox="[180,1408,181,1918]" captionTargetId="figure-24@34.[179,1408,181,1919]" captionTargetPageId="34" captionText="FIGURES 96–100. Benedictesmus aureus, n. gen., n. sp. 96. Male, dorsal view. 97. Anterior part of male, ventral view. 98. Head and anterior rings, dorsal view. 99. Midbody ring, dorsal view. 100. Pygidium, lateral view. SEM stub WS33-6." figureDoi="http://doi.org/10.5281/zenodo.4805039" httpUri="https://zenodo.org/record/4805039/files/figure.png" pageId="32" pageNumber="113">Fig. 100</figureCitation>
) slightly swollen, short, slightly decurved, with alveolate cuticle. Anterior legs crassate (
<figureCitation id="13232A04FFC99D294FF602EDDC5927BC" box="[1220,1306,2007,2033]" captionStart="FIGURES 96–100" captionStartId="34.[151,264,1943,1967]" captionTargetBox="[180,1408,181,1918]" captionTargetId="figure-24@34.[179,1408,181,1919]" captionTargetPageId="34" captionText="FIGURES 96–100. Benedictesmus aureus, n. gen., n. sp. 96. Male, dorsal view. 97. Anterior part of male, ventral view. 98. Head and anterior rings, dorsal view. 99. Midbody ring, dorsal view. 100. Pygidium, lateral view. SEM stub WS33-6." figureDoi="http://doi.org/10.5281/zenodo.4805039" httpUri="https://zenodo.org/record/4805039/files/figure.png" pageId="32" pageNumber="113">Fig. 97</figureCitation>
), tarsi with sphaerotrichomes. Gonopod (
<figureCitation id="13232A04FFCB9D2B4AD402E4DB0B27B5" box="[486,584,2014,2040]" captionStart="FIGURES 101–106" captionStartId="36.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1828]" captionTargetId="figure-24@36.[151,1436,181,1828]" captionTargetPageId="36" captionText="FIGURES 101–106. Species of Benedictesmus, n. gen. 101, Gonopods of Benedictesmus aureus, n. sp., ventral view. SEM stub WS33-6. 102–106. Benedictesmus ellenae, n. sp. 102. Head and anterior rings, dorsal view. 103. Midbody ring, dorsal view. 104. Pygidium, lateral view. 105. Gonopods in situ, lateroventral view. 106. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: p, pulvillus; pp, pulvillar process; ppf, prefemorite pore field; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805047" httpUri="https://zenodo.org/record/4805047/files/figure.png" pageId="34" pageNumber="115">Fig. 101</figureCitation>
,
<figureCitation id="13232A04FFCB9D2B496402E4DBC227B5" box="[598,641,2014,2040]" captionStart="FIGURES 133–137" captionStartId="42.[151,264,1252,1276]" captionTargetBox="[160,1429,181,1218]" captionTargetId="figure-24@42.[151,1436,181,1228]" captionTargetPageId="42" captionText="FIGURES 133–137. Drawings of gonopods of polydesmid species. 133. Right gonopod of Benedictesmus aureus, n. sp., mesal view. 134. Right gonopod of Benedictesmus ellenae,n. sp., lateral view. 135, Right gonopod of Benedictesmus yaquina, n. sp., lateral view. Fig. 136, 137, Benedictesmus timber.n. sp. 136, Right gonopod, lateral view. 137, Tip of right gonopod, mesal view. Abbreviation: p, pulvillus." figureDoi="http://doi.org/10.5281/zenodo.4805079" httpUri="https://zenodo.org/record/4805079/files/figure.png" pageId="34" pageNumber="115">133</figureCitation>
) with moderately inflated prefemorite, distal part of prefemorite with pore field, 3 or 4 setae (
<emphasis id="B96CEA93FFCA9D2A4AA905ACD88920FD" bold="true" box="[411,458,150,176]" pageId="35" pageNumber="116">ppf,</emphasis>
<figureCitation id="13232A04FFCA9D2A4AE205ADDB6920FF" box="[464,554,151,178]" captionStart="FIGURES 101–106" captionStartId="36.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1828]" captionTargetId="figure-24@36.[151,1436,181,1828]" captionTargetPageId="36" captionText="FIGURES 101–106. Species of Benedictesmus, n. gen. 101, Gonopods of Benedictesmus aureus, n. sp., ventral view. SEM stub WS33-6. 102–106. Benedictesmus ellenae, n. sp. 102. Head and anterior rings, dorsal view. 103. Midbody ring, dorsal view. 104. Pygidium, lateral view. 105. Gonopods in situ, lateroventral view. 106. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: p, pulvillus; pp, pulvillar process; ppf, prefemorite pore field; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805047" httpUri="https://zenodo.org/record/4805047/files/figure.png" pageId="35" pageNumber="116">Fig.101</figureCitation>
). Acropodite short, robust, basally flattened. Pulvillus a short tube (
<emphasis id="B96CEA93FFCA9D2A4E2105ADDC6920FC" bold="true" box="[1299,1322,151,177]" pageId="35" pageNumber="116">p,</emphasis>
<figureCitation id="13232A04FFCA9D2A4E0205ADDCCC20FC" box="[1328,1423,151,177]" captionStart="FIGURES 101–106" captionStartId="36.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1828]" captionTargetId="figure-24@36.[151,1436,181,1828]" captionTargetPageId="36" captionText="FIGURES 101–106. Species of Benedictesmus, n. gen. 101, Gonopods of Benedictesmus aureus, n. sp., ventral view. SEM stub WS33-6. 102–106. Benedictesmus ellenae, n. sp. 102. Head and anterior rings, dorsal view. 103. Midbody ring, dorsal view. 104. Pygidium, lateral view. 105. Gonopods in situ, lateroventral view. 106. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: p, pulvillus; pp, pulvillar process; ppf, prefemorite pore field; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805047" httpUri="https://zenodo.org/record/4805047/files/figure.png" pageId="35" pageNumber="116">Fig. 101</figureCitation>
). Distal, pulvillar process (
<emphasis id="B96CEA93FFCA9D2A4A840581D89F2098" bold="true" box="[438,476,187,213]" pageId="35" pageNumber="116">pp,</emphasis>
<figureCitation id="13232A04FFCA9D2A4AD60581DB072098" box="[484,580,187,213]" captionStart="FIGURES 101–106" captionStartId="36.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1828]" captionTargetId="figure-24@36.[151,1436,181,1828]" captionTargetPageId="36" captionText="FIGURES 101–106. Species of Benedictesmus, n. gen. 101, Gonopods of Benedictesmus aureus, n. sp., ventral view. SEM stub WS33-6. 102–106. Benedictesmus ellenae, n. sp. 102. Head and anterior rings, dorsal view. 103. Midbody ring, dorsal view. 104. Pygidium, lateral view. 105. Gonopods in situ, lateroventral view. 106. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: p, pulvillus; pp, pulvillar process; ppf, prefemorite pore field; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805047" httpUri="https://zenodo.org/record/4805047/files/figure.png" pageId="35" pageNumber="116">Fig. 101</figureCitation>
) extending over pulvillus as short hood. Terminal zone (
<emphasis id="B96CEA93FFCA9D2A4FFA0581DDA62098" bold="true" box="[1224,1253,187,213]" pageId="35" pageNumber="116">tz,</emphasis>
<figureCitation id="13232A04FFCA9D2A4FDF0581DC0E2098" box="[1261,1357,187,213]" captionStart="FIGURES 101–106" captionStartId="36.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1828]" captionTargetId="figure-24@36.[151,1436,181,1828]" captionTargetPageId="36" captionText="FIGURES 101–106. Species of Benedictesmus, n. gen. 101, Gonopods of Benedictesmus aureus, n. sp., ventral view. SEM stub WS33-6. 102–106. Benedictesmus ellenae, n. sp. 102. Head and anterior rings, dorsal view. 103. Midbody ring, dorsal view. 104. Pygidium, lateral view. 105. Gonopods in situ, lateroventral view. 106. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: p, pulvillus; pp, pulvillar process; ppf, prefemorite pore field; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805047" httpUri="https://zenodo.org/record/4805047/files/figure.png" pageId="35" pageNumber="116">Fig. 101</figureCitation>
) short, bifurcate. Females with 19 rings, similar to males in nonsexual characters.
</paragraph>
</subSubSection>
<caption id="DF676609FFC89D284BA502ADDA8227BA" ID-DOI="http://doi.org/10.5281/zenodo.4805029" ID-Zenodo-Dep="4805029" httpUri="https://zenodo.org/record/4805029/files/figure.png" pageId="33" pageNumber="114" startId="33.[151,264,1943,1967]" targetBox="[179,1408,181,1919]" targetPageId="33">
<paragraph id="8BA73681FFC89D284BA502ADDA8227BA" blockId="33.[151,1436,1943,2040]" pageId="33" pageNumber="114">
<emphasis id="B96CEA93FFC89D284BA502ADD80E27E2" bold="true" box="[151,333,1943,1967]" pageId="33" pageNumber="114">FIGURES 91–95</emphasis>
.
<emphasis id="B96CEA93FFC89D284A6B02ADDA0627E2" bold="true" box="[345,837,1943,1968]" pageId="33" pageNumber="114">
<taxonomicName id="4C184D02FFC89D284A6B02ADDBCD27E2" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[345,654,1943,1967]" class="Diplopoda" family="Polydesmidae" genus="Rowlandesmus" kingdom="Animalia" order="Polydesmida" pageId="33" pageNumber="114" phylum="Arthropoda" rank="species" species="dentogonopus" status="n. gen., n. sp.">
<emphasis id="B96CEA93FFC89D284A6B02ADDBCD27E2" bold="true" box="[345,654,1943,1967]" italics="true" pageId="33" pageNumber="114">Rowlandesmus dentogonopus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFC89D2849A902A2DA6527FD" box="[667,806,1944,1968]" pageId="33" pageNumber="114" rank="species">n. gen., n. sp.</taxonomicNameLabel>
91
</emphasis>
. Head and anterior rings, dorsal view.
<emphasis id="B96CEA93FFC89D284FFB02ADDDA127E2" bold="true" box="[1225,1250,1943,1967]" pageId="33" pageNumber="114">92</emphasis>
. Pygidium, lateral view.
<emphasis id="B96CEA93FFC89D284BE60281D9AE279E" bold="true" box="[212,237,1979,2003]" pageId="33" pageNumber="114">93</emphasis>
. Right gonopod, mesal view.
<emphasis id="B96CEA93FFC89D2849130281DB79279E" bold="true" box="[545,570,1979,2003]" pageId="33" pageNumber="114">94</emphasis>
. Left gonopod, lateral view.
<emphasis id="B96CEA93FFC89D2848560281DA3E279E" bold="true" box="[868,893,1979,2003]" pageId="33" pageNumber="114">95</emphasis>
. Midbody ring, dorsal view. SEM stub WS33-4. Abbreviations:
<emphasis id="B96CEA93FFC89D284A2702DAD86027B5" bold="true" box="[277,291,2016,2040]" pageId="33" pageNumber="114">k</emphasis>
, “kink” in gonopod,
<emphasis id="B96CEA93FFC89D284ACB02DADB4427B5" bold="true" box="[505,519,2016,2040]" pageId="33" pageNumber="114">p</emphasis>
, pulvillus;
<emphasis id="B96CEA93FFC89D28494802DADBE327B5" bold="true" box="[634,672,2016,2040]" pageId="33" pageNumber="114">x, y</emphasis>
, processes of terminal zone.
</paragraph>
</caption>
<caption id="DF676609FFCB9D2B4BA502ADDC7F279E" ID-DOI="http://doi.org/10.5281/zenodo.4805039" ID-Zenodo-Dep="4805039" httpUri="https://zenodo.org/record/4805039/files/figure.png" pageId="34" pageNumber="115" startId="34.[151,264,1943,1967]" targetBox="[180,1408,181,1918]" targetPageId="34">
<paragraph id="8BA73681FFCB9D2B4BA502ADDC7F279E" blockId="34.[151,1436,1943,2040]" pageId="34" pageNumber="115">
<emphasis id="B96CEA93FFCB9D2B4BA502ADD81827E2" bold="true" box="[151,347,1943,1967]" pageId="34" pageNumber="115">FIGURES 96–100</emphasis>
.
<emphasis id="B96CEA93FFCB9D2B4A5B02ADDA5127E2" bold="true" box="[361,786,1943,1968]" pageId="34" pageNumber="115">
<taxonomicName id="4C184D02FFCB9D2B4A5B02ADDB1727E2" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[361,596,1943,1967]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="34" pageNumber="115" phylum="Arthropoda" rank="species" species="aureus" status="n. gen., n. sp.">
<emphasis id="B96CEA93FFCB9D2B4A5B02ADDB1727E2" bold="true" box="[361,596,1943,1967]" italics="true" pageId="34" pageNumber="115">Benedictesmus aureus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFCB9D2B495002A2DBB127FD" box="[610,754,1944,1968]" pageId="34" pageNumber="115" rank="species">n. gen., n. sp.</taxonomicNameLabel>
96
</emphasis>
. Male, dorsal view.
<emphasis id="B96CEA93FFCB9D2B48D702ADDABD27E2" bold="true" box="[997,1022,1943,1967]" pageId="34" pageNumber="115">97</emphasis>
. Anterior part of male, ventral view.
<emphasis id="B96CEA93FFCB9D2B4E4F02ADDCD527E2" bold="true" box="[1405,1430,1943,1967]" pageId="34" pageNumber="115">98</emphasis>
. Head and anterior rings, dorsal view.
<emphasis id="B96CEA93FFCB9D2B49210281DB6F279E" bold="true" box="[531,556,1979,2003]" pageId="34" pageNumber="115">99</emphasis>
. Midbody ring, dorsal view.
<emphasis id="B96CEA93FFCB9D2B48600281DA3B279E" bold="true" box="[850,888,1979,2003]" pageId="34" pageNumber="115">100</emphasis>
. Pygidium, lateral view. SEM stub WS33-6.
</paragraph>
</caption>
<subSubSection id="C302650AFFCA9D2A4BF50438DA3F21C4" pageId="35" pageNumber="116" type="distribution">
<paragraph id="8BA73681FFCA9D2A4BF50438DAAB210C" blockId="35.[151,1436,150,393]" pageId="35" pageNumber="116">
<emphasis id="B96CEA93FFCA9D2A4BF50438D8222151" bold="true" box="[199,353,258,284]" pageId="35" pageNumber="116">Distribution.</emphasis>
Known from the
<typeStatus id="54A38823FFCA9D2A49150439DB142150" box="[551,599,259,285]" pageId="35" pageNumber="116">type</typeStatus>
locality and one site a mile away (
<quantity id="4CE09B64FFCA9D2A48E90439DD5E2150" box="[987,1053,259,285]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="35" pageNumber="116" unit="mi" value="14.0">14 mi</quantity>
east of Gold Beach), where males and females were collected by E. M. Benedict on the same day as the
<typeStatus id="54A38823FFCA9D2A4897041DDAA7210C" box="[933,996,295,321]" pageId="35" pageNumber="116">types</typeStatus>
.
</paragraph>
<paragraph id="8BA73681FFCA9D2A4BF50471DA3F21C4" blockId="35.[151,1436,150,393]" pageId="35" pageNumber="116">
<emphasis id="B96CEA93FFCA9D2A4BF50471D8522128" bold="true" box="[199,273,331,357]" pageId="35" pageNumber="116">Notes.</emphasis>
The pore field at the distal part of the gonopod prefemorite seems to be unique in this group of small polydesmids; among the pores are three or four long, curled setae.
</paragraph>
</subSubSection>
</treatment>
</document>
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