treatments-xml/data/DB/10/43/DB104374FFA3FF8D2B9388F4FEBE774E.xml

<document id="3B9E1B26CD83D55C5E149043142743C3" ID-CLB-Dataset="45535" ID-DOI="10.5281/zenodo.209217" ID-GBIF-Dataset="64526267-7e23-4f51-9fc4-c8a81b66b709" ID-ISSN="1175-5326" ID-Zenodo-Dep="209217" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460434646727" checkinUser="plazi" docAuthor="Gaviria, Santiago &amp; Defaye, Danielle" docDate="2012" docId="DB104374FFA3FF8D2B9388F4FEBE774E" docLanguage="en" docName="zt03179p038.pdf" docOrigin="Zootaxa 3179" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Attheyella (Canthosella) chocoensis Gaviria &amp; Defaye, 2012, n. sp." docType="treatment" docVersion="10" lastPageNumber="21" masterDocId="27293B0CFFA1FF992B048A05FFF37479" masterDocTitle="A new species of Attheyella (Canthosella) from Colombia and redescription of Attheyella (Delachauxiella) freyi (Copepoda: Harpacticoida: Canthocamptidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="3" updateTime="1698331928575" updateUser="ExternalLinkService">
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<mods:title id="DF2D83C0F0FB1E6AF3456161A21C36E1">A new species of Attheyella (Canthosella) from Colombia and redescription of Attheyella (Delachauxiella) freyi (Copepoda: Harpacticoida: Canthocamptidae)</mods:title>
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<mods:namePart id="6291E8C433C33576F5B564BFF7FA74AC">Gaviria, Santiago</mods:namePart>
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<mods:namePart id="37969FF65D0D4AAC68FD886191B85E3C">Defaye, Danielle</mods:namePart>
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<mods:date id="F8614572FDD0C1BDE2961B030F4CC48F">2012</mods:date>
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<treatment id="DB104374FFA3FF8D2B9388F4FEBE774E" ID-DOI="http://doi.org/10.5281/zenodo.5659227" ID-GBIF-Taxon="119431437" ID-Zenodo-Dep="5659227" LSID="urn:lsid:plazi:treatment:DB104374FFA3FF8D2B9388F4FEBE774E" httpUri="http://treatment.plazi.org/id/DB104374FFA3FF8D2B9388F4FEBE774E" lastPageId="20" lastPageNumber="21" pageId="2" pageNumber="3">
<subSubSection id="1BA3A1E9FFA3FF9B2B9388F4FD917755" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="5306F262FFA3FF9B2B9388F4FD627773" blockId="2.[151,657,752,812]" box="[151,657,752,779]" pageId="2" pageNumber="3">
<heading id="084E450EFFA3FF9B2B9388F4FD627773" bold="true" box="[151,657,752,779]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="61CD2E70FFA3FF9B2B9388F4FD627773" bold="true" box="[151,657,752,779]" pageId="2" pageNumber="3">
<taxonomicName id="94B989E1FFA3FF9B2B9388F4FDBB7772" box="[151,584,752,779]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFA3FF9B2B9388F4FEE27772" bold="true" box="[151,273,753,779]" italics="true" pageId="2" pageNumber="3">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA3FF9B2A2688F4FE407772" bold="true" box="[290,435,753,779]" italics="true" pageId="2" pageNumber="3">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA3FF9B2AC088F4FDBB7772" bold="true" box="[452,584,753,779]" italics="true" pageId="2" pageNumber="3">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFA3FF9B294B88F5FD627773" box="[591,657,752,778]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5306F262FFA3FF9B2B938911FD917755" blockId="2.[151,657,752,812]" box="[151,610,788,812]" pageId="2" pageNumber="3">
(
<figureCitation id="CB82EEE7FFA3FF9B2B9B8911FF1F7755" box="[159,236,788,812]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="2" pageNumber="3">Figs. 1</figureCitation>
–3, 5–10, 11A, 11C, 11D, 12–13)
</paragraph>
</subSubSection>
<subSubSection id="1BA3A1E9FFA3FF9B2B938959FE217179" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="5306F262FFA3FF9B2B938959FB9B77EE" blockId="2.[151,1436,860,2036]" pageId="2" pageNumber="3">
<emphasis id="61CD2E70FFA3FF9B2B938959FEF8770D" bold="true" box="[151,267,860,884]" pageId="2" pageNumber="3">
<typeStatus id="8C024CC0FFA3FF9B2B938959FEF5770D" box="[151,262,860,884]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
.
</emphasis>
ICN-MHN-CR 2220. Female, dissected on 1 slide, coll. M. Wolf and J. Betancur, 
<date id="2707D4A2FFA3FF9B2FCD8959FA6B770D" box="[1225,1432,860,884]" pageId="2" pageNumber="3" value="1999-02-19">19 February 1999</date>
, from a phytotelm of 
<taxonomicName id="94B989E1FFA3FF9B2A7A8985FE1677E1" box="[382,485,896,920]" class="Liliopsida" family="Marantaceae" genus="Calathea" kingdom="Plantae" order="Zingiberales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="61CD2E70FFA3FF9B2A7A8985FE1677E1" box="[382,485,896,920]" italics="true" pageId="2" pageNumber="3">Calathea</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="94B989E1FFA3FF9B291F897AFD5D77EE" box="[539,686,895,919]" class="Liliopsida" family="Marantaceae" kingdom="Plantae" order="Zingiberales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="family">Marantaceae</taxonomicName>
) in Nuquí, Jurubida, Chocó, 
<collectingCountry id="2BAEB2F2FFA3FF9B28F6897AFB9777EE" box="[1010,1124,895,919]" name="Colombia" pageId="2" pageNumber="3">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="5306F262FFA3FF9B2BCD89A6FA9577C5" blockId="2.[151,1436,860,2036]" box="[201,1382,931,956]" pageId="2" pageNumber="3">
<emphasis id="61CD2E70FFA3FF9B2BCD89A6FEDD77C5" bold="true" box="[201,302,931,956]" pageId="2" pageNumber="3">Allotype</emphasis>
. ICN-MHN-CR 2221. Male, dissected on 1 slide, same collectors, date and locality as 
<typeStatus id="8C024CC0FFA3FF9B2FF889A1FA9177C5" box="[1276,1378,932,956]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="5306F262FFA3FF9B2BCD89C2FE217179" blockId="2.[151,1436,860,2036]" pageId="2" pageNumber="3">
<emphasis id="61CD2E70FFA3FF9B2BCD89C2FE697799" bold="true" box="[201,410,967,992]" pageId="2" pageNumber="3">
Other 
<typeStatus id="8C024CC0FFA3FF9B2A1989CDFE667799" box="[285,405,968,992]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
.
</emphasis>
ICN-MHN-CR 2223, 
<specimenCount id="45BF39EBFFA3FF9B29AC89C2FD4577A6" box="[680,694,967,991]" pageId="2" pageNumber="3" type="female">5</specimenCount>
females and 
<specimenCount id="45BF39EBFFA3FF9B285F89C2FC4A77A6" box="[859,953,967,991]" pageId="2" pageNumber="3" type="male">4 males</specimenCount>
, undissected, ethanol preserved; 
<collectionCode id="35A86AA7FFA3FF9B2E4489CDFA6F77A6" box="[1344,1436,968,991]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NHMW</collectionCode>
20682, 
<specimenCount id="45BF39EBFFA3FF9B2BEB89E9FF0E707D" box="[239,253,1004,1028]" pageId="2" pageNumber="3" type="female">1</specimenCount>
female dissected on 1 slide; 
<collectionCode id="35A86AA7FFA3FF9B295F89E8FD44707D" box="[603,695,1005,1028]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NHMW</collectionCode>
20683 and 20684, 
<specimenCount id="45BF39EBFFA3FF9B289B89E9FC5E707D" box="[927,941,1004,1028]" pageId="2" pageNumber="3" type="male">2</specimenCount>
males each dissected on 1 slide; MNHN- Cp2296, 
<specimenCount id="45BF39EBFFA3FF9B2BF98E0AFEF8705E" box="[253,267,1039,1063]" pageId="2" pageNumber="3" type="female">1</specimenCount>
female dissected on 1 slide and MNHN-Cp2297, 
<specimenCount id="45BF39EBFFA3FF9B28398E0AFCB8705E" box="[829,843,1039,1063]" pageId="2" pageNumber="3" type="female">1</specimenCount>
female dissected on 1 slide, same locality, collectors and date as 
<typeStatus id="8C024CC0FFA3FF9B2A258E31FE747035" box="[289,391,1076,1100]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
. Phytotelm of 
<taxonomicName id="94B989E1FFA3FF9B29348E31FD647035" box="[560,663,1076,1100]" class="Liliopsida" family="Marantaceae" genus="Calathea" kingdom="Plantae" order="Zingiberales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="61CD2E70FFA3FF9B29348E31FD647035" box="[560,663,1076,1100]" italics="true" pageId="2" pageNumber="3">Calathea</emphasis>
</taxonomicName>
sp. in Utría, Chocó, 
<collectingCountry id="2BAEB2F2FFA3FF9B288E8E31FC087035" box="[906,1019,1076,1100]" name="Colombia" pageId="2" pageNumber="3">Colombia</collectingCountry>
, same collectors, 
<date id="2707D4A2FFA3FF9B2FCD8E31FA6B7035" box="[1225,1432,1076,1100]" pageId="2" pageNumber="3" value="1999-02-17">17 February 1999</date>
: ICN-MHN-CR 2222, 
<specimenCount id="45BF39EBFFA3FF9B2A898E52FE687016" box="[397,411,1111,1135]" pageId="2" pageNumber="3" type="male">1</specimenCount>
male, dissected on 1 slide; 
<collectionCode id="35A86AA7FFA3FF9B29CA8E5CFCD67009" box="[718,805,1113,1136]" pageId="2" pageNumber="3">NHHW</collectionCode>
20560, 
<specimenCount id="45BF39EBFFA3FF9B28848E52FC7D7016" box="[896,910,1111,1135]" pageId="2" pageNumber="3" type="female">1</specimenCount>
female, dissected on 1 slide; 
<collectionCode id="35A86AA7FFA3FF9B2FDC8E5CFAC77009" box="[1240,1332,1113,1136]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NHMW</collectionCode>
20561, 
<specimenCount id="45BF39EBFFA3FF9B2E8A8E52FA6F7016" box="[1422,1436,1111,1135]" pageId="2" pageNumber="3" type="male">1</specimenCount>
male dissected on 1 slide; 
<collectionCode id="35A86AA7FFA3FF9B2AB88E78FDEB70ED" box="[444,536,1149,1172]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NHMW</collectionCode>
20562, 
<specimenCount id="45BF39EBFFA3FF9B29768E79FD7370ED" box="[626,640,1148,1172]" pageId="2" pageNumber="3" type="female">8</specimenCount>
females and 
<specimenCount id="45BF39EBFFA3FF9B28108E79FC9E70ED" box="[788,877,1148,1172]" pageId="2" pageNumber="3" type="male">4 males</specimenCount>
, undissected, ethanol preserved; MNHN-Cp2299, 
<specimenCount id="45BF39EBFFA3FF9B2B938E9AFF5670CE" box="[151,165,1183,1207]" pageId="2" pageNumber="3" type="female">1</specimenCount>
female, dissected on 1 slide; MNHN-Cp2298, 
<specimenCount id="45BF39EBFFA3FF9B29CC8E9AFD2570CE" box="[712,726,1183,1207]" pageId="2" pageNumber="3" type="male">1</specimenCount>
male, dissected on 1 slide; MNHN-Cp2300, 
<specimenCount id="45BF39EBFFA3FF9B2FE78E9AFB0270CE" box="[1251,1265,1183,1207]" pageId="2" pageNumber="3" type="female">9</specimenCount>
females and 
<specimenCount id="45BF39EBFFA3FF9B2E8A8E9AFF2870A5" pageId="2" pageNumber="3" type="male">8 males</specimenCount>
, undissected, ethanol preserved. Additionally, 
<specimenCount id="45BF39EBFFA3FF9B29E88EC1FCBD70A5" box="[748,846,1220,1244]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
was treated for SEM photography and its morphology studied for comparison.
</paragraph>
</subSubSection>
<subSubSection id="1BA3A1E9FFA3FF9B2BCD8F09FE257131" pageId="2" pageNumber="3" type="etymology">
<paragraph id="5306F262FFA3FF9B2BCD8F09FE257131" blockId="2.[151,1436,860,2036]" pageId="2" pageNumber="3">
<emphasis id="61CD2E70FFA3FF9B2BCD8F09FEBC715D" bold="true" box="[201,335,1292,1316]" pageId="2" pageNumber="3">Etymology.</emphasis>
The new species is named after the Colombian State “Chocó”, where the sampling localities Nuquí and Utría are located.
</paragraph>
</subSubSection>
<subSubSection id="1BA3A1E9FFA3FF9B2BCD8F56FF187365" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="5306F262FFA3FF9B2BCD8F56FF187365" blockId="2.[151,1436,860,2036]" pageId="2" pageNumber="3">
<emphasis id="61CD2E70FFA3FF9B2BCD8F56FEB07115" bold="true" box="[201,323,1363,1388]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Female. Genital double-somite ventrally partially divided, dorsally partially divided or not, with short copulatory tube reaching the middle of the double-somite, with or without a row of spinules near the posterior margin and without spinules near the seminal receptacle. Urosomites 3 and 4 always with a posterior continuous row of long spinules ventrally. Anal operculum with 5 to 9 teeth. Caudal ramus with dorsal seta inserted on the middle of the segment, dorsal surface with a spinule row that never reaches the posterior margin, central apical seta with slightly broad base, length of outer seta more than 2 times the dorsal length of the ramus, inner seta clearly shorter than ramus. Leg 3: endopod with 2 spines of same length on inner margin. Leg 4: endopod small, 2-segmented, first segment very short. Male. Urosomites 2, 3 and 4 ventrally always with a continuous and complete row of long spinules. Posterior margin of anal urosomite ventrally with 7 spinules at each side. Anal operculum with 3 to 9 triangular teeth. Caudal rami dorsally without row of spinules. Leg 3: endopod 3-segmented, second segment with apophysis, last segment with 2 apical setae. Leg 4: endopod short, 2-segmented, outer apical seta 2 times longer than in female. Antennula: aesthetasc of segment 4 long, tip almost reaching the distal margin of segment 6.
</paragraph>
</subSubSection>
<subSubSection id="1BA3A1E9FFA3FF892BCD8D22FBA174AD" lastPageId="16" lastPageNumber="17" pageId="2" pageNumber="3" type="description">
<paragraph id="5306F262FFA3FF9B2BCD8D22FB65731D" blockId="2.[151,1436,860,2036]" pageId="2" pageNumber="3">
<emphasis id="61CD2E70FFA3FF9B2BCD8D22FE237339" bold="true" box="[201,464,1831,1856]" pageId="2" pageNumber="3">Description of female.</emphasis>
Length of 
<typeStatus id="8C024CC0FFA3FF9B294F8D2DFD5E7339" box="[587,685,1832,1856]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
428 µm, exclusive of caudal setae. Mean length of 
<typeStatus id="8C024CC0FFA3FF9B2FF78D2CFAAD7338" box="[1267,1374,1833,1857]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
from Nuquí 448 µm (n = 8, range 426–465 µm), from Utría 392 µm (n =19, range 337–465 µm).
</paragraph>
<paragraph id="5306F262FFA3FF9A2BCD8D75FBFD75D5" blockId="2.[151,1436,860,2036]" lastBlockId="3.[151,1437,151,428]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Body cylindrical (
<figureCitation id="CB82EEE7FFA3FF9B2A928D75FE1273F1" box="[406,481,1904,1928]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="2" pageNumber="3">Figs. 1</figureCitation>
A and 2A), broadest section at the posterior margin of cephalosome. Integumentary window long, narrowed anteriorly (
<figureCitation id="CB82EEE7FFA3FF9B29218D91FD9773D5" box="[549,612,1940,1964]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
A) with 2 lateral pores located at the base of the narrow section (not visible in 
<figureCitation id="CB82EEE7FFA3FF9B2BE78DBDFEB373A9" box="[227,320,1976,2000]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="2" pageNumber="3">Figure 1</figureCitation>
A and 2A). Cephalosome and body somites dorsally and laterally with sensilla. Fourth and fifth pedigerous somites and genital double-somite dorsally with ornamentation consisting of row of denticules (visible in 
<figureCitation id="CB82EEE7FFA2FF9A2BB28A92FEEB74D6" box="[182,280,151,175]" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[330,1186,210,1917]" captionTargetId="figure@4.[302,1199,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Attheyella (Canthosella) chocoensis n. sp. Female (paratype from Utria). SEM photographs. A, habitus, dorsal view; B, anal somite and caudal rami, dorsal (most right teeth of anal operculum broken); C, caudal ramus in lateral view." httpUri="https://zenodo.org/record/209232/files/figure.png" pageId="3" pageNumber="4">Figure 2</figureCitation>
A). Body somites: posterodorsal edges smooth. Genital double-somite: posterolaterally with spinules and ventrally smooth. Ventral part of the genital double-somite showing incomplete fusion and not ornamented with spinules (
<figureCitation id="CB82EEE7FFA2FF9A2A3A8ADAFE7A748E" box="[318,393,223,247]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
B and 3A). Dorsal part of the genital double-somite of one 
<typeStatus id="8C024CC0FFA2FF9A2F368AE5FB617481" box="[1074,1170,224,248]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
also shows incomplete fusion (
<figureCitation id="CB82EEE7FFA2FF9A2BE88B01FED87565" box="[236,299,260,284]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
C). Copulatory pore at the middle of the genital double somite (Fig. 3A). Urosomites 3 and 4 posterolaterally and posteroventrally with a continuous row of long spinules (
<figureCitation id="CB82EEE7FFA2FF9A28B58B22FC0F7546" box="[945,1020,295,319]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
B and 3C). Anal urosomite ventrally with a short row of 7 spinules at base of each caudal ramus, laterally with 4 spinules near posterior margin. Anal operculum (
<figureCitation id="CB82EEE7FFA2FF9A2A198B6AFE9575FE" box="[285,358,367,391]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
A and 3B) not protruding, with 9 strong marginal teeth. Female used for SEM photography (
<figureCitation id="CB82EEE7FFA2FF9A2E6B8B6AFF5575D5" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[330,1186,210,1917]" captionTargetId="figure@4.[302,1199,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Attheyella (Canthosella) chocoensis n. sp. Female (paratype from Utria). SEM photographs. A, habitus, dorsal view; B, anal somite and caudal rami, dorsal (most right teeth of anal operculum broken); C, caudal ramus in lateral view." httpUri="https://zenodo.org/record/209232/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
B) with anal operculum bearing 9 marginal teeth, the right-most tooth broken.
</paragraph>
<caption id="07C6A2EAFFA2FF9A2B938D99FD0B7396" httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="3" pageNumber="4" targetBox="[270,1303,482,1909]" targetPageId="3">
<paragraph id="5306F262FFA2FF9A2B938D99FD0B7396" blockId="3.[151,1437,1948,2031]" pageId="3" pageNumber="4">
<emphasis id="61CD2E70FFA2FF9A2B938D99FEE673C8" bold="true" box="[151,277,1948,1971]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
<taxonomicName id="94B989E1FFA2FF9A2A198D99FD6373CB" box="[285,656,1948,1970]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFA2FF9A2A198D99FE7773CB" box="[285,388,1948,1970]" italics="true" pageId="3" pageNumber="4">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA2FF9A2A918D99FDFC73CB" box="[405,527,1948,1970]" italics="true" pageId="3" pageNumber="4">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA2FF9A29258D99FD6373CB" box="[545,656,1948,1970]" italics="true" pageId="3" pageNumber="4">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFA2FF9A299D8D98FD2773CB" box="[665,724,1949,1970]" pageId="3" pageNumber="4" rank="species">
<emphasis id="61CD2E70FFA2FF9A299D8D98FD2773CB" bold="true" box="[665,724,1949,1970]" pageId="3" pageNumber="4">n. sp.</emphasis>
</taxonomicNameLabel>
Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp2299. Position of structures of C and D indicated by asterisk on B.
</paragraph>
</caption>
<caption id="07C6A2EAFFA5FF9D2B938DB5FA9A739C" httpUri="https://zenodo.org/record/209232/files/figure.png" pageId="4" pageNumber="5" targetBox="[330,1186,210,1917]" targetPageId="4">
<paragraph id="5306F262FFA5FF9D2B938DB5FA9A739C" blockId="4.[151,1436,1968,2021]" pageId="4" pageNumber="5">
<emphasis id="61CD2E70FFA5FF9D2B938DB5FEE673BF" bold="true" box="[151,277,1968,1991]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
<taxonomicName id="94B989E1FFA5FF9D2A198DB4FD7C73BE" box="[285,655,1969,1991]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFA5FF9D2A198DB4FE7073BE" box="[285,387,1969,1991]" italics="true" pageId="4" pageNumber="5">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA5FF9D2A918DB4FDFC73BE" box="[405,527,1969,1991]" italics="true" pageId="4" pageNumber="5">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA5FF9D29248DB4FD7C73BE" box="[544,655,1969,1991]" italics="true" pageId="4" pageNumber="5">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFA5FF9D299C8DB4FD2073BF" box="[664,723,1969,1990]" pageId="4" pageNumber="5" rank="species">
<emphasis id="61CD2E70FFA5FF9D299C8DB4FD2073BF" bold="true" box="[664,723,1969,1990]" pageId="4" pageNumber="5">n. sp.</emphasis>
</taxonomicNameLabel>
Female (paratype from Utria). SEM photographs. A, habitus, dorsal view; B, anal somite and caudal rami, dorsal (most right teeth of anal operculum broken); C, caudal ramus in lateral view.
</paragraph>
</caption>
<paragraph id="5306F262FFA4FF9C2B938D7BFC0273CC" blockId="5.[151,1436,1918,1973]" pageId="5" pageNumber="6">
<emphasis id="61CD2E70FFA4FF9C2B938D7BFEE073EA" bold="true" box="[151,275,1918,1941]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
<taxonomicName id="94B989E1FFA4FF9C2A1D8D7BFD7473ED" box="[281,647,1918,1940]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFA4FF9C2A1D8D7BFE7373ED" box="[281,384,1918,1940]" italics="true" pageId="5" pageNumber="6">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA4FF9C2A8B8D7BFDFA73ED" box="[399,521,1918,1940]" italics="true" pageId="5" pageNumber="6">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA4FF9C291C8D7BFD7473ED" box="[536,647,1918,1940]" italics="true" pageId="5" pageNumber="6">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFA4FF9C298A8D7AFD3B73ED" box="[654,712,1919,1940]" pageId="5" pageNumber="6" rank="species">
<emphasis id="61CD2E70FFA4FF9C298A8D7AFD3B73ED" bold="true" box="[654,712,1919,1940]" pageId="5" pageNumber="6">n. sp.</emphasis>
</taxonomicNameLabel>
Female (
<typeStatus id="8C024CC0FFA4FF9C28228D7BFC7673ED" box="[806,901,1918,1940]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
). A, genital double-somite, ventral, showing setae of 6th pair of legs; B, last abdominal somites, dorsal; C, last abdominal somites, ventral.
</paragraph>
<caption id="07C6A2EAFFA7FF9F2B938CA5FB2B72AC" httpUri="https://zenodo.org/record/209239/files/figure.png" pageId="6" pageNumber="7" targetBox="[274,1321,193,1661]" targetPageId="6">
<paragraph id="5306F262FFA7FF9F2B938CA5FB2B72AC" blockId="6.[151,1437,1696,1749]" pageId="6" pageNumber="7">
<emphasis id="61CD2E70FFA7FF9F2B938CA5FEE772CF" bold="true" box="[151,276,1696,1719]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
Females. 
<taxonomicName id="94B989E1FFA7FF9F2A7A8CA4FD3172CE" box="[382,706,1697,1719]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFA7FF9F2A7A8CA4FE1672CE" box="[382,485,1697,1719]" italics="true" pageId="6" pageNumber="7">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA7FF9F2AF18CA4FD9C72CE" box="[501,623,1697,1719]" italics="true" pageId="6" pageNumber="7">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA7FF9F297B8CA4FD3172CE" box="[639,706,1697,1719]" italics="true" pageId="6" pageNumber="7">aliena</emphasis>
</taxonomicName>
. A. caudal rami with anal operculum, dorsal; B, caudal ramus, ventral. 
<taxonomicName id="94B989E1FFA7FF9F2B938CBAFDFF72AC" box="[151,524,1727,1749]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFA7FF9F2B938CBAFF0D72AC" box="[151,254,1727,1749]" italics="true" pageId="6" pageNumber="7">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA7FF9F2A088CBAFE7572AC" box="[268,390,1727,1749]" italics="true" pageId="6" pageNumber="7">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA7FF9F2A918CBAFDFF72AC" box="[405,524,1727,1749]" italics="true" pageId="6" pageNumber="7">pilagaensis</emphasis>
</taxonomicName>
. C, caudal rami with anal operculum, dorsal; D, caudal ramus, ventral.
</paragraph>
</caption>
<paragraph id="5306F262FFA7FF9F2BCD8CF8FEA17394" blockId="6.[151,1437,1789,2029]" pageId="6" pageNumber="7">
Caudal rami subquadrate, slightly longer than wide, inner margin smooth; posterodorsally a row of 3 short spinules at the distal third (
<figureCitation id="CB82EEE7FFA7FF9F2AC78D24FDFF7340" box="[451,524,1825,1849]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="6" pageNumber="7">Figs. 1</figureCitation>
A, 2B, 3B and 5A); dorsally a longitudinal carina splitting in two, extending anteriorly to the insertion point of the dorsal seta (well visible in 
<figureCitation id="CB82EEE7FFA7FF9F283B8D40FC787324" box="[831,907,1861,1885]" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[330,1186,210,1917]" captionTargetId="figure@4.[302,1199,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Attheyella (Canthosella) chocoensis n. sp. Female (paratype from Utria). SEM photographs. A, habitus, dorsal view; B, anal somite and caudal rami, dorsal (most right teeth of anal operculum broken); C, caudal ramus in lateral view." httpUri="https://zenodo.org/record/209232/files/figure.png" pageId="6" pageNumber="7">Figs. 2</figureCitation>
B and 2C); outer margin with 2 setae, the proximal one inserted on the first third of the segment and accompanied by 1 row of 3 spinules, the distal one inserted on the distal third of the segment and accompanied by 2 spinules (
<figureCitation id="CB82EEE7FFA7FF9F289A8D88FC1073DC" box="[926,995,1933,1957]" captionStart="FIGURE 5" captionStartId="7.[151,250,1614,1637]" captionTargetBox="[238,1339,208,1584]" captionTargetId="figure@7.[227,1360,193,1593]" captionTargetPageId="7" captionText="FIGURE 5. Attheyella (Canthosella) chocoensis n. sp. Female (paratype NHMW 20682). A, caudal rami with anal operculum, dorsal; B, caudal ramus, ventral; C, caudal ramus, lateral." httpUri="https://zenodo.org/record/209245/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
C); 3 terminal setae: median seta well developed with base slightly enlarged, outer setae 2. 
<date id="2707D4A2FFA7FF9F29DF8DB4FD0C73B0" box="[731,767,1969,1993]" pageId="6" pageNumber="7">2 x</date>
longer than caudal ramus, inner seta shorter than the caudal ramus (
<figureCitation id="CB82EEE7FFA7FF9F2BEE8DD0FEDE7394" box="[234,301,2005,2029]" captionStart="FIGURE 5" captionStartId="7.[151,250,1614,1637]" captionTargetBox="[238,1339,208,1584]" captionTargetId="figure@7.[227,1360,193,1593]" captionTargetPageId="7" captionText="FIGURE 5. Attheyella (Canthosella) chocoensis n. sp. Female (paratype NHMW 20682). A, caudal rami with anal operculum, dorsal; B, caudal ramus, ventral; C, caudal ramus, lateral." httpUri="https://zenodo.org/record/209245/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
A).
</paragraph>
<caption id="07C6A2EAFFA6FF9E2B938C4BFD2E72FA" httpUri="https://zenodo.org/record/209245/files/figure.png" pageId="7" pageNumber="8" targetBox="[238,1339,208,1584]" targetPageId="7">
<paragraph id="5306F262FFA6FF9E2B938C4BFD2E72FA" blockId="7.[151,1436,1614,1667]" pageId="7" pageNumber="8">
<emphasis id="61CD2E70FFA6FF9E2B938C4BFEE1721D" bold="true" box="[151,274,1614,1637]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
<taxonomicName id="94B989E1FFA6FF9E2A1C8C4AFD77721C" box="[280,644,1615,1637]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFA6FF9E2A1C8C4AFE8C721C" box="[280,383,1615,1637]" italics="true" pageId="7" pageNumber="8">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA6FF9E2A898C4AFDF4721C" box="[397,519,1615,1637]" italics="true" pageId="7" pageNumber="8">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA6FF9E29118C4AFD77721C" box="[533,644,1615,1637]" italics="true" pageId="7" pageNumber="8">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFA6FF9E298F8C4AFD30721D" box="[651,707,1615,1636]" pageId="7" pageNumber="8" rank="species">
<emphasis id="61CD2E70FFA6FF9E298F8C4AFD30721D" bold="true" box="[651,707,1615,1636]" pageId="7" pageNumber="8">n. sp.</emphasis>
</taxonomicNameLabel>
Female (paratype NHMW 20682). A, caudal rami with anal operculum, dorsal; B, caudal ramus, ventral; C, caudal ramus, lateral.
</paragraph>
</caption>
<paragraph id="5306F262FFA6FF9E2BCD8CB5FB6C7321" blockId="7.[151,1436,1712,2024]" pageId="7" pageNumber="8">
Antennule 8-segmented (
<figureCitation id="CB82EEE7FFA6FF9E2AE08CB5FDDB72B1" box="[484,552,1712,1736]" captionStart="FIGURE 6" captionStartId="8.[151,250,1726,1749]" captionTargetBox="[161,1416,214,1704]" captionTargetId="figure@8.[151,1436,193,1705]" captionTargetPageId="8" captionText="FIGURE 6. Attheyella (Canthosella) chocoensis n. sp. Female (holotype A, B and C). A, antennule; B, leg 5; C, inner spine of exopod; D, leg 5, inner spine of exopod, paratype NHMW 20560; E, leg 5, spine of exopod, paratype MHHN-Cp 2229." httpUri="https://zenodo.org/record/209251/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
A) with setation formula: 1, 9, 4, 1 + aesthetasc (seta and aesthetasc with conjoined bases), 1, 2, 2, 5 and 1 + aesthetasc (seta and aesthetasc with conjoined bases). Antenna (
<figureCitation id="CB82EEE7FFA6FF9E2FC08CD1FAF47295" box="[1220,1287,1748,1772]" captionStart="FIGURE 7" captionStartId="9.[151,250,1726,1749]" captionTargetBox="[173,1428,215,1675]" captionTargetId="figure@9.[151,1436,193,1705]" captionTargetPageId="9" captionText="FIGURE 7. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, antenna; B, mandible; C, maxillule; D, maxilla; E, maxilliped." httpUri="https://zenodo.org/record/209260/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
A): Allobasis with 1 seta and 6 spinules on inner margin; endopod 1-segmented, inner margin of endopod with 3 proximal spinules followed by 2 spines, and 2 spinules near distal outer corner, apical margin of endopod with 2 strong, unequal spines and 3 geniculate setae. Exopod 1-segmented, with 1 inner and 3 apical setae.
</paragraph>
<paragraph id="5306F262FFA6FF9E2BCD8D61FDA573D9" blockId="7.[151,1436,1712,2024]" pageId="7" pageNumber="8">
Mandible (
<figureCitation id="CB82EEE7FFA6FF9E2A408D61FE7B7305" box="[324,392,1892,1916]" captionStart="FIGURE 7" captionStartId="9.[151,250,1726,1749]" captionTargetBox="[173,1428,215,1675]" captionTargetId="figure@9.[151,1436,193,1705]" captionTargetPageId="9" captionText="FIGURE 7. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, antenna; B, mandible; C, maxillule; D, maxilla; E, maxilliped." httpUri="https://zenodo.org/record/209260/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
B): gnathobase with 5 main strong teeth and one lateral seta, palp 1-segmented, with 3 apical setae, outer seta longer than inner setae.
</paragraph>
<paragraph id="5306F262FFA6FF9E2BCD8DA9FC327391" blockId="7.[151,1436,1712,2024]" pageId="7" pageNumber="8">
Maxillule (
<figureCitation id="CB82EEE7FFA6FF9E2A428DA9FE7873BD" box="[326,395,1964,1988]" captionStart="FIGURE 7" captionStartId="9.[151,250,1726,1749]" captionTargetBox="[173,1428,215,1675]" captionTargetId="figure@9.[151,1436,193,1705]" captionTargetPageId="9" captionText="FIGURE 7. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, antenna; B, mandible; C, maxillule; D, maxilla; E, maxilliped." httpUri="https://zenodo.org/record/209260/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
C): with arthrite of the praecoxa ending in 4 spines and bearing 4 setae, coxal endite with 2 setae; basal endite with 1 spine and 2 setae, inner margin with 1 spinule.
</paragraph>
<caption id="07C6A2EAFFA9FF912B938CBBFABF728B" httpUri="https://zenodo.org/record/209251/files/figure.png" pageId="8" pageNumber="9" targetBox="[161,1416,214,1704]" targetPageId="8">
<paragraph id="5306F262FFA9FF912B938CBBFABF728B" blockId="8.[151,1436,1726,1779]" pageId="8" pageNumber="9">
<emphasis id="61CD2E70FFA9FF912B938CBBFEE172AA" bold="true" box="[151,274,1726,1749]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
<taxonomicName id="94B989E1FFA9FF912A1C8CBBFD7572AD" box="[280,646,1726,1748]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFA9FF912A1C8CBBFE8D72AD" box="[280,382,1726,1748]" italics="true" pageId="8" pageNumber="9">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA9FF912A8A8CBBFDFB72AD" box="[398,520,1726,1748]" italics="true" pageId="8" pageNumber="9">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA9FF9129138CBBFD7572AD" box="[535,646,1726,1748]" italics="true" pageId="8" pageNumber="9">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFA9FF9129888CBAFD3672AD" box="[652,709,1727,1748]" pageId="8" pageNumber="9" rank="species">
<emphasis id="61CD2E70FFA9FF9129888CBAFD3672AD" bold="true" box="[652,709,1727,1748]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
Female (holotype A, B and C). A, antennule; B, leg 5; C, inner spine of exopod; D, leg 5, inner spine of exopod, paratype NHMW 20560; E, leg 5, spine of exopod, paratype MHHN-Cp2229.
</paragraph>
</caption>
<paragraph id="5306F262FFA9FF912BCD8D1FFD7D7303" blockId="8.[151,1436,1818,2022]" pageId="8" pageNumber="9">
Maxilla (
<figureCitation id="CB82EEE7FFA9FF912A368D1FFE8A734B" box="[306,377,1818,1842]" captionStart="FIGURE 7" captionStartId="9.[151,250,1726,1749]" captionTargetBox="[173,1428,215,1675]" captionTargetId="figure@9.[151,1436,193,1705]" captionTargetPageId="9" captionText="FIGURE 7. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, antenna; B, mandible; C, maxillule; D, maxilla; E, maxilliped." httpUri="https://zenodo.org/record/209260/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
D): composed of syncoxa with 2 endites and basis; proximal endite of syncoxa with lateral spine ending in a spiniform tip, distal endite with 1 spine and 1 seta; basis ending in spiniform tip with 1 lateral seta. Proximal margin of maxilla with 1 seta.
</paragraph>
<paragraph id="5306F262FFA9FF912BCD8D83FCFF739F" blockId="8.[151,1436,1818,2022]" pageId="8" pageNumber="9">
Maxilliped (
<figureCitation id="CB82EEE7FFA9FF912A508D83FE6B73E7" box="[340,408,1926,1950]" captionStart="FIGURE 7" captionStartId="9.[151,250,1726,1749]" captionTargetBox="[173,1428,215,1675]" captionTargetId="figure@9.[151,1436,193,1705]" captionTargetPageId="9" captionText="FIGURE 7. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, antenna; B, mandible; C, maxillule; D, maxilla; E, maxilliped." httpUri="https://zenodo.org/record/209260/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
E): with coxa ornamented with two groups of spinules, basis with an inner lateral row of 4 strong spinules and a long row of spinules, a group of subapical spinules and an external group of apical spinules; endopod 1-segmented, bearing long claw and short seta.
</paragraph>
<caption id="07C6A2EAFFA8FF902B938CBBFED5728A" httpUri="https://zenodo.org/record/209260/files/figure.png" pageId="9" pageNumber="10" targetBox="[173,1428,215,1675]" targetPageId="9">
<paragraph id="5306F262FFA8FF902B938CBBFED5728A" blockId="9.[151,1436,1726,1779]" pageId="9" pageNumber="10">
<emphasis id="61CD2E70FFA8FF902B938CBBFEE072AD" bold="true" box="[151,275,1726,1749]" pageId="9" pageNumber="10">FIGURE 7.</emphasis>
<taxonomicName id="94B989E1FFA8FF902A1E8CBBFD7B72AD" box="[282,648,1726,1748]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFA8FF902A1E8CBBFE7272AD" box="[282,385,1726,1748]" italics="true" pageId="9" pageNumber="10">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFA8FF902A948CBBFDF972AD" box="[400,522,1726,1748]" italics="true" pageId="9" pageNumber="10">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFA8FF90291D8CBBFD7B72AD" box="[537,648,1726,1748]" italics="true" pageId="9" pageNumber="10">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFA8FF90298B8CBAFD3A72AD" box="[655,713,1727,1748]" pageId="9" pageNumber="10" rank="species">
<emphasis id="61CD2E70FFA8FF90298B8CBAFD3A72AD" bold="true" box="[655,713,1727,1748]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
Female (holotype). A, antenna; B, mandible; C, maxillule; D, maxilla; E, maxilliped.
</paragraph>
</caption>
<paragraph id="5306F262FFA8FF932BCD8D25FF09748E" blockId="9.[151,1436,1823,2028]" lastBlockId="10.[151,1436,151,247]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Leg 1 (
<figureCitation id="CB82EEE7FFA8FF902A188D1AFE92734E" box="[284,353,1823,1847]" captionStart="FIGURE 8" captionStartId="10.[151,250,1412,1435]" captionTargetBox="[227,1358,319,1361]" captionTargetId="figure@10.[204,1384,301,1391]" captionTargetPageId="10" captionText="FIGURE 8. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, leg 1; B, leg 2." httpUri="https://zenodo.org/record/209267/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
A): intercoxal plate unarmed. Coxa, anterior surface with row of 3 spinules near outer margin; basis with outer spine and row of 4 spinules on anterior surface near base of exopod, 1 seta and row of spinules near base of endopod. Exopod 3-segmented; first segment with 1 bipinnate spine and row of spinules on outer margin; second segment with 1 bipinnate spine and row of spinules on outer margin as well, inner margin with 1 spinule; third segment with 1 unipinnate spine and a row of spinules on outer margin, apically with 1 unipinnate spine and 2 geniculate setae. Endopod 2-segmented, almost as long as exopod; first segment with 1 long unipinnate inner seta and a row of spinules on outer margin; second segment with 1 spinule on the middle of inner margin and 1 naked seta inserted subapically, apical margin with 1 geniculate seta and 1 naked seta, outer margin with row of spinules.
</paragraph>
<caption id="07C6A2EAFFABFF932B938F81FBB471E3" box="[151,1095,1412,1435]" httpUri="https://zenodo.org/record/209267/files/figure.png" pageId="10" pageNumber="11" targetBox="[227,1358,319,1361]" targetPageId="10">
<paragraph id="5306F262FFABFF932B938F81FBB471E3" blockId="10.[151,1095,1412,1435]" box="[151,1095,1412,1435]" pageId="10" pageNumber="11">
<emphasis id="61CD2E70FFABFF932B938F81FEE171E0" bold="true" box="[151,274,1412,1435]" pageId="10" pageNumber="11">FIGURE 8.</emphasis>
<taxonomicName id="94B989E1FFABFF932A1D8F81FD7571E3" box="[281,646,1412,1434]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFABFF932A1D8F81FE8C71E3" box="[281,383,1412,1434]" italics="true" pageId="10" pageNumber="11">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFABFF932A8A8F81FDFB71E3" box="[398,520,1412,1434]" italics="true" pageId="10" pageNumber="11">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFABFF9329138F81FD7571E3" box="[535,646,1412,1434]" italics="true" pageId="10" pageNumber="11">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFABFF9329898F80FD3571E3" box="[653,710,1413,1434]" pageId="10" pageNumber="11" rank="species">
<emphasis id="61CD2E70FFABFF9329898F80FD3571E3" bold="true" box="[653,710,1413,1434]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
Female (holotype). A, leg 1; B, leg 2.
</paragraph>
</caption>
<paragraph id="5306F262FFABFF932BCD8FC7FE7072AF" blockId="10.[151,1436,1473,2002]" pageId="10" pageNumber="11">
Leg 2 (
<figureCitation id="CB82EEE7FFABFF932A188FC4FE9371A0" box="[284,352,1473,1497]" captionStart="FIGURE 8" captionStartId="10.[151,250,1412,1435]" captionTargetBox="[227,1358,319,1361]" captionTargetId="figure@10.[204,1384,301,1391]" captionTargetPageId="10" captionText="FIGURE 8. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, leg 1; B, leg 2." httpUri="https://zenodo.org/record/209267/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
B): anterior surface of intercoxal plate with 1 spinule at base of each lateral prominence. Coxa unarmed. Basis with 1 outer spine and 1 spinule. Exopod 3-segmented; outer margin of first segment with spinules and 1 bipinnate spine; outer margin of second segment with spinules and 1 bipinnate spine and as well, inner margin with spinules and 1 long seta, unipinnate at its extremity; third segment with a few spinules on the external margin and 1 bipinnate spine inserted subapically, apical margin with 1 bipinnate spine and 2 setae, inner margin with 1 long seta unipinnate at its extremity. Endopod 2-segmented; first segment short and unarmed; second segment with spinules and 1 distal bipinnate seta on outer margin, 2 bipinnate setae apically and 1 unipinnate median seta on inner margin.
</paragraph>
<paragraph id="5306F262FFABFF932BCD8CE7FBB773AB" blockId="10.[151,1436,1473,2002]" pageId="10" pageNumber="11">
Leg 3 (
<figureCitation id="CB82EEE7FFABFF932A188CE4FE937280" box="[284,352,1761,1785]" captionStart="FIGURE 9" captionStartId="11.[151,250,1691,1714]" captionTargetBox="[193,1397,205,1662]" captionTargetId="figure@11.[181,1405,193,1670]" captionTargetPageId="11" captionText="FIGURE 9. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, leg 3; B, leg 4." httpUri="https://zenodo.org/record/209273/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
A): intercoxal plate as in leg 2. Basis with 1 long outer naked seta. Exopod 3-segmented, outer margin of first and second segments with spinules and 1 bipinnate spine and, inner margin of first segment without seta; inner margin of second segment with a short naked seta and proximal spinules, outer margin of third segment with spinules and 2 bipinnate spines, inner margin with 2 setae unipinnate in their distal part, apical margin with 2 setae, outer seta with row of strong setulae, inner seta bipinnate in its distal half. Endopod 2-segmented, first segment short and unarmed, outer margin of second segment with spinules and 1 distal bipinnate seta, apical margin with 2 partly bipinnate setae, inner margin with spinules and 2 naked similar spines.
</paragraph>
<caption id="07C6A2EAFFAAFF922B938C9EFBB472C8" box="[151,1095,1691,1714]" httpUri="https://zenodo.org/record/209273/files/figure.png" pageId="11" pageNumber="12" targetBox="[193,1397,205,1662]" targetPageId="11">
<paragraph id="5306F262FFAAFF922B938C9EFBB472C8" blockId="11.[151,1095,1691,1714]" box="[151,1095,1691,1714]" pageId="11" pageNumber="12">
<emphasis id="61CD2E70FFAAFF922B938C9EFEE172C8" bold="true" box="[151,274,1691,1714]" pageId="11" pageNumber="12">FIGURE 9.</emphasis>
<taxonomicName id="94B989E1FFAAFF922A1D8C99FD7572CB" box="[281,646,1692,1714]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFAAFF922A1D8C99FE8C72CB" box="[281,383,1692,1714]" italics="true" pageId="11" pageNumber="12">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFAAFF922A8A8C99FDFB72CB" box="[398,520,1692,1714]" italics="true" pageId="11" pageNumber="12">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFAAFF9229138C99FD7572CB" box="[535,646,1692,1714]" italics="true" pageId="11" pageNumber="12">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFAAFF9229898C99FD3572C8" box="[653,710,1692,1713]" pageId="11" pageNumber="12" rank="species">
<emphasis id="61CD2E70FFAAFF9229898C99FD3572C8" bold="true" box="[653,710,1692,1713]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
Female (holotype). A, leg 3; B, leg 4.
</paragraph>
</caption>
<paragraph id="5306F262FFAAFF922BCD8CDAFB55738B" blockId="11.[151,1436,1758,2034]" pageId="11" pageNumber="12">
Leg 4 (
<figureCitation id="CB82EEE7FFAAFF922A1C8CDBFEAF728F" box="[280,348,1758,1782]" captionStart="FIGURE 9" captionStartId="11.[151,250,1691,1714]" captionTargetBox="[193,1397,205,1662]" captionTargetId="figure@11.[181,1405,193,1670]" captionTargetPageId="11" captionText="FIGURE 9. Attheyella (Canthosella) chocoensis n. sp. Female (holotype). A, leg 3; B, leg 4." httpUri="https://zenodo.org/record/209273/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
B): intercoxal plate as in legs 2 and 3. Basis with 1 long outer naked seta and a row of spinules on anterior surface near the exopod. Exopod 3-segmented; first segment with row of spinules and 1 bipinnate spine on outer margin and spinules on anterior surface near apical border; outer margin of second segment with 1 bipinnate spine and spinules, inner margin with naked distal seta and a few proximal spinules; outer margin of third segment with 1 spine and spinules, apical margin with 2 spines (outer spine bipinnate, inner spine unipinnate) and 1 bipinnate seta, inner margin with 2 setae, proximal seta distally unipinnate, distal seta naked. Endopod 2-segmented, first segment very short and unarmed, inner margin of second segment with 2 spinules (right leg) or smooth (left leg); apical margin with 1 outer naked spine and 1 long inner seta unipinnate at its extremity.
</paragraph>
<paragraph id="5306F262FFADFF952BCD8A9DFED374AD" blockId="12.[151,1436,151,212]" pageId="12" pageNumber="13">Legs 1-4 with following formula of spines (Roman numerals) and setae (Arabic). Spinules are not included in the formula.</paragraph>
<paragraph id="5306F262FFADFF952A758AFDFB6475D3" pageId="12" pageNumber="13">
<table id="21B900C2FFAD00662B938AFDFB6475D3" box="[151,1175,248,426]" colsContinueIn="32.[151,1331,291,468]" gridcols="4" gridrows="5" pageId="12" pageNumber="13">
<tr id="ED89F020FFAD00662B938AFDFB647577" box="[151,1175,248,270]" gridrow="0" pageId="12" pageNumber="13" rowspan-0="1">
<th id="AE58995CFFAD00662A758AFDFE557577" box="[369,422,248,270]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">Coxa</th>
<th id="AE58995CFFAD006629358AFDFD9B7577" box="[561,616,248,270]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">Basis</th>
<th id="AE58995CFFAD0066281D8AFDFB647577" box="[793,1175,248,270]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">Exopod Endopod</th>
</tr>
<tr id="ED89F020FFAD00662B938B32FB647535" box="[151,1175,311,332]" gridrow="1" pageId="12" pageNumber="13">
<th id="AE58995CFFAD00662B938B32FF217535" box="[151,210,311,332]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">Leg 1</th>
<td id="AE58995CFFAD00662A758B32FE557535" box="[369,422,311,332]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">0-0</td>
<td id="AE58995CFFAD006629358B32FD9B7535" box="[561,616,311,332]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">I-1</td>
<td id="AE58995CFFAD0066281D8B32FB647535" box="[793,1175,311,332]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">I-0; I-0; I, I+2, 0 0-1; 0,2,1</td>
</tr>
<tr id="ED89F020FFAD00662B938B50FB647512" box="[151,1175,341,363]" gridrow="2" pageId="12" pageNumber="13">
<th id="AE58995CFFAD00662B938B50FF217512" box="[151,210,341,363]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">Leg 2</th>
<td id="AE58995CFFAD00662A758B50FE557512" box="[369,422,341,363]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">0-0</td>
<td id="AE58995CFFAD006629358B50FD9B7512" box="[561,616,341,363]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">I-0</td>
<td id="AE58995CFFAD0066281D8B50FB647512" box="[793,1175,341,363]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">I-0; I-1; I,I+2,1 0-0; 1,2,1</td>
</tr>
<tr id="ED89F020FFAD00662B938B71FB6475F3" box="[151,1175,372,394]" gridrow="3" pageId="12" pageNumber="13">
<th id="AE58995CFFAD00662B938B71FF2175F3" box="[151,210,372,394]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">Leg 3</th>
<td id="AE58995CFFAD00662A758B71FE5575F3" box="[369,422,372,394]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">0-0</td>
<td id="AE58995CFFAD006629358B71FD9B75F3" box="[561,616,372,394]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">1-0</td>
<td id="AE58995CFFAD0066281D8B71FB6475F3" box="[793,1175,372,394]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">I-0; I-1; II,2,2 0-0; 1,2,II</td>
</tr>
<tr id="ED89F020FFAD00662B938B90FB6475D3" box="[151,1175,405,426]" gridrow="4" pageId="12" pageNumber="13">
<th id="AE58995CFFAD00662B938B90FF2175D3" box="[151,210,405,426]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">Leg 4</th>
<td id="AE58995CFFAD00662A758B90FE5575D3" box="[369,422,405,426]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">0-0</td>
<td id="AE58995CFFAD006629358B90FD9B75D3" box="[561,616,405,426]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">1-0</td>
<td id="AE58995CFFAD0066281D8B90FB6475D3" box="[793,1175,405,426]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">I-0; I-1; I,II+1,2 0-0; 0,I+1,0</td>
</tr>
</table>
</paragraph>
<paragraph id="5306F262FFADFF952BCD8BFEFB887623" blockId="12.[151,1436,506,747]" pageId="12" pageNumber="13">
Leg 5 (
<figureCitation id="CB82EEE7FFADFF952A198BFFFE92766B" box="[285,353,506,530]" captionStart="FIGURE 6" captionStartId="8.[151,250,1726,1749]" captionTargetBox="[161,1416,214,1704]" captionTargetId="figure@8.[151,1436,193,1705]" captionTargetPageId="8" captionText="FIGURE 6. Attheyella (Canthosella) chocoensis n. sp. Female (holotype A, B and C). A, antennule; B, leg 5; C, inner spine of exopod; D, leg 5, inner spine of exopod, paratype NHMW 20560; E, leg 5, spine of exopod, paratype MHHN-Cp 2229." httpUri="https://zenodo.org/record/209251/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
B and 6C): baseoendopods separated at base, bearing 1 outer seta and 6 inner apical bipinnate setae; exopod with 1 bipinnate seta on outer margin, apically with 3 setae (two outermost naked, the third one bipinnate) and 1 inner spine, this spine modified, with 2 secondary spinules as in 
<figureCitation id="CB82EEE7FFADFF952F068847FB917623" box="[1026,1122,578,602]" captionStart="FIGURE 6" captionStartId="8.[151,250,1726,1749]" captionTargetBox="[161,1416,214,1704]" captionTargetId="figure@8.[151,1436,193,1705]" captionTargetPageId="8" captionText="FIGURE 6. Attheyella (Canthosella) chocoensis n. sp. Female (holotype A, B and C). A, antennule; B, leg 5; C, inner spine of exopod; D, leg 5, inner spine of exopod, paratype NHMW 20560; E, leg 5, spine of exopod, paratype MHHN-Cp 2229." httpUri="https://zenodo.org/record/209251/files/figure.png" pageId="12" pageNumber="13">Figure 6</figureCitation>
C.
</paragraph>
<paragraph id="5306F262FFADFF952BCD8862FD467693" blockId="12.[151,1436,506,747]" pageId="12" pageNumber="13">
Leg 6 (Fig. 3A): each reduced to a small plate, located posterior to seminal receptacle, bearing 1 seta. 
<emphasis id="61CD2E70FFADFF952BCD888FFE4E76DA" bold="true" box="[201,445,650,675]" pageId="12" pageNumber="13">Description of male.</emphasis>
Length of allotype 300 µm, exclusive of caudal setae. Mean length of 
<typeStatus id="8C024CC0FFADFF952FF5888EFAAE76DA" box="[1265,1373,651,675]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
from Nuquí 316 µm (n = 6, range 267-336 µm), mean length of 
<typeStatus id="8C024CC0FFADFF95283E88AAFC5676BE" box="[826,933,687,711]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
from Utria 274 µm (n = 15, range 238–327 µm). Differences from female are the following:
</paragraph>
<caption id="07C6A2EAFFADFF952B938DDFFAEC7389" box="[151,1311,2010,2033]" httpUri="https://zenodo.org/record/209280/files/figure.png" pageId="12" pageNumber="13" targetBox="[180,1390,802,1983]" targetPageId="12">
<paragraph id="5306F262FFADFF952B938DDFFAEC7389" blockId="12.[151,1311,2010,2033]" box="[151,1311,2010,2033]" pageId="12" pageNumber="13">
<emphasis id="61CD2E70FFADFF952B938DDFFEEC7389" bold="true" box="[151,287,2010,2033]" pageId="12" pageNumber="13">FIGURE 10.</emphasis>
<taxonomicName id="94B989E1FFADFF952A218DDFFD607389" box="[293,659,2010,2032]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFADFF952A218DDFFE7F7389" box="[293,396,2010,2032]" italics="true" pageId="12" pageNumber="13">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFADFF952A9F8DDFFDE67389" box="[411,533,2010,2032]" italics="true" pageId="12" pageNumber="13">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFADFF9529208DDFFD607389" box="[548,659,2010,2032]" italics="true" pageId="12" pageNumber="13">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFADFF95299D8DDEFD207389" box="[665,723,2011,2032]" pageId="12" pageNumber="13" rank="species">
<emphasis id="61CD2E70FFADFF95299D8DDEFD207389" bold="true" box="[665,723,2011,2032]" pageId="12" pageNumber="13">n. sp.</emphasis>
</taxonomicNameLabel>
Male (allotype). A, urosome, dorsal; B, urosome, ventral.
</paragraph>
</caption>
<caption id="07C6A2EAFFACFF942B938C8CFD5B72C7" httpUri="https://zenodo.org/record/209288/files/figure.png" pageId="13" pageNumber="14" targetBox="[224,1383,211,1639]" targetPageId="13">
<paragraph id="5306F262FFACFF942B938C8CFD5B72C7" blockId="13.[151,1436,1673,1726]" pageId="13" pageNumber="14">
<emphasis id="61CD2E70FFACFF942B938C8CFEEC72E7" bold="true" box="[151,287,1673,1696]" pageId="13" pageNumber="14">FIGURE 11.</emphasis>
Males. 
<taxonomicName id="94B989E1FFACFF942A768C8CFD4272E6" box="[370,689,1673,1695]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFACFF942A768C8CFE2A72E6" box="[370,473,1673,1695]" italics="true" pageId="13" pageNumber="14">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFACFF942AEC8C8CFD9172E6" box="[488,610,1673,1695]" italics="true" pageId="13" pageNumber="14">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFACFF9429768C8CFD4272E6" box="[626,689,1673,1695]" italics="true" pageId="13" pageNumber="14">aliena</emphasis>
</taxonomicName>
. A, antennule. 
<taxonomicName id="94B989E1FFACFF94284B8C8CFB4C72E6" box="[847,1215,1673,1695]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFACFF94284B8C8CFC4672E6" box="[847,949,1673,1695]" italics="true" pageId="13" pageNumber="14">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFACFF9428C28C8CFBB372E6" box="[966,1088,1673,1695]" italics="true" pageId="13" pageNumber="14">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFACFF942F548C8CFB4C72E6" box="[1104,1215,1673,1695]" italics="true" pageId="13" pageNumber="14">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFACFF942FC28C8FFB0D72E6" box="[1222,1278,1674,1695]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
(allotype, B, C and D). B, antennule; C, leg 5; D, leg 5, inner spine.
</paragraph>
</caption>
<paragraph id="5306F262FFACFF942BCD8CEFFEDD7317" blockId="13.[151,1436,1770,2011]" pageId="13" pageNumber="14">
Second urosomite (
<figureCitation id="CB82EEE7FFACFF942AA48CEFFE02737B" box="[416,497,1770,1794]" captionStart="FIGURE 10" captionStartId="12.[151,250,2010,2033]" captionTargetBox="[180,1390,802,1983]" captionTargetId="figure@12.[164,1423,789,2008]" captionTargetPageId="12" captionText="FIGURE 10. Attheyella (Canthosella) chocoensis n. sp. Male (allotype). A, urosome, dorsal; B, urosome, ventral." httpUri="https://zenodo.org/record/209280/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
B) posterolaterally and posteroventrally with row of long spinules. Anal somite with 2 lateral spines at each side of posterior outer corner (
<figureCitation id="CB82EEE7FFACFF9429F38D0BFCBA735F" box="[759,841,1806,1830]" captionStart="FIGURE 10" captionStartId="12.[151,250,2010,2033]" captionTargetBox="[180,1390,802,1983]" captionTargetId="figure@12.[164,1423,789,2008]" captionTargetPageId="12" captionText="FIGURE 10. Attheyella (Canthosella) chocoensis n. sp. Male (allotype). A, urosome, dorsal; B, urosome, ventral." httpUri="https://zenodo.org/record/209280/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
A). Anal operculum with 4 strongly developed triangular teeth. Caudal rami posterolaterally without row of spinules, in contrast to female, with short, poorly delimited dorsal carina.
</paragraph>
<paragraph id="5306F262FFACFF942BCD8D7FFB7E73A3" blockId="13.[151,1436,1770,2011]" pageId="13" pageNumber="14">
Antennule (
<figureCitation id="CB82EEE7FFACFF942A498D7FFE6F73EB" box="[333,412,1914,1938]" captionStart="FIGURE 11" captionStartId="13.[151,250,1673,1696]" captionTargetBox="[224,1383,211,1639]" captionTargetId="figure@13.[193,1394,193,1652]" captionTargetPageId="13" captionText="FIGURE 11. Males. Attheyella (Canthosella) aliena. A, antennule. Attheyella (Canthosella) chocoensis n. sp. (allotype, B, C and D). B, antennule; C, leg 5; D, leg 5, inner spine." httpUri="https://zenodo.org/record/209288/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
B) geniculate, 8-segmented, setation formula: 0, 3, 3, 6+aesthetasc and seta with conjoined bases, 0, 0, 0, 5+aesthetasc and seta with conjoined bases; seta and aesthetasc with conjoined bases of fourth segment almost as long as fifth and sixth segments together; fifth segment with two tubercles.
</paragraph>
<caption id="07C6A2EAFFAFFF972B938C8FFDDB72B9" httpUri="https://zenodo.org/record/209295/files/figure.png" pageId="14" pageNumber="15" targetBox="[191,1393,219,1628]" targetPageId="14">
<paragraph id="5306F262FFAFFF972B938C8FFDDB72B9" blockId="14.[151,1436,1674,1728]" pageId="14" pageNumber="15">
<emphasis id="61CD2E70FFAFFF972B938C8FFEEC72D9" bold="true" box="[151,287,1674,1697]" pageId="14" pageNumber="15">FIGURE 12.</emphasis>
<taxonomicName id="94B989E1FFAFFF972A218C8EFD6072D8" box="[293,659,1675,1697]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFAFFF972A218C8EFE7F72D8" box="[293,396,1675,1697]" italics="true" pageId="14" pageNumber="15">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFAFFF972A9F8C8EFDE672D8" box="[411,533,1675,1697]" italics="true" pageId="14" pageNumber="15">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFAFFF9729208C8EFD6072D8" box="[548,659,1675,1697]" italics="true" pageId="14" pageNumber="15">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFAFFF97299E8C8EFD2072D9" box="[666,723,1675,1696]" pageId="14" pageNumber="15" rank="species">
<emphasis id="61CD2E70FFAFFF97299E8C8EFD2072D9" bold="true" box="[666,723,1675,1696]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
Male (allotype, A and B). A, leg 1; B, leg 2; C, leg 2, first 2 segments of endopod of paratype NHMW 20684.
</paragraph>
</caption>
<paragraph id="5306F262FFAFFF972BCD8CE2FEB7735A" blockId="14.[151,1437,1767,2008]" pageId="14" pageNumber="15">Swimming legs showing differences with female, particularly in the armature of segments. Thus, they are described fully.</paragraph>
<paragraph id="5306F262FFAFFF962BCD8D35FF1C7546" blockId="14.[151,1437,1767,2008]" lastBlockId="15.[151,1437,151,1292]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Leg 1 (
<figureCitation id="CB82EEE7FFAFFF972A1A8D2AFE81733E" box="[286,370,1839,1863]" captionStart="FIGURE 12" captionStartId="14.[151,250,1674,1697]" captionTargetBox="[191,1393,219,1628]" captionTargetId="figure@14.[168,1419,193,1653]" captionTargetPageId="14" captionText="FIGURE 12. Attheyella (Canthosella) chocoensis n. sp. Male (allotype, A and B). A, leg 1; B, leg 2; C, leg 2, first 2 segments of endopod of paratype NHMW 20684." httpUri="https://zenodo.org/record/209295/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
A): intercoxal plate unarmed; coxa unarmed, except one spinule on outer margin; basis with outer unipinnate spine, anterior surface with proximal row of 4 spinules and 1 naked seta, with distal row of 6 spinules. Exopod 3-segmented; first segment with 1 bipinnate spine and row of spinules on outer margin and row of spinules near distal margin; second segment with 1 unipinnate spine and row of spinules on outer margin as well, and two spinules near distal margin; third segment with 1 unipinnate spine and a row of spinules on outer margin, apically with 1 unipinnate spine and 1 geniculate seta, inner margin with 1 geniculate seta. Endopod 2-segmented, as long as exopod; first segment with row of spinules on outer margin, with 1 long and strong bipinnate seta and 1 spinule on inner margin; second segment with row of spinules on outer margin, apical margin with 1 normally developed unipinnate seta and 1 geniculate long seta, inner margin with 1 naked seta inserted subapically and 1 spinule.
</paragraph>
<paragraph id="5306F262FFAEFF962BCD8B48FD6E7626" blockId="15.[151,1437,151,1292]" pageId="15" pageNumber="16">
Leg 2 (
<figureCitation id="CB82EEE7FFAEFF962A1D8B49FE87751D" box="[281,372,332,356]" captionStart="FIGURE 12" captionStartId="14.[151,250,1674,1697]" captionTargetBox="[191,1393,219,1628]" captionTargetId="figure@14.[168,1419,193,1653]" captionTargetPageId="14" captionText="FIGURE 12. Attheyella (Canthosella) chocoensis n. sp. Male (allotype, A and B). A, leg 1; B, leg 2; C, leg 2, first 2 segments of endopod of paratype NHMW 20684." httpUri="https://zenodo.org/record/209295/files/figure.png" pageId="15" pageNumber="16">Figs. 12</figureCitation>
B and 12C): intercoxal plate and coxa as in female. Basis with 1 outer bipinnate spine, anterior surface with 2 spinules near distal margin. Exopod 3-segmented; first segment with 1 bipinnate spine and row of spinules on outer and distal margins; second segment with 1 bipinnate spine and row of spinules on outer margin, with 1 naked seta and 2 spinules on inner margin; third segment with 1 bipinnate spine and 1 spinule on outer margin, apically with 1 bipinnate spine and 2 unipinnate setae, inner margin with 1 long seta, unipinnate at its end. Endopod 2-segmented; first segment short without armament; second segment with row of spinules and 1 naked spine inserted subapically on outer margin, with 1 long and 1 short setae (both bipinnate) on distal margin, with 1 unipinnate seta and 1 spinule on inner margin.
</paragraph>
<paragraph id="5306F262FFAEFF962BCD8868FE2A77DD" blockId="15.[151,1437,151,1292]" pageId="15" pageNumber="16">
Leg 3 (
<figureCitation id="CB82EEE7FFAEFF962A1A8869FE8976FD" box="[286,378,620,644]" captionStart="FIGURE 13" captionStartId="16.[151,250,1722,1745]" captionTargetBox="[180,1405,251,1708]" captionTargetId="figure@16.[171,1416,247,1713]" captionTargetPageId="16" captionText="FIGURE 13. Attheyella (Canthosella) chocoensis n. sp. Male (allotype, A and C). A, leg 3; C, leg 4; B, leg 3, last segment of endopod of paratype ICN-MHN 2222." httpUri="https://zenodo.org/record/209301/files/figure.png" pageId="15" pageNumber="16">Figs. 13</figureCitation>
A and 13B): intercoxal plate and coxa as in leg 2. Basis with 1 outer spine. Exopod 3-segmented; first segment with 1 outer spine and spinules, distally with two rows of surface spinules; second segment with 1 spine and spinules on outer margin, with 1 naked seta and 1 spinule on inner margin, with 1 apical surface spinule; third segment with two spines and spinules on outer margin, with 2 apical setae of different length, the inner the longest, bipinnate in its distal half, the outer unipinnate, with 2 setae on inner margin, proximal seta short, apical seta almost as long as longest terminal seta, both unipinnate at their distal part; outer spines of all exopod segments naked. Endopod 3-segmented; first segment very short and unarmed; second segment with long inner apophysis ending in a barb, as long as the terminal inner seta, third segment with 2 short apical setae, inner seta unipinnate, outer seta naked.
</paragraph>
<paragraph id="5306F262FFAEFF962BCD89B4FF1870BD" blockId="15.[151,1437,151,1292]" pageId="15" pageNumber="16">
Leg 4 (
<figureCitation id="CB82EEE7FFAEFF962A1F89AAFE9D77BE" box="[283,366,943,967]" captionStart="FIGURE 13" captionStartId="16.[151,250,1722,1745]" captionTargetBox="[180,1405,251,1708]" captionTargetId="figure@16.[171,1416,247,1713]" captionTargetPageId="16" captionText="FIGURE 13. Attheyella (Canthosella) chocoensis n. sp. Male (allotype, A and C). A, leg 3; C, leg 4; B, leg 3, last segment of endopod of paratype ICN-MHN 2222." httpUri="https://zenodo.org/record/209301/files/figure.png" pageId="15" pageNumber="16">Fig. 13</figureCitation>
C): intercoxal plate and coxa as in leg 2 and 3. Basis with one outer naked seta and 1 spinule. Exopod 3-segmented; first segment with 1 naked spine and spinules on outer margin; second segment with 1 naked spine and spinules on outer margin, with 1 naked spine and 1 spinule on inner margin, distally with two rows of surface spinules; third segment with 1 bipinnate spine on outer margin, with 2 spines (outer spine bipinnate, inner spine unipinnate) and 1 bipinnate seta on distal margin, inner margin with 2 setae, distal seta long and proximal seta short, both unipinnate. Endopod 2-segmented; first segment very short and unarmed, second segment with 1 bipinnate spine and 1 seta (unipinnate in its distal half) on apical margin, seta much shorter than corresponding seta of female.
</paragraph>
<paragraph id="5306F262FFAEFF962BCD8ED4FED37175" blockId="15.[151,1437,151,1292]" pageId="15" pageNumber="16">Legs 1–4 with following formula of spines (Roman numerals) and setae (Arabic). Spinules are not included in the formula.</paragraph>
<paragraph id="5306F262FFAEFF962A3D8F52FBAE725E" pageId="15" pageNumber="16">
<table id="21B900C2FFAE00662B938F53FBAE725E" box="[151,1117,1366,1575]" colsContinueIn="28.[151,1287,1192,1368]" gridcols="5" gridrows="5" pageId="15" pageNumber="16">
<tr id="ED89F020FFAE00662B938F53FBAE7115" box="[151,1117,1366,1388]" gridrow="0" pageId="15" pageNumber="16" rowspan-0="1">
<th id="AE58995CFFAE00662A3D8F53FE9D7115" box="[313,366,1366,1388]" gridcol="1" gridrow="0" pageId="15" pageNumber="16">Coxa</th>
<th id="AE58995CFFAE006629048F53FDC47115" box="[512,567,1366,1388]" gridcol="2" gridrow="0" pageId="15" pageNumber="16">Basis</th>
<th id="AE58995CFFAE006629BF8F53FC907115" box="[699,867,1366,1388]" gridcol="3" gridrow="0" pageId="15" pageNumber="16">Exopod</th>
<th id="AE58995CFFAE006628E18F53FBAE7115" box="[997,1117,1366,1388]" gridcol="4" gridrow="0" pageId="15" pageNumber="16">Endopod</th>
</tr>
<tr id="ED89F020FFAE00662B938FB1FBAE71B3" box="[151,1117,1460,1482]" gridrow="1" pageId="15" pageNumber="16">
<th id="AE58995CFFAE00662B938FB1FF2171B3" box="[151,210,1460,1482]" gridcol="0" gridrow="1" pageId="15" pageNumber="16">Leg 1</th>
<td id="AE58995CFFAE00662A3D8FB1FE9D71B3" box="[313,366,1460,1482]" gridcol="1" gridrow="1" pageId="15" pageNumber="16">0-0</td>
<td id="AE58995CFFAE006629048FB1FDC471B3" box="[512,567,1460,1482]" gridcol="2" gridrow="1" pageId="15" pageNumber="16">I-1</td>
<td id="AE58995CFFAE006629BF8FB1FC9071B3" box="[699,867,1460,1482]" gridcol="3" gridrow="1" pageId="15" pageNumber="16">I-0; I-0; I, I+1, 1</td>
<td id="AE58995CFFAE006628E18FB1FBAE71B3" box="[997,1117,1460,1482]" gridcol="4" gridrow="1" pageId="15" pageNumber="16">0-1; 0,2,1</td>
</tr>
<tr id="ED89F020FFAE00662B938FD6FBAE7191" box="[151,1117,1491,1512]" gridrow="2" pageId="15" pageNumber="16">
<th id="AE58995CFFAE00662B938FD6FF217191" box="[151,210,1491,1512]" gridcol="0" gridrow="2" pageId="15" pageNumber="16">Leg 2</th>
<td id="AE58995CFFAE00662A3D8FD6FE9D7191" box="[313,366,1491,1512]" gridcol="1" gridrow="2" pageId="15" pageNumber="16">0-0</td>
<td id="AE58995CFFAE006629048FD6FDC47191" box="[512,567,1491,1512]" gridcol="2" gridrow="2" pageId="15" pageNumber="16">I-0</td>
<td id="AE58995CFFAE006629BF8FD6FC907191" box="[699,867,1491,1512]" gridcol="3" gridrow="2" pageId="15" pageNumber="16">I-0; I-1; I,I+2,1</td>
<td id="AE58995CFFAE006628E18FD6FBAE7191" box="[997,1117,1491,1512]" gridcol="4" gridrow="2" pageId="15" pageNumber="16">0-0; I,2,1</td>
</tr>
<tr id="ED89F020FFAE00662B938FF7FBAE727E" box="[151,1117,1522,1543]" gridrow="3" pageId="15" pageNumber="16">
<th id="AE58995CFFAE00662B938FF7FF21727E" box="[151,210,1522,1543]" gridcol="0" gridrow="3" pageId="15" pageNumber="16">Leg 3</th>
<td id="AE58995CFFAE00662A3D8FF7FE9D727E" box="[313,366,1522,1543]" gridcol="1" gridrow="3" pageId="15" pageNumber="16">0-0</td>
<td id="AE58995CFFAE006629048FF7FDC4727E" box="[512,567,1522,1543]" gridcol="2" gridrow="3" pageId="15" pageNumber="16">I-0</td>
<td id="AE58995CFFAE006629BF8FF7FC90727E" box="[699,867,1522,1543]" gridcol="3" gridrow="3" pageId="15" pageNumber="16">I-0; I-1; II,2,2</td>
<td id="AE58995CFFAE006628E18FF7FBAE727E" box="[997,1117,1522,1543]" gridcol="4" gridrow="3" pageId="15" pageNumber="16">0-0; 0,2,0</td>
</tr>
<tr id="ED89F020FFAE00662B938C17FBAE725E" box="[151,1117,1554,1575]" gridrow="4" pageId="15" pageNumber="16">
<th id="AE58995CFFAE00662B938C17FF21725E" box="[151,210,1554,1575]" gridcol="0" gridrow="4" pageId="15" pageNumber="16">Leg 4</th>
<td id="AE58995CFFAE00662A3D8C17FE9D725E" box="[313,366,1554,1575]" gridcol="1" gridrow="4" pageId="15" pageNumber="16">0-0</td>
<td id="AE58995CFFAE006629048C17FDC4725E" box="[512,567,1554,1575]" gridcol="2" gridrow="4" pageId="15" pageNumber="16">1-0</td>
<td id="AE58995CFFAE006629BF8C17FC90725E" box="[699,867,1554,1575]" gridcol="3" gridrow="4" pageId="15" pageNumber="16">I-0; I-1; I,II+1,2</td>
<td id="AE58995CFFAE006628E18C17FBAE725E" box="[997,1117,1554,1575]" gridcol="4" gridrow="4" pageId="15" pageNumber="16">0-0; 0,I+1,0</td>
</tr>
</table>
</paragraph>
<paragraph id="5306F262FFAEFF962BCD8C7DFF1C72AE" blockId="15.[151,1437,1655,2004]" pageId="15" pageNumber="16">
Leg 5 (
<figureCitation id="CB82EEE7FFAEFF962A188C72FE8572F6" box="[284,374,1655,1679]" captionStart="FIGURE 11" captionStartId="13.[151,250,1673,1696]" captionTargetBox="[224,1383,211,1639]" captionTargetId="figure@13.[193,1394,193,1652]" captionTargetPageId="13" captionText="FIGURE 11. Males. Attheyella (Canthosella) aliena. A, antennule. Attheyella (Canthosella) chocoensis n. sp. (allotype, B, C and D). B, antennule; C, leg 5; D, leg 5, inner spine." httpUri="https://zenodo.org/record/209288/files/figure.png" pageId="15" pageNumber="16">Figs. 11</figureCitation>
C and 11 D): Baseoendopods fused at base, unarmed; exopod with 3 setae and 1 spine: outer seta short, middle and inner setae long, inner seta reaching apical margin of second urosomite; spine with lateral spinule.
</paragraph>
<paragraph id="5306F262FFAEFF892BCD8CE6FBA174AD" blockId="15.[151,1437,1655,2004]" lastBlockId="16.[151,1436,151,212]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="61CD2E70FFAEFF962BCD8CE6FEBC7285" bold="true" box="[201,335,1763,1788]" pageId="15" pageNumber="16">Variability.</emphasis>
Females. Dorsal side of genital double-somite partially separated, as is also the case ventrally (
<typeStatus id="8C024CC0FFAEFF962B9A8D02FEF27366" box="[158,257,1799,1823]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
NHMW 20560) (
<figureCitation id="CB82EEE7FFAEFF962ACF8D02FDFF7366" box="[459,524,1799,1823]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[270,1303,482,1909]" captionTargetId="figure@3.[267,1315,455,1918]" captionTargetPageId="3" captionText="FIGURE 1. Attheyella (Canthosella) chocoensis n. sp. Female (holotype): A, habitus, dorsal; B, habitus, lateral; C, genital double-somite in lateral view of paratype NHMW 20560, D, genital double-somite in lateral view of paratype MNHN-Cp 2299. Position of structures of C and D indicated by asterisk on B." httpUri="https://zenodo.org/record/209225/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
C). Anal somite with a variable number of spinules: laterally, 3, 4 as in 
<typeStatus id="8C024CC0FFAEFF962E3E8D02FA6F7366" box="[1338,1436,1799,1823]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
or 5 as in specimen of 
<figureCitation id="CB82EEE7FFAEFF962A9E8D29FE09733D" box="[410,506,1836,1860]" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[330,1186,210,1917]" captionTargetId="figure@4.[302,1199,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Attheyella (Canthosella) chocoensis n. sp. Female (paratype from Utria). SEM photographs. A, habitus, dorsal view; B, anal somite and caudal rami, dorsal (most right teeth of anal operculum broken); C, caudal ramus in lateral view." httpUri="https://zenodo.org/record/209232/files/figure.png" pageId="15" pageNumber="16">Figure 2</figureCitation>
C; ventrally, 7 as in 
<typeStatus id="8C024CC0FFAEFF9629DB8D29FCB7733D" box="[735,836,1836,1860]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, 8 as in 
<typeStatus id="8C024CC0FFAEFF9628A78D28FBF7733C" box="[931,1028,1837,1861]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
NHMW 20682 (
<figureCitation id="CB82EEE7FFAEFF962FC18D29FAFB733D" box="[1221,1288,1836,1860]" captionStart="FIGURE 5" captionStartId="7.[151,250,1614,1637]" captionTargetBox="[238,1339,208,1584]" captionTargetId="figure@7.[227,1360,193,1593]" captionTargetPageId="7" captionText="FIGURE 5. Attheyella (Canthosella) chocoensis n. sp. Female (paratype NHMW 20682). A, caudal rami with anal operculum, dorsal; B, caudal ramus, ventral; C, caudal ramus, lateral." httpUri="https://zenodo.org/record/209245/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
B), to 9 as in 
<typeStatus id="8C024CC0FFAEFF962B938D54FF047310" box="[151,247,1873,1897]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
MNHN-Cp 2296. Leg 1, second segment of exopod with 1 or 2 spinules on inner margin. Number of spines of anal operculum variable, from 5 to 10 (
<figureCitation id="CB82EEE7FFAEFF9629B08D71FCF173F5" box="[692,770,1908,1932]" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[330,1186,210,1917]" captionTargetId="figure@4.[302,1199,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Attheyella (Canthosella) chocoensis n. sp. Female (paratype from Utria). SEM photographs. A, habitus, dorsal view; B, anal somite and caudal rami, dorsal (most right teeth of anal operculum broken); C, caudal ramus in lateral view." httpUri="https://zenodo.org/record/209232/files/figure.png" pageId="15" pageNumber="16">Figs. 2</figureCitation>
B, 3B and 5A). Spine of exopod of leg 5 with variable number (2 to 4) of secondary spines (
<figureCitation id="CB82EEE7FFAEFF9629148D92FD9373D6" box="[528,608,1943,1967]" captionStart="FIGURE 6" captionStartId="8.[151,250,1726,1749]" captionTargetBox="[161,1416,214,1704]" captionTargetId="figure@8.[151,1436,193,1705]" captionTargetPageId="8" captionText="FIGURE 6. Attheyella (Canthosella) chocoensis n. sp. Female (holotype A, B and C). A, antennule; B, leg 5; C, inner spine of exopod; D, leg 5, inner spine of exopod, paratype NHMW 20560; E, leg 5, spine of exopod, paratype MHHN-Cp 2229." httpUri="https://zenodo.org/record/209251/files/figure.png" pageId="15" pageNumber="16">Figs. 6</figureCitation>
C, 6D and 6E). Males. Exopod of leg 2 of one 
<typeStatus id="8C024CC0FFAEFF962F738D9CFB2473C8" box="[1143,1239,1945,1969]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
(NHMW 20684) with 2 setae on inner margin (
<figureCitation id="CB82EEE7FFAEFF962AE98DB9FDCD73AD" box="[493,574,1980,2004]" captionStart="FIGURE 12" captionStartId="14.[151,250,1674,1697]" captionTargetBox="[191,1393,219,1628]" captionTargetId="figure@14.[168,1419,193,1653]" captionTargetPageId="14" captionText="FIGURE 12. Attheyella (Canthosella) chocoensis n. sp. Male (allotype, A and B). A, leg 1; B, leg 2; C, leg 2, first 2 segments of endopod of paratype NHMW 20684." httpUri="https://zenodo.org/record/209295/files/figure.png" pageId="15" pageNumber="16">Fig. 12</figureCitation>
C) in contrast to allotype (1 seta). Endopod of leg 3 with 1 short and 1 long seta (
<typeStatus id="8C024CC0FFB1FF892BD68A92FEC774D6" box="[210,308,151,175]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
ICN-MHN-CR 2222) (
<figureCitation id="CB82EEE7FFB1FF89293B8A92FD7C74D6" box="[575,655,151,175]" captionStart="FIGURE 13" captionStartId="16.[151,250,1722,1745]" captionTargetBox="[180,1405,251,1708]" captionTargetId="figure@16.[171,1416,247,1713]" captionTargetPageId="16" captionText="FIGURE 13. Attheyella (Canthosella) chocoensis n. sp. Male (allotype, A and C). A, leg 3; C, leg 4; B, leg 3, last segment of endopod of paratype ICN-MHN 2222." httpUri="https://zenodo.org/record/209301/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
A) instead of 2 short setae as in allotype. Number of lateral spines of anal somite varies from 2 to 5. Number of teeth of anal operculum varies from 3 to 8.
</paragraph>
</subSubSection>
<caption id="07C6A2EAFFB1FF892B938CBFFDEF7296" httpUri="https://zenodo.org/record/209301/files/figure.png" pageId="16" pageNumber="17" targetBox="[180,1405,251,1708]" targetPageId="16">
<paragraph id="5306F262FFB1FF892B938CBFFDEF7296" blockId="16.[151,1436,1722,1775]" pageId="16" pageNumber="17">
<emphasis id="61CD2E70FFB1FF892B938CBFFED372A9" bold="true" box="[151,288,1722,1745]" pageId="16" pageNumber="17">FIGURE 13.</emphasis>
<taxonomicName id="94B989E1FFB1FF892A228CBEFD6772A8" box="[294,660,1723,1745]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov." subGenus="Canthosella">
<emphasis id="61CD2E70FFB1FF892A228CBEFE7F72A8" box="[294,396,1723,1745]" italics="true" pageId="16" pageNumber="17">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB1FF892A988CBEFDE572A8" box="[412,534,1723,1745]" italics="true" pageId="16" pageNumber="17">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB1FF8929218CBEFD6772A8" box="[549,660,1723,1745]" italics="true" pageId="16" pageNumber="17">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFB1FF89299F8CBEFD2672A9" box="[667,725,1723,1744]" pageId="16" pageNumber="17" rank="species">
<emphasis id="61CD2E70FFB1FF89299F8CBEFD2672A9" bold="true" box="[667,725,1723,1744]" pageId="16" pageNumber="17">n. sp.</emphasis>
</taxonomicNameLabel>
Male (allotype, A and C). A, leg 3; C, leg 4; B, leg 3, last segment of endopod of paratype ICN-MHN 2222.
</paragraph>
</caption>
<subSubSection id="1BA3A1E9FFB1FF882BCD8D18FDAD7645" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="discussion">
<paragraph id="5306F262FFB1FF882BCD8D18FDAD7645" blockId="16.[151,1437,1820,2024]" lastBlockId="17.[151,1436,151,572]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="61CD2E70FFB1FF892BCD8D18FECE734C" bold="true" box="[201,317,1821,1845]" pageId="16" pageNumber="17">Remarks.</emphasis>
Within the subgenus 
<taxonomicName id="94B989E1FFB1FF8929368D19FD4A734D" box="[562,697,1820,1844]" class="Maxillopoda" family="Canthocamptidae" genus="Canthosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="61CD2E70FFB1FF8929368D19FD4A734D" box="[562,697,1820,1844]" italics="true" pageId="16" pageNumber="17">Canthosella</emphasis>
</taxonomicName>
, 16 species have been described so far, 11 of them from the Neotropical region and 5 from the Oriental region (
<tableCitation id="1E3BC7D9FFB1FF8929B08D45FCFA7321" box="[692,777,1856,1880]" captionStart="TABLE 1" captionStartId="17.[151,235,620,641]" captionTargetBox="[151,1436,730,1238]" captionTargetPageId="17" captionText="TABLE 1. List of species of the subgenus Canthosella, taxonomic status, known (+) and unknown (-) females and males, and geographic distribution. * species belonging to the group bearing a row of spines on the dorsal surface of the caudal ramus. ** species relocated by Janetzky et al. (1996)" httpUri="http://table.plazi.org/id/07C6A2EAFFB0FF882B938869FDB776C6" pageId="16" pageNumber="17" tableUuid="07C6A2EAFFB0FF882B938869FDB776C6">Table 1</tableCitation>
). We accept the relocation of 5 species into the subgenus 
<taxonomicName id="94B989E1FFB1FF892B938D61FEED7305" box="[151,286,1892,1916]" class="Maxillopoda" family="Canthocamptidae" genus="Canthosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="61CD2E70FFB1FF892B938D61FEED7305" box="[151,286,1892,1916]" italics="true" pageId="16" pageNumber="17">Canthosella</emphasis>
</taxonomicName>
as suggested by 
<bibRefCitation id="37288F93FFB1FF892ADB8D61FD207305" box="[479,723,1892,1916]" pageId="16" pageNumber="17" refString="Janetzky, W., Martinez Arbizu, P. &amp; Reid, J. W. (1996) Attheyella (Canthosella) mervini n. sp. (Canthocamptidae, Harpacticoida) from Jamaican bromeliads. Hydrobiologia, 339, 123 - 135." type="journal article">
Janetzky 
<emphasis id="61CD2E70FFB1FF89294C8D60FD8A7305" box="[584,633,1892,1916]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
. (1996)
</bibRefCitation>
based on the definition of the subgenus, particularly the posterior margins of the body somites being smooth, the segmentation and armature of legs 1–4 and the ornamentation of the anal operculum. The relocated species are 
<taxonomicName id="94B989E1FFB1FF8929BE8DA9FB5F73BD" authority="Noodt, 1956" authorityName="Noodt" authorityYear="1956" box="[698,1196,1964,1988]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB1FF8929BE8DA9FCD873BD" box="[698,811,1964,1988]" italics="true" pageId="16" pageNumber="17">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB1FF8928388DA9FC3073BD" box="[828,963,1964,1988]" italics="true" pageId="16" pageNumber="17">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB1FF8928D78DA9FBEA73BD" box="[979,1049,1964,1988]" italics="true" pageId="16" pageNumber="17">aliena</emphasis>
<bibRefCitation id="37288F93FFB1FF892F248DA9FB5F73BD" author="Noodt" box="[1056,1196,1964,1988]" pageId="16" pageNumber="17" refString="Noodt, W. (1956) Attheyella (Chappuisiella) aliena n. sp., ein Copepode tropischer Verwandtschaft aus Phytotelmen des Gottinger Gewasserhauses. Gewasser und Abwasser, 14, 62 - 69." type="journal article" year="1956">Noodt, 1956</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB1FF892FE68DA9FDD17391" authority="Delachaux, 1924" authorityName="Delachaux" authorityYear="1924" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="kalima" subGenus="Canthosella">
<emphasis id="61CD2E70FFB1FF892FE68DA9FAA073BD" box="[1250,1363,1964,1988]" italics="true" pageId="16" pageNumber="17">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB1FF892E608DA8FF037391" italics="true" pageId="16" pageNumber="17">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB1FF892BFB8DD5FEB87391" box="[255,331,2000,2024]" italics="true" pageId="16" pageNumber="17">kalima</emphasis>
(
<bibRefCitation id="37288F93FFB1FF892A5E8DD5FDEA7391" author="Delachaux" box="[346,537,2000,2024]" pageId="16" pageNumber="17" refString="Delachaux, T. (1924) Zur Kenntnis der Copepodenfauna von Surinam. II. Harpacticiden. Zoologischer Anzeiger, 59, 1 - 16." type="journal article" year="1924">Delachaux, 1924</bibRefCitation>
)
</taxonomicName>
from the subgenus 
<taxonomicName id="94B989E1FFB1FF8929F88DD5FC6A7391" box="[764,921,2000,2024]" class="Maxillopoda" family="Canthocamptidae" genus="Chappuisiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="61CD2E70FFB1FF8929F88DD5FC6A7391" box="[764,921,2000,2024]" italics="true" pageId="16" pageNumber="17">Chappuisiella</emphasis>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB1FF8928A38DD5FA6F7391" authority="Kiefer, 1967" authorityName="Kiefer" authorityYear="1967" box="[935,1436,2000,2024]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="siolii" subGenus="Canthosella">
<emphasis id="61CD2E70FFB1FF8928A38DD5FBEA7391" box="[935,1049,2000,2024]" italics="true" pageId="16" pageNumber="17">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB1FF892F2C8DD5FB5C7391" box="[1064,1199,2000,2024]" italics="true" pageId="16" pageNumber="17">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB1FF892FBB8DD5FB047391" box="[1215,1271,2000,2024]" italics="true" pageId="16" pageNumber="17">siolii</emphasis>
(
<bibRefCitation id="37288F93FFB1FF892E018DD5FA607391" author="Kiefer" box="[1285,1427,2000,2024]" pageId="16" pageNumber="17" refString="Kiefer, F. (1967) Neue Copepoda Harpacticoida aus dem Amazonasgebiet. Crustaceana, 13, 114 - 122." type="journal article" year="1967">Kiefer, 1967</bibRefCitation>
)
</taxonomicName>
from the genus 
<taxonomicName id="94B989E1FFB0FF882A558A9DFD5074D6" authority="Chappuis, 1929" authorityName="Chappuis" authorityYear="1929" box="[337,675,151,176]" class="Maxillopoda" family="Canthocamptidae" genus="Elaphoidella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="61CD2E70FFB0FF882A558A9DFE1174C9" box="[337,482,152,176]" italics="true" pageId="17" pageNumber="18">Elaphoidella</emphasis>
<bibRefCitation id="37288F93FFB0FF882AE98A92FD5074D6" author="Chappuis" box="[493,675,151,175]" pageId="17" pageNumber="18" refString="Chappuis, P. A. (1929) Revision du genre Canthocamptus Westwood (Note preliminaire). Buletinul Societatii de Stiinte din Cluj, 4, 2, 41 - 50." type="journal article" year="1929">Chappuis, 1929</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB0FF8829B68A9DFB2474D6" authority="Reid, 1990" authorityName="Reid" authorityYear="1990" box="[690,1239,151,176]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="striblingi" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF8829B68A9DFCD774C9" box="[690,804,152,176]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF88283C8A9DFC4C74C9" box="[824,959,152,176]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF8828D08A9DFBC874C9" box="[980,1083,152,176]" italics="true" pageId="17" pageNumber="18">striblingi</emphasis>
(
<bibRefCitation id="37288F93FFB0FF882F498A92FB3D74D6" author="Reid" box="[1101,1230,151,175]" pageId="17" pageNumber="18" refString="Reid, J. - W. (1990) Canthocamptus (Elaphoidella) striblingi, new species (Copepoda: Harpacticoida) from Costa Rica. Proceedings of the Biological Society of Washington, 103, 336 - 340." type="journal article" year="1990">Reid, 1990</bibRefCitation>
)
</taxonomicName>
originally called 
<taxonomicName id="94B989E1FFB0FF882B938AB9FD9874AD" box="[151,619,188,212]" class="Maxillopoda" family="Canthocamptidae" genus="Canthocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="striblingi" subGenus="Elaphoidella">
<emphasis id="61CD2E70FFB0FF882B938AB9FEB474AD" box="[151,327,188,212]" italics="true" pageId="17" pageNumber="18">Canthocamptus</emphasis>
(
<emphasis id="61CD2E70FFB0FF882A598AB9FE1D74AD" box="[349,494,188,212]" italics="true" pageId="17" pageNumber="18">Elaphoidella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF8829008AB9FD9874AD" box="[516,619,188,212]" italics="true" pageId="17" pageNumber="18">striblingi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="94B989E1FFB0FF8829B18AB9FEEB748E" authority="Janetzky, Martinez Arbizu Reid, 1996" authorityName="Janetzky, Martinez Arbizu Reid" authorityYear="1996" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF8829B18AB9FCD474AD" box="[693,807,188,212]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF8828398AB9FC3774AD" box="[829,964,188,212]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF8828DF8AB9FBA974AD" box="[987,1114,188,212]" italics="true" pageId="17" pageNumber="18">pilagaensis</emphasis>
<bibRefCitation id="37288F93FFB0FF882F638AB9FEEB748E" author="Janetzky" pageId="17" pageNumber="18" refString="Janetzky, W., Martinez Arbizu, P. &amp; Reid, J. W. (1996) Attheyella (Canthosella) mervini n. sp. (Canthocamptidae, Harpacticoida) from Jamaican bromeliads. Hydrobiologia, 339, 123 - 135." type="journal article" year="1996">Janetzky, Martínez Arbizu Reid, 1996</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="94B989E1FFB0FF882A208AE5FD317481" box="[292,706,224,248]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF882A208AE5FD317481" box="[292,706,224,248]" italics="true" pageId="17" pageNumber="18">Attheyella (Canthosella) pilagaensis</emphasis>
</taxonomicName>
was proposed by 
<bibRefCitation id="37288F93FFB0FF8828938ADAFB62748E" box="[919,1169,223,248]" pageId="17" pageNumber="18" refString="Janetzky, W., Martinez Arbizu, P. &amp; Reid, J. W. (1996) Attheyella (Canthosella) mervini n. sp. (Canthocamptidae, Harpacticoida) from Jamaican bromeliads. Hydrobiologia, 339, 123 - 135." type="journal article">
Janetzky 
<emphasis id="61CD2E70FFB0FF882F068AE4FBC57481" box="[1026,1078,224,248]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1996)
</bibRefCitation>
as a new name for the female specimen of 
<taxonomicName id="94B989E1FFB0FF882A878B01FCF67565" box="[387,773,260,284]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kalima" subGenus="Chappuisiella">
<emphasis id="61CD2E70FFB0FF882A878B01FE067565" box="[387,501,260,284]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF88290C8B01FD567565" box="[520,677,260,284]" italics="true" pageId="17" pageNumber="18">Chappuisiella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF8829BD8B01FCF67565" box="[697,773,260,284]" italics="true" pageId="17" pageNumber="18">kalima</emphasis>
</taxonomicName>
identified by 
<bibRefCitation id="37288F93FFB0FF8828AF8B00FB297565" author="Dussart" box="[939,1242,260,284]" pageId="17" pageNumber="18" refString="Dussart, B. H. &amp; Frutos, S. M. (1986) Sur quelques Copepodes d`Argentine. 2. Copepodes du Parana Medio. Revue d'Hydrobiologie tropical e, 19, 241 - 262." type="journal article" year="1986">Dussart and Frutos (1986)</bibRefCitation>
from 
<collectingCountry id="2BAEB2F2FFB0FF882E228B01FA6B7565" box="[1318,1432,260,284]" name="Argentina" pageId="17" pageNumber="18">Argentina</collectingCountry>
, based on the differences (according to Delachaux’s description) in the insertion point of the lateral caudal setae and in the relative lengths of several setae of the swimming legs of females. We agree with this statement because the type-material of 
<taxonomicName id="94B989E1FFB0FF882A558B75FD4475F1" box="[337,695,367,392]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kalima" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF882A558B75FE3075F1" box="[337,451,368,392]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF882AD78B75FDA975F1" box="[467,602,368,392]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF88296F8B75FD4475F1" box="[619,695,368,392]" italics="true" pageId="17" pageNumber="18">kalima</emphasis>
</taxonomicName>
from 
<collectingCountry id="2BAEB2F2FFB0FF8829F88B6AFC9475FE" box="[764,871,367,391]" name="Suriname" pageId="17" pageNumber="18">Suriname</collectingCountry>
has not been re-examined, but this synonymy will have to be confirmed subsequently with the description of new material. Both species, 
<taxonomicName id="94B989E1FFB0FF882F588B91FE1B75B6" authority="Janetzky et al., 1996" authorityName="Janetzky et al." authorityYear="1996" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF882F588B91FB3E75D5" box="[1116,1229,404,428]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF882FDA8B91FA9675D5" box="[1246,1381,404,428]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF882E718B91FF0A75A9" italics="true" pageId="17" pageNumber="18">pilagaensis</emphasis>
<bibRefCitation id="37288F93FFB0FF882A048BB2FE1B75B6" author="Janetzky" box="[256,488,439,464]" pageId="17" pageNumber="18" refString="Janetzky, W., Martinez Arbizu, P. &amp; Reid, J. W. (1996) Attheyella (Canthosella) mervini n. sp. (Canthocamptidae, Harpacticoida) from Jamaican bromeliads. Hydrobiologia, 339, 123 - 135." type="journal article" year="1996">
Janetzky 
<emphasis id="61CD2E70FFB0FF882A6D8BBCFE6875A9" box="[361,411,440,464]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
., 1996
</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB0FF88291A8BBDFC7675A9" box="[542,901,439,464]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kalima" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF88291A8BBDFD6375A9" box="[542,656,440,464]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF8829A58BBDFCDB75A9" box="[673,808,440,464]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF88283D8BBDFC7675A9" box="[825,901,440,464]" italics="true" pageId="17" pageNumber="18">kalima</emphasis>
</taxonomicName>
, are then considered as belonging to the subgenus 
<emphasis id="61CD2E70FFB0FF882BCD8BD9FEAB758D" box="[201,344,476,500]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BCD8BD9FEA7758D" box="[201,340,476,500]" class="Maxillopoda" family="Canthocamptidae" genus="Canthosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Canthosella</taxonomicName>
.
</emphasis>
The three species 
<taxonomicName id="94B989E1FFB0FF88293F8BD9FC17758D" box="[571,996,476,500]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bromelicola" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF88293F8BD9FD5E758D" box="[571,685,476,500]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF8829C68BD9FCBA758D" box="[706,841,476,500]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF88285A8BD9FC17758D" box="[862,996,476,500]" italics="true" pageId="17" pageNumber="18">bromelicola</emphasis>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB0FF8828F38BD9FA9B758D" box="[1015,1384,476,500]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="goeldii" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF8828F38BD9FB9B758D" box="[1015,1128,476,500]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF882F798BD9FAF7758D" box="[1149,1284,476,500]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF882E1E8BD9FA9B758D" box="[1306,1384,476,500]" italics="true" pageId="17" pageNumber="18">goeldii</emphasis>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB0FF882B938805FDE77661" box="[151,532,511,536]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="montana" subGenus="Canthosella">
<emphasis id="61CD2E70FFB0FF882B938805FEFB7661" box="[151,264,512,536]" italics="true" pageId="17" pageNumber="18">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB0FF882A1E8805FE527661" box="[282,417,512,536]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB0FF882AB68804FDE77661" box="[434,532,513,536]" italics="true" pageId="17" pageNumber="18">montana</emphasis>
</taxonomicName>
described by 
<bibRefCitation id="37288F93FFB0FF8829B18BFAFCB7766E" author="Ebert" box="[693,836,511,535]" pageId="17" pageNumber="18" refString="Ebert, S. (1976) Naturliches System und Biogeographie der Familie Canthocamptidae (Copepoda, Harpacticoida) am Beispiel neotropischer Taxa. Dissertation Christian-Albrechts-Universitat Kiel, 325 pp." type="book" year="1976">Ebert (1976)</bibRefCitation>
have to be considered as unavailable names because the thesis was never published (
<tableCitation id="1E3BC7D9FFB0FF882AFE8821FDBE7645" box="[506,589,548,572]" captionStart="TABLE 1" captionStartId="17.[151,235,620,641]" captionTargetBox="[151,1436,730,1238]" captionTargetPageId="17" captionText="TABLE 1. List of species of the subgenus Canthosella, taxonomic status, known (+) and unknown (-) females and males, and geographic distribution. * species belonging to the group bearing a row of spines on the dorsal surface of the caudal ramus. ** species relocated by Janetzky et al. (1996)" httpUri="http://table.plazi.org/id/07C6A2EAFFB0FF882B938869FDB776C6" pageId="17" pageNumber="18" tableUuid="07C6A2EAFFB0FF882B938869FDB776C6">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="07C6A2EAFFB0FF882B938869FDB776C6" ID-Table-UUID="07C6A2EAFFB0FF882B938869FDB776C6" httpUri="http://table.plazi.org/id/07C6A2EAFFB0FF882B938869FDB776C6" pageId="17" pageNumber="18" targetBox="[151,1436,730,1238]" targetIsTable="true" targetPageId="17">
<paragraph id="5306F262FFB0FF882B938869FDB776C6" blockId="17.[151,1436,619,703]" pageId="17" pageNumber="18">
<emphasis id="61CD2E70FFB0FF882B938869FEF676F8" bold="true" box="[151,261,620,641]" pageId="17" pageNumber="18">TABLE 1.</emphasis>
List of species of the subgenus 
<taxonomicName id="94B989E1FFB0FF8829488869FD3576FB" box="[588,710,620,642]" class="Maxillopoda" family="Canthocamptidae" genus="Canthosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="61CD2E70FFB0FF8829488869FD3576FB" box="[588,710,620,642]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
</taxonomicName>
, taxonomic status, known (+) and unknown (-) females and males, and geographic distribution. * species belonging to the group bearing a row of spines on the dorsal surface of the caudal ramus. ** species relocated by Janetzky 
<emphasis id="61CD2E70FFB0FF882ACC88AFFE0876C6" box="[456,507,681,703]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(1996)
</paragraph>
</caption>
<paragraph id="5306F262FFB0FF882B9B88DEFA9270AF" pageId="17" pageNumber="18">
<table id="21B900C2FFB000662B9388DFFA6F70AF" box="[151,1436,730,1238]" gridcols="6" gridrows="11" pageId="17" pageNumber="18">
<tr id="ED89F020FFB000662B9388DFFA6F7689" box="[151,1436,730,752]" gridrow="0" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B9388DFFF4C7689" box="[151,191,730,752]" gridcol="0" gridrow="0" pageId="17" pageNumber="18">Nr.</th>
<th id="AE58995CFFB000662BDF88DFFD3A7689" box="[219,713,730,752]" gridcol="1" gridrow="0" pageId="17" pageNumber="18">Species</th>
<th id="AE58995CFFB00066281488DFFC697689" box="[784,922,730,752]" gridcol="2" gridrow="0" pageId="17" pageNumber="18">Status</th>
<th id="AE58995CFFB0006628AB88DFFC0A7689" box="[943,1017,730,752]" gridcol="3" gridrow="0" pageId="17" pageNumber="18">Female</th>
<th id="AE58995CFFB000662F1788DFFBB47689" box="[1043,1095,730,752]" gridcol="4" gridrow="0" pageId="17" pageNumber="18">Male</th>
<th id="AE58995CFFB000662F6588DFFA6F7689" box="[1121,1436,730,752]" gridcol="5" gridrow="0" pageId="17" pageNumber="18">Distribution</th>
</tr>
<tr id="ED89F020FFB000662B938906FA6F7741" box="[151,1436,771,824]" gridrow="1" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938906FF4C7741" box="[151,191,771,824]" gridcol="0" gridrow="1" pageId="17" pageNumber="18">1</th>
<td id="AE58995CFFB000662BDF8906FD3A7741" box="[219,713,771,824]" gridcol="1" gridrow="1" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF8901FE6D7760" authority="Noodt, 1956" authorityName="Noodt" authorityYear="1956" box="[219,414,771,794]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="aliena">
<emphasis id="61CD2E70FFB0FF882BDF8901FEE97763" box="[219,282,772,794]" italics="true" pageId="17" pageNumber="18">aliena</emphasis>
Noodt, 1956
</taxonomicName>
*
</td>
<td id="AE58995CFFB0006628148906FC697741" box="[784,922,771,824]" gridcol="2" gridrow="1" pageId="17" pageNumber="18">relocated **</td>
<td id="AE58995CFFB0006628AB8906FC0A7741" box="[943,1017,771,824]" gridcol="3" gridrow="1" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F178906FBB47741" box="[1043,1095,771,824]" gridcol="4" gridrow="1" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F658906FA6F7741" box="[1121,1436,771,824]" gridcol="5" gridrow="1" pageId="17" pageNumber="18">neotropical (Brazil), introduced into Germany</td>
</tr>
<tr id="ED89F020FFB000662B93894EFA6F771B" box="[151,1436,843,866]" gridrow="2" pageId="17" pageNumber="18" rowspan-2="1">
<th id="AE58995CFFB000662B93894EFF4C771B" box="[151,191,843,866]" gridcol="0" gridrow="2" pageId="17" pageNumber="18">2</th>
<td id="AE58995CFFB000662BDF894EFD3A771B" box="[219,713,843,866]" gridcol="1" gridrow="2" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF8949FE19771B" authority="Petkovski, 1973" authorityName="Petkovski" authorityYear="1973" box="[219,490,843,866]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="antillica">
<emphasis id="61CD2E70FFB0FF882BDF8949FEDC771B" box="[219,303,844,866]" italics="true" pageId="17" pageNumber="18">antillica</emphasis>
(Petkovski, 1973)
</taxonomicName>
</td>
<td id="AE58995CFFB0006628AB894EFC0A771B" box="[943,1017,843,866]" gridcol="3" gridrow="2" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F17894EFBB4771B" box="[1043,1095,843,866]" gridcol="4" gridrow="2" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F65894EFA6F771B" box="[1121,1436,843,866]" gridcol="5" gridrow="2" pageId="17" pageNumber="18">neotropical (Cuba)</td>
</tr>
<tr id="ED89F020FFB000662B938970FA6F77F2" box="[151,1436,885,907]" gridrow="3" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938970FF4C77F2" box="[151,191,885,907]" gridcol="0" gridrow="3" pageId="17" pageNumber="18">3</th>
<td id="AE58995CFFB000662BDF8970FD3A77F2" box="[219,713,885,907]" gridcol="1" gridrow="3" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF8970FE2277F2" authority="Ebert, 1976" authorityName="Ebert" authorityYear="1976" box="[219,465,885,907]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bromelicola">
<emphasis id="61CD2E70FFB0FF882BDF8970FEA677F2" box="[219,341,885,907]" italics="true" pageId="17" pageNumber="18">bromelicola</emphasis>
Ebert, 1976
</taxonomicName>
</td>
<td id="AE58995CFFB0006628148970FC6977F2" box="[784,922,885,907]" gridcol="2" gridrow="3" pageId="17" pageNumber="18">not published</td>
<td id="AE58995CFFB0006628AB8970FC0A77F2" box="[943,1017,885,907]" gridcol="3" gridrow="3" pageId="17" pageNumber="18">-</td>
<td id="AE58995CFFB000662F178970FBB477F2" box="[1043,1095,885,907]" gridcol="4" gridrow="3" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F658970FA6F77F2" box="[1121,1436,885,907]" gridcol="5" gridrow="3" pageId="17" pageNumber="18">neotropical (Brazil)</td>
</tr>
<tr id="ED89F020FFB000662B93899BFA6F77CC" box="[151,1436,926,949]" gridrow="4" pageId="17" pageNumber="18" rowspan-2="1">
<th id="AE58995CFFB000662B93899BFF4C77CC" box="[151,191,926,949]" gridcol="0" gridrow="4" pageId="17" pageNumber="18">4</th>
<td id="AE58995CFFB000662BDF899BFD3A77CC" box="[219,713,926,949]" gridcol="1" gridrow="4" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF899AFEB977CC" box="[219,330,927,949]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="chocoensis" status="sp. nov.">
<emphasis id="61CD2E70FFB0FF882BDF899AFEB977CC" box="[219,330,927,949]" italics="true" pageId="17" pageNumber="18">chocoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AFE930BFFB0FF882A54899AFE7477CD" box="[336,391,927,948]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
*
</td>
<td id="AE58995CFFB0006628AB899BFC0A77CC" box="[943,1017,926,949]" gridcol="3" gridrow="4" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F17899BFBB477CC" box="[1043,1095,926,949]" gridcol="4" gridrow="4" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F65899BFA6F77CC" box="[1121,1436,926,949]" gridcol="5" gridrow="4" pageId="17" pageNumber="18">neotropical (Colombia)</td>
</tr>
<tr id="ED89F020FFB000662B9389CDFA6F77A7" box="[151,1436,968,990]" gridrow="5" pageId="17" pageNumber="18" rowspan-2="1">
<th id="AE58995CFFB000662B9389CDFF4C77A7" box="[151,191,968,990]" gridcol="0" gridrow="5" pageId="17" pageNumber="18">5</th>
<td id="AE58995CFFB000662BDF89CDFD3A77A7" box="[219,713,968,990]" gridcol="1" gridrow="5" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF89CDFE1377A4" authority="Chappuis, 1931" authorityName="Chappuis" authorityYear="1931" box="[219,480,968,990]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fluviatilis">
<emphasis id="61CD2E70FFB0FF882BDF89CDFEC877A7" box="[219,315,968,990]" italics="true" pageId="17" pageNumber="18">fluviatilis</emphasis>
Chappuis, 1931
</taxonomicName>
</td>
<td id="AE58995CFFB0006628AB89CDFC0A77A7" box="[943,1017,968,990]" gridcol="3" gridrow="5" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F1789CDFBB477A7" box="[1043,1095,968,990]" gridcol="4" gridrow="5" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F6589CDFA6F77A7" box="[1121,1436,968,990]" gridcol="5" gridrow="5" pageId="17" pageNumber="18">oriental (Indonesia, Sumatra)</td>
</tr>
<tr id="ED89F020FFB000662B9389F4FA6F707E" box="[151,1436,1009,1031]" gridrow="6" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B9389F4FF4C707E" box="[151,191,1009,1031]" gridcol="0" gridrow="6" pageId="17" pageNumber="18">6</th>
<td id="AE58995CFFB000662BDF89F4FD3A707E" box="[219,713,1009,1031]" gridcol="1" gridrow="6" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF89F4FE6D707E" authority="Ebert, 1976" authorityName="Ebert" authorityYear="1976" box="[219,414,1009,1031]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="goeldii">
<emphasis id="61CD2E70FFB0FF882BDF89F4FED1707E" box="[219,290,1009,1031]" italics="true" pageId="17" pageNumber="18">goeldii</emphasis>
Ebert, 1976
</taxonomicName>
*
</td>
<td id="AE58995CFFB00066281489F4FC69707E" box="[784,922,1009,1031]" gridcol="2" gridrow="6" pageId="17" pageNumber="18">not published</td>
<td id="AE58995CFFB0006628AB89F4FC0A707E" box="[943,1017,1009,1031]" gridcol="3" gridrow="6" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F1789F4FBB4707E" box="[1043,1095,1009,1031]" gridcol="4" gridrow="6" pageId="17" pageNumber="18">-</td>
<td id="AE58995CFFB000662F6589F4FA6F707E" box="[1121,1436,1009,1031]" gridcol="5" gridrow="6" pageId="17" pageNumber="18">neotropical (Brazil)</td>
</tr>
<tr id="ED89F020FFB000662B938E1FFA6F7023" box="[151,1436,1050,1114]" gridrow="7" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938E1FFF4C7023" box="[151,191,1050,1114]" gridcol="0" gridrow="7" pageId="17" pageNumber="18">7 8</th>
<td id="AE58995CFFB000662BDF8E1FFD3A7023" box="[219,713,1050,1114]" gridcol="1" gridrow="7" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF8E1FFE117048" authority="Delachaux, 1924" authorityName="Delachaux" authorityYear="1924" box="[219,482,1050,1073]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kalima">
<emphasis id="61CD2E70FFB0FF882BDF8E1FFED37049" box="[219,288,1050,1072]" italics="true" pageId="17" pageNumber="18">kalima</emphasis>
(Delachaux, 1924)
</taxonomicName>
* 
<taxonomicName id="94B989E1FFB0FF882BDF8E41FE247020" authority="Chappuis, 1931" authorityName="Chappuis" authorityYear="1931" box="[219,471,1091,1114]" class="Maxillopoda" family="Ameiridae" genus="Nitocra" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lacustris">
<emphasis id="61CD2E70FFB0FF882BDF8E41FEC17023" box="[219,306,1092,1114]" italics="true" pageId="17" pageNumber="18">lacustris</emphasis>
Chappuis, 1931
</taxonomicName>
</td>
<td id="AE58995CFFB0006628148E1FFC697023" box="[784,922,1050,1114]" gridcol="2" gridrow="7" pageId="17" pageNumber="18">relocated **</td>
<td id="AE58995CFFB0006628AB8E1FFC0A7023" box="[943,1017,1050,1114]" gridcol="3" gridrow="7" pageId="17" pageNumber="18">+ +</td>
<td id="AE58995CFFB000662F178E1FFBB47023" box="[1043,1095,1050,1114]" gridcol="4" gridrow="7" pageId="17" pageNumber="18">- -</td>
<td id="AE58995CFFB000662F658E1FFA6F7023" box="[1121,1436,1050,1114]" gridcol="5" gridrow="7" pageId="17" pageNumber="18">neotropical (Surinam) oriental (Indonesia, Sumatra)</td>
</tr>
<tr id="ED89F020FFB000662B938E68FA6F70FA" box="[151,1436,1133,1155]" gridrow="8" pageId="17" pageNumber="18" rowspan-2="1">
<th id="AE58995CFFB000662B938E68FF4C70FA" box="[151,191,1133,1155]" gridcol="0" gridrow="8" pageId="17" pageNumber="18">9</th>
<td id="AE58995CFFB000662BDF8E68FD3A70FA" box="[219,713,1133,1155]" gridcol="1" gridrow="8" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF8E6BFD3A70FA" authority="Janetzky, Martinez Arbizu &amp; Reid, 1996" authorityName="Janetzky, Martinez Arbizu &amp; Reid" authorityYear="1996" box="[219,713,1133,1155]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="mervini">
<emphasis id="61CD2E70FFB0FF882BDF8E6BFEDB70FA" box="[219,296,1134,1155]" italics="true" pageId="17" pageNumber="18">mervini</emphasis>
Janetzky, Martinez Arbizu &amp; Reid, 1996
</taxonomicName>
</td>
<td id="AE58995CFFB0006628AB8E68FC0A70FA" box="[943,1017,1133,1155]" gridcol="3" gridrow="8" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F178E68FBB470FA" box="[1043,1095,1133,1155]" gridcol="4" gridrow="8" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F658E68FA6F70FA" box="[1121,1436,1133,1155]" gridcol="5" gridrow="8" pageId="17" pageNumber="18">neotropical (Jamaica)</td>
</tr>
<tr id="ED89F020FFB000662B938E93FA6F70D4" box="[151,1436,1174,1197]" gridrow="9" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938E93FF4C70D4" box="[151,191,1174,1197]" gridcol="0" gridrow="9" pageId="17" pageNumber="18">10</th>
<td id="AE58995CFFB000662BDF8E93FD3A70D4" box="[219,713,1174,1197]" gridcol="1" gridrow="9" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF8E9DFE5C70D5" authority="Ebert, 1976" authorityName="Ebert" authorityYear="1976" box="[219,431,1174,1197]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="61CD2E70FFB0FF882BDF8E9DFEC070D4" box="[219,307,1176,1197]" italics="true" pageId="17" pageNumber="18">montana</emphasis>
Ebert, 1976
</taxonomicName>
</td>
<td id="AE58995CFFB0006628148E93FC6970D4" box="[784,922,1174,1197]" gridcol="2" gridrow="9" pageId="17" pageNumber="18">not published</td>
<td id="AE58995CFFB0006628AB8E93FC0A70D4" box="[943,1017,1174,1197]" gridcol="3" gridrow="9" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F178E93FBB470D4" box="[1043,1095,1174,1197]" gridcol="4" gridrow="9" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F658E93FA6F70D4" box="[1121,1436,1174,1197]" gridcol="5" gridrow="9" pageId="17" pageNumber="18">neotropical (Peru)</td>
</tr>
<tr id="ED89F020FFB000662B938EC5FA6F70AF" box="[151,1436,1216,1238]" gridrow="10" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938EC5FF4C70AF" box="[151,191,1216,1238]" gridcol="0" gridrow="10" pageId="17" pageNumber="18">11</th>
<td id="AE58995CFFB000662BDF8EC5FD3A70AF" box="[219,713,1216,1238]" gridcol="1" gridrow="10" pageId="17" pageNumber="18">
<taxonomicName id="94B989E1FFB0FF882BDF8EC5FE0470AF" authority="Chappuis, 1928" authorityName="Chappuis" authorityYear="1928" box="[219,503,1216,1238]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="61CD2E70FFB0FF882BDF8EC5FEB170AF" box="[219,322,1216,1238]" italics="true" pageId="17" pageNumber="18">muscicola</emphasis>
(Chappuis, 1928)
</taxonomicName>
</td>
<td id="AE58995CFFB0006628148EC5FC6970AF" box="[784,922,1216,1238]" gridcol="2" gridrow="10" pageId="17" pageNumber="18">relocated</td>
<td id="AE58995CFFB0006628AB8EC5FC0A70AF" box="[943,1017,1216,1238]" gridcol="3" gridrow="10" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F178EC5FBB470AF" box="[1043,1095,1216,1238]" gridcol="4" gridrow="10" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F658EC5FA6F70AF" box="[1121,1436,1216,1238]" gridcol="5" gridrow="10" pageId="17" pageNumber="18">oriental (Indonesia, Java)</td>
</tr>
</table>
</paragraph>
<subSubSection id="1BA3A1E9FFB0FF8D2B9B8EEFFEBE774E" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="5306F262FFB0FF882B9B8EEFFD0D7317" blockId="17.[159,1358,1257,1279]" lastBlockId="17.[151,1437,1554,2010]" pageId="17" pageNumber="18">
12 
<taxonomicName id="94B989E1FFB0FF882BDF8EECFD1D7086" authority="Janetzky, Martinez Arbizu &amp; Reid, 1996" authorityName="Janetzky, Martinez Arbizu &amp; Reid" authorityYear="1996" box="[219,750,1257,1279]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pilagaensis">
<emphasis id="61CD2E70FFB0FF882BDF8EECFEBD7086" box="[219,334,1257,1279]" italics="true" pageId="17" pageNumber="18">pilagaensis</emphasis>
<bibRefCitation id="37288F93FFB0FF882A508EECFD1D7086" author="Janetzky" box="[340,750,1257,1279]" pageId="17" pageNumber="18" refString="Janetzky, W., Martinez Arbizu, P. &amp; Reid, J. W. (1996) Attheyella (Canthosella) mervini n. sp. (Canthocamptidae, Harpacticoida) from Jamaican bromeliads. Hydrobiologia, 339, 123 - 135." type="journal article" year="1996">Janetzky, Martinez Arbizu &amp; Reid, 1996</bibRefCitation>
</taxonomicName>
* relocated ** + - neotropical (
<collectingCountry id="2BAEB2F2FFB0FF882FDB8EECFAB47086" box="[1247,1351,1257,1279]" name="Argentina" pageId="17" pageNumber="18">Argentina</collectingCountry>
) The description of the subgenus 
<taxonomicName id="94B989E1FFB0FF8829408C16FC7B7253" authority="Chappuis, 1931" authorityName="Chappuis" authorityYear="1931" box="[580,904,1554,1579]" class="Maxillopoda" family="Canthocamptidae" genus="Canthosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="61CD2E70FFB0FF8829408C16FD3F7252" box="[580,716,1555,1579]" italics="true" pageId="17" pageNumber="18">Canthosella</emphasis>
<bibRefCitation id="37288F93FFB0FF8829D18C17FC7B7253" author="Chappuis" box="[725,904,1554,1578]" pageId="17" pageNumber="18" refString="Chappuis, P. A. (1931) Copepoda Harpacticoida der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Suppl., 8, 512 - 584." type="journal article" year="1931">Chappuis, 1931</bibRefCitation>
</taxonomicName>
includes the following main characters: a not cylindrical body shape, a short rostrum, the posterior margin of the somites smooth, the caudal rami in both sexes being as long as or slightly longer than wide, with a short row of spines between second and third quarter of the inner margin. Antennula 8-segmented in male, with its fourth segment not very thickened. Endopod of antenna 1- segmented. Leg 1 with 2-segmented endopod, endopod shorter than exopod. Legs 2 to 4 of female with 2-segmented endopods, first segment of endopod of leg 4 can be very short. Second segment of endopod of leg 
<quantity id="94415F87FFB0FF882E3D8CC3FA9072A7" box="[1337,1379,1734,1758]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="17" pageNumber="18" unit="in" value="4.0">4 in</quantity>
both sexes with one apical seta. Endopod of leg 3 of the male 3-segmented, with spine of second segment transformed into an apophysis ending in a barb. Endopods of legs 2 to 4 with second segment bearing at least 1 seta on inner margin and 1 spine on outer margin. Baseoendopod of leg 5 on female strongly expanded, with 6 setae, exopod with 5 setae. Baseoendopod of male leg 5 with 2 setae.
</paragraph>
<paragraph id="5306F262FFB0FF882B9B8F16FAE871B7" pageId="17" pageNumber="18">
<table id="21B900C2FFB000662B938F17FA6F71B7" box="[151,1436,1298,1486]" gridcols="4" gridrows="4" pageId="17" pageNumber="18">
<tr id="ED89F020FFB000662B938F17FA6F7151" box="[151,1436,1298,1320]" gridrow="0" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938F17FE1B7151" box="[151,488,1298,1320]" gridcol="0" gridrow="0" pageId="17" pageNumber="18">
13 
<taxonomicName id="94B989E1FFB0FF882BDF8F17FE4E7151" authority="Loffler, 1973" authorityName="Loffler" authorityYear="1973" box="[219,445,1298,1320]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="silvicola">
<emphasis id="61CD2E70FFB0FF882BDF8F17FEC27151" box="[219,305,1298,1320]" italics="true" pageId="17" pageNumber="18">silvicola</emphasis>
Löffler, 1973
</taxonomicName>
</th>
<th id="AE58995CFFB0006628148F17FC4E7151" box="[784,957,1298,1320]" gridcol="1" gridrow="0" pageId="17" pageNumber="18">+</th>
<th id="AE58995CFFB000662F178F17FBD27151" box="[1043,1057,1298,1320]" gridcol="2" gridrow="0" pageId="17" pageNumber="18">+</th>
<th id="AE58995CFFB000662F658F17FA6F7151" box="[1121,1436,1298,1320]" gridcol="3" gridrow="0" pageId="17" pageNumber="18">oriental (Indonesia, Borneo)</th>
</tr>
<tr id="ED89F020FFB000662B938F3EFA6F712B" box="[151,1436,1339,1362]" gridrow="1" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938F3EFE1B712B" box="[151,488,1339,1362]" gridcol="0" gridrow="1" pageId="17" pageNumber="18">
14 
<taxonomicName id="94B989E1FFB0FF882BDF8F39FE507128" authority="Kiefer, 1967" authorityName="Kiefer" authorityYear="1967" box="[219,419,1339,1362]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="siolii">
<emphasis id="61CD2E70FFB0FF882BDF8F39FEFD712B" box="[219,270,1340,1362]" italics="true" pageId="17" pageNumber="18">siolii</emphasis>
(Kiefer, 1967)
</taxonomicName>
*
</th>
<td id="AE58995CFFB0006628148F3EFC4E712B" box="[784,957,1339,1362]" gridcol="1" gridrow="1" pageId="17" pageNumber="18">relocated ** -</td>
<td id="AE58995CFFB000662F178F3EFBD2712B" box="[1043,1057,1339,1362]" gridcol="2" gridrow="1" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F658F3EFA6F712B" box="[1121,1436,1339,1362]" gridcol="3" gridrow="1" pageId="17" pageNumber="18">neotropical (Brazil)</td>
</tr>
<tr id="ED89F020FFB000662B938F60FA6F71DC" box="[151,1436,1381,1445]" gridrow="2" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938F60FE1B71DC" box="[151,488,1381,1445]" gridcol="0" gridrow="2" pageId="17" pageNumber="18">
15 
<taxonomicName id="94B989E1FFB0FF882BDF8F60FE4D7105" authority="Reid, 1990" authorityName="Reid" authorityYear="1990" box="[219,446,1381,1404]" class="Maxillopoda" family="Canthocamptidae" genus="Canthocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="striblingi">
<emphasis id="61CD2E70FFB0FF882BDF8F60FECB7102" box="[219,312,1381,1403]" italics="true" pageId="17" pageNumber="18">striblingi</emphasis>
(Reid, 1990)
</taxonomicName>
16 
<taxonomicName id="94B989E1FFB0FF882BDF8F95FE3D71DD" authority="Por &amp; Hadel, 1986" authorityName="Por &amp; Hadel" authorityYear="1986" box="[219,462,1422,1445]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vera">
<emphasis id="61CD2E70FFB0FF882BDF8F95FEFB71DC" box="[219,264,1424,1445]" italics="true" pageId="17" pageNumber="18">vera</emphasis>
Por &amp; Hadel, 1986
</taxonomicName>
*
</th>
<td id="AE58995CFFB0006628148F60FC4E71DC" box="[784,957,1381,1445]" gridcol="1" gridrow="2" pageId="17" pageNumber="18">relocated ** - +</td>
<td id="AE58995CFFB000662F178F60FBD271DC" box="[1043,1057,1381,1445]" gridcol="2" gridrow="2" pageId="17" pageNumber="18">+ +</td>
<td id="AE58995CFFB000662F658F60FA6F71DC" box="[1121,1436,1381,1445]" gridcol="3" gridrow="2" pageId="17" pageNumber="18">neotropical (Costa Rica) neotropical (Brazil)</td>
</tr>
<tr id="ED89F020FFB000662B938FBDFA6F71B7" box="[151,1436,1464,1486]" gridrow="3" pageId="17" pageNumber="18">
<th id="AE58995CFFB000662B938FBDFE1B71B7" box="[151,488,1464,1486]" gridcol="0" gridrow="3" pageId="17" pageNumber="18">
17 
<taxonomicName id="94B989E1FFB0FF882BDF8FBCFE1B71B7" authority="Borutzki, 1967" authorityName="Borutzki" authorityYear="1967" box="[219,488,1464,1486]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="61CD2E70FFB0FF882BDF8FBCFEB971B7" box="[219,330,1465,1486]" italics="true" pageId="17" pageNumber="18">vietnamica</emphasis>
Borutzki, 1967
</taxonomicName>
</th>
<td id="AE58995CFFB0006628148FBDFC4E71B7" box="[784,957,1464,1486]" gridcol="1" gridrow="3" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F178FBDFBD271B7" box="[1043,1057,1464,1486]" gridcol="2" gridrow="3" pageId="17" pageNumber="18">+</td>
<td id="AE58995CFFB000662F658FBDFA6F71B7" box="[1121,1436,1464,1486]" gridcol="3" gridrow="3" pageId="17" pageNumber="18">oriental (Vietnam)</td>
</tr>
</table>
</paragraph>
<paragraph id="5306F262FFB0FF882BCD8D7EFBB373A3" blockId="17.[151,1437,1554,2010]" pageId="17" pageNumber="18">As American species were included in the subgenus, the following morphological characteristics added complementary details to the original description of the subgenus: Endopod of second to fourth legs with last segment bearing 1 or 2 armaments and baseoendopod of fifth leg of the male with 0–2 setae.</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD8A92FD57751D" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
The Asian species 
<taxonomicName id="94B989E1FFB3FF8B2AA08A9DFBF674D6" authority="Chappuis, 1928" authorityName="Chappuis" authorityYear="1928" box="[420,1029,151,176]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="muscicola" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2AA08A9DFDE574C9" box="[420,534,152,176]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B292C8A9DFD5C74C9" box="[552,687,152,176]" italics="true" pageId="18" pageNumber="19">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B29C68A9DFCC774C9" box="[706,820,152,176]" italics="true" pageId="18" pageNumber="19">muscicola</emphasis>
(
<bibRefCitation id="37288F93FFB3FF8B28418A92FC0F74D6" author="Chappuis" box="[837,1020,151,175]" pageId="18" pageNumber="19" refString="Chappuis, P. A. (1928) Neue Harpacticiden aus Java. Treubia, 10, 272 - 283." type="journal article" year="1928">Chappuis, 1928</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B2F178A9DFEBB74AD" authority="Chappuis, 1931" authorityName="Chappuis" authorityYear="1931" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fluviatilis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2F178A9DFB7674C9" box="[1043,1157,152,176]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F9C8A9DFAEC74C9" box="[1176,1311,152,176]" italics="true" pageId="18" pageNumber="19">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2E368A9DFA6F74C9" box="[1330,1436,152,176]" italics="true" pageId="18" pageNumber="19">fluviatilis</emphasis>
<bibRefCitation id="37288F93FFB3FF8B2B938AB9FEBB74AD" author="Chappuis" box="[151,328,188,212]" pageId="18" pageNumber="19" refString="Chappuis, P. A. (1931) Copepoda Harpacticoida der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Suppl., 8, 512 - 584." type="journal article" year="1931">Chappuis, 1931</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B2A508AB9FC7474AD" authority="Chappuis, 1931" authorityName="Chappuis" authorityYear="1931" box="[340,903,188,212]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lacustris" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A508AB9FE3574AD" box="[340,454,188,212]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2AD28AB9FDAE74AD" box="[470,605,188,212]" italics="true" pageId="18" pageNumber="19">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B296A8AB9FD3D74AD" box="[622,718,188,212]" italics="true" pageId="18" pageNumber="19">lacustris</emphasis>
<bibRefCitation id="37288F93FFB3FF8B29D18AB9FC7474AD" author="Chappuis" box="[725,903,188,212]" pageId="18" pageNumber="19" refString="Chappuis, P. A. (1931) Copepoda Harpacticoida der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Suppl., 8, 512 - 584." type="journal article" year="1931">Chappuis, 1931</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B28978AB9FF3C748E" authority="Borutzky, 1967" authorityName="Borutzky" authorityYear="1967" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vietnamica" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B28978AB9FBF674AD" box="[915,1029,188,212]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F118AB9FB6F74AD" box="[1045,1180,188,212]" italics="true" pageId="18" pageNumber="19">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2FA98AB8FADB74AD" box="[1197,1320,189,212]" italics="true" pageId="18" pageNumber="19">vietnamica</emphasis>
<bibRefCitation id="37288F93FFB3FF8B2E2A8AB9FF3C748E" author="Borutzky" pageId="18" pageNumber="19" refString="Borutzky, E. (1967) Presnovodny Copepoda Harpacticoida severnovo Vietnama (Freshwater Copepoda Harpacticoida of North Vietnam). Zoologicheskii Zhurnal, 46, 1015 - 1023." type="journal article" year="1967">Borutzky, 1967</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB3FF8B2A008AE5FCE6748E" authority="Loffler, 1973" authorityName="Loffler" authorityYear="1973" box="[260,789,223,248]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="silvicola" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A008AE5FE857481" box="[260,374,224,248]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2A818AE5FDFF7481" box="[389,524,224,248]" italics="true" pageId="18" pageNumber="19">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B29198AE5FD8F7481" box="[541,636,224,248]" italics="true" pageId="18" pageNumber="19">silvicola</emphasis>
<bibRefCitation id="37288F93FFB3FF8B29868ADAFCE6748E" author="Loffler" box="[642,789,223,247]" pageId="18" pageNumber="19" refString="Loffler, H. (1973) Die Harpacticidenfauna des Mt. Kinabalu (Borneo) mit besonderer Berucksichtigung der Gattung Maraenobiotus nebst Angaben zur Harpacticidenfauna des Gebietes Nuwara (Hochplateau Ceylon). Hochgebirgsforschung, 3, 5 - 28." type="journal article" year="1973">Löffler, 1973</bibRefCitation>
</taxonomicName>
have an endopod of leg 4 bearing only 1 seta, instead of 1 spine and 1 seta as in all American species (except 
<taxonomicName id="94B989E1FFB3FF8B29EF8B01FAD97565" authority="Petkovski, 1973" authorityName="Petkovski" authorityYear="1973" box="[747,1322,260,284]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="antillica" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B29EF8B01FCAF7565" box="[747,860,260,284]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B286B8B01FC057565" box="[879,1014,260,284]" italics="true" pageId="18" pageNumber="19">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2F0D8B01FB957565" box="[1033,1126,260,284]" italics="true" pageId="18" pageNumber="19">antillica</emphasis>
<bibRefCitation id="37288F93FFB3FF8B2F748B01FAD97565" author="Petkovski" box="[1136,1322,260,284]" pageId="18" pageNumber="19" refString="Petkovski, T. K. (1973) Resultats des expeditions biospeologiques cubano-roumaines a Cuba. Academiei Republicii Socialiste Romania, 1, 125 - 141." type="journal article" year="1973">Petkovski, 1973</bibRefCitation>
</taxonomicName>
). Another morphological difference of the American species is the absence of setae on the baseoendopod of leg 
<quantity id="94415F87FFB3FF8B2E018B22FAC37546" box="[1285,1328,295,319]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="18" pageNumber="19" unit="in" value="5.0">5 in</quantity>
males; in Asian species this baseoendopod bears 2 setae.
</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD8B6AFB6075B6" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
<bibRefCitation id="37288F93FFB3FF8B2BCD8B6AFE4D75FE" box="[201,446,367,392]" pageId="18" pageNumber="19" refString="Janetzky, W., Martinez Arbizu, P. &amp; Reid, J. W. (1996) Attheyella (Canthosella) mervini n. sp. (Canthocamptidae, Harpacticoida) from Jamaican bromeliads. Hydrobiologia, 339, 123 - 135." type="journal article">
Janetzky 
<emphasis id="61CD2E70FFB3FF8B2A378B74FE9F75F1" box="[307,364,368,392]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(1996)
</bibRefCitation>
proposed a division of the American species into two groups according to the presence or absence of a row of spines on the dorsal surface of the caudal ramus. 
<taxonomicName id="94B989E1FFB3FF8B28DD8B91FB1675D5" box="[985,1253,404,428]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="antillica" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B28DD8B91FBB875D5" box="[985,1099,404,428]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F588B90FB8575D5" box="[1116,1142,405,428]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2F8C8B91FB1675D5" box="[1160,1253,404,428]" italics="true" pageId="18" pageNumber="19">antillica</emphasis>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B2FF18B91FE2975B6" authority="Janetzky et al., 1996" authorityName="Janetzky et al." authorityYear="1996" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mervini" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2FF18B91FA9475D5" box="[1269,1383,404,428]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2E7D8B90FA6075D5" box="[1401,1427,405,428]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2B938BBCFF1F75A9" box="[151,236,441,464]" italics="true" pageId="18" pageNumber="19">mervini</emphasis>
<bibRefCitation id="37288F93FFB3FF8B2BF78BB2FE2975B6" author="Janetzky" box="[243,474,439,464]" pageId="18" pageNumber="19" refString="Janetzky, W., Martinez Arbizu, P. &amp; Reid, J. W. (1996) Attheyella (Canthosella) mervini n. sp. (Canthocamptidae, Harpacticoida) from Jamaican bromeliads. Hydrobiologia, 339, 123 - 135." type="journal article" year="1996">
Janetzky 
<emphasis id="61CD2E70FFB3FF8B2A588BBCFE7E75A9" box="[348,397,440,464]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
., 1996
</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB3FF8B29158BBDFD3975A9" box="[529,714,439,464]" class="Maxillopoda" family="Canthocamptidae" genus="Canthocamptus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="striblingi" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B29158BBDFDD175A9" box="[529,546,440,464]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. (
<emphasis id="61CD2E70FFB3FF8B293D8BBDFDBF75B6" box="[569,588,440,463]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.) 
<emphasis id="61CD2E70FFB3FF8B29678BBDFD3975A9" box="[611,714,440,464]" italics="true" pageId="18" pageNumber="19">striblingi</emphasis>
</taxonomicName>
belong to the group lacking such spines.
</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD8BD9FF0176FD" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
The new species 
<taxonomicName id="94B989E1FFB3FF8B2A8A8BD9FDAC758D" box="[398,607,476,500]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A8A8BD9FE6C758D" box="[398,415,476,500]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. (
<emphasis id="61CD2E70FFB3FF8B2ABC8BD8FE38758D" box="[440,459,477,500]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.) 
<emphasis id="61CD2E70FFB3FF8B2AE08BD9FDAC758D" box="[484,607,476,500]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
, as well as 
<taxonomicName id="94B989E1FFB3FF8B29E38BD9FC2E758D" box="[743,989,476,500]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B29E38BD9FCAB758D" box="[743,856,476,500]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B286F8BD8FC76758D" box="[875,901,477,500]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B28938BD9FC2E758D" box="[919,989,476,500]" italics="true" pageId="18" pageNumber="19">aliena</emphasis>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B28E88BD9FED7766E" authority="Por and Hadel, 1986" authorityName="Por and Hadel" authorityYear="1986" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vera" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B28E88BD9FBAD758D" box="[1004,1118,476,500]" italics="true" pageId="18" pageNumber="19">Attheyella</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F748BD9FB04758D" box="[1136,1271,476,500]" italics="true" pageId="18" pageNumber="19">Canthosella</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2E0D8BD8FAC8758D" box="[1289,1339,477,500]" italics="true" pageId="18" pageNumber="19">vera</emphasis>
<bibRefCitation id="37288F93FFB3FF8B2E408BD8FED7766E" author="Por" pageId="18" pageNumber="19" refString="Por, F. D. &amp; Hadel, V. F. (1986) Two new species of Attheyella (Copepoda: Harpacticoida: Canthocamptidae) from bromeliads of Serra da Jureia (Sao Paulo, Brazil). Journal of Crustacean Biology, 6, 777 - 788." type="journal article" year="1986">Por and Hadel, 1986</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B2A358805FE207661" box="[305,467,511,536]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kalima" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A358805FEB17661" box="[305,322,512,536]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. (
<emphasis id="61CD2E70FFB3FF8B2A5F8805FE86766E" box="[347,373,512,535]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2A838805FE207661" box="[391,467,512,536]" italics="true" pageId="18" pageNumber="19">kalima</emphasis>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B2AE78805FD447661" box="[483,695,511,536]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2AE78805FE087661" box="[483,507,512,536]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29098805FDD3766E" box="[525,544,512,535]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.) 
<emphasis id="61CD2E70FFB3FF8B293C8805FD447661" box="[568,695,512,536]" italics="true" pageId="18" pageNumber="19">pilagaensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B29CC8805FC987661" box="[712,875,511,536]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="goeldii" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B29CC8805FD137661" box="[712,736,512,536]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29F68805FCFF766E" box="[754,780,512,535]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B281A8805FC987661" box="[798,875,512,536]" italics="true" pageId="18" pageNumber="19">goeldii</emphasis>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB3FF8B28A18805FBC07661" box="[933,1075,512,536]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="siolii" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B28A18805FBC07661" box="[933,1075,512,536]" italics="true" pageId="18" pageNumber="19">A. (C.) siolii</emphasis>
</taxonomicName>
, belong to the group of species bearing a row of spines on the dorsal surface of the caudal ramus. Nevertheless, males of the new species lack this row of spines. The morphological differences within this group are summarized in Table 2 (females) and Table 3 (males).
</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD888AFAB27725" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
Females of the new species differ from females of the species of the group in having a genital double-somite that is ventrally and sometimes dorsally partially divided, and in the morphology and ornamentation of the caudal rami (Table 2). The insertion point of the dorsal seta of the caudal ramus is located in the middle of the segment in 
<taxonomicName id="94B989E1FFB3FF8B2B9388F9FE95776D" box="[151,358,764,788]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2B9388F9FF5C776D" box="[151,175,764,788]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2BC488F8FF29776D" box="[192,218,765,788]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2BEF88F9FE95776D" box="[235,358,764,788]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
. In contrast, this seta is inserted on the posterior quarter of the ramus in the other species, as is shown in 
<taxonomicName id="94B989E1FFB3FF8B2A078925FE697741" box="[259,410,799,824]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A078925FEE87741" box="[259,283,800,824]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2A2F8925FEB6774E" box="[299,325,800,823]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2A508925FE697741" box="[340,410,800,824]" italics="true" pageId="18" pageNumber="19">aliena</emphasis>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB3FF8B2AD48925FD527741" box="[464,673,799,824]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2AD48925FE1B7741" box="[464,488,800,824]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2AFC8925FDE1774E" box="[504,530,800,823]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B29268925FD527741" box="[546,673,800,824]" italics="true" pageId="18" pageNumber="19">pilagaensis</emphasis>
</taxonomicName>
(
<figureCitation id="CB82EEE7FFB3FF8B29B4891AFD0E774E" box="[688,765,799,823]" captionStart="FIGURE 4" captionStartId="6.[151,250,1696,1719]" captionTargetBox="[274,1321,193,1661]" captionTargetId="figure@6.[253,1334,181,1675]" captionTargetPageId="6" captionText="FIGURE 4. Females. Attheyella (Canthosella) aliena. A. caudal rami with anal operculum, dorsal; B, caudal ramus, ventral. Attheyella (Canthosella) pilagaensis. C, caudal rami with anal operculum, dorsal; D, caudal ramus, ventral." httpUri="https://zenodo.org/record/209239/files/figure.png" pageId="18" pageNumber="19">Figs. 4</figureCitation>
A and 4C). Moreover, the size of the posterodorsal spines of the new species is much smaller than in the other species (compared e.g. with 
<taxonomicName id="94B989E1FFB3FF8B2F068941FB947725" box="[1026,1127,836,860]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aliena">
<emphasis id="61CD2E70FFB3FF8B2F068941FB947725" box="[1026,1127,836,860]" italics="true" pageId="18" pageNumber="19">A. aliena</emphasis>
</taxonomicName>
(
<figureCitation id="CB82EEE7FFB3FF8B2F738941FB377725" box="[1143,1220,836,860]" captionStart="FIGURE 4" captionStartId="6.[151,250,1696,1719]" captionTargetBox="[274,1321,193,1661]" captionTargetId="figure@6.[253,1334,181,1675]" captionTargetPageId="6" captionText="FIGURE 4. Females. Attheyella (Canthosella) aliena. A. caudal rami with anal operculum, dorsal; B, caudal ramus, ventral. Attheyella (Canthosella) pilagaensis. C, caudal rami with anal operculum, dorsal; D, caudal ramus, ventral." httpUri="https://zenodo.org/record/209239/files/figure.png" pageId="18" pageNumber="19">Figs. 4</figureCitation>
A and 5A).
</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD8962FA6577DD" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
Additionally, there are small differences in the setation of leg 1. With the exception of 
<taxonomicName id="94B989E1FFB3FF8B2F84896DFABC77F9" box="[1152,1359,871,896]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2F84896DFB6A77F9" box="[1152,1177,872,896]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2FAC896DFB497706" box="[1192,1210,872,895]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.) 
<emphasis id="61CD2E70FFB3FF8B2FD5896DFABC77F9" box="[1233,1359,872,896]" italics="true" pageId="18" pageNumber="19">pilagaensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB3FF8B2E80896DFEB177DD" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2E80896DFA6F77F9" box="[1412,1436,872,896]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2BA48988FF4A77DD" box="[160,185,909,932]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2BCD8989FEB177DD" box="[201,322,908,932]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
, no other species bears spinule(s) on the inner margin of the second segment of the exopod of leg 1.
</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD89AAFBDF77B1" blockId="18.[151,1437,151,1941]" box="[201,1068,943,969]" pageId="18" pageNumber="19">Other particular similarities and differences with the species of the group are:</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD89D1FDB270D9" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
Females of the new species are close to 
<taxonomicName id="94B989E1FFB3FF8B298389D1FCD47795" box="[647,807,980,1004]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="goeldii" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B298389D1FD6C7795" box="[647,671,980,1004]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29AB89D0FD3A7795" box="[687,713,981,1004]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B29DD89D1FCD47795" box="[729,807,980,1004]" italics="true" pageId="18" pageNumber="19">goeldii</emphasis>
</taxonomicName>
from 
<collectingCountry id="2BAEB2F2FFB3FF8B286F89D1FC417795" box="[875,946,980,1004]" name="Brazil" pageId="18" pageNumber="19">Brazil</collectingCountry>
, particularly in having a similar ornamentation of the genital somite and the urosomites 3 and 4, and in the similar armature of legs 2 to 4. 
<taxonomicName id="94B989E1FFB3FF8B2FC889FDFA9F7069" box="[1228,1388,1016,1040]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="goeldii" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2FC889FDFB177069" box="[1228,1252,1016,1040]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2FF089FDFAFD7076" box="[1268,1294,1016,1039]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2E1A89FDFA9F7069" box="[1310,1388,1016,1040]" italics="true" pageId="18" pageNumber="19">goeldii</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="94B989E1FFB3FF8B2A038E19FE25704D" box="[263,470,1052,1076]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A038E19FEEC704D" box="[263,287,1052,1076]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2A348E18FEBA704D" box="[304,329,1053,1076]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2A5F8E19FE25704D" box="[347,470,1052,1076]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
in having more spines (14 instead of 
<quantity id="94415F87FFB3FF8B28858E18FC5E704D" box="[897,941,1052,1076]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="18" pageNumber="19" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="94B989E1FFB3FF8B28B08E19FB70704D" box="[948,1155,1052,1076]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B28B08E19FC3E704D" box="[948,973,1052,1076]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B28D98E18FC04704D" box="[989,1015,1053,1076]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2F0C8E19FB70704D" box="[1032,1155,1052,1076]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
) on the posterior margin of the anal somite, the proximal lateral seta of the caudal ramus is inserted in the middle of the segment (in 
<taxonomicName id="94B989E1FFB3FF8B2E548E45FEE17005" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2E548E45FA9B7021" box="[1360,1384,1088,1112]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2E7D8E45FA60702E" box="[1401,1427,1088,1111]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2B938E61FEE17005" box="[151,274,1124,1148]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
in the proximal third) and the apical inner spine on the endopod of leg 1 is much better developed. Males of 
<taxonomicName id="94B989E1FFB3FF8B2A048E8DFE5270D9" box="[256,417,1160,1184]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="goeldii" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A048E8DFEEA70D9" box="[256,281,1160,1184]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2A2D8E8DFEB070E6" box="[297,323,1160,1183]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2A578E8DFE5270D9" box="[339,417,1160,1184]" italics="true" pageId="18" pageNumber="19">goeldii</emphasis>
</taxonomicName>
are unknown.
</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD8EA9FEA9710E" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
The new species differs from females of 
<taxonomicName id="94B989E1FFB3FF8B29AB8EA9FC7A70BD" box="[687,905,1196,1220]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B29AB8EA9FD3B70BD" box="[687,712,1196,1220]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29DF8EA8FD1D70BD" box="[731,750,1197,1220]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.) 
<emphasis id="61CD2E70FFB3FF8B280E8EA9FC7A70BD" box="[778,905,1196,1220]" italics="true" pageId="18" pageNumber="19">pilagaensis</emphasis>
</taxonomicName>
particularly in the morphology of the caudal ramus. In 
<taxonomicName id="94B989E1FFB3FF8B2A0C8ED5FE2F7091" box="[264,476,1231,1256]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A0C8ED5FED27091" box="[264,289,1232,1256]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2A358ED5FEB8709E" box="[305,331,1232,1255]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2A588ED5FE2F7091" box="[348,476,1232,1256]" italics="true" pageId="18" pageNumber="19">pilagaensis</emphasis>
</taxonomicName>
it is oval (
<figureCitation id="CB82EEE7FFB3FF8B29508ECAFD51709E" box="[596,674,1231,1255]" captionStart="FIGURE 4" captionStartId="6.[151,250,1696,1719]" captionTargetBox="[274,1321,193,1661]" captionTargetId="figure@6.[253,1334,181,1675]" captionTargetPageId="6" captionText="FIGURE 4. Females. Attheyella (Canthosella) aliena. A. caudal rami with anal operculum, dorsal; B, caudal ramus, ventral. Attheyella (Canthosella) pilagaensis. C, caudal rami with anal operculum, dorsal; D, caudal ramus, ventral." httpUri="https://zenodo.org/record/209239/files/figure.png" pageId="18" pageNumber="19">Figs. 4</figureCitation>
C and 4D) and in 
<taxonomicName id="94B989E1FFB3FF8B28688ED5FBC97091" box="[876,1082,1231,1256]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B28688ED5FC777091" box="[876,900,1232,1256]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B28918ED5FC5D709E" box="[917,942,1232,1255]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B28BB8ED5FBC97091" box="[959,1082,1232,1256]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
it is subquadrate (
<figureCitation id="CB82EEE7FFB3FF8B2E0E8ECAFAA4709E" box="[1290,1367,1231,1255]" captionStart="FIGURE 5" captionStartId="7.[151,250,1614,1637]" captionTargetBox="[238,1339,208,1584]" captionTargetId="figure@7.[227,1360,193,1593]" captionTargetPageId="7" captionText="FIGURE 5. Attheyella (Canthosella) chocoensis n. sp. Female (paratype NHMW 20682). A, caudal rami with anal operculum, dorsal; B, caudal ramus, ventral; C, caudal ramus, lateral." httpUri="https://zenodo.org/record/209245/files/figure.png" pageId="18" pageNumber="19">Figs. 5</figureCitation>
A and 5B); the proximal lateral seta is inserted in the third distal part of the caudal ramus in 
<taxonomicName id="94B989E1FFB3FF8B2F708EF1FABA7175" box="[1140,1353,1268,1292]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2F708EF1FB7F7175" box="[1140,1164,1268,1292]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F9A8EF0FB427175" box="[1182,1201,1269,1292]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.) 
<emphasis id="61CD2E70FFB3FF8B2FCE8EF1FABA7175" box="[1226,1353,1268,1292]" italics="true" pageId="18" pageNumber="19">pilagaensis</emphasis>
</taxonomicName>
(in the third proximal part of the ramus in 
<taxonomicName id="94B989E1FFB3FF8B29228F1DFD067149" box="[550,757,1303,1328]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B29228F1DFDCC7149" box="[550,575,1304,1328]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29548F1DFD9A7156" box="[592,617,1304,1327]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B297E8F1DFD067149" box="[634,757,1304,1328]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
). Additionally, there are differences in the size of the apical spines of the endopod of leg 1 (the outer spine is longer than the inner one in 
<taxonomicName id="94B989E1FFB3FF8B2F0D8F39FB2B712D" box="[1033,1240,1340,1364]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2F0D8F39FBD2712D" box="[1033,1057,1340,1364]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F378F38FBBF712D" box="[1075,1100,1341,1364]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2F598F39FB2B712D" box="[1117,1240,1340,1364]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
and shorter in 
<taxonomicName id="94B989E1FFB3FF8B2E808F39FEBA7101" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pilagaensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2E808F39FA6F712D" box="[1412,1436,1340,1364]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2BA48F65FF40710E" box="[160,179,1376,1399]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.) 
<emphasis id="61CD2E70FFB3FF8B2BCE8F65FEBA7101" box="[202,329,1376,1400]" italics="true" pageId="18" pageNumber="19">pilagaensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD8F81FBB8719D" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
<taxonomicName id="94B989E1FFB3FF8B2BCD8F81FE6571E5" box="[201,406,1412,1436]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2BCD8F81FF1271E5" box="[201,225,1412,1436]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2BF58F80FEF871E5" box="[241,267,1413,1436]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2A1F8F81FE6571E5" box="[283,406,1412,1436]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
differs from females of 
<taxonomicName id="94B989E1FFB3FF8B29A28F81FCCC71E5" box="[678,831,1412,1436]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B29A28F81FD4C71E5" box="[678,703,1412,1436]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29CB8F80FD1B71E5" box="[719,744,1413,1436]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B29FD8F81FCCC71E5" box="[761,831,1412,1436]" italics="true" pageId="18" pageNumber="19">aliena</emphasis>
</taxonomicName>
particularly in the morphology of the base of the central apical seta of the caudal rami. This seta in 
<taxonomicName id="94B989E1FFB3FF8B299B8FADFCC471B9" box="[671,823,1447,1472]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B299B8FADFD4B71B9" box="[671,696,1448,1472]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29CC8FADFD1271C6" box="[712,737,1448,1471]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B29F58FADFCC471B9" box="[753,823,1448,1472]" italics="true" pageId="18" pageNumber="19">aliena</emphasis>
</taxonomicName>
has a very broad base in contrast to 
<taxonomicName id="94B989E1FFB3FF8B2FCB8FADFA6F71B9" box="[1231,1436,1447,1472]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2FCB8FADFB1471B9" box="[1231,1255,1448,1472]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2FF38FADFAE271C6" box="[1271,1297,1448,1471]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2E258FADFA6F71B9" box="[1313,1436,1448,1472]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
(
<figureCitation id="CB82EEE7FFB3FF8B2B9B8FC9FF1E719D" box="[159,237,1484,1508]" captionStart="FIGURE 4" captionStartId="6.[151,250,1696,1719]" captionTargetBox="[274,1321,193,1661]" captionTargetId="figure@6.[253,1334,181,1675]" captionTargetPageId="6" captionText="FIGURE 4. Females. Attheyella (Canthosella) aliena. A. caudal rami with anal operculum, dorsal; B, caudal ramus, ventral. Attheyella (Canthosella) pilagaensis. C, caudal rami with anal operculum, dorsal; D, caudal ramus, ventral." httpUri="https://zenodo.org/record/209239/files/figure.png" pageId="18" pageNumber="19">Figs. 4</figureCitation>
A and 5A) and to the other species of the group, and small outer apical setae.
</paragraph>
<paragraph id="5306F262FFB3FF8B2BD58FEAFB9572EE" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
Differences from females of 
<taxonomicName id="94B989E1FFB3FF8B291D8FF5FD537271" box="[537,672,1519,1544]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vera" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B291D8FF5FDC27271" box="[537,561,1520,1544]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29478FF5FDAE727E" box="[579,605,1520,1543]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B296A8FF4FD537271" box="[622,672,1521,1544]" italics="true" pageId="18" pageNumber="19">vera</emphasis>
</taxonomicName>
can be observed in the ornamentation of the urosome: 
<taxonomicName id="94B989E1FFB3FF8B2E118FF5FA687271" box="[1301,1435,1519,1544]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vera" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2E118FF5FADE7271" box="[1301,1325,1520,1544]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2E3B8FF5FAAB727E" box="[1343,1368,1520,1543]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2E6E8FF4FA687271" box="[1386,1435,1521,1544]" italics="true" pageId="18" pageNumber="19">vera</emphasis>
</taxonomicName>
has no spines on the ventral surface of urosomites 3 and 4. The mandibular palp of 
<taxonomicName id="94B989E1FFB3FF8B2F338C11FB487255" box="[1079,1211,1556,1580]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vera" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2F338C11FBBC7255" box="[1079,1103,1556,1580]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F5B8C10FB8A7255" box="[1119,1145,1557,1580]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2F8D8C10FB487255" box="[1161,1211,1557,1580]" italics="true" pageId="18" pageNumber="19">vera</emphasis>
</taxonomicName>
bears 2 setae, while that of 
<taxonomicName id="94B989E1FFB3FF8B2BE38C3DFE467229" box="[231,437,1592,1616]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2BE38C3DFF0C7229" box="[231,255,1592,1616]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2A148C3DFEDA7236" box="[272,297,1592,1615]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2A3E8C3DFE467229" box="[314,437,1592,1616]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
bears 3. According to the diagnosis of 
<bibRefCitation id="37288F93FFB3FF8B28748C3CFB957229" author="Por" box="[880,1126,1592,1616]" pageId="18" pageNumber="19" refString="Por, F. D. &amp; Hadel, V. F. (1986) Two new species of Attheyella (Copepoda: Harpacticoida: Canthocamptidae) from bromeliads of Serra da Jureia (Sao Paulo, Brazil). Journal of Crustacean Biology, 6, 777 - 788." type="journal article" year="1986">Por and Hadel (1986)</bibRefCitation>
, the endopod of leg 3 has 1 spine on the inner margin in 
<taxonomicName id="94B989E1FFB3FF8B2AE68C59FD9A720D" box="[482,617,1628,1652]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vera" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2AE68C59FE08720D" box="[482,507,1628,1652]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29088C58FDD5720D" box="[524,550,1629,1652]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B293C8C58FD9A720D" box="[568,617,1629,1652]" italics="true" pageId="18" pageNumber="19">vera</emphasis>
</taxonomicName>
, but 
<quantity id="94415F87FFB3FF8B29A28C59FD20720D" box="[678,723,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="19" unit="in" value="2.0">2 in</quantity>
the other species including 
<taxonomicName id="94B989E1FFB3FF8B2F108C59FB1B720D" box="[1044,1256,1628,1652]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2F108C59FBDE720D" box="[1044,1069,1628,1652]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F3A8C58FBAB720D" box="[1086,1112,1629,1652]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2F6E8C59FB1B720D" box="[1130,1256,1628,1652]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
.. However, it is possible that they oversighted the second spine which is located very near the first one.
</paragraph>
<paragraph id="5306F262FFB3FF8B2BCD8CA1FB5D735E" blockId="18.[151,1437,151,1941]" pageId="18" pageNumber="19">
Females of 
<taxonomicName id="94B989E1FFB3FF8B2A4A8CA1FE0372C5" box="[334,496,1700,1724]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kalima" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2A4A8CA1FEAC72C5" box="[334,351,1700,1724]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. (
<emphasis id="61CD2E70FFB3FF8B2A7C8CA0FE6172C5" box="[376,402,1701,1724]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2AA08CA1FE0372C5" box="[420,496,1700,1724]" italics="true" pageId="18" pageNumber="19">kalima</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="94B989E1FFB3FF8B297A8CA1FCBD72C5" box="[638,846,1700,1724]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B297A8CA1FD6572C5" box="[638,662,1700,1724]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29AC8CA0FD3172C5" box="[680,706,1701,1724]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B29D78CA1FCBD72C5" box="[723,846,1700,1724]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
particularly in having a longer copulatory tube that reaches to the posterior quarter of the genital segment. The description of 
<bibRefCitation id="37288F93FFB3FF8B28F98CC2FB3972A6" author="Delachaux" box="[1021,1226,1735,1759]" pageId="18" pageNumber="19" refString="Delachaux, T. (1924) Zur Kenntnis der Copepodenfauna von Surinam. II. Harpacticiden. Zoologischer Anzeiger, 59, 1 - 16." type="journal article" year="1924">Delachaux (1924)</bibRefCitation>
refers to a 1-segmented endopod of leg 4 compared with a 2-segmented endopod in 
<taxonomicName id="94B989E1FFB3FF8B28898CE9FBA9737D" box="[909,1114,1772,1796]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B28898CE9FC56737D" box="[909,933,1772,1796]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B28B18CE8FC3C737D" box="[949,975,1773,1796]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B28DB8CE9FBA9737D" box="[991,1114,1772,1796]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
; however, this author probably missed the very short first segment of the endopod. Males of 
<taxonomicName id="94B989E1FFB3FF8B286B8D15FBFD7351" box="[879,1038,1807,1832]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kalima" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B286B8D15FC737351" box="[879,896,1808,1832]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. (
<emphasis id="61CD2E70FFB3FF8B289C8D15FC42735E" box="[920,945,1808,1831]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B28C68D15FBFD7351" box="[962,1038,1808,1832]" italics="true" pageId="18" pageNumber="19">kalima</emphasis>
</taxonomicName>
are unknown.
</paragraph>
<paragraph id="5306F262FFB3FF8D2BCD8D31FD1C748E" blockId="18.[151,1437,151,1941]" lastBlockId="20.[151,1436,151,823]" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
Males of the new species differ from those of 
<taxonomicName id="94B989E1FFB3FF8B29CF8D31FC907335" box="[715,867,1844,1868]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B29CF8D31FD177335" box="[715,740,1844,1868]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29F08D30FCFE7335" box="[756,781,1845,1868]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B28198D31FC907335" box="[797,867,1844,1868]" italics="true" pageId="18" pageNumber="19">aliena</emphasis>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB3FF8B28758D31FC0F7335" box="[881,1020,1844,1868]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="siolii" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B28758D31FC797335" box="[881,906,1844,1868]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B289E8D30FC5E7335" box="[922,941,1845,1868]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.) 
<emphasis id="61CD2E70FFB3FF8B28C08D31FC0F7335" box="[964,1020,1844,1868]" italics="true" pageId="18" pageNumber="19">siolii</emphasis>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB3FF8B2F368D31FB457335" box="[1074,1206,1844,1868]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vera" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2F368D31FBB87335" box="[1074,1099,1844,1868]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2F5E8D30FB877335" box="[1114,1140,1845,1868]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2F808D30FB457335" box="[1156,1206,1845,1868]" italics="true" pageId="18" pageNumber="19">vera</emphasis>
</taxonomicName>
by the lack of a row of spinules on the dorsal surface of the caudal rami. Additionally, it was noted a size difference between the aesthetascs of segment 4 of the antennula: in 
<taxonomicName id="94B989E1FFB3FF8B296A8D79FCF573ED" box="[622,774,1916,1940]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B296A8D79FD7573ED" box="[622,646,1916,1940]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B29928D78FD4373ED" box="[662,688,1917,1940]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B29C48D79FCF573ED" box="[704,774,1916,1940]" italics="true" pageId="18" pageNumber="19">aliena</emphasis>
</taxonomicName>
this aesthetasc is short, whereas in 
<taxonomicName id="94B989E1FFB3FF8B2F908D79FA9173ED" box="[1172,1378,1916,1940]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB3FF8B2F908D79FB5E73ED" box="[1172,1197,1916,1940]" italics="true" pageId="18" pageNumber="19">A.</emphasis>
(
<emphasis id="61CD2E70FFB3FF8B2FB98D78FB2473ED" box="[1213,1239,1917,1940]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
) 
<emphasis id="61CD2E70FFB3FF8B2FE38D79FA9173ED" box="[1255,1378,1916,1940]" italics="true" pageId="18" pageNumber="19">chocoensis</emphasis>
</taxonomicName>
it is long, almost reaching the distal margin of segment 6 (
<figureCitation id="CB82EEE7FFB5FF8D28008A92FCAD74D6" box="[772,862,151,175]" captionStart="FIGURE 11" captionStartId="13.[151,250,1673,1696]" captionTargetBox="[224,1383,211,1639]" captionTargetId="figure@13.[193,1394,193,1652]" captionTargetPageId="13" captionText="FIGURE 11. Males. Attheyella (Canthosella) aliena. A, antennule. Attheyella (Canthosella) chocoensis n. sp. (allotype, B, C and D). B, antennule; C, leg 5; D, leg 5, inner spine." httpUri="https://zenodo.org/record/209288/files/figure.png" pageId="20" pageNumber="21">Figs. 11</figureCitation>
A and 11B). An additional difference with 
<taxonomicName id="94B989E1FFB5FF8D2E4A8A9DFF2E74AD" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D2E4A8A9DFA9474C9" box="[1358,1383,152,176]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2E7D8A9DFA6074D6" box="[1401,1427,152,175]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2B938AB9FF2E74AD" box="[151,221,188,212]" italics="true" pageId="20" pageNumber="21">aliena</emphasis>
</taxonomicName>
is related to the number of the apical setae of the endopod of leg 3: 
<taxonomicName id="94B989E1FFB5FF8D28DE8AB9FB8174AD" box="[986,1138,188,212]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aliena" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D28DE8AB9FC0174AD" box="[986,1010,188,212]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2F068AB8FBEF74AD" box="[1026,1052,189,212]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2F288AB9FB8174AD" box="[1068,1138,188,212]" italics="true" pageId="20" pageNumber="21">aliena</emphasis>
</taxonomicName>
bears 1 seta, 
<taxonomicName id="94B989E1FFB5FF8D2E0F8AB9FF2E7481" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D2E0F8AB9FAD074AD" box="[1291,1315,188,212]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2E308AB8FABE74AD" box="[1332,1357,189,212]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2E5A8AB9FF2E7481" italics="true" pageId="20" pageNumber="21">chocoensis</emphasis>
</taxonomicName>
bears 2. Females of 
<emphasis id="61CD2E70FFB5FF8D2AC08AE5FE2E7481" box="[452,477,224,248]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(C.) 
<taxonomicName id="94B989E1FFB5FF8D29138AE5FDBC7481" box="[535,591,224,248]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="siolii">
<emphasis id="61CD2E70FFB5FF8D29138AE5FDBC7481" box="[535,591,224,248]" italics="true" pageId="20" pageNumber="21">siolii</emphasis>
</taxonomicName>
are unknown.
</paragraph>
<paragraph id="5306F262FFB5FF8D2BCD8B01FC2875F1" blockId="20.[151,1436,151,823]" pageId="20" pageNumber="21">
The number of teeth on the anal operculum in both sexes does not constitute a useful feature to differentiate among species. Some species have been described based on only 
<specimenCount id="45BF39EBFFB5FF8D286B8B22FC497546" box="[879,954,295,319]" pageId="20" pageNumber="21" type="male">1 male</specimenCount>
and 
<specimenCount id="45BF39EBFFB5FF8D28F48B22FBA27546" box="[1008,1105,295,319]" pageId="20" pageNumber="21" type="female">1 female</specimenCount>
or 
<specimenCount id="45BF39EBFFB5FF8D2F718B22FB077546" box="[1141,1268,295,319]" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
of each sex. In mounted specimens of 
<taxonomicName id="94B989E1FFB5FF8D2A9E8B49FD9B751D" box="[410,616,332,356]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D2A9E8B49FE41751D" box="[410,434,332,356]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2AC78B48FE2F751D" box="[451,476,333,356]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2AE98B49FD9B751D" box="[493,616,332,356]" italics="true" pageId="20" pageNumber="21">chocoensis</emphasis>
</taxonomicName>
, we have found variability in the number of teeth: females 5 to 9 (n = 4), males 3 to 9 (n = 6). 
<bibRefCitation id="37288F93FFB5FF8D2A848B75FD8175FE" author="Por" box="[384,626,367,391]" pageId="20" pageNumber="21" refString="Por, F. D. &amp; Hadel, V. F. (1986) Two new species of Attheyella (Copepoda: Harpacticoida: Canthocamptidae) from bromeliads of Serra da Jureia (Sao Paulo, Brazil). Journal of Crustacean Biology, 6, 777 - 788." type="journal article" year="1986">Por and Hadel (1986)</bibRefCitation>
noted the same for 
<taxonomicName id="94B989E1FFB5FF8D28548B75FC2475F1" box="[848,983,367,392]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="vera" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D28548B75FC9B75F1" box="[848,872,368,392]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D287C8B75FC6175FE" box="[888,914,368,391]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D28A68B74FC2475F1" box="[930,983,369,392]" italics="true" pageId="20" pageNumber="21">vera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5306F262FFB5FF8D2BCD8B91FB7776DE" blockId="20.[151,1436,151,823]" pageId="20" pageNumber="21">
Although females of the new species differ from those of 
<taxonomicName id="94B989E1FFB5FF8D284F8B91FC0275D5" box="[843,1009,404,428]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mervini" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D284F8B91FC9075D5" box="[843,867,404,428]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D28778B90FC7F75D5" box="[883,908,405,428]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D28988B90FC0275D5" box="[924,1009,405,428]" italics="true" pageId="20" pageNumber="21">mervini</emphasis>
</taxonomicName>
, 
<taxonomicName id="94B989E1FFB5FF8D28FB8B91FB5D75D5" box="[1023,1198,404,428]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="antillica" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D28FB8B91FBEB75D5" box="[1023,1048,404,428]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2F238B90FBB275D5" box="[1063,1089,405,428]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2F558B91FB5D75D5" box="[1105,1198,404,428]" italics="true" pageId="20" pageNumber="21">antillica</emphasis>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB5FF8D2FE08B91FA6F75D5" box="[1252,1436,404,428]" class="Maxillopoda" family="Canthocamptidae" genus="Canthocamptus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="striblingi" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D2FE08B91FB0F75D5" box="[1252,1276,404,428]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2E088B90FAD675D5" box="[1292,1317,405,428]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2E318B91FA6F75D5" box="[1333,1436,404,428]" italics="true" pageId="20" pageNumber="21">striblingi</emphasis>
</taxonomicName>
based on the presence of a spinule row on the dorsal surface of the caudal rami, males resemble those species because they lack such spinules. Differences have been noted within males, particularly in the endopod of leg 3: in 
<taxonomicName id="94B989E1FFB5FF8D2B938805FE907661" box="[151,355,511,536]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D2B938805FF5C7661" box="[151,175,512,536]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2BBB8805FF2B766E" box="[191,216,512,535]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2BEC8805FE907661" box="[232,355,512,536]" italics="true" pageId="20" pageNumber="21">chocoensis</emphasis>
</taxonomicName>
it is 3-segmented, in 
<taxonomicName id="94B989E1FFB5FF8D29568805FD0A7661" box="[594,761,511,536]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mervini" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D29568805FD997661" box="[594,618,512,536]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D297E8805FD60766E" box="[634,659,512,535]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D29A78804FD0A7661" box="[675,761,513,536]" italics="true" pageId="20" pageNumber="21">mervini</emphasis>
</taxonomicName>
2-segmented. Additionally, the second segment of the endopod of leg 2 of 
<taxonomicName id="94B989E1FFB5FF8D2A428821FDE67645" box="[326,533,548,572]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D2A428821FEAC7645" box="[326,351,548,572]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2A6B8820FE7A7645" box="[367,393,549,572]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2A9E8821FDE67645" box="[410,533,548,572]" italics="true" pageId="20" pageNumber="21">chocoensis</emphasis>
</taxonomicName>
bears 4 setae instead of 
<quantity id="94415F87FFB5FF8D282F8821FCA57645" box="[811,854,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="20" pageNumber="21" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="94B989E1FFB5FF8D285A8821FBFB7645" box="[862,1032,548,572]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mervini" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D285A8821FC857645" box="[862,886,548,572]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D28838820FC527645" box="[903,929,549,572]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D28B68820FBFB7645" box="[946,1032,549,572]" italics="true" pageId="20" pageNumber="21">mervini</emphasis>
</taxonomicName>
and 
<taxonomicName id="94B989E1FFB5FF8D2F3B8821FB037645" box="[1087,1264,548,572]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="antillica" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D2F3B8821FBA47645" box="[1087,1111,548,572]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2F6C8820FB717645" box="[1128,1154,549,572]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2F978821FB037645" box="[1171,1264,548,572]" italics="true" pageId="20" pageNumber="21">antillica</emphasis>
</taxonomicName>
. No hump was observed at the apical end of the segment of the new species, as is present in both these species. The endopod of leg 4 is 1-segmented in 
<taxonomicName id="94B989E1FFB5FF8D2A7F8869FDDE76FD" box="[379,557,620,644]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="antillica" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D2A7F8869FE6776FD" box="[379,404,620,644]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D2AA18868FE4D76FD" box="[421,446,621,644]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2AD48869FDDE76FD" box="[464,557,620,644]" italics="true" pageId="20" pageNumber="21">antillica</emphasis>
</taxonomicName>
and 2-segmented in 
<taxonomicName id="94B989E1FFB5FF8D281E8869FC1A76FD" box="[794,1001,620,644]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D281E8869FCC176FD" box="[794,818,620,644]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D28478868FCAE76FD" box="[835,861,621,644]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D286A8869FC1A76FD" box="[878,1001,620,644]" italics="true" pageId="20" pageNumber="21">chocoensis</emphasis>
</taxonomicName>
, with 1 seta in the former and 
<quantity id="94415F87FFB5FF8D2E428869FA8176FD" box="[1350,1394,620,644]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="21" unit="in" value="2.0">2 in</quantity>
the latter. The caudal ramus in males of 
<taxonomicName id="94B989E1FFB5FF8D292A8895FD1B76D1" box="[558,744,655,680]" class="Maxillopoda" family="Canthocamptidae" genus="Canthocamptus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="striblingi" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D292A8895FDB476D1" box="[558,583,656,680]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D29538895FD8376DE" box="[599,624,656,679]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D29858895FD1B76D1" box="[641,744,656,680]" italics="true" pageId="20" pageNumber="21">striblingi</emphasis>
</taxonomicName>
is much longer than the anal somite.
</paragraph>
<paragraph id="5306F262FFB5FF8D2BCD88B1FEBE774E" blockId="20.[151,1436,151,823]" pageId="20" pageNumber="21">
A particular difference between females of 
<taxonomicName id="94B989E1FFB5FF8D29BF88B1FC9476B5" box="[699,871,692,716]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mervini" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D29BF88B1FD2076B5" box="[699,723,692,716]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D29E188B0FD0C76B5" box="[741,767,693,716]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D281588B0FC9476B5" box="[785,871,693,716]" italics="true" pageId="20" pageNumber="21">mervini</emphasis>
</taxonomicName>
and of 
<taxonomicName id="94B989E1FFB5FF8D28C588B1FB6176B5" box="[961,1170,692,716]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="chocoensis" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D28C588B1FC2976B5" box="[961,986,692,716]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D28E888B0FBF676B5" box="[1004,1029,693,716]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D2F1388B1FB6176B5" box="[1047,1170,692,716]" italics="true" pageId="20" pageNumber="21">chocoensis</emphasis>
</taxonomicName>
was noted in the ornamentation of the intercoxal plates. The new species bears 1 spine on each distal apical corner of the intercoxal plate on legs 2 and 4, instead of 2 spines in 
<taxonomicName id="94B989E1FFB5FF8D294688F9FD19776D" box="[578,746,764,788]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mervini" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D294688F9FDA9776D" box="[578,602,764,788]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D296E88F8FD77776D" box="[618,644,765,788]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D299088F8FD19776D" box="[660,746,765,788]" italics="true" pageId="20" pageNumber="21">mervini</emphasis>
</taxonomicName>
. Leg 3 of 
<taxonomicName id="94B989E1FFB5FF8D285988F9FBF5776D" box="[861,1030,764,788]" class="Maxillopoda" family="Canthocamptidae" genus="Attheyella" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mervini" subGenus="Canthosella">
<emphasis id="61CD2E70FFB5FF8D285988F9FC85776D" box="[861,886,764,788]" italics="true" pageId="20" pageNumber="21">A.</emphasis>
(
<emphasis id="61CD2E70FFB5FF8D288288F8FC6C776D" box="[902,927,765,788]" italics="true" pageId="20" pageNumber="21">C.</emphasis>
) 
<emphasis id="61CD2E70FFB5FF8D28B488F8FBF5776D" box="[944,1030,765,788]" italics="true" pageId="20" pageNumber="21">mervini</emphasis>
</taxonomicName>
has no spine there, but the new species has 1 spine.
</paragraph>
</subSubSection>
</treatment>
</document>