treatments-xml/data/03/E4/A4/03E4A47FFF88FFD1D1D1AB078A2CFC25.xml
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<document id="0E357B2647A2D2672BF31E736D0C3F69" ID-DOI="10.11646/zootaxa.5380.5.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10254481" ID-ZooBank="CC3989CD-4E87-4DC6-86AF-8BF398445CF0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701678513813" checkinUser="plazi" docAuthor="Farhangi, Seyed Vahid, Nozari, Jamasb &amp; Gharali, Babak" docDate="2023" docId="03E4A47FFF88FFD1D1D1AB078A2CFC25" docLanguage="en" docName="zootaxa.5380.5.6.pdf" docOrigin="Zootaxa 5380 (5)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5380.5.6/52403" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phthiria portokali Farhangi &amp; Nozari &amp; Gharali 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="496" masterDocId="FFDDDC07FF89FFD4D146A95C880BFFA5" masterDocTitle="A new bee fly species of the genus Phthiria (Diptera, Bombyliidae, Phthiriinae, Phthiriini) from the west of Iran" masterLastPageNumber="497" masterPageNumber="491" pageNumber="492" updateTime="1701679028319" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="85517EF54332E74F982DC97A3A6F50BC">A new bee fly species of the genus Phthiria (Diptera, Bombyliidae, Phthiriinae, Phthiriini) from the west of Iran</mods:title>
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<mods:namePart id="F3E054064551AE14A1E7DF92E24B94B8">Farhangi, Seyed Vahid</mods:namePart>
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<mods:affiliation id="45103A5D3BEE5B89D8E013D3EAF839A9">Department of Plant Protection, College of Agriculture and Natural Resources, University of Tehran, 31587 - 77871 Karaj, Iran seyedvahid. farhangi @ yahoo. com; https: // orcid. org / 0009 - 0001 - 0542 - 1950</mods:affiliation>
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<mods:namePart id="68D5D1EF82631C0CFEC70C6E46499356">Nozari, Jamasb</mods:namePart>
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<mods:affiliation id="14785E41060D2B74F5A2BB48ABEDADA6">Department of Plant Protection, College of Agriculture and Natural Resources, University of Tehran, 31587 - 77871 Karaj, Iran seyedvahid. farhangi @ yahoo. com; https: // orcid. org / 0009 - 0001 - 0542 - 1950</mods:affiliation>
<mods:nameIdentifier id="59E24B587F7DB05B0738E12595724582" type="email">seyedvahid.farhangi@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="C969C576239527F3BCE271C61A8B0D8F">Gharali, Babak</mods:namePart>
<mods:affiliation id="4860AFDAD6584133A45FE74F15EB1D83">Plant Protection Research Department, Qazvin Agricultural and Natural Resources Research and Education Center, AREEO, Qazvin, Iran</mods:affiliation>
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<emphasis id="B939C97BFF88FFD5D1D1AB0789DDFDD0" bold="true" box="[151,470,603,630]" pageId="1" pageNumber="492">
<taxonomicName id="4C4D6EEAFF88FFD5D1D1AB07897AFDD3" authority="Farhangi &amp; Nozari &amp; Gharali, 2023" authorityName="Farhangi &amp; Nozari &amp; Gharali" authorityYear="2023" box="[151,369,603,630]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="portokali" status="sp. nov.">
<emphasis id="B939C97BFF88FFD5D1D1AB07897AFDD3" bold="true" box="[151,369,603,630]" italics="true" pageId="1" pageNumber="492">Phthiria portokali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A7400FF88FFD5D03EAB0789DDFDD0" box="[376,470,603,629]" pageId="1" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BF21569FF88FFD5D1D1ABFF8D03FCA0" blockId="1.[151,1437,675,1818]" pageId="1" pageNumber="492">
<emphasis id="B939C97BFF88FFD5D1D1ABFF894AFD18" bold="true" box="[151,321,675,701]" pageId="1" pageNumber="492">Type material</emphasis>
.
<materialsCitation id="3B251F34FF88FFD5D014ABFF8D03FCA0" collectingDate="2012-08-20" collectingDateMax="2014-07-07" collectingDateMin="2012-08-20" collectionCode="IRIPP, R" country="Iran" elevation="2115" latitude="35.531666" location="Highway Razan-Avaj" longLatPrecision="19" longitude="49.197224" municipality="Provinces" pageId="1" pageNumber="492" specimenCount="14" specimenCount-female="1" specimenCount-male="6" stateProvince="Hamadan" typeStatus="holotype">
<typeStatus id="54F6ABCBFF88FFD5D014ABFF89B1FD18" box="[338,442,675,701]" pageId="1" pageNumber="492" type="holotype">Holotype</typeStatus>
female (deposited in
<collectionCode id="ED5C8DACFF88FFD5D3FEABFF8B0AFD18" box="[696,769,675,701]" pageId="1" pageNumber="492">IRIPP</collectionCode>
),
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(between
<collectingMunicipality id="6B968F13FF88FFD5D5C0ABFF8CFFFD18" box="[1158,1268,675,701]" pageId="1" pageNumber="492">Provinces</collectingMunicipality>
<collectingRegion id="4989DB8BFF88FFD5D5B8ABFF8D62FD18" box="[1278,1385,675,701]" country="Iran" name="Hamadan" pageId="1" pageNumber="492">Hamadan</collectingRegion>
and
<collectingRegion id="4989DB8BFF88FFD5D1D1AB9B88E0FD44" box="[151,235,711,737]" country="Iran" name="Qazvin" pageId="1" pageNumber="492">Qazvin</collectingRegion>
, west of
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),
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a.s.l.
</elevation>
,
<geoCoordinate id="EE7973AEFF88FFD5D36FAB948AC2FD47" box="[553,713,711,738]" degrees="35" direction="north" minutes="31" orientation="latitude" pageId="1" pageNumber="492" precision="15" seconds="54" value="35.531666">N 35° 31 54</geoCoordinate>
,
<geoCoordinate id="EE7973AEFF88FFD5D392AB948B64FD47" box="[724,879,711,738]" degrees="49" direction="east" minutes="11" orientation="longitude" pageId="1" pageNumber="492" precision="15" seconds="50" value="49.197224">E 49° 11 50</geoCoordinate>
,
<date id="FFF333A9FF88FFD5D23CAB9B8C26FD44" box="[890,1069,711,737]" pageId="1" pageNumber="492" value="2012-08-20">
<collectingDate id="EFB7CA41FF88FFD5D23CAB9B8C26FD44" box="[890,1069,711,737]" pageId="1" pageNumber="492" value="2012-08-20">20 August 2012</collectingDate>
</date>
,
<collectingMethod id="520C6D7EFF88FFD5D571AB9B8CE8FD47" box="[1079,1251,711,738]" pageId="1" pageNumber="492">white pan traps</collectingMethod>
, leg.
<collectionCode id="ED5C8DACFF88FFD5D45BAB948D3BFD47" box="[1309,1328,712,738]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="492">R</collectionCode>
Abdolahi Mesbah;
<specimenCount id="9D4BDEE0FF88FFD5D1B9ABB08974FCA3" box="[255,383,748,774]" count="7" pageId="1" pageNumber="492" type="generic" typeStatus="paratype">7 paratypes</specimenCount>
(
<specimenCount id="9D4BDEE0FF88FFD5D0D4ABB789B8FCA0" box="[402,435,747,773]" count="1" pageId="1" pageNumber="492" type="female">1♀</specimenCount>
,
<specimenCount id="9D4BDEE0FF88FFD5D086ABB789EFFCA0" box="[448,484,747,773]" count="6" pageId="1" pageNumber="492" type="male">6♂</specimenCount>
) (deposited in BG), same data of the
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except date,
<date id="FFF333A9FF88FFD5D538ABB08D0FFCA0" box="[1150,1284,747,774]" pageId="1" pageNumber="492" value="2014-07-07">
<collectingDate id="EFB7CA41FF88FFD5D538ABB08D0FFCA0" box="[1150,1284,747,774]" pageId="1" pageNumber="492" value="2014-07-07">7 July 2014</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35746E2FF88FFD5D181AA538C95FC1C" pageId="1" pageNumber="492" type="diagnosis">
<paragraph id="8BF21569FF88FFD5D181AA538C95FC1C" blockId="1.[151,1437,675,1818]" pageId="1" pageNumber="492">
<emphasis id="B939C97BFF88FFD5D181AA538949FC8C" bold="true" box="[199,322,783,809]" pageId="1" pageNumber="492">Diagnosis.</emphasis>
<taxonomicName id="4C4D6EEAFF88FFD5D00FAA538A1FFC8C" authority="Farhangi &amp; Nozari &amp; Gharali, 2023" authorityName="Farhangi &amp; Nozari &amp; Gharali" authorityYear="2023" box="[329,532,783,809]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="portokali" status="sp. nov.">
<emphasis id="B939C97BFF88FFD5D00FAA538A1FFC8C" box="[329,532,783,809]" italics="true" pageId="1" pageNumber="492">Phthiria portokali</emphasis>
</taxonomicName>
<emphasis id="B939C97BFF88FFD5D35DAA538A7EFC8C" bold="true" box="[539,629,783,809]" pageId="1" pageNumber="492">
<taxonomicNameLabel id="A20A7400FF88FFD5D35DAA538A7EFC8C" box="[539,629,783,809]" pageId="1" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a unique species among Iranian species of
<taxonomicName id="4C4D6EEAFF88FFD5D530AA538CD8FC8C" authorityName="Meigen" authorityYear="1820" box="[1142,1235,783,809]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="492" phylum="Arthropoda" rank="genus">
<emphasis id="B939C97BFF88FFD5D530AA538CD8FC8C" box="[1142,1235,783,809]" italics="true" pageId="1" pageNumber="492">Phthiria</emphasis>
</taxonomicName>
by having a large tortoise-shell-shaped orange pattern on the dorsum of mesonotum in females. In the
<bibRefCitation id="EFDC6898FF88FFD5D57AAA6F8CEBFCE8" author="Engel, O." box="[1084,1248,819,845]" pageId="1" pageNumber="492" pagination="97 - 144" refId="ref2151" refString="Engel, O. (1933) Bombyliidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. IV (3). E. Schweizerbart, Stuttgart, pp. 97 - 144." type="book chapter" year="1933">Engels (1933)</bibRefCitation>
key, females run to couplet 9 where the new species is easily separated from
<taxonomicName id="4C4D6EEAFF88FFD5D26EAA048C6AFCD4" authority="Loew, 1873" authorityName="Loew" authorityYear="1873" box="[808,1121,855,882]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="quadrinotata">
<emphasis id="B939C97BFF88FFD5D26EAA048B32FCD4" box="[808,825,856,881]" italics="true" pageId="1" pageNumber="492">P</emphasis>
.
<emphasis id="B939C97BFF88FFD5D200AA0B8BD3FCD4" box="[838,984,855,881]" italics="true" pageId="1" pageNumber="492">quadrinotata</emphasis>
<bibRefCitation id="EFDC6898FF88FFD5D299AA048C6AFCD4" author="Loew, H." box="[991,1121,855,882]" pageId="1" pageNumber="492" refId="ref2423" refString="Loew, H. (1873) Beschreibungen europaischer Dipteren. Systematische Beschreibung der bekannten europaischen zweiflugeligen Insecten. Von Johann Wilhelm Meigen. Zehnter Theil oder vierter Supplementband. Beschreibungen europaischer Dipteren. Dritter Band. H. W. Schmidt, Halle, viii + 320 pp." type="book" year="1873">Loew, 1873</bibRefCitation>
</taxonomicName>
by the large orange mark on mesonotum and the completely yellow halter. The male specimens go to the couplet 5 and it is easily distinguished by completely yellow halter (with a black spot on the halter knob in
<taxonomicName id="4C4D6EEAFF88FFD5D2D3AAFC8C99FC1C" authority="Loew" authorityName="Loew" authorityYear="1873" box="[917,1170,927,954]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="quadrinotata">
<emphasis id="B939C97BFF88FFD5D2D3AAFC8BADFC1C" box="[917,934,928,953]" italics="true" pageId="1" pageNumber="492">P</emphasis>
.
<emphasis id="B939C97BFF88FFD5D2F2AAC38C4DFC1C" box="[948,1094,927,953]" italics="true" pageId="1" pageNumber="492">quadrinotata</emphasis>
Loew
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35746E2FF88FFD1D181AA9F88C9FE51" lastPageId="5" lastPageNumber="496" pageId="1" pageNumber="492" type="description">
<paragraph id="8BF21569FF88FFD5D181AA9F8BE7FC7B" blockId="1.[151,1437,675,1818]" box="[199,1004,963,990]" pageId="1" pageNumber="492">
<emphasis id="B939C97BFF88FFD5D181AA9F895AFC78" bold="true" box="[199,337,963,989]" pageId="1" pageNumber="492">Description</emphasis>
.
<typeStatus id="54F6ABCBFF88FFD5D026AA9F89C3FC78" box="[352,456,963,989]" pageId="1" pageNumber="492" type="holotype">Holotype</typeStatus>
female. Body length:
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, wing length:
<quantity id="4CB5B88CFF88FFD5D2E2AA9F8BE3FC7B" box="[932,1000,963,990]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="492" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<paragraph id="8BF21569FF88FFD5D181AABB8D1AF8BC" blockId="1.[151,1437,675,1818]" pageId="1" pageNumber="492">
<emphasis id="B939C97BFF88FFD5D181AABB890CFBA4" bold="true" box="[199,263,999,1025]" pageId="1" pageNumber="492">Head</emphasis>
. (
<figureCitation id="137609ECFF88FFD5D059AABB8993FBA4" box="[287,408,999,1025]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[179,1407,179,1917]" captionTargetId="figure-19@2.[179,1407,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Phthiria portokali sp. nov,. Holotype female; A) lateral view; B) antennae; C) head, lateral view; D) head, dorsal view; E) pleura; F) abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254483" httpUri="https://zenodo.org/record/10254483/files/figure.png" pageId="1" pageNumber="492">Figs. 1ad</figureCitation>
) Eyes dichoptic. All facets of same size. Occiput yellow with white hairs, length of hairs subequal to combined length of two basal antennal segments; vertex yellow; frons yellow, with blackish triangular mark medially, with clear depression at base (
<figureCitation id="137609ECFF88FFD5D3D2AD738B07FBEC" box="[660,780,1071,1097]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[179,1407,179,1917]" captionTargetId="figure-19@2.[179,1407,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Phthiria portokali sp. nov,. Holotype female; A) lateral view; B) antennae; C) head, lateral view; D) head, dorsal view; E) pleura; F) abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254483" httpUri="https://zenodo.org/record/10254483/files/figure.png" pageId="1" pageNumber="492">Figs. 1a, d</figureCitation>
); parafacials yellow except two black spots, one at level of scape extended horizontally to eye margin and second at level of lower margin of eyes (
<figureCitation id="137609ECFF88FFD5D53EAD0F8CC2FBC8" box="[1144,1225,1107,1133]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[179,1407,179,1917]" captionTargetId="figure-19@2.[179,1407,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Phthiria portokali sp. nov,. Holotype female; A) lateral view; B) antennae; C) head, lateral view; D) head, dorsal view; E) pleura; F) abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254483" httpUri="https://zenodo.org/record/10254483/files/figure.png" pageId="1" pageNumber="492">Fig. 1c</figureCitation>
); oral margin with short scattered white hairs; ocellar tubercle raised, black, ocelli as equilateral triangle; scape yellow (
<figureCitation id="137609ECFF88FFD5D5A2AD2B8D3FFB34" box="[1252,1332,1143,1169]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[179,1407,179,1917]" captionTargetId="figure-19@2.[179,1407,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Phthiria portokali sp. nov,. Holotype female; A) lateral view; B) antennae; C) head, lateral view; D) head, dorsal view; E) pleura; F) abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254483" httpUri="https://zenodo.org/record/10254483/files/figure.png" pageId="1" pageNumber="492">Fig. 1b</figureCitation>
), slightly longer than wide; pedicel brown, cylindrical, wider than long; first antennal flagellomere, dark brown, cylindrical, with long dorsal process, longer than width of first flagellomere apically; style transparent, about subequal to antennal process; mentum yellow; palpi yellow, with few short white hairs; proboscis long, 1.8 times head length, black.
<emphasis id="B939C97BFF88FFD5D1A5AC5B8937FA84" bold="true" box="[227,316,1287,1313]" pageId="1" pageNumber="492">Thorax</emphasis>
(
<figureCitation id="137609ECFF88FFD5D00AAC5B8994FA84" box="[332,415,1287,1313]" captionStart="FIGURE 2" captionStartId="3.[152,255,1944,1968]" captionTargetBox="[176,1412,181,1920]" captionTargetId="figure-18@3.[176,1412,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Phthiria portokali sp. nov, (AC &amp; F). compared with Phthiria socotrae Greathead &amp; Evenhuis (D &amp; E). A) mesonotum in dorsal view, holotype; B) mesonotum in dorsal view, paratype female; C) mesonotum in dorsal view, paratype male; D) mesonotum in dorsal view, paratype female; E) female paratype lateral view." figureDoi="http://doi.org/10.5281/zenodo.10254485" httpUri="https://zenodo.org/record/10254485/files/figure.png" pageId="1" pageNumber="492">Fig. 2a</figureCitation>
). Mesonotum yellow, with short whitish hairs, and with large tortoise-shell-shaped orange pattern on the dorsum; with black coloring as follows: two black spots next to transverse suture, two round blackish spots next to scutellum, two black marks on anterior margin and next to scutellum connecting with trace of black line, lateral margins yellow; postpronotal lobes yellow, with long white hairs; scutellum yellow, black ventrally; anepisternum (
<figureCitation id="137609ECFF88FFD5D079ACCB899BFA14" box="[319,400,1431,1457]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[179,1407,179,1917]" captionTargetId="figure-19@2.[179,1407,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Phthiria portokali sp. nov,. Holotype female; A) lateral view; B) antennae; C) head, lateral view; D) head, dorsal view; E) pleura; F) abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254483" httpUri="https://zenodo.org/record/10254483/files/figure.png" pageId="1" pageNumber="492">Fig. 1e</figureCitation>
) with long white hairs, lower half blackish, upper half yellow; anepimeron yellow, except antero-ventrally with blackish mark; katepisternum upper half yellow, lower half black, upper half yellow; meron posteriorly black, anteriorly yellow; halter stem and knob yellow.
<emphasis id="B939C97BFF88FFD5D23EAC838BAEFA5C" bold="true" box="[888,933,1503,1529]" pageId="1" pageNumber="492">Leg</emphasis>
. Coxa and femur yellowish, densely covered with white hairs; tibia yellowish; tarsal segments with short bristles laterally, basitarsus yellowish, as long as other segments combined, other segments black.
<emphasis id="B939C97BFF88FFD5D338AF7B8AF8F9E4" bold="true" box="[638,755,1575,1601]" pageId="1" pageNumber="492">Abdomen</emphasis>
(
<figureCitation id="137609ECFF88FFD5D244AF7B8B44F9E4" box="[770,847,1575,1601]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[179,1407,179,1917]" captionTargetId="figure-19@2.[179,1407,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Phthiria portokali sp. nov,. Holotype female; A) lateral view; B) antennae; C) head, lateral view; D) head, dorsal view; E) pleura; F) abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254483" httpUri="https://zenodo.org/record/10254483/files/figure.png" pageId="1" pageNumber="492">Fig. 1f</figureCitation>
). Tergites yellow, with long white hairs, with row of black spots medially; sternites yellow.
<emphasis id="B939C97BFF88FFD5D304AF178A88F9C0" bold="true" box="[578,643,1611,1637]" pageId="1" pageNumber="492">Wing</emphasis>
. Hyaline, uniformly microtrichose; veins blackish brown; costa reaches vein CuA1+A1; R originates before level of mm crossvein; crossvein rm beyond middle of discal cell; anal cell closed before wing margin, stalked; anal lobe broad. (
<figureCitation id="137609ECFF88FFD5D3B8AFCF8B4CF908" box="[766,839,1683,1709]" captionStart="FIGURE 2" captionStartId="3.[152,255,1944,1968]" captionTargetBox="[176,1412,181,1920]" captionTargetId="figure-18@3.[176,1412,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Phthiria portokali sp. nov, (AC &amp; F). compared with Phthiria socotrae Greathead &amp; Evenhuis (D &amp; E). A) mesonotum in dorsal view, holotype; B) mesonotum in dorsal view, paratype female; C) mesonotum in dorsal view, paratype male; D) mesonotum in dorsal view, paratype female; E) female paratype lateral view." figureDoi="http://doi.org/10.5281/zenodo.10254485" httpUri="https://zenodo.org/record/10254485/files/figure.png" pageId="1" pageNumber="492">Fig.2f</figureCitation>
).
<emphasis id="B939C97BFF88FFD5D21DAFCF8BD5F908" bold="true" box="[859,990,1683,1709]" pageId="1" pageNumber="492">Terminalia</emphasis>
(
<figureCitation id="137609ECFF88FFD5D2A9AFCF8C68F908" box="[1007,1123,1683,1709]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[179,1407,182,1918]" captionTargetId="figure-19@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Genitalia of Phthiria portokali sp. nov, A) female genitalia; B) spermathecal; C) gonocoxae in dorsal view; D) gonostylus; E) phallic complex in dorsal view; F) epandrium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254487" httpUri="https://zenodo.org/record/10254487/files/figure.png" pageId="1" pageNumber="492">Figs. 3a,b</figureCitation>
). Genital fork consisting of two parallel bars; common spermathecal duct short, membranous; basal ducts membranous; sperm pump as wide as apical ducts, striated, sclerotized, longer than apical ducts; apical ducts slightly sclerotized; spermathecal reservoir membranous, round, apical process well sclerotized, curved, long, more than 3.5 times capsule length.
</paragraph>
<paragraph id="8BF21569FF88FFD5D025AFDD8976F92A" blockId="1.[151,1437,675,1818]" box="[355,381,1665,1679]" pageId="1" pageNumber="492">
<subScript id="17C9172CFF88FFD5D025AFDD8976F92A" attach="left" box="[355,381,1665,1679]" fontSize="6" pageId="1" pageNumber="492">4+5</subScript>
</paragraph>
<caption id="DF3245E1FF8BFFD6D1DEAECB8A49F876" ID-DOI="http://doi.org/10.5281/zenodo.10254483" ID-Zenodo-Dep="10254483" httpUri="https://zenodo.org/record/10254483/files/figure.png" pageId="2" pageNumber="493" startId="2.[152,255,1943,1967]" targetBox="[179,1407,179,1917]" targetPageId="2" targetType="figure">
<paragraph id="8BF21569FF8BFFD6D1DEAECB8A49F876" blockId="2.[151,1436,1943,2004]" pageId="2" pageNumber="493">
<emphasis id="B939C97BFF8BFFD6D1DEAECB8912F80A" bold="true" box="[152,281,1943,1967]" pageId="2" pageNumber="493">FIGURE 1.</emphasis>
<taxonomicName id="4C4D6EEAFF8BFFD6D059AECB89DCF80A" authority="Farhangi &amp; Nozari &amp; Gharali, 2023" authorityName="Farhangi &amp; Nozari &amp; Gharali" authorityYear="2023" box="[287,471,1943,1967]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="493" phylum="Arthropoda" rank="species" species="portokali" status="sp. nov.">
<emphasis id="B939C97BFF8BFFD6D059AECB89DCF80A" box="[287,471,1943,1967]" italics="true" pageId="2" pageNumber="493">Phthiria portokali</emphasis>
</taxonomicName>
<emphasis id="B939C97BFF8BFFD6D098AEC48A24F815" bold="true" box="[478,559,1944,1968]" pageId="2" pageNumber="493">
<taxonomicNameLabel id="A20A7400FF8BFFD6D098AEC48A21F815" box="[478,554,1944,1968]" pageId="2" pageNumber="493" rank="species">sp. nov</taxonomicNameLabel>
,
</emphasis>
. Holotype female; A) lateral view; B) antennae; C) head, lateral view; D) head, dorsal view; E) pleura; F) abdomen, dorsal view.
</paragraph>
</caption>
<caption id="DF3245E1FF8AFFD7D1DEAEC48C0AF85D" ID-DOI="http://doi.org/10.5281/zenodo.10254485" ID-Zenodo-Dep="10254485" httpUri="https://zenodo.org/record/10254485/files/figure.png" pageId="3" pageNumber="494" startId="3.[152,255,1944,1968]" targetBox="[176,1412,181,1920]" targetPageId="3" targetType="figure">
<paragraph id="8BF21569FF8AFFD7D1DEAEC48C0AF85D" blockId="3.[151,1436,1944,2041]" pageId="3" pageNumber="494">
<emphasis id="B939C97BFF8AFFD7D1DEAEC48910F815" bold="true" box="[152,283,1944,1968]" pageId="3" pageNumber="494">FIGURE 2.</emphasis>
<taxonomicName id="4C4D6EEAFF8AFFD7D062AEC489D5F815" authority="Farhangi &amp; Nozari &amp; Gharali, 2023" authorityName="Farhangi &amp; Nozari &amp; Gharali" authorityYear="2023" box="[292,478,1944,1968]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="species" species="portokali" status="sp. nov.">
<emphasis id="B939C97BFF8AFFD7D062AEC489D5F815" box="[292,478,1944,1968]" italics="true" pageId="3" pageNumber="494">Phthiria portokali</emphasis>
</taxonomicName>
<emphasis id="B939C97BFF8AFFD7D0A1AEC48A30F815" bold="true" box="[487,571,1944,1968]" pageId="3" pageNumber="494">
<taxonomicNameLabel id="A20A7400FF8AFFD7D0A1AEC48A3DF815" box="[487,566,1944,1968]" pageId="3" pageNumber="494" rank="species">sp. nov</taxonomicNameLabel>
,
</emphasis>
(AC &amp; F). compared with
<taxonomicName id="4C4D6EEAFF8AFFD7D22FAEC48D19F815" authority="Greathead &amp; Evenhuis" authorityName="Greathead &amp; Evenhuis" authorityYear="2001" box="[873,1298,1944,1968]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="species" species="socotrae">
<emphasis id="B939C97BFF8AFFD7D22FAEC48C15F815" box="[873,1054,1944,1968]" italics="true" pageId="3" pageNumber="494">Phthiria socotrae</emphasis>
Greathead &amp; Evenhuis
</taxonomicName>
(D &amp; E). A) mesonotum in dorsal view, holotype; B) mesonotum in dorsal view, paratype female; C) mesonotum in dorsal view, paratype male; D) mesonotum in dorsal view, paratype female; E) female paratype lateral view.
</paragraph>
</caption>
<caption id="DF3245E1FF8DFFD0D1DEAECB8B90F876" ID-DOI="http://doi.org/10.5281/zenodo.10254487" ID-Zenodo-Dep="10254487" httpUri="https://zenodo.org/record/10254487/files/figure.png" pageId="4" pageNumber="495" startId="4.[152,255,1943,1967]" targetBox="[179,1407,182,1918]" targetPageId="4" targetType="figure">
<paragraph id="8BF21569FF8DFFD0D1DEAECB8B90F876" blockId="4.[151,1436,1943,2004]" pageId="4" pageNumber="495">
<emphasis id="B939C97BFF8DFFD0D1DEAECB8911F80A" bold="true" box="[152,282,1943,1967]" pageId="4" pageNumber="495">FIGURE 3.</emphasis>
Genitalia of
<taxonomicName id="4C4D6EEAFF8DFFD0D0E5AECB8A56F80A" authority="Farhangi &amp; Nozari &amp; Gharali, 2023" authorityName="Farhangi &amp; Nozari &amp; Gharali" authorityYear="2023" box="[419,605,1943,1967]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="495" phylum="Arthropoda" rank="species" species="portokali" status="sp. nov.">
<emphasis id="B939C97BFF8DFFD0D0E5AECB8A56F80A" box="[419,605,1943,1967]" italics="true" pageId="4" pageNumber="495">Phthiria portokali</emphasis>
</taxonomicName>
<emphasis id="B939C97BFF8DFFD0D322AEC48ABCF815" bold="true" box="[612,695,1944,1968]" pageId="4" pageNumber="495">
<taxonomicNameLabel id="A20A7400FF8DFFD0D322AEC48AB9F815" box="[612,690,1944,1968]" pageId="4" pageNumber="495" rank="species">sp. nov</taxonomicNameLabel>
,
</emphasis>
A) female genitalia; B) spermathecal; C) gonocoxae in dorsal view; D) gonostylus; E) phallic complex in dorsal view; F) epandrium in dorsal view.
</paragraph>
</caption>
<paragraph id="8BF21569FF8CFFD1D181A9CA88C9FE51" blockId="5.[151,1437,150,897]" pageId="5" pageNumber="496">
<emphasis id="B939C97BFF8CFFD1D181A9CA8AC7FF15" bold="true" box="[199,716,150,176]" pageId="5" pageNumber="496">Male. Similar to female except for: Head. C</emphasis>
ompletely black; frons black, gray dusted, with long white hairs; parafacials completely black, with long white hairs; mentum gray dusted, with sparse long white hairs; ocellar tubercle prominent black, with few white hairs; eyes holoptic for entire length of frons, contact portion of eyes medially depressed, facets of upper half of eyes enlarged.Antennae similar to female; palpi blackish brown, proboscis black. Thorax (
<figureCitation id="137609ECFF8CFFD1D007A87A8998FEE5" box="[321,403,294,320]" captionStart="FIGURE 2" captionStartId="3.[152,255,1944,1968]" captionTargetBox="[176,1412,181,1920]" captionTargetId="figure-18@3.[176,1412,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Phthiria portokali sp. nov, (AC &amp; F). compared with Phthiria socotrae Greathead &amp; Evenhuis (D &amp; E). A) mesonotum in dorsal view, holotype; B) mesonotum in dorsal view, paratype female; C) mesonotum in dorsal view, paratype male; D) mesonotum in dorsal view, paratype female; E) female paratype lateral view." figureDoi="http://doi.org/10.5281/zenodo.10254485" httpUri="https://zenodo.org/record/10254485/files/figure.png" pageId="5" pageNumber="496">Fig. 2c</figureCitation>
). Mesonotum black except lateral margins and two round large spots at prescutellum yellow, scutellum yellow; pleura completely black, halter stem and knob white.
<emphasis id="B939C97BFF8CFFD1D2F0A8168C32FEC1" bold="true" box="[950,1081,330,356]" pageId="5" pageNumber="496">Terminalia</emphasis>
. Epandrium triangular in dorsal view, with small invagination apically (
<figureCitation id="137609ECFF8CFFD1D308A8328A9DFE2D" box="[590,662,366,392]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[179,1407,182,1918]" captionTargetId="figure-19@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Genitalia of Phthiria portokali sp. nov, A) female genitalia; B) spermathecal; C) gonocoxae in dorsal view; D) gonostylus; E) phallic complex in dorsal view; F) epandrium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254487" httpUri="https://zenodo.org/record/10254487/files/figure.png" pageId="5" pageNumber="496">Fig.3f</figureCitation>
); gonocoxites ovoid, fused medially, apicomesal process smaller than gonostylus, with small bristles (
<figureCitation id="137609ECFF8CFFD1D346A8CE8A59FE09" box="[512,594,402,428]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[179,1407,182,1918]" captionTargetId="figure-19@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Genitalia of Phthiria portokali sp. nov, A) female genitalia; B) spermathecal; C) gonocoxae in dorsal view; D) gonostylus; E) phallic complex in dorsal view; F) epandrium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254487" httpUri="https://zenodo.org/record/10254487/files/figure.png" pageId="5" pageNumber="496">Fig. 3c</figureCitation>
); gonostylus oval, apically curved, pointed, hollowed basally (
<figureCitation id="137609ECFF8CFFD1D45DA8CE8D85FE09" box="[1307,1422,402,428]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[179,1407,182,1918]" captionTargetId="figure-19@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Genitalia of Phthiria portokali sp. nov, A) female genitalia; B) spermathecal; C) gonocoxae in dorsal view; D) gonostylus; E) phallic complex in dorsal view; F) epandrium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254487" httpUri="https://zenodo.org/record/10254487/files/figure.png" pageId="5" pageNumber="496">Figs.3c, d</figureCitation>
); basal aedeagal process narrow in dorsal view, lateral aedeagal apodemes narrow; aedeagus apically curved (
<figureCitation id="137609ECFF8CFFD1D429A8EA88BEFE51" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[179,1407,182,1918]" captionTargetId="figure-19@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Genitalia of Phthiria portokali sp. nov, A) female genitalia; B) spermathecal; C) gonocoxae in dorsal view; D) gonostylus; E) phallic complex in dorsal view; F) epandrium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10254487" httpUri="https://zenodo.org/record/10254487/files/figure.png" pageId="5" pageNumber="496">Fig. 3e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35746E2FF8CFFD1D181A8A28BF2FD99" pageId="5" pageNumber="496" type="distribution">
<paragraph id="8BF21569FF8CFFD1D181A8A28BF2FD99" blockId="5.[151,1437,150,897]" pageId="5" pageNumber="496">
<emphasis id="B939C97BFF8CFFD1D181A8A2896AFDBD" bold="true" box="[199,353,510,536]" pageId="5" pageNumber="496">Distribution.</emphasis>
This species is currently known from mountainous rangeland (the location called Soltan bag= king garden) at the border of two provinces,
<collectingRegion id="4989DB8BFF8CFFD1D3CDAB7E8AD7FD99" box="[651,732,546,572]" country="Iran" name="Qazvin" pageId="5" pageNumber="496">Qazvin</collectingRegion>
and
<collectingRegion id="4989DB8BFF8CFFD1D254AB7E8B76FD99" box="[786,893,546,572]" country="Iran" name="Hamadan" pageId="5" pageNumber="496">Hamedan</collectingRegion>
provinces.
</paragraph>
</subSubSection>
<subSubSection id="C35746E2FF8CFFD1D181AB1A8A2CFC25" pageId="5" pageNumber="496" type="etymology">
<paragraph id="8BF21569FF8CFFD1D181AB1A8C74FD21" blockId="5.[151,1437,150,897]" pageId="5" pageNumber="496">
<emphasis id="B939C97BFF8CFFD1D181AB1A8945FDC5" bold="true" box="[199,334,582,608]" pageId="5" pageNumber="496">Etymology.</emphasis>
The name of new species,
<emphasis id="B939C97BFF8CFFD1D331AB1A8BA8FDC5" bold="true" box="[631,931,582,609]" pageId="5" pageNumber="496">
<taxonomicName id="4C4D6EEAFF8CFFD1D331AB1A8B4EFDC4" authority="Farhangi &amp; Nozari &amp; Gharali, 2023" authorityName="Farhangi &amp; Nozari &amp; Gharali" authorityYear="2023" box="[631,837,582,609]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="portokali" status="sp. nov.">
<emphasis id="B939C97BFF8CFFD1D331AB1A8B4EFDC4" bold="true" box="[631,837,582,609]" italics="true" pageId="5" pageNumber="496">Phthiria portokali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A7400FF8CFFD1D20DAB1A8BA8FDC5" box="[843,931,582,608]" pageId="5" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is derived from the Greek noun “πορτοκαλί” meaning orange and referring to the large orange mark on the mesonotum of the females.
</paragraph>
<paragraph id="8BF21569FF8CFFD1D181ABD28A2CFC25" blockId="5.[151,1437,150,897]" pageId="5" pageNumber="496">
<emphasis id="B939C97BFF8CFFD1D181ABD28AC2FD0D" bold="true" box="[199,713,654,681]" pageId="5" pageNumber="496">
Taxonomic note.
<taxonomicName id="4C4D6EEAFF8CFFD1D0D3ABD28A6DFD0C" authority="Farhangi &amp; Nozari &amp; Gharali, 2023" authorityName="Farhangi &amp; Nozari &amp; Gharali" authorityYear="2023" box="[405,614,654,681]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="portokali" status="sp. nov.">
<emphasis id="B939C97BFF8CFFD1D0D3ABD28A6DFD0C" bold="true" box="[405,614,654,681]" italics="true" pageId="5" pageNumber="496">Phthiria portokali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A7400FF8CFFD1D329ABD28AC2FD0D" box="[623,713,654,680]" pageId="5" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only species, in
<collectingCountry id="F35A55F9FF8CFFD1D289ABD38BF7FD0C" box="[975,1020,655,681]" name="Iran" pageId="5" pageNumber="496">Iran</collectingCountry>
and the Palaearctic region, in which females have a large orange mark on the dorsum of mesonotum. Interestingly on the dorsum of mesonotum, in the female
<typeStatus id="54F6ABCBFF8CFFD1D05DAB8A898BFD55" box="[283,384,726,752]" pageId="5" pageNumber="496" type="holotype">holotype</typeStatus>
, there are fine black striations seen similarly in one of the
<typeStatus id="54F6ABCBFF8CFFD1D57AAB8B8CACFD54" box="[1084,1191,727,753]" pageId="5" pageNumber="496" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C4D6EEAFF8CFFD1D592AB8A89D6FCB1" authority="Greathead &amp; Evenhuis, 2001" authorityName="Greathead &amp; Evenhuis" authorityYear="2001" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="socotrae">
<emphasis id="B939C97BFF8CFFD1D592AB8A8D97FD55" box="[1236,1436,726,752]" italics="true" pageId="5" pageNumber="496">Phthiria socotrae</emphasis>
<bibRefCitation id="EFDC6898FF8CFFD1D1D1ABA689D6FCB1" author="Greathead, D. J. &amp; Evenhuis, N." box="[151,477,762,788]" pageId="5" pageNumber="496" pagination="1 - 11" refId="ref2371" refString="Greathead, D. J. &amp; Evenhuis, N. L (2001) Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae) from the island of Sokotra. Zootaxa, 14 (1), 1 - 11. https: // doi. org / 10.11646 / zotaxa. 14.1.1" type="journal article" year="2001">Greathead &amp; Evenhuis, 2001</bibRefCitation>
</taxonomicName>
described from
<collectingCountry id="F35A55F9FF8CFFD1D3D2ABA68AE1FCB1" box="[660,746,762,788]" name="Yemen" pageId="5" pageNumber="496">Socotra</collectingCountry>
island (Afrotropical Region) (
<figureCitation id="137609ECFF8CFFD1D579ABA68C98FCB1" box="[1087,1171,762,788]" captionStart="FIGURE 2" captionStartId="3.[152,255,1944,1968]" captionTargetBox="[176,1412,181,1920]" captionTargetId="figure-18@3.[176,1412,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Phthiria portokali sp. nov, (AC &amp; F). compared with Phthiria socotrae Greathead &amp; Evenhuis (D &amp; E). A) mesonotum in dorsal view, holotype; B) mesonotum in dorsal view, paratype female; C) mesonotum in dorsal view, paratype male; D) mesonotum in dorsal view, paratype female; E) female paratype lateral view." figureDoi="http://doi.org/10.5281/zenodo.10254485" httpUri="https://zenodo.org/record/10254485/files/figure.png" pageId="5" pageNumber="496">Fig. 2d</figureCitation>
). As this striation is not seen in the female
<typeStatus id="54F6ABCBFF8CFFD1D02EAA4389C3FC9C" box="[360,456,799,825]" pageId="5" pageNumber="496" type="paratype">paratype</typeStatus>
of the Iranian species and in some
<typeStatus id="54F6ABCBFF8CFFD1D215AA438BB5FC9C" box="[851,958,799,825]" pageId="5" pageNumber="496" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C4D6EEAFF8CFFD1D2A5AA428C7FFC9C" authorityName="Greathead &amp; Evenhuis" authorityYear="2001" box="[995,1140,798,825]" class="Insecta" family="Bombyliidae" genus="Phthiria" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="socotrae">
<emphasis id="B939C97BFF8CFFD1D2A5AA428C7FFC9C" box="[995,1140,798,825]" italics="true" pageId="5" pageNumber="496">Ph. Socotrae</emphasis>
</taxonomicName>
, we are considering it as a morphological variation. In the female
<typeStatus id="54F6ABCBFF8CFFD1D30BAA1F8AA6FCF8" box="[589,685,835,861]" pageId="5" pageNumber="496" type="paratype">paratype</typeStatus>
of the Iranian species, the blackness of the mesonotum is extended more than in the
<typeStatus id="54F6ABCBFF8CFFD1D013AA3A89BCFC25" box="[341,439,870,896]" pageId="5" pageNumber="496" type="holotype">holotype</typeStatus>
(
<figureCitation id="137609ECFF8CFFD1D080AA3A8A11FC25" box="[454,538,870,896]" captionStart="FIGURE 2" captionStartId="3.[152,255,1944,1968]" captionTargetBox="[176,1412,181,1920]" captionTargetId="figure-18@3.[176,1412,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Phthiria portokali sp. nov, (AC &amp; F). compared with Phthiria socotrae Greathead &amp; Evenhuis (D &amp; E). A) mesonotum in dorsal view, holotype; B) mesonotum in dorsal view, paratype female; C) mesonotum in dorsal view, paratype male; D) mesonotum in dorsal view, paratype female; E) female paratype lateral view." figureDoi="http://doi.org/10.5281/zenodo.10254485" httpUri="https://zenodo.org/record/10254485/files/figure.png" pageId="5" pageNumber="496">Fig. 2b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>