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<document id="394093A9A65A94B690CEA58813283E94" ID-CLB-Dataset="46579" ID-DOI="10.11646/zootaxa.3755.5.3" ID-GBIF-Dataset="0396d331-6a2c-469f-a95a-a5b07cb2a91e" ID-ISSN="1175-5326" ID-Zenodo-Dep="225020" ID-ZooBank="B06C2DCA-78B6-46F9-8D4F-1716DD20C4A5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460374767504" checkinUser="plazi" docAuthor="Grismer, L. Lee, Belabut, Daicus M., Quah, Evan S. H., Onn, Chan Kin, Wood, Perry L. &amp; Hasim, Rosli" docDate="2014" docId="C03587DFFF9BFFECFF28FA03739EBA45" docLanguage="en" docName="zt03755p446.pdf" docOrigin="Zootaxa 3755 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus guakanthanensis Grismer, Belabut, Quah, Onn, Wood &amp; Hasim, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="445" masterDocId="3C0CFFA7FF9EFFE7FFBFFFBE7757BB27" masterDocTitle="A new species of karst forest-adapted Bent-toed Gecko (genus Cyrtodactylus Gray, 1827) belonging to the C. sworderi complex from a threatened karst forest in Perak, Peninsular Malaysia" masterLastPageNumber="446" masterPageNumber="434" pageNumber="439" updateTime="1698296198246" updateUser="ExternalLinkService">
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<mods:title id="9DF15FC306F2D912951362469B0C8A2E">A new species of karst forest-adapted Bent-toed Gecko (genus Cyrtodactylus Gray, 1827) belonging to the C. sworderi complex from a threatened karst forest in Perak, Peninsular Malaysia</mods:title>
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<mods:namePart id="9791DDD9525EFBC5FD4C7733BDF3ED55">Grismer, L. Lee</mods:namePart>
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<mods:namePart id="22F65DF2BFFFE95164AA05E8C911D380">Belabut, Daicus M.</mods:namePart>
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<mods:namePart id="DA5DB20B5588ACB5C076083ED7FCDE0B">Quah, Evan S. H.</mods:namePart>
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<mods:namePart id="9D5911515B574168A3064052AA170C94">Onn, Chan Kin</mods:namePart>
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<mods:namePart id="64EDC3375F2171B8F77F42B9E5DDDF82">Wood, Perry L.</mods:namePart>
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<mods:namePart id="2CA3BB84D39F8E30228C81B2C927E386">Hasim, Rosli</mods:namePart>
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<treatment id="C03587DFFF9BFFECFF28FA03739EBA45" ID-DOI="http://doi.org/10.5281/zenodo.6141125" ID-GBIF-Taxon="119412887" ID-Zenodo-Dep="6141125" LSID="urn:lsid:plazi:treatment:C03587DFFF9BFFECFF28FA03739EBA45" httpUri="http://treatment.plazi.org/id/C03587DFFF9BFFECFF28FA03739EBA45" lastPageId="11" lastPageNumber="445" pageId="5" pageNumber="439">
<subSubSection id="00866542FF9BFFE2FF28FA037643BD67" pageId="5" pageNumber="439" type="nomenclature">
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<heading id="136B81A5FF9BFFE2FF28FA03752FBEF0" bold="true" box="[151,632,1469,1495]" fontSize="11" level="1" pageId="5" pageNumber="439" reason="1">
<emphasis id="7AE8EADBFF9BFFE2FF28FA03752FBEF0" bold="true" box="[151,632,1469,1495]" pageId="5" pageNumber="439">
<taxonomicName id="8F9C4D4AFF9BFFE2FF28FA037544BEF0" box="[151,531,1469,1495]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="439" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF9BFFE2FF28FA037544BEF0" bold="true" box="[151,531,1469,1495]" italics="true" pageId="5" pageNumber="439">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF9BFFE2FDA5FA03752FBEF0" box="[538,632,1469,1495]" pageId="5" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="482336C9FF9BFFE2FF28FA5E7623BD3A" blockId="5.[151,632,1469,1600]" pageId="5" pageNumber="439">
<collectingCountry id="308B7659FF9BFFE2FF28FA5E7791BEDF" box="[151,198,1504,1528]" name="India" pageId="5" pageNumber="439">Gua</collectingCountry>
Kanthan Bent-toed Gecko Cicak
<collectingCountry id="308B7659FF9BFFE2FF60F9BA7659BD3B" box="[223,270,1540,1564]" name="India" pageId="5" pageNumber="439">Gua</collectingCountry>
Kanthan
</paragraph>
<paragraph id="482336C9FF9BFFE2FF28F9997643BD67" blockId="5.[151,632,1469,1600]" box="[151,276,1575,1600]" pageId="5" pageNumber="439">
<figureCitation id="D0A72A4CFF9BFFE2FF28F999778ABD67" box="[151,221,1575,1600]" captionStart="FIGURE 3" captionStartId="7.[151,250,1953,1975]" captionTargetBox="[327,1263,193,1921]" captionTargetId="figure@7.[327,1264,193,1921]" captionTargetPageId="7" captionText="FIGURE 3. Upper: adult male holotype of Cyrtodactylus guakanthanensis sp. nov. (LSUHC 11322) from Gua Kanthan, Perak, Peninsular Malaysia. Middle: adult male paratype of C. guakanthanensis sp. nov. (LSUHC 11322). Lower: hatchling C. guakanthanensis sp. nov. (LSUDPC 8175)" httpUri="https://zenodo.org/record/225023/files/figure.png" pageId="5" pageNumber="439">Figs 3</figureCitation>
,
<figureCitation id="D0A72A4CFF9BFFE2FF55F99677ADBD67" box="[234,250,1576,1600]" captionStart="FIGURE 4" captionStartId="8.[151,250,1927,1949]" captionTargetBox="[151,1436,193,1905]" captionTargetId="figure@8.[151,1436,193,1906]" captionTargetPageId="8" captionText="FIGURE 4. Habitat at the type locality, Gua Kanthan, Perak, Peninsular Malaysia. Upper: eroded and cracked limestone wall that provides refuge for Cyrtodactylus guakanthanensis sp. nov. Lower: structure of the limestone forest of the type locality." httpUri="https://zenodo.org/record/225024/files/figure.png" pageId="5" pageNumber="439">4</figureCitation>
, 6
</paragraph>
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<typeStatus id="9727886BFF9BFFE2FF28F9CD7653BDAC" box="[151,260,1651,1675]" pageId="5" pageNumber="439" type="holotype">
<emphasis id="7AE8EADBFF9BFFE2FF28F9CD7653BDAC" bold="true" box="[151,260,1651,1675]" pageId="5" pageNumber="439">Holotype</emphasis>
</typeStatus>
. Adult male (
<collectionCode id="2E8DAE0CFF9BFFE2FE77F9CD7571BDAC" box="[456,550,1651,1675]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="5" pageNumber="439">LSUHC</collectionCode>
11322) from
<collectingCountry id="308B7659FF9BFFE2FD55F9CD744EBDAC" box="[746,793,1651,1675]" name="India" pageId="5" pageNumber="439">Gua</collectingCountry>
Kanthan, Perak, Peninsular
<collectingCountry id="308B7659FF9BFFE2FB2FF9CC73A1BDAC" box="[1168,1270,1650,1675]" name="Malaysia" pageId="5" pageNumber="439">Malaysia</collectingCountry>
(
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,
<geoCoordinate id="2DA8500EFF9BFFE2FF28F9247664BD95" box="[151,307,1686,1714]" direction="east" orientation="longitude" pageId="5" pageNumber="439" precision="1" value="101.12203">101o07.322E</geoCoordinate>
;
<quantity id="8F649B2CFF9BFFE2FEFFF927762CBD95" box="[320,379,1689,1714]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="439" unit="m" value="45.0">45 m</quantity>
) collected by Daicus M. Belabut, Chan K. Onn, Evan S. H. Quah and L. Lee Grismer on
<date id="3C221009FF9BFFE2FA3FF924765FBDF1" pageId="5" pageNumber="439" value="2013-07-14">14 July 2013</date>
.
</paragraph>
<paragraph id="482336C9FF9BFFE2FF78F95F7201BDDD" blockId="5.[151,1436,1647,2038]" box="[199,1366,1761,1786]" pageId="5" pageNumber="439">
<emphasis id="7AE8EADBFF9BFFE2FF78F95F7611BDDE" bold="true" box="[199,326,1761,1785]" pageId="5" pageNumber="439">
<typeStatus id="9727886BFF9BFFE2FF78F95F7616BDDE" box="[199,321,1761,1785]" pageId="5" pageNumber="439" type="paratype">Paratypes</typeStatus>
.
</emphasis>
All
<typeStatus id="9727886BFF9BFFE2FEC8F95C76B5BDDD" box="[375,482,1762,1786]" pageId="5" pageNumber="439" type="paratype">paratypes</typeStatus>
(
<collectionCode id="2E8DAE0CFF9BFFE2FE4EF95C7518BDDD" box="[497,591,1762,1786]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="5" pageNumber="439">LSUHC</collectionCode>
11321, 11323, 1132536, 11339) bear the same data as the
<typeStatus id="9727886BFF9BFFE2FB53F95F7205BDDD" box="[1260,1362,1761,1786]" pageId="5" pageNumber="439" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="00866542FF9BFFE1FF78F8BB77F0B8C6" lastPageId="6" lastPageNumber="440" pageId="5" pageNumber="439" type="diagnosis">
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<emphasis id="7AE8EADBFF9BFFE2FF78F8BB766DBC39" bold="true" box="[199,314,1797,1822]" pageId="5" pageNumber="439">Diagnosis</emphasis>
.
<taxonomicName id="8F9C4D4AFF9BFFE2FEF6F8BB75FCBC3A" box="[329,683,1797,1821]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="439" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF9BFFE2FEF6F8BB75FCBC3A" box="[329,683,1797,1821]" italics="true" pageId="5" pageNumber="439">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
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<emphasis id="7AE8EADBFF9BFFE2FD0CF8B8745ABC39" bold="true" box="[691,781,1798,1822]" pageId="5" pageNumber="439">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from all other Sundaland species by having the following suite of character states: adult SVL
<quantity id="8F649B2CFF9BFFE2FC89F894748ABC66" box="[822,989,1834,1858]" metricMagnitude="-2" metricUnit="m" metricValue="7.995" metricValueMax="8.22" metricValueMin="7.77" pageId="5" pageNumber="439" unit="mm" value="79.95" valueMax="82.2" valueMin="77.7">77.782.2 mm</quantity>
adult SVL; moderately sized, conical, weakly keeled, body tubercles; tubercles present on occiput, nape, and limbs, and extend posteriorly beyond base of tail; 3744 ventral scales; no transversely enlarged, median, subcaudal scales; proximal subdigital lamellae transversely expanded; 1921 subdigital lamellae on fourth toe; abrupt transition between posterior and ventral femoral scales; enlarged femoral scales; no femoral or precloacal pores; precloacal groove absent; wide, dark postorbital stripes from each eye extending posteriorly to the anterior margin of the shoulder region thence forming a transverse band across the anterior margin of the forelimbs; and body bearing five (rarely four) wide, bold, dark bands. The meristic characters are scored against
<emphasis id="7AE8EADBFF98FFE1FD71FF0274BABBF3" box="[718,1005,188,212]" italics="true" pageId="6" pageNumber="440">
<taxonomicName id="8F9C4D4AFF98FFE1FD71FF027402BBF3" box="[718,853,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="sworderi">C. sworderi</taxonomicName>
,
<taxonomicName id="8F9C4D4AFF98FFE1FCDCFF0274BABBF3" box="[867,1005,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="tebuensis">C. tebuensis</taxonomicName>
</emphasis>
and
<taxonomicName id="8F9C4D4AFF98FFE1FB98FF0273A7BBF3" box="[1063,1264,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="7AE8EADBFF98FFE1FB98FF0273A7BBF3" box="[1063,1264,188,212]" italics="true" pageId="6" pageNumber="440">C. quadrivirgatus</emphasis>
</taxonomicName>
in
<tableCitation id="051E0372FF98FFE1FAA9FF02723CBBF2" box="[1302,1387,188,213]" captionStart="TABLE 5" captionStartId="6.[151,239,296,319]" captionTargetBox="[165,1420,389,939]" captionTargetPageId="6" captionText="TABLE 5. Diagnostic characters (in bold) separating the species of the C. sworderi complex from Cyrtodactylus guakanthanensis sp. nov." httpUri="http://table.plazi.org/id/1CE36641FF98FFE1FF28FE9676F1BA45" pageId="6" pageNumber="440" tableUuid="1CE36641FF98FFE1FF28FE9676F1BA45">Table 5</tableCitation>
and against all other Sundaland species in
<bibRefCitation id="2C0D4B38FF98FFE1FDFCFF61746FBBDF" author="Grismer" box="[579,824,223,248]" pageId="6" pageNumber="440" refString="Grismer, L. L., Wood, P. L. &amp; Lim, K. K. P. (2012 b) Cyrtodactylus majulah, a new species of Bent-toed Gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. Raffles Bulletin of Zoology, 60, 487 - 499." type="journal article" year="2012" yearSuffix="b">
Grismer
<emphasis id="7AE8EADBFF98FFE1FD1AFF5F758ABBDF" box="[677,733,224,248]" italics="true" pageId="6" pageNumber="440">et al.</emphasis>
(2012b
</bibRefCitation>
:
<tableCitation id="051E0372FF98FFE1FCFDFF6174C0BBDF" box="[834,919,223,248]" captionStart="TABLE 5" captionStartId="6.[151,239,296,319]" captionTargetBox="[165,1420,389,939]" captionTargetPageId="6" captionText="TABLE 5. Diagnostic characters (in bold) separating the species of the C. sworderi complex from Cyrtodactylus guakanthanensis sp. nov." httpUri="http://table.plazi.org/id/1CE36641FF98FFE1FF28FE9676F1BA45" pageId="6" pageNumber="440" tableUuid="1CE36641FF98FFE1FF28FE9676F1BA45">Table 5</tableCitation>
).
</paragraph>
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<paragraph id="482336C9FF98FFE1FF28FE9676F1BA45" blockId="6.[151,1435,295,354]" pageId="6" pageNumber="440">
<emphasis id="7AE8EADBFF98FFE1FF28FE967647BA18" bold="true" box="[151,272,296,319]" pageId="6" pageNumber="440">TABLE 5.</emphasis>
Diagnostic characters (in bold) separating the species of the
<taxonomicName id="8F9C4D4AFF98FFE1FC6FFE967305BA18" box="[976,1106,295,319]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="sworderi">
<emphasis id="7AE8EADBFF98FFE1FC6FFE967305BA18" box="[976,1106,295,319]" italics="true" pageId="6" pageNumber="440">C. sworderi</emphasis>
</taxonomicName>
complex from
<taxonomicName id="8F9C4D4AFF98FFE1FAB9FE99761DBA45" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF98FFE1FAB9FE99761DBA45" italics="true" pageId="6" pageNumber="440">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF98FFE1FEEEFEF576F1BA45" box="[337,422,331,354]" pageId="6" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="482336C9FF98FFE1FDE6FE3B7219B88C" pageId="6" pageNumber="440">
<table id="3A9CC469FF980018FF1AFE3B72DBB88C" box="[165,1420,389,939]" gridcols="3" gridrows="12" pageId="6" pageNumber="440">
<tr id="F6AC348BFF980018FF1AFE3B72DBBABC" box="[165,1420,389,411]" gridrow="0" pageId="6" pageNumber="440" rowspan-0="1">
<th id="B57D5DF7FF980018FDE7FE3B73CDBABC" box="[600,1178,389,411]" gridcol="1" gridrow="0" pageId="6" pageNumber="440">
<taxonomicName id="8F9C4D4AFF98FFE1FDE6FE3B7454BABC" box="[601,771,389,411]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF98FFE1FDE6FE3B7454BABC" box="[601,771,389,411]" italics="true" pageId="6" pageNumber="440">guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF98FFE1FCB6FE3B740BBABC" box="[777,860,389,411]" pageId="6" pageNumber="440" rank="species">
<emphasis id="7AE8EADBFF98FFE1FCB6FE3B740BBABC" bold="true" box="[777,860,389,411]" pageId="6" pageNumber="440">sp. nov.</emphasis>
</taxonomicNameLabel>
<emphasis id="7AE8EADBFF98FFE1FCC6FE3B73CDBABC" box="[889,1178,389,411]" italics="true" pageId="6" pageNumber="440">
<taxonomicName id="8F9C4D4AFF98FFE1FCC6FE3B7480BABC" box="[889,983,389,411]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="tebuensis">tebuensis</taxonomicName>
<taxonomicName id="8F9C4D4AFF98FFE1FBFFFE3B73CDBABC" box="[1088,1178,389,411]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="sworderi">sworderi</taxonomicName>
</emphasis>
</th>
<th id="B57D5DF7FF980018FB4AFE3B72DBBABC" box="[1269,1420,389,411]" gridcol="2" gridrow="0" pageId="6" pageNumber="440">
<taxonomicName id="8F9C4D4AFF98FFE1FB4AFE3B72DBBABC" box="[1269,1420,389,411]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="440" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="7AE8EADBFF98FFE1FB4AFE3B72DBBABC" box="[1269,1420,389,411]" italics="true" pageId="6" pageNumber="440">quadrivirgatus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="F6AC348BFF980018FF1AFE0F72DBBAE0" box="[165,1420,433,455]" gridrow="1" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFE0F7549BAE0" box="[165,542,433,455]" gridcol="0" gridrow="1" pageId="6" pageNumber="440">Supralabials</th>
<td id="B57D5DF7FF980018FDE7FE0F73CDBAE0" box="[600,1178,433,455]" gridcol="1" gridrow="1" pageId="6" pageNumber="440">
9,10
<emphasis id="7AE8EADBFF98FFE1FCC6FE0F74EFBAE0" bold="true" box="[889,952,433,455]" pageId="6" pageNumber="440">1113</emphasis>
10
<emphasis id="7AE8EADBFF98FFE1FBE5FE0F7331BAE0" bold="true" box="[1114,1126,433,455]" pageId="6" pageNumber="440"></emphasis>
13
</td>
<td id="B57D5DF7FF980018FB4AFE0F72DBBAE0" box="[1269,1420,433,455]" gridcol="2" gridrow="1" pageId="6" pageNumber="440">
8
<emphasis id="7AE8EADBFF98FFE1FABDFE0F7259BAE0" bold="true" box="[1282,1294,433,455]" pageId="6" pageNumber="440"></emphasis>
11
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFE6372DBBAD4" box="[165,1420,477,499]" gridrow="2" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFE637549BAD4" box="[165,542,477,499]" gridcol="0" gridrow="2" pageId="6" pageNumber="440">infralaials</th>
<td id="B57D5DF7FF980018FDE7FE6373CDBAD4" box="[600,1178,477,499]" gridcol="1" gridrow="2" pageId="6" pageNumber="440">
7,8 8
<emphasis id="7AE8EADBFF98FFE1FC39FE6374C4BAD4" bold="true" box="[902,915,477,499]" pageId="6" pageNumber="440"></emphasis>
10 10
</td>
<td id="B57D5DF7FF980018FB4AFE6372DBBAD4" box="[1269,1420,477,499]" gridcol="2" gridrow="2" pageId="6" pageNumber="440">
8
<emphasis id="7AE8EADBFF98FFE1FABDFE637259BAD4" bold="true" box="[1282,1294,477,499]" pageId="6" pageNumber="440"></emphasis>
11
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFDB772DBB938" box="[165,1420,521,543]" gridrow="3" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFDB77549B938" box="[165,542,521,543]" gridcol="0" gridrow="3" pageId="6" pageNumber="440">No. of paravertebral tubercles</th>
<td id="B57D5DF7FF980018FDE7FDB773CDB938" box="[600,1178,521,543]" gridcol="1" gridrow="3" pageId="6" pageNumber="440">
32
<emphasis id="7AE8EADBFF98FFE1FDCDFDB77528B938" bold="true" box="[626,639,521,543]" pageId="6" pageNumber="440"></emphasis>
34 32
<emphasis id="7AE8EADBFF98FFE1FC2CFDB774C8B938" bold="true" box="[915,927,521,543]" pageId="6" pageNumber="440"></emphasis>
39
<emphasis id="7AE8EADBFF98FFE1FBFFFDB77328B938" bold="true" box="[1088,1151,521,543]" pageId="6" pageNumber="440">3638</emphasis>
</td>
<td id="B57D5DF7FF980018FB4AFDB772DBB938" box="[1269,1420,521,543]" gridcol="2" gridrow="3" pageId="6" pageNumber="440">
32
<emphasis id="7AE8EADBFF98FFE1FAB0FDB7724BB938" bold="true" box="[1295,1308,521,543]" pageId="6" pageNumber="440"></emphasis>
39
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFD8B72DBB96C" box="[165,1420,565,587]" gridrow="4" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFD8B7549B96C" box="[165,542,565,587]" gridcol="0" gridrow="4" pageId="6" pageNumber="440">No. of ventral scales</th>
<td id="B57D5DF7FF980018FDE7FD8B73CDB96C" box="[600,1178,565,587]" gridcol="1" gridrow="4" pageId="6" pageNumber="440">
37
<emphasis id="7AE8EADBFF98FFE1FDCDFD8B7528B96C" bold="true" box="[626,639,565,587]" pageId="6" pageNumber="440"></emphasis>
44 43
<emphasis id="7AE8EADBFF98FFE1FC2CFD8B74C8B96C" bold="true" box="[915,927,565,587]" pageId="6" pageNumber="440"></emphasis>
51 40
<emphasis id="7AE8EADBFF98FFE1FBE5FD8B7331B96C" bold="true" box="[1114,1126,565,587]" pageId="6" pageNumber="440"></emphasis>
49
</td>
<td id="B57D5DF7FF980018FB4AFD8B72DBB96C" box="[1269,1420,565,587]" gridcol="2" gridrow="4" pageId="6" pageNumber="440">
28
<emphasis id="7AE8EADBFF98FFE1FAB0FD8B724BB96C" bold="true" box="[1295,1308,565,587]" pageId="6" pageNumber="440"></emphasis>
40
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFDDF72DBB984" box="[165,1420,609,675]" gridrow="5" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFDDF7549B984" box="[165,542,609,675]" gridcol="0" gridrow="5" pageId="6" pageNumber="440">Postfemoral scale transition abrupt No. of 4th toe lamella</th>
<td id="B57D5DF7FF980018FDE7FDDF73CDB984" box="[600,1178,609,675]" gridcol="1" gridrow="5" pageId="6" pageNumber="440">
yes yes
<emphasis id="7AE8EADBFF98FFE1FBFFFDDF730CB950" bold="true" box="[1088,1115,609,631]" pageId="6" pageNumber="440">no</emphasis>
19
<emphasis id="7AE8EADBFF98FFE1FDCDFD337528B984" bold="true" box="[626,639,653,675]" pageId="6" pageNumber="440"></emphasis>
21 17
<emphasis id="7AE8EADBFF98FFE1FC2CFD3374C8B984" bold="true" box="[915,927,653,675]" pageId="6" pageNumber="440"></emphasis>
21 16
<emphasis id="7AE8EADBFF98FFE1FBE5FD337331B984" bold="true" box="[1114,1126,653,675]" pageId="6" pageNumber="440"></emphasis>
20
</td>
<td id="B57D5DF7FF980018FB4AFDDF72DBB984" box="[1269,1420,609,675]" gridcol="2" gridrow="5" pageId="6" pageNumber="440">
<emphasis id="7AE8EADBFF98FFE1FB4AFDDF7247B950" bold="true" box="[1269,1296,609,631]" pageId="6" pageNumber="440">no</emphasis>
18
<emphasis id="7AE8EADBFF98FFE1FAB0FD33724BB984" bold="true" box="[1295,1308,653,675]" pageId="6" pageNumber="440"></emphasis>
23
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFD0772DBB9E8" box="[165,1420,697,719]" gridrow="6" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFD077549B9E8" box="[165,542,697,719]" gridcol="0" gridrow="6" pageId="6" pageNumber="440">Precloacal and /or femoral pores</th>
<td id="B57D5DF7FF980018FDE7FD0773CDB9E8" box="[600,1178,697,719]" gridcol="1" gridrow="6" pageId="6" pageNumber="440">
absent absent
<emphasis id="7AE8EADBFF98FFE1FBFFFD0773C7B9E8" bold="true" box="[1088,1168,697,719]" pageId="6" pageNumber="440">present</emphasis>
</td>
<td id="B57D5DF7FF980018FB4AFD0772DBB9E8" box="[1269,1420,697,719]" gridcol="2" gridrow="6" pageId="6" pageNumber="440">
<emphasis id="7AE8EADBFF98FFE1FB4AFD077219B9E8" bold="true" box="[1269,1358,697,719]" pageId="6" pageNumber="440">variable</emphasis>
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFD5B72DBB9DC" box="[165,1420,741,763]" gridrow="7" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFD5B7549B9DC" box="[165,542,741,763]" gridcol="0" gridrow="7" pageId="6" pageNumber="440">No. of precloacal pores</th>
<td id="B57D5DF7FF980018FDE7FD5B73CDB9DC" box="[600,1178,741,763]" gridcol="1" gridrow="7" pageId="6" pageNumber="440">
0 0
<emphasis id="7AE8EADBFF98FFE1FBFFFD5B7331B9DC" bold="true" box="[1088,1126,741,763]" pageId="6" pageNumber="440">59</emphasis>
</td>
<td id="B57D5DF7FF980018FB4AFD5B72DBB9DC" box="[1269,1420,741,763]" gridcol="2" gridrow="7" pageId="6" pageNumber="440">
<emphasis id="7AE8EADBFF98FFE1FB4AFD5B727FB9DC" bold="true" box="[1269,1320,741,763]" pageId="6" pageNumber="440">012</emphasis>
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFCAF72DBB800" box="[165,1420,785,807]" gridrow="8" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFCAF7549B800" box="[165,542,785,807]" gridcol="0" gridrow="8" pageId="6" pageNumber="440">No. of enlarged femorocloacal scales</th>
<td id="B57D5DF7FF980018FDE7FCAF73CDB800" box="[600,1178,785,807]" gridcol="1" gridrow="8" pageId="6" pageNumber="440">
36
<emphasis id="7AE8EADBFF98FFE1FDCDFCAF7528B800" bold="true" box="[626,639,785,807]" pageId="6" pageNumber="440"></emphasis>
41 31
<emphasis id="7AE8EADBFF98FFE1FC2CFCAF74C8B800" bold="true" box="[915,927,785,807]" pageId="6" pageNumber="440"></emphasis>
37
<emphasis id="7AE8EADBFF98FFE1FBFFFCAF731AB800" bold="true" box="[1088,1101,785,807]" pageId="6" pageNumber="440">0</emphasis>
</td>
<td id="B57D5DF7FF980018FB4AFCAF72DBB800" box="[1269,1420,785,807]" gridcol="2" gridrow="8" pageId="6" pageNumber="440">
31
<emphasis id="7AE8EADBFF98FFE1FAB0FCAF724BB800" bold="true" box="[1295,1308,785,807]" pageId="6" pageNumber="440"></emphasis>
44
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFC8372DBB874" box="[165,1420,829,851]" gridrow="9" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFC837549B874" box="[165,542,829,851]" gridcol="0" gridrow="9" pageId="6" pageNumber="440">Light caudal bands</th>
<td id="B57D5DF7FF980018FDE7FC8373CDB874" box="[600,1178,829,851]" gridcol="1" gridrow="9" pageId="6" pageNumber="440">
wide
<emphasis id="7AE8EADBFF98FFE1FCC6FC8373C7B874" bold="true" box="[889,1168,829,851]" pageId="6" pageNumber="440">narrow narrow</emphasis>
</td>
<td id="B57D5DF7FF980018FB4AFC8372DBB874" box="[1269,1420,829,851]" gridcol="2" gridrow="9" pageId="6" pageNumber="440">wide</td>
</tr>
<tr id="F6AC348BFF980018FF1AFCD472DBB8A7" box="[165,1420,874,896]" gridrow="10" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFCD47549B8A7" box="[165,542,874,896]" gridcol="0" gridrow="10" pageId="6" pageNumber="440">No. light bands on original tail</th>
<td id="B57D5DF7FF980018FDE7FCD473CDB8A7" box="[600,1178,874,896]" gridcol="1" gridrow="10" pageId="6" pageNumber="440">
9
<emphasis id="7AE8EADBFF98FFE1FCC6FCD474EFB8A7" bold="true" box="[889,952,874,896]" pageId="6" pageNumber="440">1014</emphasis>
9
<emphasis id="7AE8EADBFF98FFE1FBF2FCD4730DB8A7" bold="true" box="[1101,1114,874,896]" pageId="6" pageNumber="440"></emphasis>
15
</td>
<td id="B57D5DF7FF980018FB4AFCD472DBB8A7" box="[1269,1420,874,896]" gridcol="2" gridrow="10" pageId="6" pageNumber="440">
9
<emphasis id="7AE8EADBFF98FFE1FABDFCD47259B8A7" bold="true" box="[1282,1294,874,896]" pageId="6" pageNumber="440"></emphasis>
15
</td>
</tr>
<tr id="F6AC348BFF980018FF1AFC2B72DBB88C" box="[165,1420,917,939]" gridrow="11" pageId="6" pageNumber="440">
<th id="B57D5DF7FF980018FF1AFC2B7549B88C" box="[165,542,917,939]" gridcol="0" gridrow="11" pageId="6" pageNumber="440">Dorsal pattern</th>
<td id="B57D5DF7FF980018FDE7FC2B73CDB88C" box="[600,1178,917,939]" gridcol="1" gridrow="11" pageId="6" pageNumber="440">banded Striped/spotted spotted</td>
<td id="B57D5DF7FF980018FB4AFC2B72DBB88C" box="[1269,1420,917,939]" gridcol="2" gridrow="11" pageId="6" pageNumber="440">variable</td>
</tr>
</table>
</paragraph>
<paragraph id="482336C9FF98FFE1FF28FC7477F0B8C6" blockId="6.[151,1436,970,1570]" box="[151,167,970,993]" pageId="6" pageNumber="440">=</paragraph>
</subSubSection>
<subSubSection id="00866542FF98FFEEFF78FC537454BE5F" lastPageId="9" lastPageNumber="443" pageId="6" pageNumber="440" type="description">
<paragraph id="482336C9FF98FFE1FF78FC537690BD05" blockId="6.[151,1436,970,1570]" pageId="6" pageNumber="440">
<emphasis id="7AE8EADBFF98FFE1FF78FC537606BF21" bold="true" box="[199,337,1005,1030]" pageId="6" pageNumber="440">Description</emphasis>
. Adult male SVL
<quantity id="8F649B2CFF98FFE1FD97FC5075C6BF22" box="[552,657,1006,1030]" metricMagnitude="-2" metricUnit="m" metricValue="7.97" pageId="6" pageNumber="440" unit="mm" value="79.7">79.7 mm</quantity>
; head large, moderate in length (HL/SVL 0.29) and width (HW/HL 0.65), somewhat depressed (HD/HL 0.40), distinct from neck, and triangular in dorsal profile; lores weakly inflated, prefrontal region slightly concave; canthus rostralis smoothly rounded; snout elongate (ES/HL 0.44) and rounded in dorsal profile; eye large (ED/HL 0.22); ear opening elliptical and small (EL/HL 0.09); eye-to-ear distance greater than diameter of eye; rostral subrectangular with a deep dorsomedial furrow containing a postrostral; rostral partially divided dorsally, bordered posteriorly by large left and right supranasals, one large medial postrostral (=internasal) and one small azygous postrostral; external nares bordered anteriorly by rostral, dorsally by two supranasals (anterior one largest), posteriorly by five postnasals and ventrally by first supralabial; nine (R, L) square to rectangular supralabials extending to and tapering smoothly below posterior margin of orbit; eight (R), seven (L) infralabials tapering smoothly posteriorly to below posterior margin of orbit; scales of rostrum, lores, top of head, and occiput small and granular; scales on top of occiput intermixed with slightly enlarged tubercles; dorsal and ventral superciliaries rectangular; mental triangular, bordered laterally by first infralabials and posteriorly by left and right rectangular postmentals contacting medially for approximately 50% of their length posterior to mental; one enlarged row of sublabials extending posteriorly to 4th or 5th infralabial; gular scales small and raised, grading posteriorly into slightly larger, flatter, throat scales, and thence into large, flat, imbricate pectoral and ventral scales.
</paragraph>
<caption id="1CE36641FF99FFE0FF28F81F751EBCD3" httpUri="https://zenodo.org/record/225023/files/figure.png" pageId="7" pageNumber="441" targetBox="[327,1263,193,1921]" targetPageId="7">
<paragraph id="482336C9FF99FFE0FF28F81F751EBCD3" blockId="7.[151,1436,1953,2036]" pageId="7" pageNumber="441">
<emphasis id="7AE8EADBFF99FFE0FF28F81F7645BC90" bold="true" box="[151,274,1953,1975]" pageId="7" pageNumber="441">FIGURE 3.</emphasis>
Upper: adult male holotype of
<taxonomicName id="8F9C4D4AFF99FFE0FDF3F81F74DDBC90" box="[588,906,1953,1975]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="441" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF99FFE0FDF3F81F74DDBC90" box="[588,906,1953,1975]" italics="true" pageId="7" pageNumber="441">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF99FFE0FC2FF81F74B7BC90" box="[912,992,1953,1975]" pageId="7" pageNumber="441" rank="species">
<emphasis id="7AE8EADBFF99FFE0FC2FF81F74B7BC90" bold="true" box="[912,992,1953,1975]" pageId="7" pageNumber="441">sp. nov.</emphasis>
</taxonomicNameLabel>
(LSUHC 11322) from Gua Kanthan, Perak, Peninsular Malaysia. Middle: adult male paratype of
<taxonomicName id="8F9C4D4AFF99FFE0FD6DF87E74C9BCF2" box="[722,926,1983,2005]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="441" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF99FFE0FD6DF87E74C9BCF2" box="[722,926,1983,2005]" italics="true" pageId="7" pageNumber="441">C. guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF99FFE0FC16F80174A9BCF2" box="[937,1022,1983,2005]" pageId="7" pageNumber="441" rank="species">
<emphasis id="7AE8EADBFF99FFE0FC16F80174A9BCF2" bold="true" box="[937,1022,1983,2005]" pageId="7" pageNumber="441">sp. nov.</emphasis>
</taxonomicNameLabel>
(LSUHC 11322). Lower: hatchling
<taxonomicName id="8F9C4D4AFF99FFE0FA3BF87E7616BCD3" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="441" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF99FFE0FA3BF87E7616BCD3" italics="true" pageId="7" pageNumber="441">C. guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF99FFE0FEF8F86076C0BCD3" box="[327,407,2014,2036]" pageId="7" pageNumber="441" rank="species">
<emphasis id="7AE8EADBFF99FFE0FEF8F86076C0BCD3" bold="true" box="[327,407,2014,2036]" pageId="7" pageNumber="441">sp. nov.</emphasis>
</taxonomicNameLabel>
(LSUDPC 8175)
</paragraph>
</caption>
<caption id="1CE36641FF96FFEFFF28F83972D4BC9B" httpUri="https://zenodo.org/record/225024/files/figure.png" pageId="8" pageNumber="442" targetBox="[151,1436,193,1905]" targetPageId="8">
<paragraph id="482336C9FF96FFEFFF28F83972D4BC9B" blockId="8.[151,1436,1927,1980]" pageId="8" pageNumber="442">
<emphasis id="7AE8EADBFF96FFEFFF28F8397644BCBA" bold="true" box="[151,275,1927,1949]" pageId="8" pageNumber="442">FIGURE 4.</emphasis>
Habitat at the type locality, Gua Kanthan, Perak, Peninsular Malaysia. Upper: eroded and cracked limestone wall that provides refuge for
<taxonomicName id="8F9C4D4AFF96FFEFFE34F818759EBC9B" box="[395,713,1958,1980]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="442" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF96FFEFFE34F818759EBC9B" box="[395,713,1958,1980]" italics="true" pageId="8" pageNumber="442">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF96FFEFFD6FF8187477BC9B" box="[720,800,1958,1980]" pageId="8" pageNumber="442" rank="species">
<emphasis id="7AE8EADBFF96FFEFFD6FF8187477BC9B" bold="true" box="[720,800,1958,1980]" pageId="8" pageNumber="442">sp. nov.</emphasis>
</taxonomicNameLabel>
Lower: structure of the limestone forest of the type locality.
</paragraph>
</caption>
<paragraph id="482336C9FF97FFEEFF78FF29723FBA67" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">Body relatively short (AG/SVL 0.42) with weak, tuberculate ventrolateral folds; dorsal scales small and granular, interspersed with larger, conical, semi-regularly arranged, weakly keeled tubercles; tubercles extending from occiput to anterior one-third of tail; tubercles on occiput and nape relatively small, those on body largest; approximately 17 longitudinal rows of dorsal tubercles and 32 paravertebral tubercles; 37 flat, imbricate ventrals, ventrals much larger than dorsals; patch of enlarged precloacal scales lacking pores; precloacal groove absent.</paragraph>
<paragraph id="482336C9FF97FFEEFF78FEF274C3BAF7" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">Forelimbs moderate in stature, relatively short (FL/SVL 0.17); granular scales of forearm larger than those of body and interspersed with small tubercles; palmar scales rounded; digits well-developed, inflected at basal interphalangeal joints; subdigital lamellae slightly enlarged proximal to joint inflections, digits narrower distal to joints; claws well-developed, sheathed by a dorsal and ventral scale.</paragraph>
<paragraph id="482336C9FF97FFEEFF78FE6273B2B9D7" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">
<collectingCountry id="308B7659FF97FFEEFF78FE6277A8BAD2" box="[199,255,476,501]" name="India" pageId="9" pageNumber="443">Hind</collectingCountry>
limbs more robust than forelimbs, moderate in length (TBL/SVL 0.18), covered dorsally by granular scales interspersed with larger tubercles and anteriorly by granular scales; ventral scales of femora flat and larger than dorsals; ventral tibial scales flat and imbricate; a single row of enlarged femoral scales extend medially from proximal region of femur making contact with large precloacal scales; this enlarged row is bordered anteriorly by two rows of slightly smaller scales; femoral pores absent; dorsal and ventral femoral scales meeting abruptly on posteroventral margin of thigh; plantar scales low and slightly rounded; digits well-developed, inflected at basal interphalangeal joints; subdigital lamellae enlarged proximal to inflected joints, digits narrower distal to joints; 20 (L,R) subdigital lamellae on 4th toe; claws well-developed, sheathed by a dorsal and ventral scale.
</paragraph>
<paragraph id="482336C9FF97FFEEFF78FD427355B8A7" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">Original tail widest at base, tapering to a point, approximately last one-fifth of tail missing; dorsal scales on base of tail granular, becoming flatter posteriorly; no median row of transversely enlarged subcaudal scales; caudal scales arranged in semi-whorls; three enlarged tubercles on both sides of base of tail; base of tail bearing lateral, bulbous swellings; all postcloacal scales moderately sized, flat, and imbricate.</paragraph>
<paragraph id="482336C9FF97FFEEFF78FC357397BF87" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">
<emphasis id="7AE8EADBFF97FFEEFF78FC3576AFB883" bold="true" box="[199,504,907,932]" pageId="9" pageNumber="443">
Coloration in life (
<figureCitation id="D0A72A4CFF97FFEEFE1CFC3576BEB883" box="[419,489,907,932]" captionStart="FIGURE 3" captionStartId="7.[151,250,1953,1975]" captionTargetBox="[327,1263,193,1921]" captionTargetId="figure@7.[327,1264,193,1921]" captionTargetPageId="7" captionText="FIGURE 3. Upper: adult male holotype of Cyrtodactylus guakanthanensis sp. nov. (LSUHC 11322) from Gua Kanthan, Perak, Peninsular Malaysia. Middle: adult male paratype of C. guakanthanensis sp. nov. (LSUHC 11322). Lower: hatchling C. guakanthanensis sp. nov. (LSUDPC 8175)" httpUri="https://zenodo.org/record/225023/files/figure.png" pageId="9" pageNumber="443">Fig. 3</figureCitation>
).
</emphasis>
Ground color of head, body, limbs and anterior portion of tail beige; snout and top of head brown bearing darker mottling; wide, dark brown postorbital stripes from each eye extend posteriorly to the anterior margin of the shoulder region, thence connecting medially and forming a transverse band across the base of the anterior margin of the forelimbs; single wide dark brown chevron-shaped marking on nape and base of occiput; four wide, dark brown body bands between limb insertions that extend laterally one-half way down flanks; single wide, dark brown sacral band; five dark caudal bands becoming darker and wider posteriorly; dorsal surface of limbs beige bearing darker mottling; ventral surface of head, body, and limbs immaculate off-white except for fine black stippling in scales; subcaudal region nearly black with six light bands encircling tail.
</paragraph>
<paragraph id="482336C9FF97FFEEFF78FB157454BE5F" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">
<emphasis id="7AE8EADBFF97FFEEFF78FB157668BFE3" bold="true" box="[199,319,1195,1220]" pageId="9" pageNumber="443">Variation.</emphasis>
The
<typeStatus id="9727886BFF97FFEEFEC3FB1376B0BFE2" box="[380,487,1197,1221]" pageId="9" pageNumber="443" type="paratype">paratypes</typeStatus>
closely resemble the
<typeStatus id="9727886BFF97FFEEFD5EFB127414BFE2" box="[737,835,1196,1221]" pageId="9" pageNumber="443" type="holotype">holotype</typeStatus>
in all aspects of coloration and pattern (
<figureCitation id="D0A72A4CFF97FFEEFAACFB12720EBFE3" box="[1299,1369,1196,1221]" captionStart="FIGURE 3" captionStartId="7.[151,250,1953,1975]" captionTargetBox="[327,1263,193,1921]" captionTargetId="figure@7.[327,1264,193,1921]" captionTargetPageId="7" captionText="FIGURE 3. Upper: adult male holotype of Cyrtodactylus guakanthanensis sp. nov. (LSUHC 11322) from Gua Kanthan, Perak, Peninsular Malaysia. Middle: adult male paratype of C. guakanthanensis sp. nov. (LSUHC 11322). Lower: hatchling C. guakanthanensis sp. nov. (LSUDPC 8175)" httpUri="https://zenodo.org/record/225023/files/figure.png" pageId="9" pageNumber="443">Fig. 3</figureCitation>
). The body bands in LSUHC 1132526, and 11339 are oblique as opposed to being transverse. The regenerated tails of LSUHC 11321, 11323, 11326, and 11339 are mottled and lack banding. Hatchlings and juveniles (LSUDPC 8175 79) are more boldly marked, having a nearly immaculate ground color and generally darker bands and the posterior one-third of the tail is white (
<figureCitation id="D0A72A4CFF97FFEEFE5DFA827574BE73" box="[482,547,1340,1365]" captionStart="FIGURE 3" captionStartId="7.[151,250,1953,1975]" captionTargetBox="[327,1263,193,1921]" captionTargetId="figure@7.[327,1264,193,1921]" captionTargetPageId="7" captionText="FIGURE 3. Upper: adult male holotype of Cyrtodactylus guakanthanensis sp. nov. (LSUHC 11322) from Gua Kanthan, Perak, Peninsular Malaysia. Middle: adult male paratype of C. guakanthanensis sp. nov. (LSUHC 11322). Lower: hatchling C. guakanthanensis sp. nov. (LSUDPC 8175)" httpUri="https://zenodo.org/record/225023/files/figure.png" pageId="9" pageNumber="443">Fig. 3</figureCitation>
)—a character noted in other karst-dwelling species of
<taxonomicName id="8F9C4D4AFF97FFEEFB2DFA827278BE73" box="[1170,1327,1340,1364]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="443" phylum="Chordata" rank="genus">
<emphasis id="7AE8EADBFF97FFEEFB2DFA827278BE73" box="[1170,1327,1340,1364]" italics="true" pageId="9" pageNumber="443">Cyrtodactylus</emphasis>
</taxonomicName>
(
<bibRefCitation id="2C0D4B38FF97FFEEFA81FA827648BE5F" author="Grismer" pageId="9" pageNumber="443" refString="Grismer, L. L., Wood, P. L. Jr., Quah, E. S. H., Shahrul, A., Muin, M. A. Sumontha, M., Norhayati, A., Bauer, A. M., Wangkulangkul, S., Grismer, J. L. &amp; Pauwels, O. S. G. (2012 a) A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa, 3520, 1 - 55." type="journal article" year="2012" yearSuffix="a">
Grismer
<emphasis id="7AE8EADBFF97FFEEFF28FADF7798BE5F" box="[151,207,1376,1400]" italics="true" pageId="9" pageNumber="443">et al.</emphasis>
2012a
</bibRefCitation>
). Meristic variation is presented in
<tableCitation id="051E0372FF97FFEEFD16FAE175A8BE5F" box="[681,767,1375,1400]" captionStart="TABLE 6" captionStartId="10.[151,239,371,394]" captionTargetBox="[151,1390,424,1736]" captionTargetPageId="10" captionText="TABLE 6. Morphological characters of type series of Cyrtodactylus guakanthanensis sp. nov. / = data unavailable." httpUri="http://table.plazi.org/id/1CE36641FF94FFEDFF28FECD720FBAAD" pageId="9" pageNumber="443" tableUuid="1CE36641FF94FFEDFF28FECD720FBAAD">Table 6</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="00866542FF97FFEEFF78FA3D7408BC22" pageId="9" pageNumber="443" type="distribution">
<paragraph id="482336C9FF97FFEEFF78FA3D7308BEE7" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">
<emphasis id="7AE8EADBFF97FFEEFF78FA3D7637BEBB" bold="true" box="[199,352,1411,1436]" pageId="9" pageNumber="443">Distribution.</emphasis>
Currently,
<taxonomicName id="8F9C4D4AFF97FFEEFE5FFA3A7416BEBB" box="[480,833,1412,1436]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="443" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF97FFEEFE5FFA3A7416BEBB" box="[480,833,1412,1436]" italics="true" pageId="9" pageNumber="443">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF97FFEEFCF6FA3A74F4BEBB" box="[841,931,1412,1436]" pageId="9" pageNumber="443" rank="species">
<emphasis id="7AE8EADBFF97FFEEFCF6FA3A74F4BEBB" bold="true" box="[841,931,1412,1436]" pageId="9" pageNumber="443">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the
<typeStatus id="9727886BFF97FFEEFB04FA3B73BCBEBA" box="[1211,1259,1413,1437]" pageId="9" pageNumber="443">type</typeStatus>
locality of
<collectingCountry id="308B7659FF97FFEEFAD2FA3A72CBBEBB" box="[1389,1436,1412,1436]" name="India" pageId="9" pageNumber="443">Gua</collectingCountry>
Kanthan in the Gunung Kanthan limestone forest, Perak, Peninsular
<collectingCountry id="308B7659FF97FFEEFC29FA1974AABEE7" box="[918,1021,1447,1472]" name="Malaysia" pageId="9" pageNumber="443">Malaysia</collectingCountry>
(
<figureCitation id="D0A72A4CFF97FFEEFBB3FA197319BEE7" box="[1036,1102,1447,1472]" captionStart="FIGURE 1" captionStartId="1.[151,250,1775,1797]" captionTargetBox="[452,1112,916,1730]" captionTargetId="figure@1.[452,1136,916,1752]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the species of the Cyrotdactylus sworderi complex in Peninsular Malaysia. Cyrtodactylus quadrivirgatus was omitted owing to its ubiquitous distribution throughout Peninsular Malayisa (Grismer 2011 a)." httpUri="https://zenodo.org/record/225021/files/figure.png" pageId="9" pageNumber="443">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="482336C9FF97FFEEFF78FA727408BC22" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">
<emphasis id="7AE8EADBFF97FFEEFF78FA7276DBBEC3" bold="true" box="[199,396,1483,1508]" pageId="9" pageNumber="443">Natural history.</emphasis>
All lizards specimens were active at night on the limestone walls, adjacent boulders surrounded by limestone forest, and the limestone forest vegetation. Lizards only occurred in areas where cliff faces were eroded and exfoliated, providing cracks and holes in which to take refuge during the day (
<figureCitation id="D0A72A4CFF97FFEEFAA2F9AA7233BD0B" box="[1309,1380,1556,1581]" captionStart="FIGURE 4" captionStartId="8.[151,250,1927,1949]" captionTargetBox="[151,1436,193,1905]" captionTargetId="figure@8.[151,1436,193,1906]" captionTargetPageId="8" captionText="FIGURE 4. Habitat at the type locality, Gua Kanthan, Perak, Peninsular Malaysia. Upper: eroded and cracked limestone wall that provides refuge for Cyrtodactylus guakanthanensis sp. nov. Lower: structure of the limestone forest of the type locality." httpUri="https://zenodo.org/record/225024/files/figure.png" pageId="9" pageNumber="443">Fig. 4</figureCitation>
). No lizards were found in areas where the cliff faces were smooth and bearing no surface irregularities. Many of the specimens observed were found in the adjacent vegetation between
<quantity id="8F649B2CFF97FFEEFC36F9E274B3BD53" box="[905,996,1628,1653]" metricMagnitude="0" metricUnit="m" metricValue="2.25" metricValueMax="4.0" metricValueMin="0.5" pageId="9" pageNumber="443" unit="m" value="2.25" valueMax="4.0" valueMin="0.5">0.54 m</quantity>
above the ground on the trunks of trees or on vertical and horizontal branches and vines, indicating that the surrounding forest itself is an integral part of this new species habitat preference. One specimen was observed deep within the cave where vegetation was absent. Hatchlings as small as
<quantity id="8F649B2CFF97FFEEFE4BF976750DBDC7" box="[500,602,1736,1760]" metricMagnitude="-2" metricUnit="m" metricValue="3.38" pageId="9" pageNumber="443" unit="mm" value="33.8">33.8 mm</quantity>
SVL (LSUDPC 8175;
<figureCitation id="D0A72A4CFF97FFEEFCDAF97974F0BDC7" box="[869,935,1735,1760]" captionStart="FIGURE 3" captionStartId="7.[151,250,1953,1975]" captionTargetBox="[327,1263,193,1921]" captionTargetId="figure@7.[327,1264,193,1921]" captionTargetPageId="7" captionText="FIGURE 3. Upper: adult male holotype of Cyrtodactylus guakanthanensis sp. nov. (LSUHC 11322) from Gua Kanthan, Perak, Peninsular Malaysia. Middle: adult male paratype of C. guakanthanensis sp. nov. (LSUHC 11322). Lower: hatchling C. guakanthanensis sp. nov. (LSUDPC 8175)" httpUri="https://zenodo.org/record/225023/files/figure.png" pageId="9" pageNumber="443">Fig. 3</figureCitation>
) were observed but no gravid females were found, suggesting that the reproductive season occurred in July.
</paragraph>
</subSubSection>
<subSubSection id="00866542FF97FFECFF78F8AE739EBA45" lastPageId="11" lastPageNumber="445" pageId="9" pageNumber="443" type="etymology">
<paragraph id="482336C9FF97FFEEFF78F8AE740DBC6A" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="443">
<emphasis id="7AE8EADBFF97FFEEFF78F8AE7619BC0F" bold="true" box="[199,334,1808,1832]" pageId="9" pageNumber="443">Etymology.</emphasis>
The specific epithet
<taxonomicName id="8F9C4D4AFF97FFEEFD81F8AE75ADBC0F" box="[574,762,1808,1832]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="443" phylum="Chordata" rank="species" species="guakanthanensis">
<emphasis id="7AE8EADBFF97FFEEFD81F8AE75ADBC0F" box="[574,762,1808,1832]" italics="true" pageId="9" pageNumber="443">guakanthanensis</emphasis>
</taxonomicName>
is an adjective and refers to the
<typeStatus id="9727886BFF97FFEEFBCEF8AF73F6BC0E" box="[1137,1185,1809,1833]" pageId="9" pageNumber="443">type</typeStatus>
locality
<collectingCountry id="308B7659FF97FFEEFAB9F8AE7262BC0F" box="[1286,1333,1808,1832]" name="India" pageId="9" pageNumber="443">Gua</collectingCountry>
Kanthan (=Kanthan Cave) within the Gunung Kanthan limestone forest.
</paragraph>
<paragraph id="482336C9FF97FFEDFF78F8E97622BA67" blockId="9.[151,1436,151,2013]" lastBlockId="10.[151,1436,151,320]" lastPageId="10" lastPageNumber="444" pageId="9" pageNumber="443">
<emphasis id="7AE8EADBFF97FFEEFF78F8E9763ABC57" bold="true" box="[199,365,1879,1904]" pageId="9" pageNumber="443">Comparisons.</emphasis>
Within the
<taxonomicName id="8F9C4D4AFF97FFEEFDB2F8E6744ABC57" box="[525,797,1880,1904]" genus="Cyrotdactylus" pageId="9" pageNumber="443" rank="species" species="sworderi">
<emphasis id="7AE8EADBFF97FFEEFDB2F8E6744ABC57" box="[525,797,1880,1904]" italics="true" pageId="9" pageNumber="443">Cyrotdactylus sworderi</emphasis>
</taxonomicName>
complex,
<taxonomicName id="8F9C4D4AFF97FFEEFC19F8E673DBBC57" box="[934,1164,1880,1904]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="443" phylum="Chordata" rank="species" species="guakanthanensis">
<emphasis id="7AE8EADBFF97FFEEFC19F8E673DBBC57" box="[934,1164,1880,1904]" italics="true" pageId="9" pageNumber="443">C. guakanthanensis</emphasis>
</taxonomicName>
is separated from
<taxonomicName id="8F9C4D4AFF97FFEEFA3DF8E6766ABCB3" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="443" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="7AE8EADBFF97FFEEFA3DF8E6766ABCB3" italics="true" pageId="9" pageNumber="443">C. quadrivirgatus</emphasis>
</taxonomicName>
by having an abrupt transition between small and large postfemoral scales and a banded dorsal pattern. From
<taxonomicName id="8F9C4D4AFF97FFEEFE80F81F769FBC9F" box="[319,456,1952,1976]" genus="Cyrotdactylus" pageId="9" pageNumber="443" rank="species" species="sworderi">
<emphasis id="7AE8EADBFF97FFEEFE80F81F769FBC9F" box="[319,456,1952,1976]" italics="true" pageId="9" pageNumber="443">C. sworderi</emphasis>
</taxonomicName>
it differs in having fewer paravertebral tubercles (3234 versus 3638); lacking as opposed to having precloacal pores; having as opposed to lacking enlarged femoral scales; having wide as opposed to narrow whitish caudal bands (
<figureCitation id="D0A72A4CFF94FFEDFDBEFF297515BB97" box="[513,578,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1775,1797]" captionTargetBox="[452,1112,916,1730]" captionTargetId="figure@1.[452,1136,916,1752]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the species of the Cyrotdactylus sworderi complex in Peninsular Malaysia. Cyrtodactylus quadrivirgatus was omitted owing to its ubiquitous distribution throughout Peninsular Malayisa (Grismer 2011 a)." httpUri="https://zenodo.org/record/225021/files/figure.png" pageId="10" pageNumber="444">Fig. 1</figureCitation>
); and having a banded as opposed to spotted dorsal pattern (
<figureCitation id="D0A72A4CFF94FFEDFB5CFF297273BB97" box="[1251,1316,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1775,1797]" captionTargetBox="[452,1112,916,1730]" captionTargetId="figure@1.[452,1136,916,1752]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the species of the Cyrotdactylus sworderi complex in Peninsular Malaysia. Cyrtodactylus quadrivirgatus was omitted owing to its ubiquitous distribution throughout Peninsular Malayisa (Grismer 2011 a)." httpUri="https://zenodo.org/record/225021/files/figure.png" pageId="10" pageNumber="444">Fig. 1</figureCitation>
). It differs from
<taxonomicName id="8F9C4D4AFF94FFEDFF67FF027632BBF3" box="[216,357,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="444" phylum="Chordata" rank="species" species="tebuensis">
<emphasis id="7AE8EADBFF94FFEDFF67FF027632BBF3" box="[216,357,188,212]" italics="true" pageId="10" pageNumber="444">C. tebuensis</emphasis>
</taxonomicName>
in potentially having few suprlabials (nine or 10 versus 1113); having wide as opposed to narrow whitish caudal bands which are fewer in number (nine versus 1014;
<figureCitation id="D0A72A4CFF94FFEDFC46FF61736EBBDF" box="[1017,1081,223,248]" captionStart="FIGURE 1" captionStartId="1.[151,250,1775,1797]" captionTargetBox="[452,1112,916,1730]" captionTargetId="figure@1.[452,1136,916,1752]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the species of the Cyrotdactylus sworderi complex in Peninsular Malaysia. Cyrtodactylus quadrivirgatus was omitted owing to its ubiquitous distribution throughout Peninsular Malayisa (Grismer 2011 a)." httpUri="https://zenodo.org/record/225021/files/figure.png" pageId="10" pageNumber="444">Fig. 1</figureCitation>
); and having a banded versus a spotted or striped dorsal pattern (
<figureCitation id="D0A72A4CFF94FFEDFDB2FEBA7518BA3B" box="[525,591,260,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1775,1797]" captionTargetBox="[452,1112,916,1730]" captionTargetId="figure@1.[452,1136,916,1752]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the species of the Cyrotdactylus sworderi complex in Peninsular Malaysia. Cyrtodactylus quadrivirgatus was omitted owing to its ubiquitous distribution throughout Peninsular Malayisa (Grismer 2011 a)." httpUri="https://zenodo.org/record/225021/files/figure.png" pageId="10" pageNumber="444">Fig. 1</figureCitation>
). These differences are compared between all members of the
<taxonomicName id="8F9C4D4AFF94FFEDFAA7FEBA72CBBA3B" box="[1304,1436,260,284]" genus="Cyrotdactylus" pageId="10" pageNumber="444" rank="species" species="sworderi">
<emphasis id="7AE8EADBFF94FFEDFAA7FEBA72CBBA3B" box="[1304,1436,260,284]" italics="true" pageId="10" pageNumber="444">C. sworderi</emphasis>
</taxonomicName>
complex in
<tableCitation id="051E0372FF94FFEDFEA4FE997626BA67" box="[283,369,295,320]" captionStart="TABLE 5" captionStartId="6.[151,239,296,319]" captionTargetBox="[165,1420,389,939]" captionTargetPageId="6" captionText="TABLE 5. Diagnostic characters (in bold) separating the species of the C. sworderi complex from Cyrtodactylus guakanthanensis sp. nov." httpUri="http://table.plazi.org/id/1CE36641FF98FFE1FF28FE9676F1BA45" pageId="10" pageNumber="444" tableUuid="1CE36641FF98FFE1FF28FE9676F1BA45">Table 5</tableCitation>
.
</paragraph>
<caption id="1CE36641FF94FFEDFF28FECD720FBAAD" ID-Table-UUID="1CE36641FF94FFEDFF28FECD720FBAAD" box="[151,1368,370,395]" httpUri="http://table.plazi.org/id/1CE36641FF94FFEDFF28FECD720FBAAD" pageId="10" pageNumber="444" targetBox="[151,1390,424,1736]" targetIsTable="true" targetPageId="10">
<paragraph id="482336C9FF94FFEDFF28FECD720FBAAD" blockId="10.[151,1368,370,395]" box="[151,1368,370,395]" pageId="10" pageNumber="444">
<emphasis id="7AE8EADBFF94FFEDFF28FECD765DBAAD" bold="true" box="[151,266,371,394]" pageId="10" pageNumber="444">TABLE 6.</emphasis>
Morphological characters of type series of
<taxonomicName id="8F9C4D4AFF94FFEDFD65FECC737EBAAD" box="[730,1065,370,394]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="444" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF94FFEDFD65FECC737EBAAD" box="[730,1065,370,394]" italics="true" pageId="10" pageNumber="444">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF94FFEDFB8FFECD73D2BAAD" box="[1072,1157,371,394]" pageId="10" pageNumber="444" rank="species">
<emphasis id="7AE8EADBFF94FFEDFB8FFECD73D2BAAD" bold="true" box="[1072,1157,371,394]" pageId="10" pageNumber="444">sp. nov.</emphasis>
</taxonomicNameLabel>
/=data unavailable.
</paragraph>
</caption>
<paragraph id="482336C9FF94FFEDFDF1FE16764CBDEF" pageId="10" pageNumber="444">
<table id="3A9CC469FF940018FF28FE167239BDEF" box="[151,1390,424,1736]" gridcols="6" gridrows="30" pageId="10" pageNumber="444">
<tr id="F6AC348BFF940018FF28FE167239BA99" box="[151,1390,424,446]" gridrow="0" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FE1675F2BA99" box="[151,677,424,446]" gridcol="0" gridrow="0" pageId="10" pageNumber="444">LSUHC</th>
<th id="B57D5DF7FF940018FD63FE167463BA99" box="[732,820,424,446]" gridcol="1" gridrow="0" pageId="10" pageNumber="444">LSUHC</th>
<th id="B57D5DF7FF940018FCD4FE167495BA99" box="[875,962,424,446]" gridcol="2" gridrow="0" pageId="10" pageNumber="444">LSUHC</th>
<th id="B57D5DF7FF940018FC45FE167306BA99" box="[1018,1105,424,446]" gridcol="3" gridrow="0" pageId="10" pageNumber="444">LSUHC</th>
<th id="B57D5DF7FF940018FB36FE1673B7BA99" box="[1161,1248,424,446]" gridcol="4" gridrow="0" pageId="10" pageNumber="444">LSUHC</th>
<th id="B57D5DF7FF940018FAA8FE167239BA99" box="[1303,1390,424,446]" gridcol="5" gridrow="0" pageId="10" pageNumber="444">LSUHC</th>
</tr>
<tr id="F6AC348BFF940018FF28FE6F7239BAC0" box="[151,1390,465,487]" gridrow="1" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FE6F75F2BAC0" box="[151,677,465,487]" gridcol="0" gridrow="1" pageId="10" pageNumber="444">11321</th>
<td id="B57D5DF7FF940018FD63FE6F7463BAC0" box="[732,820,465,487]" gridcol="1" gridrow="1" pageId="10" pageNumber="444">11322</td>
<td id="B57D5DF7FF940018FCD4FE6F7495BAC0" box="[875,962,465,487]" gridcol="2" gridrow="1" pageId="10" pageNumber="444">11323</td>
<td id="B57D5DF7FF940018FC45FE6F7306BAC0" box="[1018,1105,465,487]" gridcol="3" gridrow="1" pageId="10" pageNumber="444">11325</td>
<td id="B57D5DF7FF940018FB36FE6F73B7BAC0" box="[1161,1248,465,487]" gridcol="4" gridrow="1" pageId="10" pageNumber="444">11326</td>
<td id="B57D5DF7FF940018FAA8FE6F7239BAC0" box="[1303,1390,465,487]" gridcol="5" gridrow="1" pageId="10" pageNumber="444">11339</td>
</tr>
<tr id="F6AC348BFF940018FF28FE447239B937" box="[151,1390,506,528]" gridrow="2" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FE4475F2B937" box="[151,677,506,528]" gridcol="0" gridrow="2" pageId="10" pageNumber="444">paratype</th>
<td id="B57D5DF7FF940018FD63FE447463B937" box="[732,820,506,528]" gridcol="1" gridrow="2" pageId="10" pageNumber="444">holotype</td>
<td id="B57D5DF7FF940018FCD4FE447495B937" box="[875,962,506,528]" gridcol="2" gridrow="2" pageId="10" pageNumber="444">paratype</td>
<td id="B57D5DF7FF940018FC45FE447306B937" box="[1018,1105,506,528]" gridcol="3" gridrow="2" pageId="10" pageNumber="444">paratype</td>
<td id="B57D5DF7FF940018FB36FE4473B7B937" box="[1161,1248,506,528]" gridcol="4" gridrow="2" pageId="10" pageNumber="444">paratype</td>
<td id="B57D5DF7FF940018FAA8FE447239B937" box="[1303,1390,506,528]" gridcol="5" gridrow="2" pageId="10" pageNumber="444">paratype</td>
</tr>
<tr id="F6AC348BFF940018FF28FD9A7239B91D" box="[151,1390,548,570]" gridrow="3" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FD9A75F2B91D" box="[151,677,548,570]" gridcol="0" gridrow="3" pageId="10" pageNumber="444">sex f</th>
<td id="B57D5DF7FF940018FD63FD9A7463B91D" box="[732,820,548,570]" gridcol="1" gridrow="3" pageId="10" pageNumber="444">m</td>
<td id="B57D5DF7FF940018FCD4FD9A7495B91D" box="[875,962,548,570]" gridcol="2" gridrow="3" pageId="10" pageNumber="444">m</td>
<td id="B57D5DF7FF940018FC45FD9A7306B91D" box="[1018,1105,548,570]" gridcol="3" gridrow="3" pageId="10" pageNumber="444">m</td>
<td id="B57D5DF7FF940018FB36FD9A73B7B91D" box="[1161,1248,548,570]" gridcol="4" gridrow="3" pageId="10" pageNumber="444">m</td>
<td id="B57D5DF7FF940018FAA8FD9A7239B91D" box="[1303,1390,548,570]" gridcol="5" gridrow="3" pageId="10" pageNumber="444">m</td>
</tr>
<tr id="F6AC348BFF940018FF28FDF37239B944" box="[151,1390,589,611]" gridrow="4" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FDF375F2B944" box="[151,677,589,611]" gridcol="0" gridrow="4" pageId="10" pageNumber="444">supralabials 10</th>
<td id="B57D5DF7FF940018FD63FDF37463B944" box="[732,820,589,611]" gridcol="1" gridrow="4" pageId="10" pageNumber="444">9</td>
<td id="B57D5DF7FF940018FCD4FDF37495B944" box="[875,962,589,611]" gridcol="2" gridrow="4" pageId="10" pageNumber="444">10</td>
<td id="B57D5DF7FF940018FC45FDF37306B944" box="[1018,1105,589,611]" gridcol="3" gridrow="4" pageId="10" pageNumber="444">9</td>
<td id="B57D5DF7FF940018FB36FDF373B7B944" box="[1161,1248,589,611]" gridcol="4" gridrow="4" pageId="10" pageNumber="444">10</td>
<td id="B57D5DF7FF940018FAA8FDF37239B944" box="[1303,1390,589,611]" gridcol="5" gridrow="4" pageId="10" pageNumber="444">10</td>
</tr>
<tr id="F6AC348BFF940018FF28FDC87239B9AB" box="[151,1390,630,652]" gridrow="5" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FDC875F2B9AB" box="[151,677,630,652]" gridcol="0" gridrow="5" pageId="10" pageNumber="444">infralabials 7</th>
<td id="B57D5DF7FF940018FD63FDC87463B9AB" box="[732,820,630,652]" gridcol="1" gridrow="5" pageId="10" pageNumber="444">8</td>
<td id="B57D5DF7FF940018FCD4FDC87495B9AB" box="[875,962,630,652]" gridcol="2" gridrow="5" pageId="10" pageNumber="444">8</td>
<td id="B57D5DF7FF940018FC45FDC87306B9AB" box="[1018,1105,630,652]" gridcol="3" gridrow="5" pageId="10" pageNumber="444">8</td>
<td id="B57D5DF7FF940018FB36FDC873B7B9AB" box="[1161,1248,630,652]" gridcol="4" gridrow="5" pageId="10" pageNumber="444">8</td>
<td id="B57D5DF7FF940018FAA8FDC87239B9AB" box="[1303,1390,630,652]" gridcol="5" gridrow="5" pageId="10" pageNumber="444">7</td>
</tr>
<tr id="F6AC348BFF940018FF28FD1E7239B991" box="[151,1390,672,694]" gridrow="6" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FD1E75F2B991" box="[151,677,672,694]" gridcol="0" gridrow="6" pageId="10" pageNumber="444">No. of paravertebral tubercles 34</th>
<td id="B57D5DF7FF940018FD63FD1E7463B991" box="[732,820,672,694]" gridcol="1" gridrow="6" pageId="10" pageNumber="444">32</td>
<td id="B57D5DF7FF940018FCD4FD1E7495B991" box="[875,962,672,694]" gridcol="2" gridrow="6" pageId="10" pageNumber="444">33</td>
<td id="B57D5DF7FF940018FC45FD1E7306B991" box="[1018,1105,672,694]" gridcol="3" gridrow="6" pageId="10" pageNumber="444">32</td>
<td id="B57D5DF7FF940018FB36FD1E73B7B991" box="[1161,1248,672,694]" gridcol="4" gridrow="6" pageId="10" pageNumber="444">33</td>
<td id="B57D5DF7FF940018FAA8FD1E7239B991" box="[1303,1390,672,694]" gridcol="5" gridrow="6" pageId="10" pageNumber="444">33</td>
</tr>
<tr id="F6AC348BFF940018FF28FD777239B9F8" box="[151,1390,713,735]" gridrow="7" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FD7775F2B9F8" box="[151,677,713,735]" gridcol="0" gridrow="7" pageId="10" pageNumber="444">No. of ventral scales 44</th>
<td id="B57D5DF7FF940018FD63FD777463B9F8" box="[732,820,713,735]" gridcol="1" gridrow="7" pageId="10" pageNumber="444">37</td>
<td id="B57D5DF7FF940018FCD4FD777495B9F8" box="[875,962,713,735]" gridcol="2" gridrow="7" pageId="10" pageNumber="444">42</td>
<td id="B57D5DF7FF940018FC45FD777306B9F8" box="[1018,1105,713,735]" gridcol="3" gridrow="7" pageId="10" pageNumber="444">42</td>
<td id="B57D5DF7FF940018FB36FD7773B7B9F8" box="[1161,1248,713,735]" gridcol="4" gridrow="7" pageId="10" pageNumber="444">41</td>
<td id="B57D5DF7FF940018FAA8FD777239B9F8" box="[1303,1390,713,735]" gridcol="5" gridrow="7" pageId="10" pageNumber="444">38</td>
</tr>
<tr id="F6AC348BFF940018FF28FD4C7239B82C" box="[151,1390,754,779]" gridrow="8" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FD4C75F2B82C" box="[151,677,754,779]" gridcol="0" gridrow="8" pageId="10" pageNumber="444">4th toe lamellae 21</th>
<td id="B57D5DF7FF940018FD63FD4C7463B82C" box="[732,820,754,779]" gridcol="1" gridrow="8" pageId="10" pageNumber="444">20</td>
<td id="B57D5DF7FF940018FCD4FD4C7495B82C" box="[875,962,754,779]" gridcol="2" gridrow="8" pageId="10" pageNumber="444">20</td>
<td id="B57D5DF7FF940018FC45FD4C7306B82C" box="[1018,1105,754,779]" gridcol="3" gridrow="8" pageId="10" pageNumber="444">21</td>
<td id="B57D5DF7FF940018FB36FD4C73B7B82C" box="[1161,1248,754,779]" gridcol="4" gridrow="8" pageId="10" pageNumber="444">21</td>
<td id="B57D5DF7FF940018FAA8FD4C7239B82C" box="[1303,1390,754,779]" gridcol="5" gridrow="8" pageId="10" pageNumber="444">19</td>
</tr>
<tr id="F6AC348BFF940018FF28FCA07239B813" box="[151,1390,798,820]" gridrow="9" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FCA075F2B813" box="[151,677,798,820]" gridcol="0" gridrow="9" pageId="10" pageNumber="444">Preclacal pores present (1) or absent (0) 0</th>
<td id="B57D5DF7FF940018FD63FCA07463B813" box="[732,820,798,820]" gridcol="1" gridrow="9" pageId="10" pageNumber="444">0</td>
<td id="B57D5DF7FF940018FCD4FCA07495B813" box="[875,962,798,820]" gridcol="2" gridrow="9" pageId="10" pageNumber="444">0</td>
<td id="B57D5DF7FF940018FC45FCA07306B813" box="[1018,1105,798,820]" gridcol="3" gridrow="9" pageId="10" pageNumber="444">0</td>
<td id="B57D5DF7FF940018FB36FCA073B7B813" box="[1161,1248,798,820]" gridcol="4" gridrow="9" pageId="10" pageNumber="444">0</td>
<td id="B57D5DF7FF940018FAA8FCA07239B813" box="[1303,1390,798,820]" gridcol="5" gridrow="9" pageId="10" pageNumber="444">0</td>
</tr>
<tr id="F6AC348BFF940018FF28FCF97239B87A" box="[151,1390,839,861]" gridrow="10" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FCF975F2B87A" box="[151,677,839,861]" gridcol="0" gridrow="10" pageId="10" pageNumber="444">No. of enlarged femorocloacal scales 37</th>
<td id="B57D5DF7FF940018FD63FCF97463B87A" box="[732,820,839,861]" gridcol="1" gridrow="10" pageId="10" pageNumber="444">39</td>
<td id="B57D5DF7FF940018FCD4FCF97495B87A" box="[875,962,839,861]" gridcol="2" gridrow="10" pageId="10" pageNumber="444">36</td>
<td id="B57D5DF7FF940018FC45FCF97306B87A" box="[1018,1105,839,861]" gridcol="3" gridrow="10" pageId="10" pageNumber="444">41</td>
<td id="B57D5DF7FF940018FB36FCF973B7B87A" box="[1161,1248,839,861]" gridcol="4" gridrow="10" pageId="10" pageNumber="444">39</td>
<td id="B57D5DF7FF940018FAA8FCF97239B87A" box="[1303,1390,839,861]" gridcol="5" gridrow="10" pageId="10" pageNumber="444">36</td>
</tr>
<tr id="F6AC348BFF940018FF28FCCF7239B8A0" box="[151,1390,881,903]" gridrow="11" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FCCF75F2B8A0" box="[151,677,881,903]" gridcol="0" gridrow="11" pageId="10" pageNumber="444">No. of light bands on original tail /</th>
<td id="B57D5DF7FF940018FD63FCCF7463B8A0" box="[732,820,881,903]" gridcol="1" gridrow="11" pageId="10" pageNumber="444">/</td>
<td id="B57D5DF7FF940018FCD4FCCF7495B8A0" box="[875,962,881,903]" gridcol="2" gridrow="11" pageId="10" pageNumber="444">/</td>
<td id="B57D5DF7FF940018FC45FCCF7306B8A0" box="[1018,1105,881,903]" gridcol="3" gridrow="11" pageId="10" pageNumber="444">9</td>
<td id="B57D5DF7FF940018FB36FCCF73B7B8A0" box="[1161,1248,881,903]" gridcol="4" gridrow="11" pageId="10" pageNumber="444">/</td>
<td id="B57D5DF7FF940018FAA8FCCF7239B8A0" box="[1303,1390,881,903]" gridcol="5" gridrow="11" pageId="10" pageNumber="444">/</td>
</tr>
<tr id="F6AC348BFF940018FF28FC247239B897" box="[151,1390,922,944]" gridrow="12" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FC2475F2B897" box="[151,677,922,944]" gridcol="0" gridrow="12" pageId="10" pageNumber="444">No. of body bands 5</th>
<td id="B57D5DF7FF940018FD63FC247463B897" box="[732,820,922,944]" gridcol="1" gridrow="12" pageId="10" pageNumber="444">4</td>
<td id="B57D5DF7FF940018FCD4FC247495B897" box="[875,962,922,944]" gridcol="2" gridrow="12" pageId="10" pageNumber="444">5</td>
<td id="B57D5DF7FF940018FC45FC247306B897" box="[1018,1105,922,944]" gridcol="3" gridrow="12" pageId="10" pageNumber="444">5</td>
<td id="B57D5DF7FF940018FB36FC2473B7B897" box="[1161,1248,922,944]" gridcol="4" gridrow="12" pageId="10" pageNumber="444">5</td>
<td id="B57D5DF7FF940018FAA8FC247239B897" box="[1303,1390,922,944]" gridcol="5" gridrow="12" pageId="10" pageNumber="444">5</td>
</tr>
<tr id="F6AC348BFF940018FF28FC7A7239B8FD" box="[151,1390,964,986]" gridrow="13" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FC7A75F2B8FD" box="[151,677,964,986]" gridcol="0" gridrow="13" pageId="10" pageNumber="444">SVL 82.2</th>
<td id="B57D5DF7FF940018FD63FC7A7463B8FD" box="[732,820,964,986]" gridcol="1" gridrow="13" pageId="10" pageNumber="444">79.7</td>
<td id="B57D5DF7FF940018FCD4FC7A7495B8FD" box="[875,962,964,986]" gridcol="2" gridrow="13" pageId="10" pageNumber="444">79.8</td>
<td id="B57D5DF7FF940018FC45FC7A7306B8FD" box="[1018,1105,964,986]" gridcol="3" gridrow="13" pageId="10" pageNumber="444">78.2</td>
<td id="B57D5DF7FF940018FB36FC7A73B7B8FD" box="[1161,1248,964,986]" gridcol="4" gridrow="13" pageId="10" pageNumber="444">77.7</td>
<td id="B57D5DF7FF940018FAA8FC7A7239B8FD" box="[1303,1390,964,986]" gridcol="5" gridrow="13" pageId="10" pageNumber="444">81.8</td>
</tr>
<tr id="F6AC348BFF940018FF28FC537239BF24" box="[151,1390,1005,1027]" gridrow="14" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FC5375F2BF24" box="[151,677,1005,1027]" gridcol="0" gridrow="14" pageId="10" pageNumber="444">TL 91.5</th>
<td id="B57D5DF7FF940018FD63FC537463BF24" box="[732,820,1005,1027]" gridcol="1" gridrow="14" pageId="10" pageNumber="444">78.3</td>
<td id="B57D5DF7FF940018FCD4FC537495BF24" box="[875,962,1005,1027]" gridcol="2" gridrow="14" pageId="10" pageNumber="444">83.4</td>
<td id="B57D5DF7FF940018FC45FC537306BF24" box="[1018,1105,1005,1027]" gridcol="3" gridrow="14" pageId="10" pageNumber="444">97.9</td>
<td id="B57D5DF7FF940018FB36FC5373B7BF24" box="[1161,1248,1005,1027]" gridcol="4" gridrow="14" pageId="10" pageNumber="444">85.7</td>
<td id="B57D5DF7FF940018FAA8FC537239BF24" box="[1303,1390,1005,1027]" gridcol="5" gridrow="14" pageId="10" pageNumber="444">89.2</td>
</tr>
<tr id="F6AC348BFF940018FF28FBA87239BF0B" box="[151,1390,1046,1068]" gridrow="15" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FBA875F2BF0B" box="[151,677,1046,1068]" gridcol="0" gridrow="15" pageId="10" pageNumber="444">TW 7</th>
<td id="B57D5DF7FF940018FD63FBA87463BF0B" box="[732,820,1046,1068]" gridcol="1" gridrow="15" pageId="10" pageNumber="444">7.4</td>
<td id="B57D5DF7FF940018FCD4FBA87495BF0B" box="[875,962,1046,1068]" gridcol="2" gridrow="15" pageId="10" pageNumber="444">7.3</td>
<td id="B57D5DF7FF940018FC45FBA87306BF0B" box="[1018,1105,1046,1068]" gridcol="3" gridrow="15" pageId="10" pageNumber="444">7.1</td>
<td id="B57D5DF7FF940018FB36FBA873B7BF0B" box="[1161,1248,1046,1068]" gridcol="4" gridrow="15" pageId="10" pageNumber="444">7.2</td>
<td id="B57D5DF7FF940018FAA8FBA87239BF0B" box="[1303,1390,1046,1068]" gridcol="5" gridrow="15" pageId="10" pageNumber="444">6.8</td>
</tr>
<tr id="F6AC348BFF940018FF28FB817239BF72" box="[151,1390,1087,1109]" gridrow="16" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FB8175F2BF72" box="[151,677,1087,1109]" gridcol="0" gridrow="16" pageId="10" pageNumber="444">FL 12.9</th>
<td id="B57D5DF7FF940018FD63FB817463BF72" box="[732,820,1087,1109]" gridcol="1" gridrow="16" pageId="10" pageNumber="444">13.3</td>
<td id="B57D5DF7FF940018FCD4FB817495BF72" box="[875,962,1087,1109]" gridcol="2" gridrow="16" pageId="10" pageNumber="444">13</td>
<td id="B57D5DF7FF940018FC45FB817306BF72" box="[1018,1105,1087,1109]" gridcol="3" gridrow="16" pageId="10" pageNumber="444">13.2</td>
<td id="B57D5DF7FF940018FB36FB8173B7BF72" box="[1161,1248,1087,1109]" gridcol="4" gridrow="16" pageId="10" pageNumber="444">12.6</td>
<td id="B57D5DF7FF940018FAA8FB817239BF72" box="[1303,1390,1087,1109]" gridcol="5" gridrow="16" pageId="10" pageNumber="444">12.7</td>
</tr>
<tr id="F6AC348BFF940018FF28FBD77239BF58" box="[151,1390,1129,1151]" gridrow="17" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FBD775F2BF58" box="[151,677,1129,1151]" gridcol="0" gridrow="17" pageId="10" pageNumber="444">TBL 14.7</th>
<td id="B57D5DF7FF940018FD63FBD77463BF58" box="[732,820,1129,1151]" gridcol="1" gridrow="17" pageId="10" pageNumber="444">14.7</td>
<td id="B57D5DF7FF940018FCD4FBD77495BF58" box="[875,962,1129,1151]" gridcol="2" gridrow="17" pageId="10" pageNumber="444">15.5</td>
<td id="B57D5DF7FF940018FC45FBD77306BF58" box="[1018,1105,1129,1151]" gridcol="3" gridrow="17" pageId="10" pageNumber="444">15.4</td>
<td id="B57D5DF7FF940018FB36FBD773B7BF58" box="[1161,1248,1129,1151]" gridcol="4" gridrow="17" pageId="10" pageNumber="444">14.5</td>
<td id="B57D5DF7FF940018FAA8FBD77239BF58" box="[1303,1390,1129,1151]" gridcol="5" gridrow="17" pageId="10" pageNumber="444">16</td>
</tr>
<tr id="F6AC348BFF940018FF28FB2C7239BF8F" box="[151,1390,1170,1192]" gridrow="18" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FB2C75F2BF8F" box="[151,677,1170,1192]" gridcol="0" gridrow="18" pageId="10" pageNumber="444">AG 36.2</th>
<td id="B57D5DF7FF940018FD63FB2C7463BF8F" box="[732,820,1170,1192]" gridcol="1" gridrow="18" pageId="10" pageNumber="444">33.3</td>
<td id="B57D5DF7FF940018FCD4FB2C7495BF8F" box="[875,962,1170,1192]" gridcol="2" gridrow="18" pageId="10" pageNumber="444">34.1</td>
<td id="B57D5DF7FF940018FC45FB2C7306BF8F" box="[1018,1105,1170,1192]" gridcol="3" gridrow="18" pageId="10" pageNumber="444">35</td>
<td id="B57D5DF7FF940018FB36FB2C73B7BF8F" box="[1161,1248,1170,1192]" gridcol="4" gridrow="18" pageId="10" pageNumber="444">34.5</td>
<td id="B57D5DF7FF940018FAA8FB2C7239BF8F" box="[1303,1390,1170,1192]" gridcol="5" gridrow="18" pageId="10" pageNumber="444">37.8</td>
</tr>
<tr id="F6AC348BFF940018FF28FB027239BFF5" box="[151,1390,1212,1234]" gridrow="19" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FB0275F2BFF5" box="[151,677,1212,1234]" gridcol="0" gridrow="19" pageId="10" pageNumber="444">HL 23</th>
<td id="B57D5DF7FF940018FD63FB027463BFF5" box="[732,820,1212,1234]" gridcol="1" gridrow="19" pageId="10" pageNumber="444">22.3</td>
<td id="B57D5DF7FF940018FCD4FB027495BFF5" box="[875,962,1212,1234]" gridcol="2" gridrow="19" pageId="10" pageNumber="444">22.8</td>
<td id="B57D5DF7FF940018FC45FB027306BFF5" box="[1018,1105,1212,1234]" gridcol="3" gridrow="19" pageId="10" pageNumber="444">22</td>
<td id="B57D5DF7FF940018FB36FB0273B7BFF5" box="[1161,1248,1212,1234]" gridcol="4" gridrow="19" pageId="10" pageNumber="444">21.3</td>
<td id="B57D5DF7FF940018FAA8FB027239BFF5" box="[1303,1390,1212,1234]" gridcol="5" gridrow="19" pageId="10" pageNumber="444">23</td>
</tr>
<tr id="F6AC348BFF940018FF28FB5B7239BFDC" box="[151,1390,1253,1275]" gridrow="20" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FB5B75F2BFDC" box="[151,677,1253,1275]" gridcol="0" gridrow="20" pageId="10" pageNumber="444">HW 15.1</th>
<td id="B57D5DF7FF940018FD63FB5B7463BFDC" box="[732,820,1253,1275]" gridcol="1" gridrow="20" pageId="10" pageNumber="444">14.5</td>
<td id="B57D5DF7FF940018FCD4FB5B7495BFDC" box="[875,962,1253,1275]" gridcol="2" gridrow="20" pageId="10" pageNumber="444">15.1</td>
<td id="B57D5DF7FF940018FC45FB5B7306BFDC" box="[1018,1105,1253,1275]" gridcol="3" gridrow="20" pageId="10" pageNumber="444">15</td>
<td id="B57D5DF7FF940018FB36FB5B73B7BFDC" box="[1161,1248,1253,1275]" gridcol="4" gridrow="20" pageId="10" pageNumber="444">13.3</td>
<td id="B57D5DF7FF940018FAA8FB5B7239BFDC" box="[1303,1390,1253,1275]" gridcol="5" gridrow="20" pageId="10" pageNumber="444">15.1</td>
</tr>
<tr id="F6AC348BFF940018FF28FAB07239BE03" box="[151,1390,1294,1316]" gridrow="21" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FAB075F2BE03" box="[151,677,1294,1316]" gridcol="0" gridrow="21" pageId="10" pageNumber="444">HD 9.5</th>
<td id="B57D5DF7FF940018FD63FAB07463BE03" box="[732,820,1294,1316]" gridcol="1" gridrow="21" pageId="10" pageNumber="444">9</td>
<td id="B57D5DF7FF940018FCD4FAB07495BE03" box="[875,962,1294,1316]" gridcol="2" gridrow="21" pageId="10" pageNumber="444">9.6</td>
<td id="B57D5DF7FF940018FC45FAB07306BE03" box="[1018,1105,1294,1316]" gridcol="3" gridrow="21" pageId="10" pageNumber="444">9.5</td>
<td id="B57D5DF7FF940018FB36FAB073B7BE03" box="[1161,1248,1294,1316]" gridcol="4" gridrow="21" pageId="10" pageNumber="444">8.3</td>
<td id="B57D5DF7FF940018FAA8FAB07239BE03" box="[1303,1390,1294,1316]" gridcol="5" gridrow="21" pageId="10" pageNumber="444">9.7</td>
</tr>
<tr id="F6AC348BFF940018FF28FA897239BE6A" box="[151,1390,1335,1357]" gridrow="22" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FA8975F2BE6A" box="[151,677,1335,1357]" gridcol="0" gridrow="22" pageId="10" pageNumber="444">DE 5.5</th>
<td id="B57D5DF7FF940018FD63FA897463BE6A" box="[732,820,1335,1357]" gridcol="1" gridrow="22" pageId="10" pageNumber="444">5</td>
<td id="B57D5DF7FF940018FCD4FA897495BE6A" box="[875,962,1335,1357]" gridcol="2" gridrow="22" pageId="10" pageNumber="444">5.5</td>
<td id="B57D5DF7FF940018FC45FA897306BE6A" box="[1018,1105,1335,1357]" gridcol="3" gridrow="22" pageId="10" pageNumber="444">5.2</td>
<td id="B57D5DF7FF940018FB36FA8973B7BE6A" box="[1161,1248,1335,1357]" gridcol="4" gridrow="22" pageId="10" pageNumber="444">4.7</td>
<td id="B57D5DF7FF940018FAA8FA897239BE6A" box="[1303,1390,1335,1357]" gridcol="5" gridrow="22" pageId="10" pageNumber="444">5.6</td>
</tr>
<tr id="F6AC348BFF940018FF28FADF7239BE50" box="[151,1390,1377,1399]" gridrow="23" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FADF75F2BE50" box="[151,677,1377,1399]" gridcol="0" gridrow="23" pageId="10" pageNumber="444">EE 6.5</th>
<td id="B57D5DF7FF940018FD63FADF7463BE50" box="[732,820,1377,1399]" gridcol="1" gridrow="23" pageId="10" pageNumber="444">6.3</td>
<td id="B57D5DF7FF940018FCD4FADF7495BE50" box="[875,962,1377,1399]" gridcol="2" gridrow="23" pageId="10" pageNumber="444">6.2</td>
<td id="B57D5DF7FF940018FC45FADF7306BE50" box="[1018,1105,1377,1399]" gridcol="3" gridrow="23" pageId="10" pageNumber="444">6.3</td>
<td id="B57D5DF7FF940018FB36FADF73B7BE50" box="[1161,1248,1377,1399]" gridcol="4" gridrow="23" pageId="10" pageNumber="444">5.8</td>
<td id="B57D5DF7FF940018FAA8FADF7239BE50" box="[1303,1390,1377,1399]" gridcol="5" gridrow="23" pageId="10" pageNumber="444">6.5</td>
</tr>
<tr id="F6AC348BFF940018FF28FA347239BE87" box="[151,1390,1418,1440]" gridrow="24" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FA3475F2BE87" box="[151,677,1418,1440]" gridcol="0" gridrow="24" pageId="10" pageNumber="444">ES 9.9</th>
<td id="B57D5DF7FF940018FD63FA347463BE87" box="[732,820,1418,1440]" gridcol="1" gridrow="24" pageId="10" pageNumber="444">9.8</td>
<td id="B57D5DF7FF940018FCD4FA347495BE87" box="[875,962,1418,1440]" gridcol="2" gridrow="24" pageId="10" pageNumber="444">10</td>
<td id="B57D5DF7FF940018FC45FA347306BE87" box="[1018,1105,1418,1440]" gridcol="3" gridrow="24" pageId="10" pageNumber="444">10.1</td>
<td id="B57D5DF7FF940018FB36FA3473B7BE87" box="[1161,1248,1418,1440]" gridcol="4" gridrow="24" pageId="10" pageNumber="444">10.1</td>
<td id="B57D5DF7FF940018FAA8FA347239BE87" box="[1303,1390,1418,1440]" gridcol="5" gridrow="24" pageId="10" pageNumber="444">10.2</td>
</tr>
<tr id="F6AC348BFF940018FF28FA0A7239BEED" box="[151,1390,1460,1482]" gridrow="25" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FA0A75F2BEED" box="[151,677,1460,1482]" gridcol="0" gridrow="25" pageId="10" pageNumber="444">EN 7.8</th>
<td id="B57D5DF7FF940018FD63FA0A7463BEED" box="[732,820,1460,1482]" gridcol="1" gridrow="25" pageId="10" pageNumber="444">7.5</td>
<td id="B57D5DF7FF940018FCD4FA0A7495BEED" box="[875,962,1460,1482]" gridcol="2" gridrow="25" pageId="10" pageNumber="444">7.6</td>
<td id="B57D5DF7FF940018FC45FA0A7306BEED" box="[1018,1105,1460,1482]" gridcol="3" gridrow="25" pageId="10" pageNumber="444">7.6</td>
<td id="B57D5DF7FF940018FB36FA0A73B7BEED" box="[1161,1248,1460,1482]" gridcol="4" gridrow="25" pageId="10" pageNumber="444">7.5</td>
<td id="B57D5DF7FF940018FAA8FA0A7239BEED" box="[1303,1390,1460,1482]" gridcol="5" gridrow="25" pageId="10" pageNumber="444">7.8</td>
</tr>
<tr id="F6AC348BFF940018FF28FA637239BED4" box="[151,1390,1501,1523]" gridrow="26" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28FA6375F2BED4" box="[151,677,1501,1523]" gridcol="0" gridrow="26" pageId="10" pageNumber="444">IO 4.2</th>
<td id="B57D5DF7FF940018FD63FA637463BED4" box="[732,820,1501,1523]" gridcol="1" gridrow="26" pageId="10" pageNumber="444">3.9</td>
<td id="B57D5DF7FF940018FCD4FA637495BED4" box="[875,962,1501,1523]" gridcol="2" gridrow="26" pageId="10" pageNumber="444">4.2</td>
<td id="B57D5DF7FF940018FC45FA637306BED4" box="[1018,1105,1501,1523]" gridcol="3" gridrow="26" pageId="10" pageNumber="444">3.8</td>
<td id="B57D5DF7FF940018FB36FA6373B7BED4" box="[1161,1248,1501,1523]" gridcol="4" gridrow="26" pageId="10" pageNumber="444">3.8</td>
<td id="B57D5DF7FF940018FAA8FA637239BED4" box="[1303,1390,1501,1523]" gridcol="5" gridrow="26" pageId="10" pageNumber="444">4</td>
</tr>
<tr id="F6AC348BFF940018FF28F9B87239BD3B" box="[151,1390,1542,1564]" gridrow="27" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28F9B875F2BD3B" box="[151,677,1542,1564]" gridcol="0" gridrow="27" pageId="10" pageNumber="444">EL 1.7</th>
<td id="B57D5DF7FF940018FD63F9B87463BD3B" box="[732,820,1542,1564]" gridcol="1" gridrow="27" pageId="10" pageNumber="444">1.9</td>
<td id="B57D5DF7FF940018FCD4F9B87495BD3B" box="[875,962,1542,1564]" gridcol="2" gridrow="27" pageId="10" pageNumber="444">1.7</td>
<td id="B57D5DF7FF940018FC45F9B87306BD3B" box="[1018,1105,1542,1564]" gridcol="3" gridrow="27" pageId="10" pageNumber="444">2.3</td>
<td id="B57D5DF7FF940018FB36F9B873B7BD3B" box="[1161,1248,1542,1564]" gridcol="4" gridrow="27" pageId="10" pageNumber="444">1.5</td>
<td id="B57D5DF7FF940018FAA8F9B87239BD3B" box="[1303,1390,1542,1564]" gridcol="5" gridrow="27" pageId="10" pageNumber="444">1.9</td>
</tr>
<tr id="F6AC348BFF940018FF28F9917239BD62" box="[151,1390,1583,1605]" gridrow="28" pageId="10" pageNumber="444">
<th id="B57D5DF7FF940018FF28F99175F2BD62" box="[151,677,1583,1605]" gridcol="0" gridrow="28" pageId="10" pageNumber="444">IN 2.5</th>
<td id="B57D5DF7FF940018FD63F9917463BD62" box="[732,820,1583,1605]" gridcol="1" gridrow="28" pageId="10" pageNumber="444">2.6</td>
<td id="B57D5DF7FF940018FCD4F9917495BD62" box="[875,962,1583,1605]" gridcol="2" gridrow="28" pageId="10" pageNumber="444">2.5</td>
<td id="B57D5DF7FF940018FC45F9917306BD62" box="[1018,1105,1583,1605]" gridcol="3" gridrow="28" pageId="10" pageNumber="444">2.4</td>
<td id="B57D5DF7FF940018FB36F99173B7BD62" box="[1161,1248,1583,1605]" gridcol="4" gridrow="28" pageId="10" pageNumber="444">2.4</td>
<td id="B57D5DF7FF940018FAA8F9917239BD62" box="[1303,1390,1583,1605]" gridcol="5" gridrow="28" pageId="10" pageNumber="444">2.4</td>
</tr>
<tr id="F6AC348BFF940018FF28F9107239BDEF" box="[151,1390,1710,1736]" gridrow="29" pageId="10" pageNumber="444" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="B57D5DF7FF940018FF28F91075F2BDEF" box="[151,677,1710,1736]" gridcol="0" gridrow="29" pageId="10" pageNumber="444">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="482336C9FF94FFEDFF28F94B75CABCC1" blockId="10.[151,1436,1781,2022]" pageId="10" pageNumber="444">
The Gunung Kanthan limestone forest surrounds 400 million year old limestone formations that harbor a number of unique plants and animals. Owing to the purity of its limestone, the area is scheduled to be quarried. Our data indicate that
<taxonomicName id="8F9C4D4AFF94FFEDFE96F88375DDBC72" box="[297,650,1853,1877]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="444" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF94FFEDFE96F88375DDBC72" box="[297,650,1853,1877]" italics="true" pageId="10" pageNumber="444">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF94FFEDFD2DF88375BABC72" box="[658,749,1853,1877]" pageId="10" pageNumber="444" rank="species">
<emphasis id="7AE8EADBFF94FFEDFD2DF88375BABC72" bold="true" box="[658,749,1853,1877]" pageId="10" pageNumber="444">sp. nov.</emphasis>
</taxonomicNameLabel>
does not range continuously throughout the limestone forest but appears to be restricted to areas that have an eroded complex limestone microhabitat. One of the quarry zones targeted is
<collectingCountry id="308B7659FF94FFEDFEAEF8387617BCB9" box="[273,320,1926,1950]" name="India" pageId="10" pageNumber="444">Gua</collectingCountry>
Kanthan, the
<typeStatus id="9727886BFF94FFEDFE62F838755ABCB9" box="[477,525,1926,1950]" pageId="10" pageNumber="444">type</typeStatus>
locality of
<taxonomicName id="8F9C4D4AFF94FFEDFD31F838743DBCBA" box="[654,874,1925,1949]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="444" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF94FFEDFD31F838743DBCBA" box="[654,874,1925,1949]" italics="true" pageId="10" pageNumber="444">C. guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF94FFEDFCCDF83B749CBCBA" box="[882,971,1925,1949]" pageId="10" pageNumber="444" rank="species">
<emphasis id="7AE8EADBFF94FFEDFCCDF83B749CBCBA" bold="true" box="[882,971,1925,1949]" pageId="10" pageNumber="444">sp. nov.</emphasis>
</taxonomicNameLabel>
Grinding down the limestone in this area and cutting the surrounding forest will destroy the microhabitat to which this new species has adapted and our data indicate this species may not occur elsewhere.
</paragraph>
<paragraph id="482336C9FF95FFECFF78FF29739EBA45" blockId="11.[151,1436,151,354]" pageId="11" pageNumber="445">
The discovery of
<taxonomicName id="8F9C4D4AFF95FFECFE25FF2675A8BB97" box="[410,767,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="445" phylum="Chordata" rank="species" species="guakanthanensis" status="sp. nov.">
<emphasis id="7AE8EADBFF95FFECFE25FF2675A8BB97" box="[410,767,152,176]" italics="true" pageId="11" pageNumber="445">Cyrtodactylus guakanthanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DB57A0FF95FFECFCB3FF26743DBB97" box="[780,874,152,176]" pageId="11" pageNumber="445" rank="species">
<emphasis id="7AE8EADBFF95FFECFCB3FF26743DBB97" bold="true" box="[780,874,152,176]" pageId="11" pageNumber="445">sp. nov.</emphasis>
</taxonomicNameLabel>
adds to a growing body of evidence that karst regions should be protected and better studied. If reptiles are an indication of the hidden diversity within these unique habitats, then limestone forests may be some of the most biotically rich habitats in Peninsular
<collectingCountry id="308B7659FF95FFECFB42FF607233BBD0" box="[1277,1380,222,247]" name="Malaysia" pageId="11" pageNumber="445">Malaysia</collectingCountry>
with a level of herpetological endemism approaching that of Malaysias islands (see Chan
<emphasis id="7AE8EADBFF95FFECFBD3FEBD73F1BA3D" box="[1132,1190,258,282]" italics="true" pageId="11" pageNumber="445">et al.</emphasis>
2010; Grismer 2008, 2011b;
<bibRefCitation id="2C0D4B38FF95FFECFF52FE9B7684BA19" author="Grismer" box="[237,467,293,318]" pageId="11" pageNumber="445" refString="Grismer, L. L., Grismer, J. L., Wood, P. L. Jr., Ngo, V. T. &amp; Chan, K. O. (2011) Herpetology on the fringes of the Sunda Shelf: a discussion of discovery, taxonomy, and biogeography. Bonner Zoologische Monographien, 57, 57 - 97." type="journal article" year="2011">
Grismer
<emphasis id="7AE8EADBFF95FFECFEECFE9976D9BA19" box="[339,398,294,318]" italics="true" pageId="11" pageNumber="445">et al.</emphasis>
2011
</bibRefCitation>
). Terminating this species before it is discovered, described, and studied is not only illogical, it is tantamount to discarding a wrapped gift before it is opened and its value assessed.
</paragraph>
</subSubSection>
</treatment>
</document>