160 lines
19 KiB
XML
160 lines
19 KiB
XML
<document id="E9E9A531DCA6B0D71963CE9B3CFC3F98" ID-DOI="10.5281/zenodo.170508" ID-GBIF-Dataset="0d830f6b-b32d-4552-a4b0-f11046271145" ID-ISSN="11755326" ID-Zenodo-Dep="170508" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749171970" checkinUser="plazi" docAuthor="Menon, Federica" docDate="2005" docId="03F8E03FFFC0FFF8FE82F981FB82FBD7" docLanguage="en" docName="zt01087p058.pdf" docOrigin="Zootaxa 1087" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tettagalma striata Menon, new species" docType="treatment" docVersion="8" lastPageNumber="56" masterDocId="FFC19847FFC1FFFBFF8AFFDDFFD6FFAA" masterDocTitle="New record of Tettigarctidae (Insecta, Hemiptera, Cicadoidea) from the Lower Cretaceous of Brazil" masterLastPageNumber="58" masterPageNumber="53" pageNumber="54" updateTime="1698211349253" updateUser="plazi">
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<mods:mods id="25C4AE974120CB0EC0AE0C3FBAE004E5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C6B3A264F50D3ABC5CC659AF2B11B2B5">
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<mods:title id="4814D2BE8CDE681044D5B39E8202C326">New record of Tettigarctidae (Insecta, Hemiptera, Cicadoidea) from the Lower Cretaceous of Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="DA3AE1A0FB2EF6595B973A1260231B72" type="personal">
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<mods:role id="225771FD3C9F542216A6C2BC604143C2">
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<mods:roleTerm id="EF26AFAB0F1991B617D00424F1B081D2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E47B13B2F1B745E4C423BDCF99690DB">Menon, Federica</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="17C4470080F96708826A8620D5D8BF8C">text</mods:typeOfResource>
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<mods:relatedItem id="164F22DD9F4E9B33CCE0B9A96375D30D" type="host">
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<mods:titleInfo id="6C1264B282E7673ED5C87E93595C2C59">
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<mods:title id="75297F6B5BEFB1887B5F8392A0015368">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B22B1B133F3901F3ABBF28E14D4839E9">
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<mods:date id="F64C1076FDD0A5BD7AFCB4AB93793D9B">2005</mods:date>
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<mods:detail id="2E01DB1D58402BC1DC2E28C6415F3076" type="volume">
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<mods:number id="F435E6C9C23961923B0D1E1568D17336">1087</mods:number>
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</mods:detail>
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<mods:extent id="156DEDA3167BE3FD71269DDE602C7CFF" unit="page">
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<mods:start id="F17B34D6EE3647F136DBCDE69A8D011C">53</mods:start>
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<mods:end id="658ED540CA41DAD13DAD682FC1D54228">58</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="89C1666CDFDAFBA3968A639514C0080D">journal article</mods:classification>
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<mods:identifier id="1683521125D718BFE412E6BA4A961CB7" type="DOI">10.5281/zenodo.170508</mods:identifier>
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<mods:identifier id="B771CEB1326F6FD91E199AEF2548295E" type="GBIF-Dataset">0d830f6b-b32d-4552-a4b0-f11046271145</mods:identifier>
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<mods:identifier id="EA75002D9505C88658D3B91169F84CC3" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="4B55055B9AA583AC1770B68A2F3FC8D1" type="Zenodo-Dep">170508</mods:identifier>
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</mods:mods>
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<treatment id="03F8E03FFFC0FFF8FE82F981FB82FBD7" ID-DOI="http://doi.org/10.5281/zenodo.6265312" ID-GBIF-Taxon="119337312" ID-Zenodo-Dep="6265312" LSID="urn:lsid:plazi:treatment:03F8E03FFFC0FFF8FE82F981FB82FBD7" httpUri="http://treatment.plazi.org/id/03F8E03FFFC0FFF8FE82F981FB82FBD7" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54">
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<subSubSection id="C34B02A2FFC0FFFAFE82F981FEB8F933" pageId="1" pageNumber="54" type="nomenclature">
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<paragraph id="8BEE5129FFC0FFFAFE82F981FD09F9DC" blockId="1.[264,735,1628,1689]" box="[264,735,1628,1654]" pageId="1" pageNumber="54">
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<heading id="D0A6E645FFC0FFFAFE82F981FD09F9DC" bold="true" box="[264,735,1628,1654]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
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<emphasis id="B9258D3BFFC0FFFAFE82F981FD09F9DC" bold="true" box="[264,735,1628,1654]" pageId="1" pageNumber="54">
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<taxonomicName id="4C512AAAFFC0FFFAFE82F981FD94F9DC" ID-CoL="8JFPM" authority="Menon" authorityName="Menon" box="[264,578,1628,1654]" class="Insecta" family="Tettigarctidae" genus="Tettagalma" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="striata" status="sp. nov.">
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<emphasis id="B9258D3BFFC0FFFAFE82F981FE37F9DC" bold="true" box="[264,481,1628,1654]" italics="true" pageId="1" pageNumber="54">Tettagalma striata</emphasis>
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Menon
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2163040FFC0FFFAFDC5F981FD09F9DC" box="[591,735,1628,1654]" pageId="1" pageNumber="54" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEE5129FFC0FFFAFE82F9A2FEB8F933" blockId="1.[264,735,1628,1689]" box="[264,366,1663,1689]" pageId="1" pageNumber="54">
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<figureCitation id="136A4DACFFC0FFFAFE82F9A2FEB8F933" box="[264,366,1663,1689]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[264,368,1273,1297]" captionStartId-1="3.[264,368,1772,1796]" captionTargetBox-0="[304,1282,593,1246]" captionTargetBox-1="[425,1149,1253,1654]" captionTargetId-0="figure@2.[287,1294,591,1246]" captionTargetId-1="figure@3.[397,1189,1209,1745]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Tettagalma striata Menon, new genus and species; photograph of the holotype SMNS 65507. Scale bar represents 10 mm." captionText-1="FIGURE 2. Tettagalma striata Menon, new genus and species; in line drawing of the holotype SMNS 65507. Scale bar represents 5 mm." httpUri-0="https://zenodo.org/record/170509/files/figure.png" httpUri-1="https://zenodo.org/record/170510/files/figure.png" pageId="1" pageNumber="54">Figs 1–2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B02A2FFC0FFFAFE82F913FDFBF943" box="[264,557,1742,1769]" pageId="1" pageNumber="54" type="diagnosis">
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<paragraph id="8BEE5129FFC0FFFAFE82F913FDFBF943" blockId="1.[264,1323,1742,1849]" box="[264,557,1742,1769]" pageId="1" pageNumber="54">
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<emphasis id="B9258D3BFFC0FFFAFE82F913FE5EF942" bold="true" box="[264,392,1742,1768]" pageId="1" pageNumber="54">Diagnosis.</emphasis>
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<collectingCountry id="F34611B9FFC0FFFAFE05F912FE66F943" box="[399,432,1743,1769]" name="American Samoa" pageId="1" pageNumber="54">As</collectingCountry>
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for genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B02A2FFC0FFF8FEB2F92BFD14FDDC" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54" type="description">
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<paragraph id="8BEE5129FFC0FFFAFEB2F92BFC1FF893" blockId="1.[264,1323,1742,1849]" pageId="1" pageNumber="54">
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<emphasis id="B9258D3BFFC0FFFAFEB2F92BFE19F8BA" bold="true" box="[312,463,1782,1808]" pageId="1" pageNumber="54">Description.</emphasis>
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Adult in ventral aspect, with right forewing almost complete and left forewing largely missing; hind wings not preserved (
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<figureCitation id="136A4DACFFC0FFFAFCF9F8C2FC6EF893" box="[883,952,1823,1849]" captionStart="FIGURE 1" captionStartId="2.[264,368,1273,1297]" captionTargetBox="[304,1282,593,1246]" captionTargetId="figure@2.[287,1294,591,1246]" captionTargetPageId="2" captionText="FIGURE 1. Tettagalma striata Menon, new genus and species; photograph of the holotype SMNS 65507. Scale bar represents 10 mm." httpUri="https://zenodo.org/record/170509/files/figure.png" pageId="1" pageNumber="54">Fig. 1</figureCitation>
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).
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</paragraph>
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<paragraph id="8BEE5129FFC3FFF9FEB2FEC1FC27FE54" blockId="2.[264,1324,284,510]" pageId="2" pageNumber="55">
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Body. Head and thorax partly obscured by thin film of sediment. Head small,
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<quantity id="4CA9FCCCFFC3FFF9FB6FFEC1FAFAFE9C" box="[1253,1324,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="55" unit="mm" value="1.0">1 mm</quantity>
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long,
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<quantity id="4CA9FCCCFFC3FFF9FEC5FE99FE41FEF4" box="[335,407,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="55" unit="mm" value="3.0">3 mm</quantity>
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wide, bearing small, lateral eyes; antennae short, inserted before the eyes. Rostrum and most of cuticle not preserved. Thorax
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<quantity id="4CA9FCCCFFC3FFF9FC06FEB1FC03FE2C" box="[908,981,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="55" unit="mm" value="8.0">8 mm</quantity>
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long,
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<quantity id="4CA9FCCCFFC3FFF9FBA3FEB1FBA4FE2C" box="[1065,1138,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="55" unit="mm" value="9.0">9 mm</quantity>
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wide; legs not preserved. Abdomen
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<quantity id="4CA9FCCCFFC3FFF9FD93FE49FDA0FE04" box="[537,630,404,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="55" unit="mm" value="14.0">14 mm</quantity>
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long robust, elongated, pointed posteriorly; abdomen composed of nine visible segments of slightly different size (
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<figureCitation id="136A4DACFFC3FFF9FC65FE61FBE1FE7C" box="[1007,1079,444,470]" captionStart="FIGURE 1" captionStartId="2.[264,368,1273,1297]" captionTargetBox="[304,1282,593,1246]" captionTargetId="figure@2.[287,1294,591,1246]" captionTargetPageId="2" captionText="FIGURE 1. Tettagalma striata Menon, new genus and species; photograph of the holotype SMNS 65507. Scale bar represents 10 mm." httpUri="https://zenodo.org/record/170509/files/figure.png" pageId="2" pageNumber="55">Fig. 1</figureCitation>
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): second segment
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<quantity id="4CA9FCCCFFC3FFF9FA96FE61FEE0FE54" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="55" unit="mm" value="1.0">1 mm</quantity>
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long,
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<quantity id="4CA9FCCCFFC3FFF9FEF5FE39FE15FE54" box="[383,451,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="55" unit="mm" value="9.0">9 mm</quantity>
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wide; last segment 1.5 mm long, 2.5 mm wide.
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</paragraph>
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<caption id="DF2E01A1FFC3FFF9FE82FB24FD18FA99" httpUri="https://zenodo.org/record/170509/files/figure.png" pageId="2" pageNumber="55" targetBox="[304,1282,593,1246]" targetPageId="2">
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<paragraph id="8BEE5129FFC3FFF9FE82FB24FD18FA99" blockId="2.[264,1323,1273,1331]" pageId="2" pageNumber="55">
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<emphasis id="B9258D3BFFC3FFF9FE82FB24FE47FABB" bold="true" box="[264,401,1273,1297]" pageId="2" pageNumber="55">FIGURE 1.</emphasis>
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<taxonomicName id="4C512AAAFFC3FFF9FE15FB24FD13FABA" authority="Menon" authorityName="Menon" box="[415,709,1273,1296]" class="Insecta" family="Tettigarctidae" genus="Tettagalma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="striata" status="gen. nov.">
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<emphasis id="B9258D3BFFC3FFF9FE15FB24FDBCFABA" box="[415,618,1273,1296]" italics="true" pageId="2" pageNumber="55">Tettagalma striata</emphasis>
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Menon
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2163040FFC3FFF9FD52FB24FC86FABA" box="[728,848,1273,1296]" pageId="2" pageNumber="55" rank="genus">new genus</taxonomicNameLabel>
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and species; photograph of the holotype SMNS65507. Scale bar represents 10 mm.
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</paragraph>
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</caption>
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<paragraph id="8BEE5129FFC3FFF8FEB2FA54FD14FDDC" blockId="2.[264,1323,1417,1883]" lastBlockId="3.[264,1326,284,1150]" lastPageId="3" lastPageNumber="56" pageId="2" pageNumber="55">
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Forewings.
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<quantity id="4CA9FCCCFFC3FFF9FE48FA54FDC3FA09" box="[450,533,1417,1443]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="55" unit="mm" value="25.0">25 mm</quantity>
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long, elongated, with rounded apex, highly sclerotized, covered by thin layer of sediment, at least over basal area. Costal margin prominent, indented at nodus and thinning toward apex. Costal space slightly wider basally. Nodus situated
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<quantity id="4CA9FCCCFFC3FFF9FA81FA04FEE0F9B1" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="55" unit="mm" value="12.0">12 mm</quantity>
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from wing base. Sc+R origin at wing base, straight, prominent, bifurcating at approximately a quarter of wing length (
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<quantity id="4CA9FCCCFFC3FFF9FD6FF9F4FCFCF9E9" box="[741,810,1577,1603]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="55" unit="mm" value="6.0">6 mm</quantity>
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from wing base); R simple, almost parallel to costal margin, with slight indentation at nodal line, reaching the margin anteriorly to apex. Rs with three branches, basally straight, slightly sigmoidal distally, first two branches reaching margin anterior to apex (
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<figureCitation id="136A4DACFFC3FFF9FCA3F97CFCA1F911" box="[809,887,1697,1723]" captionStart="FIGURE 2" captionStartId="3.[264,368,1772,1796]" captionTargetBox="[425,1149,1253,1654]" captionTargetId="figure@3.[397,1189,1209,1745]" captionTargetPageId="3" captionText="FIGURE 2. Tettagalma striata Menon, new genus and species; in line drawing of the holotype SMNS 65507. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/170510/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
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). Crossvein rsma straight, simple, situated on distal half of wing, with anterior tip tilted toward wing base. M+CuA origin apparently from R, near wing base, straight, bifurcating at intersection with cross vein scp+rm+cua; the latter long, vertical, strongly prominent; M branches, MA and MP, originating before nodal line; MA forking well after nodal line; MA slightly indented at level of mamp1 crossvein; MP slightly sigmoid, with two apical branches. CuA simple, initially straight, curving anteriorly near nodal line, deflected abruptly along nodal line and curving posterorly to margin. Crossveins mpcua long, prominent, distally placed. CuP apparently straight and anal area not visible, obscured by body. Cross venation between the main longitudinal veins forming eight closed apical cells and four closed anteapical cells (
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<figureCitation id="136A4DACFFC2FFF8FEC7FE39FE43FE54" box="[333,405,484,510]" captionStart="FIGURE 2" captionStartId="3.[264,368,1772,1796]" captionTargetBox="[425,1149,1253,1654]" captionTargetId="figure@3.[397,1189,1209,1745]" captionTargetPageId="3" captionText="FIGURE 2. Tettagalma striata Menon, new genus and species; in line drawing of the holotype SMNS 65507. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/170510/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
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). Marginal membrane relatively wide, 0.40.
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<quantity id="4CA9FCCCFFC2FFF8FC1CFE39FC0AFE54" box="[918,988,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="56" unit="mm" value="6.0">6 mm</quantity>
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, bearing prominent striation. Striae equidistant, approximately 0.2 mm. Hairs and setae have left an impression basally on wing costal area. Original colour pattern across radial, medial, and cubital systems preserved as a dark brown coloration.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B02A2FFC2FFF8FEB2FD59FE38FD6C" pageId="3" pageNumber="56" type="materials_examined">
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<paragraph id="8BEE5129FFC2FFF8FEB2FD59FE38FD6C" blockId="3.[264,1326,284,1150]" pageId="3" pageNumber="56">
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<typeStatus id="54EAEF8BFFC2FFF8FEB2FD59FE7CFD34" box="[312,426,644,670]" pageId="3" pageNumber="56" type="holotype">
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<emphasis id="B9258D3BFFC2FFF8FEB2FD59FE7CFD34" bold="true" box="[312,426,644,670]" pageId="3" pageNumber="56">Holotype</emphasis>
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</typeStatus>
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.
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<collectionCode id="ED40C9ECFFC2FFF8FE4BFD59FDC7FD34" LSID="urn:lsid:biocol.org:col:34840" box="[449,529,644,670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="3" pageNumber="56">SMNS</collectionCode>
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65507, deposited in the Staatlisches Museum für Naturkunde Stuttgart,
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<collectingCountry id="F34611B9FFC2FFF8FEF1FD71FE3CFD6C" box="[379,490,684,710]" name="Germany" pageId="3" pageNumber="56">Germany</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B02A2FFC2FFF8FEB2FD09FE5AFCBC" pageId="3" pageNumber="56" type="etymology">
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<paragraph id="8BEE5129FFC2FFF8FEB2FD09FE5AFCBC" blockId="3.[264,1326,284,1150]" pageId="3" pageNumber="56">
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<emphasis id="B9258D3BFFC2FFF8FEB2FD09FE12FD44" bold="true" box="[312,452,724,750]" pageId="3" pageNumber="56">Etymology.</emphasis>
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From the Latin
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<emphasis id="B9258D3BFFC2FFF8FD01FD09FD09FD47" box="[651,735,724,749]" italics="true" pageId="3" pageNumber="56">
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<taxonomicName id="4C512AAAFFC2FFF8FD01FD09FD0CFD47" box="[651,730,724,749]" class="Insecta" family="Tettigarctidae" genus="Tettagalma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="striata">striata</taxonomicName>
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,
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</emphasis>
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in reference to the parallel striae in the marginal membrane.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B02A2FFC2FFF8FEB2FCF9FB82FBD7" pageId="3" pageNumber="56" type="discussion">
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<paragraph id="8BEE5129FFC2FFF8FEB2FCF9FB82FBD7" blockId="3.[264,1326,284,1150]" pageId="3" pageNumber="56">
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<emphasis id="B9258D3BFFC2FFF8FEB2FCF9FE67FC94" bold="true" box="[312,433,804,830]" pageId="3" pageNumber="56">Remarks.</emphasis>
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This genus and species is most similar to the extant
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<taxonomicName id="4C512AAAFFC2FFF8FBD8FCF9FB06FC97" box="[1106,1232,804,829]" class="Insecta" family="Tettigarctidae" genus="Tettigarcta" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B9258D3BFFC2FFF8FBD8FCF9FB06FC97" box="[1106,1232,804,829]" italics="true" pageId="3" pageNumber="56">Tettigarcta</emphasis>
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</taxonomicName>
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and to
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<emphasis id="B9258D3BFFC2FFF8FE82FC91FE04FCCF" box="[264,466,844,869]" italics="true" pageId="3" pageNumber="56">Liassotectigarcta</emphasis>
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from the Liassic of
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<collectingCountry id="F34611B9FFC2FFF8FD4AFC91FCF9FCCC" box="[704,815,844,870]" name="Germany" pageId="3" pageNumber="56">Germany</collectingCountry>
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. It differs from the Recent genus by having more branches of MA, the fork of M being placed more distally, closer to nodal line, and by the basal venation with the origin of M closer to the base. The most noticeable difference from
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<emphasis id="B9258D3BFFC2FFF8FE5FFC19FD76FC77" box="[469,672,964,989]" italics="true" pageId="3" pageNumber="56">Liassotectigarcta</emphasis>
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is the wing length (42.7 mm in
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<emphasis id="B9258D3BFFC2FFF8FBDEFC19FACFFC77" box="[1108,1305,964,989]" italics="true" pageId="3" pageNumber="56">Liassotettigarcta</emphasis>
|
||
). Furthermore, the new species differs also from this species mainly by the forewings being wider with rounded apex, and by having a wider marginal membrane. CuA in
|
||
<emphasis id="B9258D3BFFC2FFF8FE82FBE1FE1BFBFF" box="[264,461,1084,1109]" italics="true" pageId="3" pageNumber="56">Liassotettigarcta</emphasis>
|
||
is not deflected anteriorly at the nodal line as in
|
||
<emphasis id="B9258D3BFFC2FFF8FB85FBE1FB44FBFF" box="[1039,1170,1084,1109]" italics="true" pageId="3" pageNumber="56">Tettagalma</emphasis>
|
||
; this may be caused by the nodal line being more distally placed in
|
||
<emphasis id="B9258D3BFFC2FFF8FC0CFBB9FB82FBD7" box="[902,1108,1124,1149]" italics="true" pageId="3" pageNumber="56">Liassotettigarcta.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |