464 lines
57 KiB
XML
464 lines
57 KiB
XML
<document id="4E0095EB45DB0FD0281AB802CA174000" ID-DOI="10.11646/zootaxa.3980.4.6" ID-GBIF-Dataset="2596acca-ff76-4a22-b3c2-7d4e6b3f7e7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="232824" ID-ZooBank="C13F5A20-2E38-44BC-BC3E-4FDB6ECBB806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461244349238" checkinUser="plazi" docAuthor="Missassi, Alexandre F. R. & Prudente, Ana L. C." docDate="2015" docId="C01087BF5248BE24F397F8E4AE718B0F" docLanguage="en" docName="zt03980p574.pdf" docOrigin="Zootaxa 3980 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Imantodes guane Missassi & Prudente, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="571" masterDocId="3C29FFC75249BE2DF300FFD5AC318A6A" masterDocTitle="A new species of Imantodes Duméril, 1853 (Serpentes, Dipsadidae) from the Eastern Cordillera of Colombia" masterLastPageNumber="574" masterPageNumber="562" pageNumber="563" updateTime="1698605933454" updateUser="plazi">
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<mods:mods id="F950755CCD189C4CE479D27ECE933E0F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B7163863C29BBBE56F8B13B03C3D82E4">
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<mods:title id="58F73D14091288F5C4FE5436F1D8A542">A new species of Imantodes Duméril, 1853 (Serpentes, Dipsadidae) from the Eastern Cordillera of Colombia</mods:title>
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<mods:name id="AD003704C5841B58FB31F11FEEDB1ABB" type="personal">
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<mods:roleTerm id="D54EFE8F97A0B0A238EBFD00576C313E">Author</mods:roleTerm>
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<mods:namePart id="9773AE4879168980998044ABF4527F10">Missassi, Alexandre F. R.</mods:namePart>
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<mods:name id="1AF8608222356F7E9A8A8AC5AFC82B84" type="personal">
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<mods:roleTerm id="AA8F9BA875C2FFDCDE0B7951DD328081">Author</mods:roleTerm>
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<mods:namePart id="7844861C89D1CC8BFED27FF7AB93610E">Prudente, Ana L. C.</mods:namePart>
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<mods:typeOfResource id="A668EF924C11E0561903ECD56FB46343">text</mods:typeOfResource>
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<mods:title id="9859834F90777F7FC2817EC4757E0B5E">Zootaxa</mods:title>
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<mods:part id="E1A76CC9489130B004BA450F289F1A9B">
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<mods:date id="A71517E52D6FBCB24570A6197E929B4E">2015</mods:date>
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<mods:number id="8DEF70E9D09449FF52C69D11FCDC7BB3">3980</mods:number>
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<mods:number id="FCC88DCA733509665B662040585D0A24">4</mods:number>
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<mods:classification id="EC76B2DF12CACB3BEC778FC46D90ABC8">journal article</mods:classification>
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<mods:identifier id="1A03507CA7F7ECEFED76BEAA2CA18DD8" type="DOI">10.11646/zootaxa.3980.4.6</mods:identifier>
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<mods:identifier id="AA2F72412A6EA17E4235ED0080BC4A02" type="GBIF-Dataset">2596acca-ff76-4a22-b3c2-7d4e6b3f7e7e</mods:identifier>
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<mods:identifier id="DBA533B224E87B5EE6A4D8428FB0E7B3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="C01087BF5248BE24F397F8E4AE718B0F" ID-DOI="http://doi.org/10.5281/zenodo.5661756" ID-GBIF-Taxon="119634441" ID-Zenodo-Dep="5661756" LSID="urn:lsid:plazi:treatment:C01087BF5248BE24F397F8E4AE718B0F" httpUri="http://treatment.plazi.org/id/C01087BF5248BE24F397F8E4AE718B0F" lastPageId="9" lastPageNumber="571" pageId="1" pageNumber="563">
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<subSubSection id="00A365225248BE2CF397F8E4AD208D06" pageId="1" pageNumber="563" type="nomenclature">
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<paragraph id="480636A95248BE2CF397F8E4ADFC8D21" blockId="1.[151,461,1841,1901]" box="[151,461,1841,1867]" pageId="1" pageNumber="563">
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<heading id="134E81C55248BE2CF397F8E4ADFC8D21" bold="true" box="[151,461,1841,1867]" fontSize="11" level="1" pageId="1" pageNumber="563" reason="1">
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<emphasis id="7ACDEABB5248BE2CF397F8E4ADFC8D21" bold="true" box="[151,461,1841,1867]" pageId="1" pageNumber="563">
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<taxonomicName id="8FB94D2A5248BE2CF397F8E4AD598D21" box="[151,360,1841,1867]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="563" phylum="Chordata" rank="species" species="guane" status="sp. nov.">
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<emphasis id="7ACDEABB5248BE2CF397F8E4AD598D21" bold="true" box="[151,360,1841,1867]" italics="true" pageId="1" pageNumber="563">Imantodes guane</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="61FE57C05248BE2CF26FF8E4ADFC8D21" box="[367,461,1841,1867]" pageId="1" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="480636A95248BE2CF397F881AD208D06" blockId="1.[151,461,1841,1901]" box="[151,273,1876,1901]" pageId="1" pageNumber="563">
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||
(
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<figureCitation id="D0822A2C5248BE2CF39FF881AD388D06" box="[159,265,1876,1901]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="5.[151,249,1955,1977]" captionStartId-1="6.[151,249,1954,1976]" captionTargetBox-0="[193,1389,216,1928]" captionTargetBox-1="[232,1370,199,1929]" captionTargetId-0="figure@5.[172,1414,193,1933]" captionTargetId-1="figure@6.[196,1391,193,1933]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 2. Dorsal (A) and ventral views (B) of the holotype of Imantodes guane (ICN 5730) from Charalá, Departament of Santander, Colombia. Scale bar = 10 mm. Photo by D. R. Padilla-Rivera." captionText-1="FIGURE 3. Dorsal (A), ventral (B), and lateral views (C) of head of the holotype of Imantodes guane (ICN 5730) from Charalá, departament of Santander, Colombia. Scale bar = 5 mm." httpUri-0="https://zenodo.org/record/232826/files/figure.png" httpUri-1="https://zenodo.org/record/232827/files/figure.png" pageId="1" pageNumber="563">Figs. 2–3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="00A365225248BE2EF397F849A8BC8ABF" lastPageId="3" lastPageNumber="565" pageId="1" pageNumber="563" type="materials_examined">
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<paragraph id="480636A95248BE2CF397F849A9338DB3" blockId="1.[151,1436,1948,2009]" pageId="1" pageNumber="563">
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<emphasis id="7ACDEABB5248BE2CF397F849AD3A8DDF" bold="true" box="[151,267,1948,1973]" pageId="1" pageNumber="563">
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<typeStatus id="9702880B5248BE2CF397F849AD378DDF" box="[151,262,1948,1973]" pageId="1" pageNumber="563" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult male,
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<collectionCode id="2EA8AE6C5248BE2CF2A4F849ADE58DDE" box="[420,468,1948,1972]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="1" pageNumber="563">ICN</collectionCode>
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5730, collected by R. Hernández in 1981 at headwater of the Luisito River, Virolín (
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<geoCoordinate id="2D8D506E5248BE2CF3A0F815AD2A8DB3" box="[160,283,1984,2009]" direction="north" orientation="latitude" pageId="1" pageNumber="563" precision="15" value="6.3050003">6º18’18”N</geoCoordinate>
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,
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<geoCoordinate id="2D8D506E5248BE2CF227F815AD848DB3" box="[295,437,1984,2009]" direction="west" orientation="longitude" pageId="1" pageNumber="563" precision="15" value="-73.17361">73º10’25”W</geoCoordinate>
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;
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<quantity id="8F419B4C5248BE2CF2C1F815AE278DB2" box="[449,534,1984,2009]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="563" unit="m" value="1750.0">1750 m</quantity>
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asl), municipality of Charalá, department of Santander,
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<collectingCountry id="30AE76395248BE2CF78CF815A8CF8DB3" box="[1164,1278,1984,2009]" name="Colombia" pageId="1" pageNumber="563">Colombia</collectingCountry>
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.
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</paragraph>
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<caption id="1CC66621524BBE2FF37DFD93A89788DB" ID-Table-UUID="1CC66621524BBE2FF37DFD93A89788DB" httpUri="http://table.plazi.org/id/1CC66621524BBE2FF37DFD93A89788DB" pageId="2" pageNumber="564" targetBox="[125,1813,725,1612]" targetIsTable="true" targetPageId="2">
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<paragraph id="480636A9524BBE2FF37DFD93A89788DB" blockId="2.[125,1827,582,689]" pageId="2" pageNumber="564">
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<emphasis id="7ACDEABB524BBE2FF37DFD93ACD88836" bold="true" box="[125,233,582,604]" pageId="2" pageNumber="564">TABLE 1.</emphasis>
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Selected qualitative and quantitative morphological characters for the genus
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<taxonomicName id="8FB94D2A524BBE2FF0DEFD93A8758831" box="[990,1092,582,603]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="genus">
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<emphasis id="7ACDEABB524BBE2FF0DEFD93A8758831" box="[990,1092,582,603]" italics="true" pageId="2" pageNumber="564">Imantodes</emphasis>
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</taxonomicName>
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. Numbers in parenthesis represent the number of specimens examined and additional data was obtained or augmented from the following literature: for
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<taxonomicName id="8FB94D2A524BBE2FF06FFDB6AFDF881D" box="[879,1006,610,632]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="chocoensis">
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<emphasis id="7ACDEABB524BBE2FF06FFDB6AFDF881D" box="[879,1006,610,632]" italics="true" pageId="2" pageNumber="564">I. chocoensis</emphasis>
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</taxonomicName>
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data from Torres-Carvajal
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<emphasis id="7ACDEABB524BBE2FF7FBFDB6A91C881D" box="[1275,1325,610,632]" italics="true" pageId="2" pageNumber="564">et al.</emphasis>
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(2012) and Jaramillo-Martinéz
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<emphasis id="7ACDEABB524BBE2FF566FDB6AAA9881D" box="[1638,1688,610,632]" italics="true" pageId="2" pageNumber="564">et al.</emphasis>
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(2013); for
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<taxonomicName id="8FB94D2A524BBE2FF40FFDB6ACFE88FE" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="cenchoa">
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<emphasis id="7ACDEABB524BBE2FF40FFDB6ACFE88FE" italics="true" pageId="2" pageNumber="564">I. cenchoa</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="8FB94D2A524BBE2FF3DBFDAAAD7C88FE" box="[219,333,639,660]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="inornatus">
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<emphasis id="7ACDEABB524BBE2FF3DBFDAAAD7C88FE" box="[219,333,639,660]" italics="true" pageId="2" pageNumber="564">I. inornatus</emphasis>
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</taxonomicName>
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,
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<emphasis id="7ACDEABB524BBE2FF25AFDAAADE088FE" box="[346,465,639,660]" italics="true" pageId="2" pageNumber="564">
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<taxonomicName id="8FB94D2A524BBE2FF25AFDAAADFC88FE" box="[346,461,639,660]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="lentiferus">I. lentiferus</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName id="8FB94D2A524BBE2FF100FDAAAEB088FE" box="[512,641,639,660]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="phantasma">
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<emphasis id="7ACDEABB524BBE2FF100FDAAAEB088FE" box="[512,641,639,660]" italics="true" pageId="2" pageNumber="564">I. phantasma</emphasis>
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</taxonomicName>
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data from Myers (1982); for
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<taxonomicName id="8FB94D2A524BBE2FF0A2FDAAA80B88FE" box="[930,1082,639,660]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="gemmistratus">
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<emphasis id="7ACDEABB524BBE2FF0A2FDAAA80B88FE" box="[930,1082,639,660]" italics="true" pageId="2" pageNumber="564">I. gemmistratus</emphasis>
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</taxonomicName>
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data from Zweifel (1959) and Myers (1982); and for
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<taxonomicName id="8FB94D2A524BBE2FF548FDAAAAFE88FE" box="[1608,1743,639,660]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="tenuissimus">
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<emphasis id="7ACDEABB524BBE2FF548FDAAAAFE88FE" box="[1608,1743,639,660]" italics="true" pageId="2" pageNumber="564">I. tenuissimus</emphasis>
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</taxonomicName>
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data from Cope (1866). The abbreviations are as following: GeA=Anterior chinshield; GeP=Posterior chinshield.
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</paragraph>
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</caption>
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<paragraph id="480636A9524BBE2FF37DFD00AA968FD5" pageId="2" pageNumber="564">
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<table id="3AB9C409524B41D2F37DFD00AB248C26" box="[125,1813,725,1612]" gridcols="7" gridrows="8" pageId="2" pageNumber="564">
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||
<tr id="F68934EB524B41D2F37DFD00AB24894E" box="[125,1813,725,804]" gridrow="0" pageId="2" pageNumber="564">
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||
<th id="B5585D97524B41D2F37DFD00AD82894E" box="[125,435,725,804]" gridcol="0" gridrow="0" pageId="2" pageNumber="564">
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Character
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<taxonomicName id="8FB94D2A524BBE2FF23AFD00ADB3894E" authority="Loreal Present" authorityName="Loreal Present" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="guane">
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<emphasis id="7ACDEABB524BBE2FF23AFD00AD48896D" bold="true" box="[314,421,725,775]" italics="true" pageId="2" pageNumber="564">Imantodes guane</emphasis>
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Loreal Present
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</taxonomicName>
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||
</th>
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||
<th id="B5585D97524B41D2F2F6FD00AE4B894E" box="[502,634,725,804]" gridcol="1" gridrow="0" pageId="2" pageNumber="564">
|
||
<taxonomicName id="8FB94D2A524BBE2FF2F6FD00AE0A894E" authority="Absent" authorityName="Absent" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="chocoensis">
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||
<emphasis id="7ACDEABB524BBE2FF2F6FD00AE448880" box="[502,629,725,746]" italics="true" pageId="2" pageNumber="564">I. chocoensis</emphasis>
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Absent
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</taxonomicName>
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||
</th>
|
||
<th id="B5585D97524B41D2F1B2FD00AF72894E" box="[690,835,725,804]" gridcol="2" gridrow="0" pageId="2" pageNumber="564">
|
||
<taxonomicName id="8FB94D2A524BBE2FF1B2FD00AECB894E" authority="Present" authorityName="Present" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="cenchoa">
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||
<emphasis id="7ACDEABB524BBE2FF1B2FD00AF288880" box="[690,793,725,746]" italics="true" pageId="2" pageNumber="564">I. cenchoa</emphasis>
|
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Present
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="B5585D97524B41D2F06FFD00A836894E" box="[879,1031,725,804]" gridcol="3" gridrow="0" pageId="2" pageNumber="564">
|
||
<taxonomicName id="8FB94D2A524BBE2FF06FFD00AF86894E" authority="Present" authorityName="Present" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="gemmistratus">
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||
<emphasis id="7ACDEABB524BBE2FF06FFD00A8368880" box="[879,1031,725,746]" italics="true" pageId="2" pageNumber="564">I. gemmistratus</emphasis>
|
||
Present
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="B5585D97524B41D2F72DFD00A94F894E" box="[1069,1406,725,804]" gridcol="4" gridrow="0" pageId="2" pageNumber="564">
|
||
<emphasis id="7ACDEABB524BBE2FF72DFD00A9508880" box="[1069,1377,725,746]" italics="true" pageId="2" pageNumber="564">
|
||
<taxonomicName id="8FB94D2A524BBE2FF72DFD00A8918880" box="[1069,1184,725,746]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="inornatus">I. inornatus</taxonomicName>
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||
<taxonomicName id="8FB94D2A524BBE2FF7F0FD00A9508880" authority="Present Present" authorityName="Present Present" box="[1264,1377,725,746]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="lentiferus">I. lentiferus</taxonomicName>
|
||
</emphasis>
|
||
Present Present
|
||
</th>
|
||
<th id="B5585D97524B41D2F6D1FD00AA6F894E" box="[1489,1630,725,804]" gridcol="5" gridrow="0" pageId="2" pageNumber="564">
|
||
<taxonomicName id="8FB94D2A524BBE2FF6D1FD00AA28894E" authority="Present" authorityName="Present" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="phantasma">
|
||
<emphasis id="7ACDEABB524BBE2FF6D1FD00AA638880" box="[1489,1618,725,746]" italics="true" pageId="2" pageNumber="564">I.phantasma</emphasis>
|
||
Present
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="B5585D97524B41D2F58FFD00AB24894E" box="[1679,1813,725,804]" gridcol="6" gridrow="0" pageId="2" pageNumber="564">
|
||
<taxonomicName id="8FB94D2A524BBE2FF58FFD00AAE6894E" authority="Present" authorityName="Present" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="564" phylum="Chordata" rank="species" species="tenuissimus">
|
||
<emphasis id="7ACDEABB524BBE2FF58FFD00AB248880" box="[1679,1813,725,746]" italics="true" pageId="2" pageNumber="564">I.tenuissimus</emphasis>
|
||
Present
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="F68934EB524B41D2F37DFCE4AB24892D" box="[125,1813,817,839]" gridrow="1" pageId="2" pageNumber="564">
|
||
<th id="B5585D97524B41D2F37DFCE4AD82892D" box="[125,435,817,839]" gridcol="0" gridrow="1" pageId="2" pageNumber="564">Supralabials 8–9 (2)</th>
|
||
<td id="B5585D97524B41D2F2F6FCE4AE4B892D" box="[502,634,817,839]" gridcol="1" gridrow="1" pageId="2" pageNumber="564">9 (11)</td>
|
||
<td id="B5585D97524B41D2F1B2FCE4AF72892D" box="[690,835,817,839]" gridcol="2" gridrow="1" pageId="2" pageNumber="564">7–10 (263)</td>
|
||
<td id="B5585D97524B41D2F06FFCE4A836892D" box="[879,1031,817,839]" gridcol="3" gridrow="1" pageId="2" pageNumber="564">6–9 (8)</td>
|
||
<td id="B5585D97524B41D2F72DFCE4A94F892D" box="[1069,1406,817,839]" gridcol="4" gridrow="1" pageId="2" pageNumber="564">7–9 (18) 7–9 (27)</td>
|
||
<td id="B5585D97524B41D2F6D1FCE4AA6F892D" box="[1489,1630,817,839]" gridcol="5" gridrow="1" pageId="2" pageNumber="564">9 (2)</td>
|
||
<td id="B5585D97524B41D2F58FFCE4AB24892D" box="[1679,1813,817,839]" gridcol="6" gridrow="1" pageId="2" pageNumber="564">8 (1)</td>
|
||
</tr>
|
||
<tr id="F68934EB524B41D2F37DFC81AB248900" box="[125,1813,852,874]" gridrow="2" pageId="2" pageNumber="564">
|
||
<th id="B5585D97524B41D2F37DFC81AD828900" box="[125,435,852,874]" gridcol="0" gridrow="2" pageId="2" pageNumber="564">Infralabials 12–13 (2)</th>
|
||
<td id="B5585D97524B41D2F2F6FC81AE4B8900" box="[502,634,852,874]" gridcol="1" gridrow="2" pageId="2" pageNumber="564">12–15 (11)</td>
|
||
<td id="B5585D97524B41D2F1B2FC81AF728900" box="[690,835,852,874]" gridcol="2" gridrow="2" pageId="2" pageNumber="564">8–11 (263)</td>
|
||
<td id="B5585D97524B41D2F06FFC81A8368900" box="[879,1031,852,874]" gridcol="3" gridrow="2" pageId="2" pageNumber="564">9–10 (8)</td>
|
||
<td id="B5585D97524B41D2F72DFC81A94F8900" box="[1069,1406,852,874]" gridcol="4" gridrow="2" pageId="2" pageNumber="564">8–11 (18) 10–12 (26)</td>
|
||
<td id="B5585D97524B41D2F6D1FC81AA6F8900" box="[1489,1630,852,874]" gridcol="5" gridrow="2" pageId="2" pageNumber="564">11–12 (2)</td>
|
||
<td id="B5585D97524B41D2F58FFC81AB248900" box="[1679,1813,852,874]" gridcol="6" gridrow="2" pageId="2" pageNumber="564">—</td>
|
||
</tr>
|
||
<tr id="F68934EB524B41D2F37DFCA2AB248994" box="[125,1813,887,1022]" gridrow="3" pageId="2" pageNumber="564">
|
||
<th id="B5585D97524B41D2F37DFCA2AD828994" box="[125,435,887,1022]" gridcol="0" gridrow="3" pageId="2" pageNumber="564">Infralabials 6 (2) contacting GeA Infralabials 6–7 (2) contacting GeP Temporal stripe Present</th>
|
||
<td id="B5585D97524B41D2F2F6FCA2AE4B8994" box="[502,634,887,1022]" gridcol="1" gridrow="3" pageId="2" pageNumber="564">6 (4) 6–8 (4) Absent</td>
|
||
<td id="B5585D97524B41D2F1B2FCA2AF728994" box="[690,835,887,1022]" gridcol="2" gridrow="3" pageId="2" pageNumber="564">4–6 (263) 5–7 (263) Absent</td>
|
||
<td id="B5585D97524B41D2F06FFCA2A8368994" box="[879,1031,887,1022]" gridcol="3" gridrow="3" pageId="2" pageNumber="564">4–5 (8) 5–6 (8) Absent</td>
|
||
<td id="B5585D97524B41D2F72DFCA2A94F8994" box="[1069,1406,887,1022]" gridcol="4" gridrow="3" pageId="2" pageNumber="564">5 (2) 5–6 (26) 5–6 (2) 6–7 (26) Absent Absent</td>
|
||
<td id="B5585D97524B41D2F6D1FCA2AA6F8994" box="[1489,1630,887,1022]" gridcol="5" gridrow="3" pageId="2" pageNumber="564">6 (2) 6–7 (2) Absent</td>
|
||
<td id="B5585D97524B41D2F58FFCA2AB248994" box="[1679,1813,887,1022]" gridcol="6" gridrow="3" pageId="2" pageNumber="564">— — —</td>
|
||
</tr>
|
||
<tr id="F68934EB524B41D2F37DFBDFAB248EA0" box="[125,1813,1034,1226]" gridrow="4" pageId="2" pageNumber="564">
|
||
<th id="B5585D97524B41D2F37DFBDFAD828EA0" box="[125,435,1034,1226]" gridcol="0" gridrow="4" pageId="2" pageNumber="564">Anterior dorsal 17 (2) scale rows Midbody dorsal 17 (2) scale rows Posterior Dorsal 15 (2) scale rows Ventrals 227–236 (2)</th>
|
||
<td id="B5585D97524B41D2F2F6FBDFAE4B8EA0" box="[502,634,1034,1226]" gridcol="1" gridrow="4" pageId="2" pageNumber="564">17 (11) 17 (11) 17 (11) 232–251 (11)</td>
|
||
<td id="B5585D97524B41D2F1B2FBDFAF728EA0" box="[690,835,1034,1226]" gridcol="2" gridrow="4" pageId="2" pageNumber="564">19 or 17 (263) 17 (263) 17 or 15 (263) 235–282 (263)</td>
|
||
<td id="B5585D97524B41D2F06FFBDFA8368EA0" box="[879,1031,1034,1226]" gridcol="3" gridrow="4" pageId="2" pageNumber="564">19 or 17 (8) 17 (8) 17 (8) 217–253 (48)</td>
|
||
<td id="B5585D97524B41D2F72DFBDFA94F8EA0" box="[1069,1406,1034,1226]" gridcol="4" gridrow="4" pageId="2" pageNumber="564">17 or 15 (18) 15 (27) 19 or 17 (18) 15 (27) 17, 15 or 13 (18) 15 (27) 196–218 (18) 222–240 (27)</td>
|
||
<td id="B5585D97524B41D2F6D1FBDFAA6F8EA0" box="[1489,1630,1034,1226]" gridcol="5" gridrow="4" pageId="2" pageNumber="564">17 (2) 17 (2) 17 or 15 (2) 235–236 (2)</td>
|
||
<td id="B5585D97524B41D2F58FFBDFAB248EA0" box="[1679,1813,1034,1226]" gridcol="6" gridrow="4" pageId="2" pageNumber="564">— 17 (1) — 250 (1)</td>
|
||
</tr>
|
||
<tr id="F68934EB524B41D2F37DFB02AB248E87" box="[125,1813,1239,1261]" gridrow="5" pageId="2" pageNumber="564">
|
||
<th id="B5585D97524B41D2F37DFB02AD828E87" box="[125,435,1239,1261]" gridcol="0" gridrow="5" pageId="2" pageNumber="564">Subcaudals 147–148 (2)</th>
|
||
<td id="B5585D97524B41D2F2F6FB02AE4B8E87" box="[502,634,1239,1261]" gridcol="1" gridrow="5" pageId="2" pageNumber="564">140–161 (9)</td>
|
||
<td id="B5585D97524B41D2F1B2FB02AF728E87" box="[690,835,1239,1261]" gridcol="2" gridrow="5" pageId="2" pageNumber="564">137–192 (223)</td>
|
||
<td id="B5585D97524B41D2F06FFB02A8368E87" box="[879,1031,1239,1261]" gridcol="3" gridrow="5" pageId="2" pageNumber="564">109–170 (32)</td>
|
||
<td id="B5585D97524B41D2F72DFB02A94F8E87" box="[1069,1406,1239,1261]" gridcol="4" gridrow="5" pageId="2" pageNumber="564">108–132 (16) 138–163 (22)</td>
|
||
<td id="B5585D97524B41D2F6D1FB02AA6F8E87" box="[1489,1630,1239,1261]" gridcol="5" gridrow="5" pageId="2" pageNumber="564">156–161 (2)</td>
|
||
<td id="B5585D97524B41D2F58FFB02AB248E87" box="[1679,1813,1239,1261]" gridcol="6" gridrow="5" pageId="2" pageNumber="564">157 (1)</td>
|
||
</tr>
|
||
<tr id="F68934EB524B41D2F37DFB2FAB248F7A" box="[125,1813,1274,1296]" gridrow="6" pageId="2" pageNumber="564">
|
||
<th id="B5585D97524B41D2F37DFB2FAD828F7A" box="[125,435,1274,1296]" gridcol="0" gridrow="6" pageId="2" pageNumber="564">Cloacal plate Divided</th>
|
||
<td id="B5585D97524B41D2F2F6FB2FAE4B8F7A" box="[502,634,1274,1296]" gridcol="1" gridrow="6" pageId="2" pageNumber="564">Entire</td>
|
||
<td id="B5585D97524B41D2F1B2FB2FAF728F7A" box="[690,835,1274,1296]" gridcol="2" gridrow="6" pageId="2" pageNumber="564">Divided</td>
|
||
<td id="B5585D97524B41D2F06FFB2FA8368F7A" box="[879,1031,1274,1296]" gridcol="3" gridrow="6" pageId="2" pageNumber="564">Divided</td>
|
||
<td id="B5585D97524B41D2F72DFB2FA94F8F7A" box="[1069,1406,1274,1296]" gridcol="4" gridrow="6" pageId="2" pageNumber="564">Divided/entire Divided/entire</td>
|
||
<td id="B5585D97524B41D2F6D1FB2FAA6F8F7A" box="[1489,1630,1274,1296]" gridcol="5" gridrow="6" pageId="2" pageNumber="564">Entire/divided</td>
|
||
<td id="B5585D97524B41D2F58FFB2FAB248F7A" box="[1679,1813,1274,1296]" gridcol="6" gridrow="6" pageId="2" pageNumber="564">Divided</td>
|
||
</tr>
|
||
<tr id="F68934EB524B41D2F37DFAC9AB248C26" box="[125,1813,1308,1612]" gridrow="7" pageId="2" pageNumber="564">
|
||
<th id="B5585D97524B41D2F37DFAC9AD828C26" box="[125,435,1308,1612]" gridcol="0" gridrow="7" pageId="2" pageNumber="564">Expanded area Present on the distal portion of the sulcus spermaticus Hemipenis 10 (1) length of everted organs in relation to the number of subcaudals</th>
|
||
<td id="B5585D97524B41D2F2F6FAC9AE4B8C26" box="[502,634,1308,1612]" gridcol="1" gridrow="7" pageId="2" pageNumber="564">Absent 6 (2)</td>
|
||
<td id="B5585D97524B41D2F1B2FAC9AF728C26" box="[690,835,1308,1612]" gridcol="2" gridrow="7" pageId="2" pageNumber="564">Absent 4–6 (18)</td>
|
||
<td id="B5585D97524B41D2F06FFAC9A8368C26" box="[879,1031,1308,1612]" gridcol="3" gridrow="7" pageId="2" pageNumber="564">Absent 5 (2)</td>
|
||
<td id="B5585D97524B41D2F72DFAC9A94F8C26" box="[1069,1406,1308,1612]" gridcol="4" gridrow="7" pageId="2" pageNumber="564">Absent Absent 6–8 (6) 6–7 (4)</td>
|
||
<td id="B5585D97524B41D2F6D1FAC9AA6F8C26" box="[1489,1630,1308,1612]" gridcol="5" gridrow="7" pageId="2" pageNumber="564">Absent 6 (1)</td>
|
||
<td id="B5585D97524B41D2F58FFAC9AB248C26" box="[1679,1813,1308,1612]" gridcol="6" gridrow="7" pageId="2" pageNumber="564">— —</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="480636A9524ABE2EF3C7FF42A8BC8ABF" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="565">
|
||
<emphasis id="7ACDEABB524ABE2EF3C7FF42AD0A8ADA" bold="true" box="[199,315,151,176]" pageId="3" pageNumber="565">
|
||
<typeStatus id="9702880B524ABE2EF3C7FF42AD078ADA" box="[199,310,151,176]" pageId="3" pageNumber="565" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Juvenile female, UIS-R 1705, Cerro de La Paz (
|
||
<geoCoordinate id="2D8D506E524ABE2EF05EFF42AFEB8ADA" box="[862,986,151,176]" direction="north" orientation="latitude" pageId="3" pageNumber="565" precision="15" value="6.9722223">6º58’20”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2D8D506E524ABE2EF0E5FF42A84D8ADA" box="[997,1148,151,176]" direction="west" orientation="longitude" pageId="3" pageNumber="565" precision="15" value="-73.43833">73º26’ 18”W</geoCoordinate>
|
||
;
|
||
<quantity id="8F419B4C524ABE2EF787FF4DA8EC8AC5" box="[1159,1245,152,176]" metricMagnitude="3" metricUnit="m" metricValue="2.324" pageId="3" pageNumber="565" unit="m" value="2324.0">2324 m</quantity>
|
||
asl) collected by E. Brisceño at municipality of Zapatoca, department of Santander,
|
||
<collectingCountry id="30AE7639524ABE2EF083FF69AFC08ABF" box="[899,1009,188,213]" name="Colombia" pageId="3" pageNumber="565">Colombia</collectingCountry>
|
||
(
|
||
<figureCitation id="D0822A2C524ABE2EF701FF69A8608ABE" box="[1025,1105,188,213]" captionStart="FIGURE 4" captionStartId="7.[151,249,858,880]" captionTargetBox="[162,1417,195,816]" captionTargetId="figure@7.[151,1436,193,837]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal (A) and ventral views (B) of the paratype of Imantodes guane (UIS-R 1705) from Zapatoca, departament of Santander, Colombia. Scale bar = 10 mm." httpUri="https://zenodo.org/record/232828/files/figure.png" pageId="3" pageNumber="565">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="D0822A2C524ABE2EF75CFF69A85A8ABF" box="[1116,1131,188,213]" captionStart="FIGURE 5" captionStartId="7.[151,249,1890,1912]" captionTargetBox="[162,1421,975,1858]" captionTargetId="figure@7.[150,1436,961,1869]" captionTargetPageId="7" captionText="FIGURE 5. Schematic representation of the color patterns of holotype (A) and paratype (B) of the Imantodes guane and I. chocoensis (C - ICN 434)." httpUri="https://zenodo.org/record/232829/files/figure.png" pageId="3" pageNumber="565">5</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00A36522524ABE2EF3C7FF0AA8CC890A" pageId="3" pageNumber="565" type="diagnosis">
|
||
<paragraph id="480636A9524ABE2EF3C7FF0AA82D8BC5" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="565">
|
||
<emphasis id="7ACDEABB524ABE2EF3C7FF0AAD708A92" bold="true" box="[199,321,223,248]" pageId="3" pageNumber="565">Diagnosis.</emphasis>
|
||
<taxonomicName id="8FB94D2A524ABE2EF248FF35AE388A92" box="[328,521,224,248]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB524ABE2EF248FF35AE388A92" box="[328,521,224,248]" italics="true" pageId="3" pageNumber="565">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
is distinguish from all congeners by the combination of the following characters: smooth dorsal scale rows 17/17/15; apical pits absent; infralabials 12–13; ventrals 227–236; subcaudals 147–148; loreal scale present; cloacal plate divided; dark brown temporal stripe between the lower edge of temporal scales and upper region of supralabials; dorsum of body light brown with dark brown transversal streaks, weakly evident in lateral view; hemipenis in situ extending to the level of 11th subcaudal and reaches the 10th subcaudal when everted; hemipenis with sulcus spermaticus laterally expanded inside capitulum.
|
||
</paragraph>
|
||
<paragraph id="480636A9524ABE2EF3C7FE6FA8CC890A" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="565">
|
||
<emphasis id="7ACDEABB524ABE2EF3C7FE6FAD5C8BB9" bold="true" box="[199,365,442,467]" pageId="3" pageNumber="565">Comparisons.</emphasis>
|
||
<taxonomicName id="8FB94D2A524ABE2EF274FE6EAE048BB8" box="[372,565,443,467]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB524ABE2EF274FE6EAE048BB8" box="[372,565,443,467]" italics="true" pageId="3" pageNumber="565">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="8FB94D2A524ABE2EF1CBFE6EAF7C8BB9" box="[715,845,443,467]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="lentiferus">
|
||
<emphasis id="7ACDEABB524ABE2EF1CBFE6EAF7C8BB9" box="[715,845,443,467]" italics="true" pageId="3" pageNumber="565">I. lentiferus</emphasis>
|
||
</taxonomicName>
|
||
by having 17 dorsal scale rows anteriorly and at the midbody (vs. 15); from
|
||
<emphasis id="7ACDEABB524ABE2EF2ACFE35AED28B9D" box="[428,739,479,503]" italics="true" pageId="3" pageNumber="565">
|
||
<taxonomicName id="8FB94D2A524ABE2EF2ACFE35AE148B9D" box="[428,549,479,503]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="cenchoa">I. cenchoa</taxonomicName>
|
||
,
|
||
<taxonomicName id="8FB94D2A524ABE2EF132FE35AED28B9D" box="[562,739,480,503]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="gemmistratus">I. gemmistratus</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="8FB94D2A524ABE2EF024FE35AFF18B9D" box="[804,960,480,503]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="tenuissimus">
|
||
<emphasis id="7ACDEABB524ABE2EF024FE35AFF18B9D" box="[804,960,480,503]" italics="true" pageId="3" pageNumber="565">I. tenuissimus</emphasis>
|
||
</taxonomicName>
|
||
by having color pattern light brown with dark brown transversal streaks, weakly evident in lateral view (vs. well defined body blotches with shape of wide saddles); differs from
|
||
<taxonomicName id="8FB94D2A524ABE2EF294FDFDAE3A8855" box="[404,523,551,575]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="cenchoa">
|
||
<emphasis id="7ACDEABB524ABE2EF294FDFDAE3A8855" box="[404,523,551,575]" italics="true" pageId="3" pageNumber="565">I. cenchoa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8FB94D2A524ABE2EF11BFDFDAEFA8855" box="[539,715,552,575]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="gemmistratus">
|
||
<emphasis id="7ACDEABB524ABE2EF11BFDFDAEFA8855" box="[539,715,552,575]" italics="true" pageId="3" pageNumber="565">I. gemmistratus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="8FB94D2A524ABE2EF00CFDFDAFA08855" box="[780,913,552,575]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="7ACDEABB524ABE2EF00CFDFDAFA08855" box="[780,913,552,575]" italics="true" pageId="3" pageNumber="565">I. inornatus</emphasis>
|
||
</taxonomicName>
|
||
by having 12–13 infralabials (vs.
|
||
<date id="3C071069524ABE2EF61DFDF2A9678855" box="[1309,1366,551,575]" pageId="3" pageNumber="565" value="2011-08">8–11</date>
|
||
,
|
||
<date id="3C071069524ABE2EF664FDF2A9408855" box="[1380,1393,551,575]" pageId="3" pageNumber="565" value="2011-09-10" valueMax="2011-09-08" valueMin="2011-09-10">9</date>
|
||
–10 and 8–11, respectively); differs from
|
||
<taxonomicName id="8FB94D2A524ABE2EF13CFD9EAEF18808" box="[572,704,587,610]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="7ACDEABB524ABE2EF13CFD9EAEF18808" box="[572,704,587,610]" italics="true" pageId="3" pageNumber="565">I. inornatus</emphasis>
|
||
</taxonomicName>
|
||
by having 227–236 ventrals and 147–148 subcaudals scales (vs. 196–218 ventrals and <140 subcaudals); differs from
|
||
<taxonomicName id="8FB94D2A524ABE2EF1F2FDA5AFB288ED" box="[754,899,623,647]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="phantasma">
|
||
<emphasis id="7ACDEABB524ABE2EF1F2FDA5AFB288ED" box="[754,899,623,647]" italics="true" pageId="3" pageNumber="565">I. phantasma</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a dark brown temporal stripe (vs. absence of a dark pigmentation on temporal region), dorsum of the head light brown with dark brown blotches (vs. dorsum of the head orangish brown mottled with grayish brown), and 147–148 subcaudals (vs.> 150 subcaudals); differs from
|
||
<taxonomicName id="8FB94D2A524ABE2EF2C5FD0EAE6C8899" box="[453,605,731,755]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="chocoensis">
|
||
<emphasis id="7ACDEABB524ABE2EF2C5FD0EAE6C8899" box="[453,605,731,755]" italics="true" pageId="3" pageNumber="565">I. chocoensis</emphasis>
|
||
</taxonomicName>
|
||
by presence of loreal (vs. absence of loreal), presence of dark brown temporal stripe (vs. absence of a dark brown temporal stripe), dorsal scale rows with posterior reduction to 15 rows (vs. dorsal scales without reduction), and cloacal plate divided (vs. cloacal plate entire). Refers to
|
||
<tableCitation id="053B0312524ABE2EF61AFCF7A9418951" box="[1306,1392,802,827]" captionStart="TABLE 1" captionStartId="2.[125,208,582,604]" captionTargetBox="[125,1813,725,1612]" captionTargetPageId="2" captionText="TABLE 1. Selected qualitative and quantitative morphological characters for the genus Imantodes. Numbers in parenthesis represent the number of specimens examined and additional data was obtained or augmented from the following literature: for I. chocoensis data from Torres-Carvajal et al. (2012) and Jaramillo-Martinéz et al. (2013); for I. cenchoa, I. inornatus, I. lentiferus, and I. phantasma data from Myers (1982); for I. gemmistratus data from Zweifel (1959) and Myers (1982); and for I. tenuissimus data from Cope (1866). The abbreviations are as following: GeA = Anterior chinshield; GeP = Posterior chinshield." httpUri="http://table.plazi.org/id/1CC66621524BBE2FF37DFD93A89788DB" pageId="3" pageNumber="565" tableUuid="1CC66621524BBE2FF37DFD93A89788DB">Table 1</tableCitation>
|
||
for additional morphological characters differing
|
||
<taxonomicName id="8FB94D2A524ABE2EF199FC92AF6B8935" box="[665,858,839,863]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="565" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB524ABE2EF199FC92AF6B8935" box="[665,858,839,863]" italics="true" pageId="3" pageNumber="565">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
from remaining species in the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00A36522524ABE25F3C7FCBFA8998D05" lastPageId="8" lastPageNumber="570" pageId="3" pageNumber="565" type="description">
|
||
<paragraph id="480636A9524ABE2EF3C7FCBFAF8A8C64" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="565">
|
||
<emphasis id="7ACDEABB524ABE2EF3C7FCBFAEE789E9" bold="true" box="[199,726,874,899]" pageId="3" pageNumber="565">
|
||
Description of the
|
||
<typeStatus id="9702880B524ABE2EF2ACFCBFAE2289E9" box="[428,531,874,899]" pageId="3" pageNumber="565" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="D0822A2C524ABE2EF125FCBFAEA489E9" box="[549,661,874,899]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="5.[151,249,1955,1977]" captionStartId-1="6.[151,249,1954,1976]" captionTargetBox-0="[193,1389,216,1928]" captionTargetBox-1="[232,1370,199,1929]" captionTargetId-0="figure@5.[172,1414,193,1933]" captionTargetId-1="figure@6.[196,1391,193,1933]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 2. Dorsal (A) and ventral views (B) of the holotype of Imantodes guane (ICN 5730) from Charalá, Departament of Santander, Colombia. Scale bar = 10 mm. Photo by D. R. Padilla-Rivera." captionText-1="FIGURE 3. Dorsal (A), ventral (B), and lateral views (C) of head of the holotype of Imantodes guane (ICN 5730) from Charalá, departament of Santander, Colombia. Scale bar = 5 mm." httpUri-0="https://zenodo.org/record/232826/files/figure.png" httpUri-1="https://zenodo.org/record/232827/files/figure.png" pageId="3" pageNumber="565">Figs. 2–3</figureCitation>
|
||
,
|
||
<figureCitation id="D0822A2C524ABE2EF1A4FCBFAE8289E9" box="[676,691,874,899]" captionStart="FIGURE 5" captionStartId="7.[151,249,1890,1912]" captionTargetBox="[162,1421,975,1858]" captionTargetId="figure@7.[150,1436,961,1869]" captionTargetPageId="7" captionText="FIGURE 5. Schematic representation of the color patterns of holotype (A) and paratype (B) of the Imantodes guane and I. chocoensis (C - ICN 434)." httpUri="https://zenodo.org/record/232829/files/figure.png" pageId="3" pageNumber="565">5</figureCitation>
|
||
A).
|
||
</emphasis>
|
||
Adult male; SVL
|
||
<quantity id="8F419B4C524ABE2EF0B2FCBEA82489E8" box="[946,1045,875,899]" metricMagnitude="-1" metricUnit="m" metricValue="7.4799999999999995" pageId="3" pageNumber="565" unit="mm" value="748.0">748 mm</quantity>
|
||
; CL
|
||
<quantity id="8F419B4C524ABE2EF751FCBEA88089E8" box="[1105,1201,875,899]" metricMagnitude="-1" metricUnit="m" metricValue="3.4799999999999995" pageId="3" pageNumber="565" unit="mm" value="348.0">348 mm</quantity>
|
||
(46.5% SVL); head sharply distinct from the neck, head length 15.3 mm (2% SVL) and width 9.8 mm (64% head length); distance between the nostrils 3.3 mm (34% head width); snout short and truncated, 4.5 mm length (29.5% head length); eyes large, rounded, and protuberant; eyes diameter 4.2 mm (27% head length); canthus rostralis conspicuous; rostral scale trapezoidal, 2.1 mm wide, 3.1 mm high; internasal and prefrontal scales slightly wider than long; frontal pentagonal, anterior region 1.3 times wider than posterior; supraocular and parietal scales wider than long; nine supralabial scales with fourth to sixth contacting eye; six anterior supralabial shorter than seventh and eighth; ninth supralabial wider and similar in height to first six supralabials; nasal entire, contacting rostral, internasal, prefrontal, loreal, and first two supralabials; loreal 0.7 mm wide, 1.4 mm high, contacting nasal, prefrontal, preocular, second and third supralabials; preocular single; postoculars three on both sides; upper postocular taller than the two lower postoculars; ventral apex of lower postocular inserted between dorsal portion of sixth and seventh supralabials; temporals 2+2+2/2+2+3; symphysial triangular, 2.3 mm wide, 1.3 mm high; infralabials 12/ 12, first six contacting anterior chinshield, sixth and seventh contacting posterior chinshields; seventh infralabial wider and taller than other infralabials; first pair of infralabial scales in contact after symphysial, preventing contact between symphysial and anterior chinshields; anterior chinshield longer and wider than posterior; presence of tubercles on symphysial, first to tenth infralabial, chinshields, and gular scales; smooth dorsal scale rows 17/17/15, reduction at the level of 196th (right) and 199th (left) ventrals; conspicuous dorsal vertebral scale row, 2.3 times wider than adjacent paravertebral scales; preventral one, ventrals 227, subcaudals 147 (terminal spine absent). Maxillary with 18/19 prediastemal teeth and single postdiastemal tooth.
|
||
</paragraph>
|
||
<paragraph id="480636A9524ABE2EF3C7F9CFAE378D88" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="565">Dorsum of head light brown, except for grey frontal scale. Head dorsally with small brown spots at snout region; brown blotches covering the suture region of supraoculars, frontal, and parietals; parietals with two small brown blotches on the anterior region of each scale and two dark brown stripes posteriorly, extending to occipital region. Supralabials light brown, clearest at labial border; region around the orbit blackish; dark brown temporal stripe on the suture between inferior edge of temporals and superior region of seventh, eighth, and ninth supralabials. Ventral surface of head creamish yellow with brownish red spots on the anterior border of symphysial, first and second infralabials; remain infralabials, chinshields, gulars, and preventral immaculate creamish yellow. Background of body light brown with sixty-nine dark brown diffused dorsal blotches, almost restricted to vertebral region; lateral portion of body with dark brown transversal streaks, weakly evident in lateral view; belly creamish yellow scattered with small dark spots concentrated in the lateral edges of ventral scales, forming two irregular lateral stripes more conspicuous on the posterior region of the body; dorsal surface of tail with six evident dorsal blotches and posteriorly small irregular spots; ventral surface of tail creamish yellow with brown spots (dark base and light distal border of scales).</paragraph>
|
||
<caption id="1CC66621524DBE29F397F871A83A8DB3" httpUri="https://zenodo.org/record/232825/files/figure.png" pageId="4" pageNumber="566" targetBox="[155,1432,195,1930]" targetPageId="4">
|
||
<paragraph id="480636A9524DBE29F397F871A83A8DB3" blockId="4.[151,1435,1956,2009]" pageId="4" pageNumber="566">
|
||
<emphasis id="7ACDEABB524DBE29F397F871AD228DD3" bold="true" box="[151,275,1956,1978]" pageId="4" pageNumber="566">FIGURE 1.</emphasis>
|
||
Dorsal (A) and lateral views (B) of the head and dorsal (C) and ventral views (D) of the body of holotype of the
|
||
<taxonomicName id="8FB94D2A524DBE29F397F816AD958DB3" box="[151,420,1987,2009]" class="Squamata" family="Colubridae" genus="Himantodes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="platycephalus">
|
||
<emphasis id="7ACDEABB524DBE29F397F816AD958DB3" box="[151,420,1987,2009]" italics="true" pageId="4" pageNumber="566">Himantodes platycephalus</emphasis>
|
||
</taxonomicName>
|
||
(AMNH 38400). Scale bar = 5 mm. Photo by L. Vonnahme.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1CC66621524CBE28F397F876AF4A8DBD" httpUri="https://zenodo.org/record/232826/files/figure.png" pageId="5" pageNumber="567" targetBox="[193,1389,216,1928]" targetPageId="5">
|
||
<paragraph id="480636A9524CBE28F397F876AF4A8DBD" blockId="5.[151,1436,1955,2007]" pageId="5" pageNumber="567">
|
||
<emphasis id="7ACDEABB524CBE28F397F876AD228DD2" bold="true" box="[151,275,1955,1977]" pageId="5" pageNumber="567">FIGURE 2.</emphasis>
|
||
Dorsal (A) and ventral views (B) of the holotype of
|
||
<taxonomicName id="8FB94D2A524CBE28F034F876AFD28DD3" box="[820,995,1955,1977]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="567" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB524CBE28F034F876AFD28DD3" box="[820,995,1955,1977]" italics="true" pageId="5" pageNumber="567">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
(ICN 5730) from Charalá, Departament of Santander, Colombia. Scale bar = 10 mm. Photo by D. R. Padilla-Rivera.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1CC66621524FBE2BF397F877AF1D8DBD" httpUri="https://zenodo.org/record/232827/files/figure.png" pageId="6" pageNumber="568" targetBox="[232,1370,199,1929]" targetPageId="6">
|
||
<paragraph id="480636A9524FBE2BF397F877AF1D8DBD" blockId="6.[151,1436,1954,2007]" pageId="6" pageNumber="568">
|
||
<emphasis id="7ACDEABB524FBE2BF397F877AD278DDD" bold="true" box="[151,278,1954,1976]" pageId="6" pageNumber="568">FIGURE 3.</emphasis>
|
||
Dorsal (A), ventral (B), and lateral views (C) of head of the holotype of
|
||
<taxonomicName id="8FB94D2A524FBE2BF72DF877A8EF8DD2" box="[1069,1246,1954,1976]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="568" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB524FBE2BF72DF877A8EF8DD2" box="[1069,1246,1954,1976]" italics="true" pageId="6" pageNumber="568">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
(ICN 5730) from Charalá, departament of Santander, Colombia. Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1CC66621524EBE2AF397FC8FAE0E89E5" httpUri="https://zenodo.org/record/232828/files/figure.png" pageId="7" pageNumber="569" targetBox="[162,1417,195,816]" targetPageId="7">
|
||
<paragraph id="480636A9524EBE2AF397FC8FAE0E89E5" blockId="7.[151,1436,858,911]" pageId="7" pageNumber="569">
|
||
<emphasis id="7ACDEABB524EBE2AF397FC8FAD208905" bold="true" box="[151,273,858,880]" pageId="7" pageNumber="569">FIGURE 4.</emphasis>
|
||
Dorsal (A) and ventral views (B) of the paratype of
|
||
<taxonomicName id="8FB94D2A524EBE2AF020FC8FAFFC891A" box="[800,973,858,880]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="569" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB524EBE2AF020FC8FAFFC891A" box="[800,973,858,880]" italics="true" pageId="7" pageNumber="569">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
(UIS-R 1705) from Zapatoca, departament of Santander, Colombia. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1CC66621524EBE2AF397F8B7AD938DFD" httpUri="https://zenodo.org/record/232829/files/figure.png" pageId="7" pageNumber="569" targetBox="[162,1421,975,1858]" targetPageId="7">
|
||
<paragraph id="480636A9524EBE2AF397F8B7AD938DFD" blockId="7.[151,1437,1890,1943]" pageId="7" pageNumber="569">
|
||
<emphasis id="7ACDEABB524EBE2AF397F8B7AD258D1D" bold="true" box="[151,276,1890,1912]" pageId="7" pageNumber="569">FIGURE 5.</emphasis>
|
||
Schematic representation of the color patterns of holotype (A) and paratype (B) of the
|
||
<taxonomicName id="8FB94D2A524EBE2AF7A8F8B7A9698D12" box="[1192,1368,1890,1912]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="569" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB524EBE2AF7A8F8B7A9698D12" box="[1192,1368,1890,1912]" italics="true" pageId="7" pageNumber="569">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8FB94D2A524EBE2AF68DF8B6AD378DFD" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="569" phylum="Chordata" rank="species" species="chocoensis">
|
||
<emphasis id="7ACDEABB524EBE2AF68DF8B6AD378DFD" italics="true" pageId="7" pageNumber="569">I. chocoensis</emphasis>
|
||
</taxonomicName>
|
||
(C - ICN 434).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1CC666215241BE25F397FB0CAD608F64" httpUri="https://zenodo.org/record/232830/files/figure.png" pageId="8" pageNumber="570" targetBox="[151,1421,206,1205]" targetPageId="8">
|
||
<paragraph id="480636A95241BE25F397FB0CAD608F64" blockId="8.[151,1435,1241,1294]" pageId="8" pageNumber="570">
|
||
<emphasis id="7ACDEABB5241BE25F397FB0CAD258E84" bold="true" box="[151,276,1241,1263]" pageId="8" pageNumber="570">FIGURE 6.</emphasis>
|
||
Sulcate (A), lateral (B), and asulcate views (C) of hemipenis of the holotype (ICN 5730) of
|
||
<taxonomicName id="8FB94D2A5241BE25F7E5FB0CA9A78E85" box="[1253,1430,1241,1263]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="570" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB5241BE25F7E5FB0CA9A78E85" box="[1253,1430,1241,1263]" italics="true" pageId="8" pageNumber="570">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="480636A95241BE25F3C7FAEDA9BD8C3A" blockId="8.[151,1437,1333,2012]" pageId="8" pageNumber="570">
|
||
<emphasis id="7ACDEABB5241BE25F3C7FAEDAE478F3B" bold="true" box="[199,630,1336,1361]" pageId="8" pageNumber="570">
|
||
Hemipenis of the
|
||
<typeStatus id="9702880B5241BE25F2A4FAEDAE3A8F3B" box="[420,523,1336,1361]" pageId="8" pageNumber="570" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="D0822A2C5241BE25F11DFAEDAE568F3B" box="[541,615,1336,1361]" captionStart="FIGURE 6" captionStartId="8.[151,249,1241,1263]" captionTargetBox="[151,1421,206,1205]" captionTargetId="figure@8.[151,1435,193,1220]" captionTargetPageId="8" captionText="FIGURE 6. Sulcate (A), lateral (B), and asulcate views (C) of hemipenis of the holotype (ICN 5730) of Imantodes guane. Scale bar = 5 mm." httpUri="https://zenodo.org/record/232830/files/figure.png" pageId="8" pageNumber="570">Fig. 6</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Right organ in situ extending to the level of 11th subcaudal (
|
||
<quantity id="8F419B4C5241BE25F648FAEDA9AD8F3A" box="[1352,1436,1336,1360]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="570" unit="mm" value="22.0">22 mm</quantity>
|
||
length) and reaches the 10th subcaudal when everted. Hemipenis unilobated, capitate, and calyculate; calyces in transverse rows, conspicuous in sulcate, lateral, and asulcate sides of capitulum; asulcate side with small naked pocket located at the base of capitulum; asulcate side with two rows of small spines separating capitulum from the hemipenial body; capitular portion of sulcus spermaticus laterally expanded; lateral expansion of sulcus spermaticus begin at the base and extending the mid-distal region of capitulum; sulcus spermaticus margins narrow and bordered with spinules from the base of hemipenial body until beginning of capitulum; hemipenial body with four rows of large spines disposed transversally and dispersed spinules concentrated on the basal region of organ.
|
||
</paragraph>
|
||
<paragraph id="480636A95241BE25F3C7F98EA8998D05" blockId="8.[151,1437,1333,2012]" pageId="8" pageNumber="570">
|
||
<emphasis id="7ACDEABB5241BE25F3C7F98EAE348C1E" bold="true" box="[199,517,1627,1652]" pageId="8" pageNumber="570">
|
||
Variation of the
|
||
<typeStatus id="9702880B5241BE25F292F98EADCE8C1E" box="[402,511,1627,1652]" pageId="8" pageNumber="570" type="paratype">paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Two postocular scales on both sides; 2+2+3/2+2+3 temporal scales; infralabial scales 12/13; vertebral scale row 1.9 times wider than paravertebral scales; ventrals 236 and subcaudals 148. Dorsum of the head with dark brown blotches bordering internasals and two small rounded blotches at the suture of prefrontals; blotches covering region of suture among supraoculars and frontal connected to small circular blotches on the parietals; suture of parietals with dark brown blotch followed by two small blotches at posterior region of each scale; posterior region of parietals with two dark brown stripes transversally connected on the neck. Sides of head with dark brown spots on first to third supralabials; dark brown spots also covering postoculars and temporals. Dorsum of body with eighty dark brown blotches; tail with twenty-eight blotches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00A365225241BE24F3C7F8AEAE718B0F" lastPageId="9" lastPageNumber="571" pageId="8" pageNumber="570" type="etymology">
|
||
<paragraph id="480636A95241BE24F3C7F8AEAF6E8B76" blockId="8.[151,1437,1333,2012]" lastBlockId="9.[151,1436,151,357]" lastPageId="9" lastPageNumber="571" pageId="8" pageNumber="570">
|
||
<emphasis id="7ACDEABB5241BE25F3C7F8AEAD7F8DFE" bold="true" box="[199,334,1915,1940]" pageId="8" pageNumber="570">Etymology.</emphasis>
|
||
The specific epithet
|
||
<emphasis id="7ACDEABB5241BE25F146F8AEAEAB8DF8" box="[582,666,1915,1938]" italics="true" pageId="8" pageNumber="570">
|
||
“
|
||
<taxonomicName id="8FB94D2A5241BE25F14EF8AEAEAB8DF8" box="[590,666,1915,1938]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="570" phylum="Chordata" rank="species" species="guane">guane</taxonomicName>
|
||
</emphasis>
|
||
” alludes to the name given to indigenous people that lived in the northeast region of the
|
||
<collectingCountry id="30AE76395241BE25F2ACF84BAE2C8DDD" box="[428,541,1950,1975]" name="Colombia" pageId="8" pageNumber="570">Colombia</collectingCountry>
|
||
, where the new specimens were collected. This native Indians occupied the margin of the Fonce and Suárez Rivers, to the left margin Sogamoso river, in Cañon of Chicamocha (
|
||
<bibRefCitation id="2C284B585241BE25F603F816A9A68DB6" author="Gomez" box="[1283,1431,1987,2012]" pageId="8" pageNumber="570" refString="Gomez, J. M. (2005) Tejedores Indigenas de la Montana Santandereana. Boletin de Historia y Antiguedades, 831, 745 - 767" type="journal article" year="2005">Gómez 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2C284B585240BE24F397FF4DAD9A8ADA" author="Corona" box="[151,427,152,176]" pageId="9" pageNumber="571" refString="Corona, A. N. & Moreno, M. E. M. (2010) Los Guane y el Arte Rupestre Xerirense. Fundacion el Libro Total, Gobernacion de Santander, 641 pp." type="book" year="2010">Corona & Moreno 2010</bibRefCitation>
|
||
). The men of the tribe would have their skulls deformed from the birth to the first year of life by using splints in the frontal and posterior surface of the skull (
|
||
<bibRefCitation id="2C284B585240BE24F0B0FF69A88B8ABE" author="Correal" box="[944,1210,188,213]" pageId="9" pageNumber="571" refString="Correal, U. G. & Florez, I. (1992) Estudio de las momias Guane de la Mesa de Los Santos (Santander, Colombia). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 70, 283 - 289." type="journal article" year="1992">Correal & Flórez 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2C284B585240BE24F7C8FF69ACE58A92" author="Corona" pageId="9" pageNumber="571" refString="Corona, A. N. & Moreno, M. E. M. (2010) Los Guane y el Arte Rupestre Xerirense. Fundacion el Libro Total, Gobernacion de Santander, 641 pp." type="book" year="2010">Corona & Moreno 2010</bibRefCitation>
|
||
). The resulting cranial shape denominated “tabular oblique deformation” (sensu
|
||
<bibRefCitation id="2C284B585240BE24F787FF0AA9A58A92" author="Correal" box="[1159,1428,223,248]" pageId="9" pageNumber="571" refString="Correal, U. G. & Florez, I. (1992) Estudio de las momias Guane de la Mesa de Los Santos (Santander, Colombia). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 70, 283 - 289." type="journal article" year="1992">Correal & Flórez 1992</bibRefCitation>
|
||
) reminded us the elongated head typical of the genus
|
||
<taxonomicName id="8FB94D2A5240BE24F1E3FED1AF668B76" box="[739,855,260,284]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="571" phylum="Chordata" rank="genus">
|
||
<emphasis id="7ACDEABB5240BE24F1E3FED1AF668B76" box="[739,855,260,284]" italics="true" pageId="9" pageNumber="571">Imantodes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="480636A95240BE24F3C7FEF2AE718B0F" blockId="9.[151,1436,151,357]" pageId="9" pageNumber="571">
|
||
<emphasis id="7ACDEABB5240BE24F3C7FEF2AE788B2A" bold="true" box="[199,585,295,320]" pageId="9" pageNumber="571">
|
||
Geographic distribution (
|
||
<figureCitation id="D0822A2C5240BE24F2F5FEF2AE0A8B2A" box="[501,571,295,320]" captionStart="FIGURE 7" captionStartId="9.[151,249,1318,1340]" captionTargetBox="[154,1425,419,1290]" captionTargetId="figure@9.[151,1436,411,1296]" captionTargetPageId="9" captionText="FIGURE 7. Geographic distribution of Imantodes guane and I. chocoensis. Black star represent the type locality and black circle the additional record of I. guane. White star represent the type locality and white circles the additional records of I. chocoensis." httpUri="https://zenodo.org/record/232831/files/figure.png" pageId="9" pageNumber="571">Fig. 7</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<taxonomicName id="8FB94D2A5240BE24F14FFEFDAF3E8B2A" box="[591,783,296,320]" class="Reptilia" family="Colubridae" genus="Imantodes" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="571" phylum="Chordata" rank="species" species="guane">
|
||
<emphasis id="7ACDEABB5240BE24F14FFEFDAF3E8B2A" box="[591,783,296,320]" italics="true" pageId="9" pageNumber="571">Imantodes guane</emphasis>
|
||
</taxonomicName>
|
||
is known from the municipalities of Charalá and Zapatoca in the eastern Cordillera of
|
||
<collectingCountry id="30AE76395240BE24F2CAFE99AE0D8B0F" box="[458,572,332,357]" name="Colombia" pageId="9" pageNumber="571">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |