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<document id="ED5A3A1D6BC5EE34D2F8E7C2E55E3000" ID-DOI="10.2478/if-2019-0023" ID-ISSN="2533-4069" ID-Zenodo-Dep="5457521" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630294627634" checkinUser="diego" docAuthor="Hartman, Julian, Van De Weerd, Andrew A., Bruijn, Hans De &amp; Wessels, Wilma" docDate="2019" docId="03E6B8178C3A617FFF9DFE0D7FE7FBD3" docLanguage="en" docName="06_Hartman.pdf" docOrigin="Fossil Imprint 75 (3 - 4)" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Sayimys WOOD 1937" docType="treatment" docVersion="4" lastPageNumber="373" masterDocId="FFDFC06F8C346171FFE7FFC67F71FFDB" masterDocTitle="An Exceptional Large Sample Of The Early Miocene Ctenodactyline Rodent Sayimys Giganteus, Specific Variation And Taxonomic Implications" masterLastPageNumber="382" masterPageNumber="359" pageNumber="373" updateTime="1699218057821" updateUser="plazi">
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<mods:title id="B28F68186287E6519C06C2B6BF3D4730">An Exceptional Large Sample Of The Early Miocene Ctenodactyline Rodent Sayimys Giganteus, Specific Variation And Taxonomic Implications</mods:title>
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<mods:namePart id="73302CB3D1452051585F65065545F066">Hartman, Julian</mods:namePart>
<mods:affiliation id="BC8899CF4D284E64AC482F522CD0755D">Oranjepoldererf 36, 2807 NE Gouda, the Netherlands; e-mail: juulhartman @ gmail. com. &amp; Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:namePart id="879FAD929584B74F75AA6D1681F896CF">Van De Weerd, Andrew A.</mods:namePart>
<mods:affiliation id="416F23A779BB9CD7BFEA383E37FFC829">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:namePart id="7BBE7F9ADEE47A0ADD94015DF68ED3B8">Bruijn, Hans De</mods:namePart>
<mods:affiliation id="1FB5C8462A62181E04792D7488E2CC41">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:namePart id="658CD029A9F9FB83B6D777435888AEDD">Wessels, Wilma</mods:namePart>
<mods:affiliation id="2B14F6274FC0640148E50276FA8FE0D3">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<treatment id="03E6B8178C3A617FFF9DFE0D7FE7FBD3" ID-DOI="http://doi.org/10.5281/zenodo.5495920" ID-Zenodo-Dep="5495920" LSID="urn:lsid:plazi:treatment:03E6B8178C3A617FFF9DFE0D7FE7FBD3" httpUri="http://treatment.plazi.org/id/03E6B8178C3A617FFF9DFE0D7FE7FBD3" lastPageNumber="373" pageId="14" pageNumber="373">
<subSubSection id="C3555A8A8C3A617FFF9DFE0D7DA9FDC8" pageId="14" pageNumber="373" type="nomenclature">
<paragraph id="8BF009018C3A617FFF9DFE0D7DA9FE38" blockId="14.[122,728,459,483]" box="[122,728,459,483]" pageId="14" pageNumber="373">
<heading id="D0B8BE6D8C3A617FFF9DFE0D7DA9FE38" box="[122,728,459,483]" centered="true" fontSize="8" level="2" pageId="14" pageNumber="373" reason="2">
<emphasis id="B93BD5138C3A617FFF9DFE0D7EB1FE38" bold="true" box="[122,448,459,483]" pageId="14" pageNumber="373">
<emphasis id="B93BD5138C3A617FFF9DFE0D7E1FFE38" bold="true" box="[122,366,459,483]" italics="true" pageId="14" pageNumber="373">
<taxonomicName id="4C4F72828C3A617FFF9DFE0D7FA5FE38" authorityName="WOOD" authorityYear="1937" box="[122,212,459,483]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="373" phylum="Chordata" rank="genus">Sayimys</taxonomicName>
assarrarensis
</emphasis>
LÓPEZ-
</emphasis>
<emphasis id="B93BD5138C3A617FFE26FE0D7DA9FE38" bold="true" box="[449,728,459,483]" pageId="14" pageNumber="373">ANTOÑANZAS et SEN, 2004</emphasis>
</heading>
</paragraph>
<paragraph id="8BF009018C3A617FFF77FE3D7DA9FDC8" blockId="14.[144,728,507,531]" box="[144,728,507,531]" pageId="14" pageNumber="373">H o l o t y p e. A fragmentary left maxilla with P4M2.</paragraph>
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<subSubSection id="C3555A8A8C3A617FFF77FDED7FE7FBD3" pageId="14" pageNumber="373" type="description">
<paragraph id="8BF009018C3A617FFF77FDED7D02FD99" blockId="14.[144,627,554,579]" box="[144,627,554,579]" pageId="14" pageNumber="373">P a r a t y p e s. Some 18 isolated cheek teeth.</paragraph>
<paragraph id="8BF009018C3A617FFF77FD9D7E38FD68" blockId="14.[106,745,602,691]" pageId="14" pageNumber="373">
Ty p e l o c a l i t y. As-Sarrar, Dam Formation,
<collectingCountry id="F35849918C3A617FFD4BFD9C7FC7FD48" name="Saudi Arabia" pageId="14" pageNumber="373">Saudi Arabia</collectingCountry>
; assigned age of the site: later part of the early Miocene, 1517 Ma.
</paragraph>
<paragraph id="8BF009018C3A617FFF77FD0D7FE7FBD3" blockId="14.[106,746,715,1032]" pageId="14" pageNumber="373">
D i s c u s s i o n. This small species is similar in size to
<taxonomicName id="4C4F72828C3A617FFF6EFD2A7E18FCD8" baseAuthorityName="SEN et THOMAS" baseAuthorityYear="1979" box="[137,361,747,771]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="373" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B93BD5138C3A617FFF6EFD2A7E18FCD8" box="[137,361,747,771]" italics="true" pageId="14" pageNumber="373">Sayimys intermedius</emphasis>
</taxonomicName>
. The As Sarrar locality is slightly older than the
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locality of
<taxonomicName id="4C4F72828C3A617FFE21FCCA7D12FCFF" baseAuthorityName="SEN et THOMAS" baseAuthorityYear="1979" box="[454,611,780,804]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="373" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B93BD5138C3A617FFE21FCCA7EA2FCF8" box="[454,467,780,803]" italics="true" pageId="14" pageNumber="373">S</emphasis>
.
<emphasis id="B93BD5138C3A617FFE02FCCA7D12FCFF" box="[485,611,780,804]" italics="true" pageId="14" pageNumber="373">intermedius</emphasis>
</taxonomicName>
. The single reasonably preserved dp4 of
<emphasis id="B93BD5138C3A617FFE46FCEB7EDFFC9F" box="[417,430,813,836]" italics="true" pageId="14" pageNumber="373">S</emphasis>
.
<emphasis id="B93BD5138C3A617FFE5AFCEB7D3CFC9F" box="[445,589,813,836]" italics="true" pageId="14" pageNumber="373">assarrarensis</emphasis>
shows a welldeveloped mesolophid and a slightly elongated anteroconid. Considering the large variation in dental morphology and size of large
<taxonomicName id="4C4F72828C3A617FFF24FC487E6BFC7E" authorityName="WOOD" authorityYear="1937" box="[195,282,910,933]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B93BD5138C3A617FFF24FC487E6BFC7E" box="[195,282,910,933]" italics="true" pageId="14" pageNumber="373">Sayimys</emphasis>
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assemblages, we consider
<emphasis id="B93BD5138C3A617FFDDAFC487D3BFC7E" box="[573,586,910,933]" italics="true" pageId="14" pageNumber="373">S</emphasis>
.
<emphasis id="B93BD5138C3A617FFDBFFC487D99FC7E" box="[600,744,910,933]" italics="true" pageId="14" pageNumber="373">assarrarensis</emphasis>
a junior synonym of
<taxonomicName id="4C4F72828C3A617FFEA2FC697EADFC1D" baseAuthorityName="SEN et THOMAS" baseAuthorityYear="1979" box="[325,476,942,966]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="373" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B93BD5138C3A617FFEA2FC697E23FC1D" box="[325,338,943,966]" italics="true" pageId="14" pageNumber="373">S</emphasis>
.
<emphasis id="B93BD5138C3A617FFEB9FC687EADFC1D" box="[350,476,942,966]" italics="true" pageId="14" pageNumber="373">intermedius</emphasis>
</taxonomicName>
. More material is needed from the
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localities to ascertain specific differences, if any.
</paragraph>
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