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<document id="EE92BC29291094FAA07B6A4742150F0D" ID-CLB-Dataset="22402" ID-DOI="10.5852/ejt.2015.128" ID-GBIF-Dataset="44b592de-b878-417f-80e1-a651f83183ad" ID-ISSN="2118-9773" ID-Zenodo-Dep="3785153" ID-ZooBank="FBAF2DED-5FD6-4F22-8A3A-E94733A92D73" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588612901601" checkinUser="carolina" docAuthor="Constant, Jérôme" docDate="2015" docId="03EFE724FFAF333E25DF970F1FAB8D76" docLanguage="en" docName="ejt-2015-128.pdf.imf" docOrigin="European Journal of Taxonomy 128" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pyrops effusus" docType="treatment" docVersion="6" lastPageNumber="6" masterDocId="FFD69F5CFFAC333B253C915F1A378828" masterDocTitle="Review of the effusus group of the Lanternfly genus Pyrops Spinola, 1839, with one new species and notes on trophobiosis (Hemiptera: Fulgoromorpha: Fulgoridae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="4" updateTime="1698838393947" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E82E8C66D9727112A4CDE1AD2D9CECCA">Review of the effusus group of the Lanternfly genus Pyrops Spinola, 1839, with one new species and notes on trophobiosis (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<treatment id="03EFE724FFAF333E25DF970F1FAB8D76" ID-DOI="http://doi.org/10.5281/zenodo.3794763" ID-GBIF-Taxon="163887569" ID-Zenodo-Dep="3794763" LSID="urn:lsid:plazi:treatment:03EFE724FFAF333E25DF970F1FAB8D76" httpUri="http://treatment.plazi.org/id/03EFE724FFAF333E25DF970F1FAB8D76" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<subSubSection id="C35C05B9FFAF333825DF970F18168E42" box="[227,545,1615,1642]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BF95632FFAF333825DF970F18168E42" blockId="3.[227,545,1615,1642]" box="[227,545,1615,1642]" pageId="3" pageNumber="4">
<heading id="D0B1E15EFFAF333825DF970F18168E42" bold="true" box="[227,545,1615,1642]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B9328A20FFAF333825DF970F18168E42" bold="true" box="[227,545,1615,1642]" pageId="3" pageNumber="4">
The
<taxonomicName id="4C462DB1FFAF3338242797101B458E41" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[283,370,1615,1641]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFAF3338242797101B458E41" bold="true" box="[283,370,1615,1641]" italics="true" pageId="3" pageNumber="4">effusus</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFAF333825DF972118678F0D" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BF95632FFAF333825DF972118678F0D" blockId="3.[227,1436,1662,1830]" pageId="3" pageNumber="4">
The characters of this very consistent group mainly follow
<bibRefCitation id="EFD72BC3FFAF333826A197211EBE8EB0" author="Baker's" box="[925,1161,1662,1689]" firstAuthor="Baker's" pageId="3" pageNumber="4" pagination="343 - 364" refId="ref9357" refString="Baker C. F. 1925. Remarks on certain Indo-Malayan Fulgora, with special reference to Philippines species. Philippines Journal of Science 28: 343 - 364." type="journal article" year="1925">Bakers (1925: 348)</bibRefCitation>
definition: (1) medium sized species; (2) cephalic process very short, clavate, bent in middle, strongly curved upwards, and with at least apical half blue or green; (3) tegmina green or blue basally, with yellow dots or markings; (4) hind wings blue basally and broadly black apically, the black colour extending towards the base along the main veins; (5) legs brown.
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFAF333E25DF96131EDA8947" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BF95632FFAF333825DF96131ECE8FDC" blockId="3.[227,1437,1868,2036]" pageId="3" pageNumber="4">
As compared to the list of included species given by
<bibRefCitation id="EFD72BC3FFAF3338260696131E718F4F" author="Nagai &amp; Porion" box="[826,1094,1868,1895]" firstAuthor="Nagai" pageId="3" pageNumber="4" refId="ref10327" refString="Nagai S. &amp; Porion T. 1996. Fulgoridae 2: Catalogue illustre des faunes asiatique et australienne. Sciences Nat, Compiegne." type="book" year="1996">Nagai &amp; Porion (1996)</bibRefCitation>
,
<taxonomicName id="4C462DB1FFAF3338216C96121F058F4E" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1845" box="[1104,1330,1869,1894]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cyanirostris">
<emphasis id="B9328A20FFAF3338216C96121F058F4E" box="[1104,1330,1869,1894]" italics="true" pageId="3" pageNumber="4">Pyrops cyanirostris</emphasis>
</taxonomicName>
(Guérin- Méneville, 1845) and
<taxonomicName id="4C462DB1FFAF333824D8962F19618FA2" authority="(Baker, 1925)" baseAuthorityName="Baker" baseAuthorityYear="1925" box="[484,854,1903,1930]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="maquilinganus">
<emphasis id="B9328A20FFAF333824D8962F18998FA1" box="[484,686,1903,1929]" italics="true" pageId="3" pageNumber="4">P. maquilinganus</emphasis>
(
<bibRefCitation id="EFD72BC3FFAF33382782963019798FA2" author="Baker" box="[702,846,1903,1930]" firstAuthor="Baker" pageId="3" pageNumber="4" pagination="343 - 364" refId="ref9357" refString="Baker C. F. 1925. Remarks on certain Indo-Malayan Fulgora, with special reference to Philippines species. Philippines Journal of Science 28: 343 - 364." type="journal article" year="1925">Baker, 1925</bibRefCitation>
)
</taxonomicName>
are here removed from the
<taxonomicName id="4C462DB1FFAF333821A5962F1EDD8FA2" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[1177,1258,1904,1930]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFAF333821A5962F1EDD8FA2" box="[1177,1258,1904,1930]" italics="true" pageId="3" pageNumber="4">effusus</emphasis>
</taxonomicName>
group, the first because it does not match characters (3) and (4), the latter because it does not match characters (2), (4) and (5).
<taxonomicName id="4C462DB1FFAF3338247896E818398FF8" baseAuthorityName="Baker" baseAuthorityYear="1925" box="[324,526,1974,2000]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="maquilinganus">
<emphasis id="B9328A20FFAF3338247896E818398FF8" box="[324,526,1974,2000]" italics="true" pageId="3" pageNumber="4">P. maquilinganus</emphasis>
</taxonomicName>
is returned to the
<taxonomicName id="4C462DB1FFAF333827E396E919698FF8" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[735,862,1974,2000]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="B9328A20FFAF333827E396E919698FF8" box="[735,862,1974,2000]" italics="true" pageId="3" pageNumber="4">candelaria</emphasis>
</taxonomicName>
group as originally proposed by
<bibRefCitation id="EFD72BC3FFAF333821E196E91F4C8FF9" author="Baker" box="[1245,1403,1974,2001]" firstAuthor="Baker" pageId="3" pageNumber="4" pagination="343 - 364" refId="ref9357" refString="Baker C. F. 1925. Remarks on certain Indo-Malayan Fulgora, with special reference to Philippines species. Philippines Journal of Science 28: 343 - 364." type="journal article" year="1925">Baker (1925)</bibRefCitation>
;
<taxonomicName id="4C462DB1FFAF333820BB96E81B598FDB" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1845" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cyanirostris">
<emphasis id="B9328A20FFAF333820BB96E81B598FDB" italics="true" pageId="3" pageNumber="4">P. cyanirostris</emphasis>
</taxonomicName>
cannot be attributed to any of the currently defined species groups in
<taxonomicName id="4C462DB1FFAF3338219C96851EC58FDB" authorityName="Spinola" authorityYear="1839" box="[1184,1266,2010,2035]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFAF3338219C96851EC58FDB" box="[1184,1266,2010,2035]" italics="true" pageId="3" pageNumber="4">Pyrops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF95632FFA8333F25DF906E190489BA" blockId="4.[227,1437,305,402]" pageId="4" pageNumber="5">
All 4 species included here in the group are restricted to Borneo and adjacent islands. Noteworthy,
<taxonomicName id="4C462DB1FFA8333F20BB906D1B598946" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1845" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cyanirostris">
<emphasis id="B9328A20FFA8333F20BB906D1B598946" italics="true" pageId="4" pageNumber="5">P. cyanirostris</emphasis>
</taxonomicName>
is recorded from Java (
<bibRefCitation id="EFD72BC3FFA8333F27BC900B19A08947" author="Guerin-Meneville" box="[640,919,340,367]" firstAuthor="Guerin-Meneville" pageId="4" pageNumber="5" pagination="96" refId="ref9895" refString="Guerin-Meneville F. E. 1845. (Fulgora cyanirostris. Guer.). Bulletin de la Societe Entomologique de France. (Series 2) 3: 96." type="journal article" year="1845">Guérin-Méneville 1845</bibRefCitation>
) and Sumatra (
<bibRefCitation id="EFD72BC3FFA8333F2176900B1F718947" author="Nagai &amp; Porion" box="[1098,1350,340,367]" firstAuthor="Nagai" pageId="4" pageNumber="5" refId="ref10327" refString="Nagai S. &amp; Porion T. 1996. Fulgoridae 2: Catalogue illustre des faunes asiatique et australienne. Sciences Nat, Compiegne." type="book" year="1996">Nagai &amp; Porion 1996</bibRefCitation>
) and
<taxonomicName id="4C462DB1FFA8333F20BB900A1BA689BA" baseAuthorityName="Baker" baseAuthorityYear="1925" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maquilinganus">
<emphasis id="B9328A20FFA8333F20BB900A1BA689BA" italics="true" pageId="4" pageNumber="5">P. maquilinganus</emphasis>
</taxonomicName>
from the
<collectingCountry id="F35116A2FFA8333F2738902818BF89B9" box="[516,648,375,401]" name="Philippines" pageId="4" pageNumber="5">Philippines</collectingCountry>
(
<bibRefCitation id="EFD72BC3FFA8333F27AB9028191189BA" author="Baker" box="[663,806,375,402]" firstAuthor="Baker" pageId="4" pageNumber="5" pagination="343 - 364" refId="ref9357" refString="Baker C. F. 1925. Remarks on certain Indo-Malayan Fulgora, with special reference to Philippines species. Philippines Journal of Science 28: 343 - 364." type="journal article" year="1925">Baker 1925</bibRefCitation>
).
</paragraph>
<paragraph id="8BF95632FFA8333F25DF90E618CC8A48" blockId="4.[227,1436,441,608]" pageId="4" pageNumber="5">
The group seems to be close to the
<emphasis id="B9328A20FFA8333F274390E6192889FB" box="[639,799,441,467]" italics="true" pageId="4" pageNumber="5">pyrorhynchus</emphasis>
species group which shares characters (3) and (4), but contains large sized species with an elongate cephalic process. However, the cephalic process in the latter group is also bent in the middle even if not so strongly curved upwards and slightly dilated apically like in
<taxonomicName id="4C462DB1FFA8333F249F937C1BC38A15" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[419,500,547,573]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA8333F249F937C1BC38A15" box="[419,500,547,573]" italics="true" pageId="4" pageNumber="5">effusus</emphasis>
</taxonomicName>
group. In some way the members of the
<taxonomicName id="4C462DB1FFA8333F26C8937C1E728A15" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[1012,1093,547,573]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA8333F26C8937C1E728A15" box="[1012,1093,547,573]" italics="true" pageId="4" pageNumber="5">effusus</emphasis>
</taxonomicName>
group could be regarded as dwarfed members of the
<emphasis id="B9328A20FFA8333F27349319189F8A48" box="[520,680,582,608]" italics="true" pageId="4" pageNumber="5">pyrorhynchus</emphasis>
group.
</paragraph>
<caption id="DF3906BAFFA8333F25DF96EF18688039" ID-DOI="http://doi.org/10.5281/zenodo.3785155" ID-Zenodo-Dep="3785155" httpUri="https://zenodo.org/record/3785155/files/figure.png" pageId="4" pageNumber="5" startId="4.[227,270,1968,1994]" targetBox="[233,1434,713,1927]" targetPageId="4">
<paragraph id="8BF95632FFA8333F25DF96EF1FAA8FC5" blockId="4.[227,1437,1968,2030]" pageId="4" pageNumber="5">
<emphasis id="B9328A20FFA8333F25DF96EF1B038FE2" bold="true" box="[227,308,1968,1994]" pageId="4" pageNumber="5">Fig. 1.</emphasis>
<taxonomicName id="4C462DB1FFA8333F240396EE183B8FE2" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[319,524,1969,1994]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="B9328A20FFA8333F240396EE183B8FE2" box="[319,524,1969,1994]" italics="true" pageId="4" pageNumber="5">Pyrops intricatus</emphasis>
</taxonomicName>
♀, Kalimantan, Mt Bawan (RBINS).
<emphasis id="B9328A20FFA8333F26DE96EF19C08FE2" bold="true" box="[994,1015,1968,1994]" pageId="4" pageNumber="5">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B9328A20FFA8333F202896EF1F1F8FE2" bold="true" box="[1300,1320,1968,1994]" pageId="4" pageNumber="5">B</emphasis>
. Habitus, ventral view.
<emphasis id="B9328A20FFA8333F24B8968C1BAE8FC5" bold="true" box="[388,409,2003,2029]" pageId="4" pageNumber="5">C</emphasis>
. Habitus, lateral view.
<emphasis id="B9328A20FFA8333F2793968B18F38FC6" bold="true" box="[687,708,2004,2030]" pageId="4" pageNumber="5">D</emphasis>
. Head and thorax, lateral view.
<emphasis id="B9328A20FFA8333F217F968B1E608FC6" bold="true" box="[1091,1111,2004,2030]" pageId="4" pageNumber="5">E</emphasis>
. Frons, normal view. Scale
</paragraph>
<paragraph id="8BF95632FFA8333F25DF96A918688039" blockId="4.[227,607,2038,2065]" box="[227,607,2038,2065]" pageId="4" pageNumber="5">
bar =
<quantity id="4CBEFBD7FFA8333F241B96A81B4D8039" box="[295,378,2039,2065]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="5" unit="mm" value="30.0">30 mm</quantity>
(DE not to scale).
</paragraph>
</caption>
<paragraph id="8BF95632FFA9333E25DF906E1EDA8947" blockId="5.[227,1436,305,367]" pageId="5" pageNumber="6">
Noteworthy, of all members of the
<emphasis id="B9328A20FFA9333E274A906E19218963" box="[630,790,305,331]" italics="true" pageId="5" pageNumber="6">pyrorhynchus</emphasis>
group, the Bornean species
<taxonomicName id="4C462DB1FFA9333E2160906D1FAB8964" authority="(Walker, 1857)" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[1116,1436,305,332]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="B9328A20FFA9333E2160906D1ED08963" box="[1116,1255,306,331]" italics="true" pageId="5" pageNumber="6">P. intricatus</emphasis>
(Walker, 1857)
</taxonomicName>
(
<figureCitation id="137D4AB7FFA9333E25D7900B1B188946" box="[235,303,340,366]" captionStart="Fig" captionStartId="4.[227,270,1968,1994]" captionTargetBox="[233,1434,713,1927]" captionTargetId="figure@4.[227,1436,711,1927]" captionTargetPageId="4" captionText="Fig. 1. Pyrops intricatus ♀, Kalimantan, Mt Bawan (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785155" httpUri="https://zenodo.org/record/3785155/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
) shows the greatest affinities in the pattern of the tegmina with the
<taxonomicName id="4C462DB1FFA9333E2174900B1EAE8946" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[1096,1177,340,366]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA9333E2174900B1EAE8946" box="[1096,1177,340,366]" italics="true" pageId="5" pageNumber="6">effusus</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFA9333E25DF90C9195F8998" box="[227,872,405,432]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BF95632FFA9333E25DF90C9195F8998" blockId="5.[227,872,405,432]" box="[227,872,405,432]" pageId="5" pageNumber="6">
<heading id="D0B1E15EFFA9333E25DF90C9195F8998" bold="true" box="[227,872,405,432]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B9328A20FFA9333E25DF90C9195F8998" bold="true" box="[227,872,405,432]" pageId="5" pageNumber="6">
Identification key to the species of the
<taxonomicName id="4C462DB1FFA9333E278390CA19218987" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[703,790,405,431]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA9333E278390CA19218987" bold="true" box="[703,790,405,431]" italics="true" pageId="5" pageNumber="6">effusus</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFA9333E25DF909A1B8C8A0D" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BF95632FFA9333E25DF909A1B8C8A0D" blockId="5.[227,1436,452,549]" pageId="5" pageNumber="6">
A Bornean species pertaining to the
<emphasis id="B9328A20FFA9333E27AA909B190189F6" box="[662,822,452,478]" italics="true" pageId="5" pageNumber="6">pyrorhynchus</emphasis>
group, but sharing a number of characters with the species of the
<taxonomicName id="4C462DB1FFA9333E24AF90B71BD38A2A" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[403,484,488,514]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA9333E24AF90B71BD38A2A" box="[403,484,488,514]" italics="true" pageId="5" pageNumber="6">effusus</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C462DB1FFA9333E277990B718E28A29" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[581,725,488,513]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="B9328A20FFA9333E277990B718E28A29" box="[581,725,488,513]" italics="true" pageId="5" pageNumber="6">P. intricatus</emphasis>
</taxonomicName>
(cf. above), is included in the key and illustrated to avoid misidentifications.
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFA9333E25DF93131FAB8D76" pageId="5" pageNumber="6" type="key">
<key id="FFD70103FFA9333E25DF93131FAB8D76" pageId="5" pageNumber="6">
<keyStep id="30B24E97FFA9333E25DF93131FAA8B3E" pageId="5" pageNumber="6">
<paragraph id="8BF95632FFA9333E25DF93131FAA8A85" blockId="5.[227,1437,588,791]" pageId="5" pageNumber="6">
<keyLead id="30B7F507FFA9333E25DF93131FAA8A85" pageId="5" pageNumber="6">
1 Head with cephalic process longer than half of tegmen (
<figureCitation id="137D4AB7FFA9333E26AC93131E278A4E" box="[912,1040,588,614]" captionStart="Fig" captionStartId="4.[227,270,1968,1994]" captionTargetBox="[233,1434,713,1927]" captionTargetId="figure@4.[227,1436,711,1927]" captionTargetPageId="4" captionText="Fig. 1. Pyrops intricatus ♀, Kalimantan, Mt Bawan (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785155" httpUri="https://zenodo.org/record/3785155/files/figure.png" pageId="5" pageNumber="6">Fig. 1A, C</figureCitation>
); cephalic process slightly curved upwards, brown with bright yellow apex, without blue or green (
<figureCitation id="137D4AB7FFA9333E211B93301E5B8AA1" box="[1063,1132,623,649]" captionStart="Fig" captionStartId="4.[227,270,1968,1994]" captionTargetBox="[233,1434,713,1927]" captionTargetId="figure@4.[227,1436,711,1927]" captionTargetPageId="4" captionText="Fig. 1. Pyrops intricatus ♀, Kalimantan, Mt Bawan (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785155" httpUri="https://zenodo.org/record/3785155/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
DE); ratio LPr/LF&gt; 4; LPr/BPrH&gt; 9 …………………………………
<taxonomicName id="4C462DB1FFA9333E260493CC1EBA8A85" ID-CoL="78PT6" authority="(Walker, 1857)" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[824,1165,658,685]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="B9328A20FFA9333E260493CC19FA8A85" bold="true" box="[824,973,659,685]" italics="true" pageId="5" pageNumber="6">P. intricatus</emphasis>
(Walker, 1857)
</taxonomicName>
[
<emphasis id="B9328A20FFA9333E219F93CC1F748A85" box="[1187,1347,659,685]" italics="true" pageId="5" pageNumber="6">pyrorhynchus</emphasis>
group]
</keyLead>
</paragraph>
<subSubSection id="C35C05B9FFA9333E25DF93E91FAA8B3E" pageId="5" pageNumber="6" type="description">
<paragraph id="8BF95632FFA9333E25DF93E91FAA8B3E" blockId="5.[227,1437,588,791]" pageId="5" pageNumber="6">
<keyLead id="30B7F507FFA9333E25DF93E91FAA8B3E" pageId="5" pageNumber="6">
Head with cephalic process shorter than half of tegmen (
<figureCitation id="137D4AB7FFA9333E26B093E91E398AF8" box="[908,1038,694,720]" captionStart="Fig" captionStartId="6.[227,270,1968,1994]" captionTargetBox="[229,1434,758,1923]" captionTargetId="figure@6.[227,1436,757,1928]" captionTargetPageId="6" captionText="Fig. 2. Pyrops effusus ♀, Kalimantan, Batulicin (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785157" httpUri="https://zenodo.org/record/3785157/files/figure.png" pageId="5" pageNumber="6">Figs 2A, C</figureCitation>
,
<figureCitation id="137D4AB7FFA9333E212593E91E568AF8" box="[1049,1121,694,720]" captionStart="Fig" captionStartId="11.[227,270,1968,1994]" captionTargetBox="[235,1430,747,1929]" captionTargetId="figure@11.[227,1436,739,1929]" captionTargetPageId="11" captionText="Fig. 4. Pyrops synavei sp. nov. ♀, holotype (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785161" httpUri="https://zenodo.org/record/3785161/files/figure.png" pageId="5" pageNumber="6">4A, C</figureCitation>
); cephalic process strongly curved upwards, at least partly green or blue (
<figureCitation id="137D4AB7FFA9333E2630938619648ADB" box="[780,851,729,755]" captionStart="Fig" captionStartId="6.[227,270,1968,1994]" captionTargetBox="[229,1434,758,1923]" captionTargetId="figure@6.[227,1436,757,1928]" captionTargetPageId="6" captionText="Fig. 2. Pyrops effusus ♀, Kalimantan, Batulicin (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785157" httpUri="https://zenodo.org/record/3785157/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
DE, 4DE); ratio LPr / LF &lt;2.5; LPr/BPrH &lt;7… ………………………………………………………………………………………………………
<emphasis id="B9328A20FFA9333E20B293A31FAA8B3E" bold="true" box="[1422,1437,764,790]" pageId="5" pageNumber="6">2</emphasis>
</keyLead>
</paragraph>
</subSubSection>
</keyStep>
<keyStep id="30B24E97FFA9333E25DF92611FAA8BB6" pageId="5" pageNumber="6">
<paragraph id="8BF95632FFA9333E25DF92611FAA8B53" blockId="5.[227,1437,830,927]" pageId="5" pageNumber="6">
<keyLead id="30B7F507FFA9333E25DF92611FAA8B53" pageId="5" pageNumber="6">
2 Basal half of head yellow-brown, strongly contrasting with green or blue apical half of cephalic process (
<figureCitation id="137D4AB7FFA9333E2457923E1BD98B53" box="[363,494,865,891]" captionStart="Fig" captionStartId="6.[227,270,1968,1994]" captionTargetBox="[229,1434,758,1923]" captionTargetId="figure@6.[227,1436,757,1928]" captionTargetPageId="6" captionText="Fig. 2. Pyrops effusus ♀, Kalimantan, Batulicin (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785157" httpUri="https://zenodo.org/record/3785157/files/figure.png" pageId="5" pageNumber="6">Fig. 2A, C</figureCitation>
) …………………………………………………………………………………
<emphasis id="B9328A20FFA9333E20B2923E1FAA8B53" bold="true" box="[1422,1437,865,891]" pageId="5" pageNumber="6">3</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF95632FFA9333E25DF92DA1FAA8BB6" blockId="5.[227,1437,830,927]" box="[227,1437,900,927]" pageId="5" pageNumber="6">
<keyLead id="30B7F507FFA9333E25DF92DA1FAA8BB6" box="[227,1437,900,927]" pageId="5" pageNumber="6">
Head unicolorous blue or green, at best slightly paler basally (
<figureCitation id="137D4AB7FFA9333E26E592DB1E6A8BB6" box="[985,1117,900,926]" captionStart="Fig" captionStartId="11.[227,270,1968,1994]" captionTargetBox="[235,1430,747,1929]" captionTargetId="figure@11.[227,1436,739,1929]" captionTargetPageId="11" captionText="Fig. 4. Pyrops synavei sp. nov. ♀, holotype (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785161" httpUri="https://zenodo.org/record/3785161/files/figure.png" pageId="5" pageNumber="6">Figs 4A, C</figureCitation>
,
<figureCitation id="137D4AB7FFA9333E215592DB1E878BB6" box="[1129,1200,900,927]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="5" pageNumber="6">5A, C</figureCitation>
) …………………
<emphasis id="B9328A20FFA9333E20B292DB1FAA8BB6" bold="true" box="[1422,1437,900,926]" pageId="5" pageNumber="6">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<subSubSection id="C35C05B9FFA9333E25DF92991FAB8CB8" pageId="5" pageNumber="6" type="reference_group">
<keyStep id="30B24E97FFA9333E25DF92991FAB8CB8" pageId="5" pageNumber="6">
<paragraph id="8BF95632FFA9333E25DF92991FAA8C0E" blockId="5.[227,1437,965,1168]" pageId="5" pageNumber="6">
<keyLead id="30B7F507FFA9333E25DF92991FAA8C0E" pageId="5" pageNumber="6">
3 Tegmina green on basal half, getting progressively more yellow on membrane, infuscate along apical margin; spots on tegmina yellow and rather large, visible on membrane (
<figureCitation id="137D4AB7FFA9333E218B92B61F218C2B" box="[1207,1302,1001,1027]" captionStart="Fig" captionStartId="6.[227,270,1968,1994]" captionTargetBox="[229,1434,758,1923]" captionTargetId="figure@6.[227,1436,757,1928]" captionTargetPageId="6" captionText="Fig. 2. Pyrops effusus ♀, Kalimantan, Batulicin (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785157" httpUri="https://zenodo.org/record/3785157/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
) ………… ………………………………………………………………………………
<taxonomicName id="4C462DB1FFA9333E214D95531FAA8C0E" ID-CoL="4QV9M" authority="(Distant, 1891)" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[1137,1437,1035,1062]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA9333E214D95531ED38C0D" bold="true" box="[1137,1252,1035,1062]" italics="true" pageId="5" pageNumber="6">P. effusus</emphasis>
(
<bibRefCitation id="EFD72BC3FFA9333E21CF95531FA38C0E" author="Distant" box="[1267,1428,1036,1062]" firstAuthor="Distant" pageId="5" pageNumber="6" pagination="517 - 519" refId="ref9767" refString="Distant W. L. 1891. XXIII. Descriptions of four new species of the genus Fulgora. Transactions of the Entomological Society of London 1891: 517 - 519." type="journal article" year="1891">Distant, 1891</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF95632FFA9333E25DF956F1FAB8CB8" blockId="5.[227,1437,965,1168]" pageId="5" pageNumber="6">
<keyLead id="30B7F507FFA9333E25DF956F1FAB8CB8" pageId="5" pageNumber="6">
Tegmina with basal half dark blue-green followed by a contrasting large oval yellow patch and broadly dark brown along apical margin; spots on tegmina pale yellow-white and rather small, getting indistinct on membrane (
<figureCitation id="137D4AB7FFA9333E27DC952A19778CB8" box="[736,832,1141,1168]" captionStart="Fig" captionStartId="9.[227,272,1966,1992]" captionTargetBox="[227,1435,803,1924]" captionTargetId="figure@9.[225,1436,801,1924]" captionTargetPageId="9" captionText="Fig. 3. Pyrops gunjii ♂, E Kalimantan, Samarinda, Mt Beratus (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)" figureDoi="http://doi.org/10.5281/zenodo.3785159" httpUri="https://zenodo.org/record/3785159/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
) …………………
<taxonomicName id="4C462DB1FFA9333E211D95291FAB8CB8" ID-CoL="9F54Z" authority="(Sato &amp; Nagai, 1994)" baseAuthorityName="Sato &amp; Nagai" baseAuthorityYear="1994" box="[1057,1436,1141,1168]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gunjii">
<emphasis id="B9328A20FFA9333E211D95291EBE8CA7" bold="true" box="[1057,1161,1141,1168]" italics="true" pageId="5" pageNumber="6">P. gunjii</emphasis>
(
<bibRefCitation id="EFD72BC3FFA9333E21A295291FA38CB8" author="Sato &amp; Nagai" box="[1182,1428,1141,1168]" firstAuthor="Sato" pageId="5" pageNumber="6" pagination="307 - 314" refId="ref10496" refString="Sato M. &amp; Nagai S. 1994. Taxonomic notes on the Southeast Asian species of the genus Fulgora (Homoptera: Fulgoridae). Transactions of the Shikoku Entomological Society 20 (3 - 4): 307 - 314." type="journal article" year="1994">Satô &amp; Nagai, 1994</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<keyStep id="30B24E97FFA9333E25DF95E81FAB8D76" pageId="5" pageNumber="6">
<subSubSection id="C35C05B9FFA9333E25DF95E81FAA8CDD" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BF95632FFA9333E25DF95E81FAA8CDD" blockId="5.[227,1437,1207,1374]" pageId="5" pageNumber="6">
<keyLead id="30B7F507FFA9333E25DF95E81FAA8CDD" pageId="5" pageNumber="6">
4 Tegmina with transverse yellow bands on corium, the basal one complete (
<figureCitation id="137D4AB7FFA9333E214895E81EE38CF9" box="[1140,1236,1207,1233]" captionStart="Fig" captionStartId="11.[227,270,1968,1994]" captionTargetBox="[235,1430,747,1929]" captionTargetId="figure@11.[227,1436,739,1929]" captionTargetPageId="11" captionText="Fig. 4. Pyrops synavei sp. nov. ♀, holotype (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785161" httpUri="https://zenodo.org/record/3785161/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
); pro- and mesonotum concolorous (
<figureCitation id="137D4AB7FFA9333E24CF958518658CDC" box="[499,594,1242,1268]" captionStart="Fig" captionStartId="11.[227,270,1968,1994]" captionTargetBox="[235,1430,747,1929]" captionTargetId="figure@11.[227,1436,739,1929]" captionTargetPageId="11" captionText="Fig. 4. Pyrops synavei sp. nov. ♀, holotype (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785161" httpUri="https://zenodo.org/record/3785161/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
); head dark olivaceous blue-green (
<figureCitation id="137D4AB7FFA9333E26C995851E648CDC" box="[1013,1107,1242,1268]" captionStart="Fig" captionStartId="11.[227,270,1968,1994]" captionTargetBox="[235,1430,747,1929]" captionTargetId="figure@11.[227,1436,739,1929]" captionTargetPageId="11" captionText="Fig. 4. Pyrops synavei sp. nov. ♀, holotype (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785161" httpUri="https://zenodo.org/record/3785161/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
, DE) …
<taxonomicName id="4C462DB1FFA9333E21F895841F0C8CDC" ID-CoL="4QVBK" authorityName="Constant" authorityYear="2015" box="[1220,1339,1242,1269]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="synavei" status="sp. nov.">
<emphasis id="B9328A20FFA9333E21F895841F0C8CDC" bold="true" box="[1220,1339,1242,1269]" italics="true" pageId="5" pageNumber="6">P. synavei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201375BFFA9333E207E95841FAA8CDD" box="[1346,1437,1243,1269]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFA9333E25DF95A11FAB8D76" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BF95632FFA9333E25DF95A11FAB8D76" blockId="5.[227,1437,1207,1374]" pageId="5" pageNumber="6">
<keyLead id="30B7F507FFA9333E25DF95A11FAB8D76" pageId="5" pageNumber="6">
<emphasis id="B9328A20FFA9333E25DF95A11AC58D30" bold="true" box="[227,242,1278,1304]" pageId="5" pageNumber="6"></emphasis>
Tegmina with small round yellow spots (
<figureCitation id="137D4AB7FFA9333E27C595A2196B8D30" box="[761,860,1277,1304]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="5" pageNumber="6">Fig. 5A</figureCitation>
); pronotum yellow to orange, mesonotum dark blue-green (
<figureCitation id="137D4AB7FFA9333E24AB947F1BCA8D13" box="[407,509,1312,1339]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="5" pageNumber="6">Fig. 5A</figureCitation>
); head bright blue (often faded in collection specimens) (
<figureCitation id="137D4AB7FFA9333E21DF947F1F7F8D13" box="[1251,1352,1312,1339]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (DE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="5" pageNumber="6">Fig. 5A</figureCitation>
, DE) …………………………………………………………………………
<taxonomicName id="4C462DB1FFA9333E210A941B1FAB8D76" ID-CoL="6JQHY" authority="(Distant, 1889)" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[1078,1436,1347,1374]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA9333E210A941B1EEC8D76" bold="true" box="[1078,1243,1347,1374]" italics="true" pageId="5" pageNumber="6">P. whiteheadi</emphasis>
(
<bibRefCitation id="EFD72BC3FFA9333E21D3941B1FA38D76" author="Distant" box="[1263,1428,1348,1374]" firstAuthor="Distant" pageId="5" pageNumber="6" pagination="419 - 422" refId="ref9724" refString="Distant W. L. 1889. XLIX. Descriptions of new species of Rhynchota collected on or near the Kina Balu Mountain, North Borneo. Annals and Magazine of Natural History (Series 6) 3: 419 - 422." type="journal article" year="1889">Distant, 1889</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</subSubSection>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>