197 lines
21 KiB
XML
197 lines
21 KiB
XML
<document id="3C2A7AF8F1B7CF21FC5C802C04840491" ID-CLB-Dataset="27700" ID-DOI="10.11646/zootaxa.4543.3.2" ID-GBIF-Dataset="7520908e-3595-4ed3-96f8-9a0fffb45439" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617841" ID-ZooBank="58142B27-5E25-46EE-A1B0-71BB31DFB3E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928404141" checkinUser="plazi" docAuthor="Davis, Steven R., Brav-Cubitt, Talia, Buckley, Thomas R. & Leschen, Richard A. B." docDate="2019" docId="03DEC54AFFE5C91DFBA847D8FCD3F975" docLanguage="en" docName="zootaxa.4543.3.2.pdf" docOrigin="Zootaxa 4543 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Etheophanus optandus Broun 1921" docType="treatment" docVersion="8" lastPageNumber="355" masterDocId="FFE7BD32FFEBC913FB3F442FFFFDFFAF" masterDocTitle="Systematics of the New Zealand Weevil Etheophanus Broun (Curculionidae: Molytinae)" masterLastPageNumber="374" masterPageNumber="341" pageNumber="355" updateTime="1698715414758" updateUser="ExternalLinkService">
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<mods:titleInfo id="C2E4E7999FFC09298115A7C083A6A899">
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<mods:title id="C213DE03162C42A83588708E544C587D">Systematics of the New Zealand Weevil Etheophanus Broun (Curculionidae: Molytinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C0B7BDFFFE93678C1B369F562F99E36D" type="personal">
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<mods:namePart id="84BAD4EED0EB761AFB0EF1CEE7498560">Davis, Steven R.</mods:namePart>
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<mods:roleTerm id="A80C40C5436169D073A3B5A1D0D6A749">Author</mods:roleTerm>
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<mods:namePart id="AB4F18AB3DC3BFD6F0A84BE862F92450">Brav-Cubitt, Talia</mods:namePart>
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<mods:roleTerm id="956E325E5CAD1FCEA8A3766265C1214E">Author</mods:roleTerm>
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<mods:namePart id="D6A500FFA45C2A5431C921028C5FBC8D">Buckley, Thomas R.</mods:namePart>
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<mods:name id="7A4277DE6662F49B531C5827E2BEE383" type="personal">
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<mods:roleTerm id="DD804810D95373BE302E0E92BAF62670">Author</mods:roleTerm>
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<mods:namePart id="CB31493FBABAC734296EEBC8A13273FE">Leschen, Richard A. B.</mods:namePart>
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<mods:typeOfResource id="8219F496845B662E955161993AFAF74B">text</mods:typeOfResource>
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<mods:title id="87F50DF52F330AA7EDA6F53147766FBF">Zootaxa</mods:title>
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<mods:part id="2578E3A09F64EA21179CBC7CA36C8C22">
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<mods:date id="06D42261D64287E08D2BE38A8FFD7700">2019</mods:date>
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<mods:number id="1A67437EB76FC7EA281D53405218E5B3">2019-01-08</mods:number>
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<mods:number id="DD83C783DEF9B97BC72C63018C7CD0C0">4543</mods:number>
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<treatment id="03DEC54AFFE5C91DFBA847D8FCD3F975" ID-DOI="http://doi.org/10.5281/zenodo.5944061" ID-GBIF-Taxon="154529842" ID-Zenodo-Dep="5944061" LSID="urn:lsid:plazi:treatment:03DEC54AFFE5C91DFBA847D8FCD3F975" httpUri="http://treatment.plazi.org/id/03DEC54AFFE5C91DFBA847D8FCD3F975" lastPageNumber="355" pageId="14" pageNumber="355">
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<subSubSection id="C36D27D7FFE5C91DFBA847D8FDB3FBBD" box="[151,590,1015,1042]" pageId="14" pageNumber="355" type="nomenclature">
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<paragraph id="8BC8745CFFE5C91DFBA847D8FDB3FBBD" blockId="14.[151,590,1015,1075]" box="[151,590,1015,1042]" pageId="14" pageNumber="355">
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<heading id="D080C330FFE5C91DFBA847D8FDB3FBBD" bold="true" box="[151,590,1015,1042]" fontSize="11" level="1" pageId="14" pageNumber="355" reason="1">
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<taxonomicName id="4C770FDFFFE5C91DFBA847D8FDB3FBBD" ID-CoL="3BKZS" authority="Broun, 1923" authorityName="Broun" authorityYear="1921" box="[151,590,1015,1042]" class="Insecta" family="Curculionidae" genus="Etheophanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="optandus">
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<emphasis id="B903A84EFFE5C91DFBA847D8FDB3FBBD" bold="true" box="[151,590,1015,1042]" pageId="14" pageNumber="355">
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<emphasis id="B903A84EFFE5C91DFBA847D8FE50FBBE" bold="true" box="[151,429,1015,1041]" italics="true" pageId="14" pageNumber="355">Etheophanus optandus</emphasis>
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<bibRefCitation id="EFE609ADFFE5C91DFA8B47D7FDB3FBBD" author="Broun, T." box="[436,590,1016,1042]" pageId="14" pageNumber="355" pagination="667 - 708" refId="ref10401" refString="Broun, T. (1923) Descriptions of new genera and species of Coleoptera. Part VIII. Bulletin of the New Zealand Institute, 1, 667 - 708." type="journal article" year="1923">Broun, 1923</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C36D27D7FFE5C91DFBA84035FE4FFB9C" box="[151,434,1050,1075]" pageId="14" pageNumber="355" type="description">
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<paragraph id="8BC8745CFFE5C91DFBA84035FE4FFB9C" blockId="14.[151,590,1015,1075]" box="[151,434,1050,1075]" pageId="14" pageNumber="355">
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(
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<figureCitation id="134C68D9FFE5C91DFBA04035FF04FB9C" box="[159,249,1050,1075]" captionStart="FIGURES 1" captionStartId="3.[151,264,1973,1995]" captionTargetBox="[278,1316,213,1942]" captionTargetId="figure@3.[260,1328,195,1956]" captionTargetPageId="3" captionText="FIGURES 1A–G. Dorsal habiti of Etheophanus spp. A, E. kuscheli sp. n.; B, E. nitidellus; C, E. optandus; D, E. pingius; E, E. striatus." httpUri="https://zenodo.org/record/2617843/files/figure.png" pageId="14" pageNumber="355">Figs 1C</figureCitation>
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,
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<figureCitation id="134C68D9FFE5C91DFA394034FED4FB9C" box="[262,297,1051,1075]" captionStart="FIGURES 2" captionStartId="4.[151,264,1962,1984]" captionTargetBox="[189,1414,253,1889]" captionTargetId="figure@4.[151,1435,230,1908]" captionTargetPageId="4" captionText="FIGURES 2A–G. Lateral habiti of Etheophanus spp. A, E. kuscheli sp. n.; B, E. nitidellus; C, E. optandus; D, E. pingius; E, E. striatus." httpUri="https://zenodo.org/record/2617845/files/figure.png" pageId="14" pageNumber="355">2C</figureCitation>
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,
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<figureCitation id="134C68D9FFE5C91DFA0A4035FEA5FB9C" box="[309,344,1050,1075]" captionStart="FIGURES 5" captionStartId="7.[151,264,1881,1903]" captionTargetBox="[174,1417,713,1866]" captionTargetId="figure@7.[151,1436,690,1870]" captionTargetPageId="7" captionText="FIGURES 5A–G. Key characters of Etheophanus spp. A, E. kuscheli sp. n., apical half of median lobe (dorsal view); B, E. nitidellus, apical half of median lobe (dorsal view); C, E. optandus, apical half of median lobe (dorsal view); D, E. pingius, apical half of median lobe (dorsal view); E, E. striatus, apical half of median lobe (dorsal view); F, E. nitidellus (female), internal view of abdominal ventrites showing pair of apodemes on fifth ventrite; G, E. kuscheli, showing semi-lamellate process on procoxa (arrow)." httpUri="https://zenodo.org/record/2617851/files/figure.png" pageId="14" pageNumber="355">5C</figureCitation>
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,
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<figureCitation id="134C68D9FFE5C91DFA5C4034FE7CFB9C" box="[355,385,1051,1075]" captionStart="FIGURES 10" captionStartId="15.[151,264,1956,1978]" captionTargetBox="[150,1436,260,1864]" captionTargetId="figure@15.[150,1436,260,1864]" captionTargetPageId="15" captionText="FIGURES 10A–D. Etheophanus optandus, male terminalia. A, aedeagus, dorsal view; B, aedeagus, lateral view; C, tegmen, dorsal view; D, sternites VIII and IX, showing spiculum relictum." httpUri="https://zenodo.org/record/2617861/files/figure.png" pageId="14" pageNumber="355">10</figureCitation>
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,
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<figureCitation id="134C68D9FFE5C91DFAB24034FE55FB9C" box="[397,424,1051,1075]" captionStart="FIGURES 11" captionStartId="16.[151,264,1996,2018]" captionTargetBox="[151,1434,210,1966]" captionTargetId="figure@16.[151,1435,210,1966]" captionTargetPageId="16" captionText="FIGURES 11A–D. Etheophanus optandus, female terminalia. A, sternite VIII; B, tergite VIII; C, gonocoxites; D, spermatheca." httpUri="https://zenodo.org/record/2617863/files/figure.png" pageId="14" pageNumber="355">11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D27D7FFE5C91DFBA8404DFC0BFBD7" box="[151,1014,1122,1144]" pageId="14" pageNumber="355" type="reference_group">
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<paragraph id="8BC8745CFFE5C91DFBA8404DFC0BFBD7" blockId="14.[151,1014,1122,1144]" box="[151,1014,1122,1144]" pageId="14" pageNumber="355">
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<treatmentCitationGroup id="AB675372FFE5C91DFBA8404DFC0BFBD7" box="[151,1014,1122,1144]" pageId="14" pageNumber="355">
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<taxonomicName id="4C770FDFFFE5C91DFBA8404DFDC4FBD7" ID-CoL="3BKZS" authority="Broun, 1923: 702" authorityName="Broun" authorityPageNumber="702" authorityYear="1921" box="[151,569,1122,1144]" class="Insecta" family="Curculionidae" genus="Etheophanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="optandus">
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<emphasis id="B903A84EFFE5C91DFBA8404DFE83FBD7" box="[151,382,1122,1144]" italics="true" pageId="14" pageNumber="355">Etheophanus optandus</emphasis>
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<treatmentCitation id="0AD6524DFFE5C91DFABB404CFDC4FBD7" author="Broun, T." box="[388,569,1122,1144]" page="702" pageId="14" pageNumber="355" year="1923">
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<bibRefCitation id="EFE609ADFFE5C91DFABB404CFDFBFBD7" author="Broun, T." box="[388,518,1122,1144]" pageId="14" pageNumber="355" pagination="667 - 708" refId="ref10401" refString="Broun, T. (1923) Descriptions of new genera and species of Coleoptera. Part VIII. Bulletin of the New Zealand Institute, 1, 667 - 708." type="journal article" year="1923">Broun, 1923</bibRefCitation>
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: 702
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</treatmentCitation>
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||
</taxonomicName>
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||
.
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<typeStatus id="54CCCAFEFFE5C91DF97C404CFD88FBD7" box="[579,629,1123,1144]" pageId="14" pageNumber="355">Type</typeStatus>
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locality: Hollyford, southwest
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<collectingRegion id="49B3BABEFFE5C91DF88E404DFC0FFBD7" box="[945,1010,1122,1144]" country="New Zealand" name="Otago" pageId="14" pageNumber="355">Otago</collectingRegion>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36D27D7FFE5C91DFBA8408AFA60FA39" pageId="14" pageNumber="355" type="diagnosis">
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<paragraph id="8BC8745CFFE5C91DFBA8408AFA60FA39" blockId="14.[151,1437,1189,1754]" pageId="14" pageNumber="355">
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<emphasis id="B903A84EFFE5C91DFBA8408AFEECFB11" bold="true" box="[151,273,1189,1214]" pageId="14" pageNumber="355">Diagnosis.</emphasis>
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Dorsal surfaces apunctate, alveolate microsculpture present. Pronotal vibrissae present. Procoxa without semi-lamellate process, primary dorsal setae adpressed. Eighth sternite of male with each sclerite bearing ca. 5 setae along apical margins; spiculum relictum slender. Aedeagus with setae present at apical ¼ of parameroid lobes; median lobe in dorsal view not narrowing to apex and rounded apically with simple edges, in lateral view narrow and elongate and not strongly curved ventrally near apex; apico-lateral margin of median lobe bearing a bundle of ca. 7 to 8 setae at each side. Female tergite VIII apically evenly arcuate; eighth sternite with arms of base parallel-sided, apices with 3 setae. Gonostylus with 4–5 apical setae. Spermatheca falciform without globular base.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D27D7FFE5C91DFBF8418EFAF1FA15" box="[199,1292,1441,1466]" pageId="14" pageNumber="355" type="discussion">
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<paragraph id="8BC8745CFFE5C91DFBF8418EFAF1FA15" blockId="14.[151,1437,1189,1754]" box="[199,1292,1441,1466]" pageId="14" pageNumber="355">
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<emphasis id="B903A84EFFE5C91DFBF8418EFEB2FA15" bold="true" box="[199,335,1441,1466]" pageId="14" pageNumber="355">Comments.</emphasis>
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Broun based the description on a single specimen that we confirmed as the
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<typeStatus id="54CCCAFEFFE5C91DFF9D418EFAF5FA15" box="[1186,1288,1441,1466]" pageId="14" pageNumber="355" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D27D7FFE5C91DFBF841EAFCFDFA71" box="[199,768,1477,1502]" pageId="14" pageNumber="355" type="distribution">
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<paragraph id="8BC8745CFFE5C91DFBF841EAFCFDFA71" blockId="14.[151,1437,1189,1754]" box="[199,768,1477,1502]" pageId="14" pageNumber="355">
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<emphasis id="B903A84EFFE5C91DFBF841EAFE9DFA71" bold="true" box="[199,352,1477,1502]" pageId="14" pageNumber="355">Distribution.</emphasis>
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South Island: MK, OL, WD, FD, SI.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D27D7FFE5C91DFBF841C6FCD3F975" pageId="14" pageNumber="355" type="materials_examined">
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<paragraph id="8BC8745CFFE5C91DFBF841C6FB0CF9C1" blockId="14.[151,1437,1189,1754]" pageId="14" pageNumber="355">
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<emphasis id="B903A84EFFE5C91DFBF841C6FE0AF9AD" bold="true" box="[199,503,1513,1538]" pageId="14" pageNumber="355">Type material examined.</emphasis>
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<materialsCitation id="3B1F7E01FFE5C91DF93D41C6FB0CF9C1" ID-GBIF-Occurrence="2147524958" collectionCode="BMNH" collectorName="Coll. Brit. Mus." country="New Zealand" county="Broun's" determinerName="R. Craw" location="Type" municipality="Broun's" pageId="14" pageNumber="355" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="54CCCAFEFFE5C91DF93D41C6FD97F9AD" box="[514,618,1513,1538]" pageId="14" pageNumber="355" type="holotype">Holotype</typeStatus>
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||
(
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<collectionCode id="ED66EC99FFE5C91DF94241C5FD25F9AD" box="[637,728,1514,1538]" pageId="14" pageNumber="355">BMNH</collectionCode>
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): card-mounted, “
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<location id="8EA82287FFE5C91DF89941C5FC1DF9AD" LSID="urn:lsid:plazi:treatment:03DEC54AFFE5C91DFBA847D8FCD3F975:8EA82287FFE5C91DF89941C5FC1DF9AD" box="[934,992,1514,1538]" country="New Zealand" county="Broun's" municipality="Broun's" name="Type" pageId="14" pageNumber="355">Type</location>
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[round label with red border] // 4311 [
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<location id="8EA82287FFE5C91DFBA24221FF1BF98A" LSID="urn:lsid:plazi:treatment:03DEC54AFFE5C91DFBA847D8FCD3F975:8EA82287FFE5C91DFBA24221FF1BF98A" box="[157,230,1550,1573]" country="New Zealand" county="Broun's" municipality="Broun's" name="Broun" pageId="14" pageNumber="355">Broun</location>
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number in pencil, unknown hand] //
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<collectingCountry id="F36034CCFFE5C91DF9A14221FCC6F989" box="[670,827,1549,1574]" name="New Zealand" pageId="14" pageNumber="355">New Zealand</collectingCountry>
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. [red underline] Broun
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<collectorName id="2682118AFFE5C91DFF714222FAFBF98A" box="[1102,1286,1549,1574]" pageId="14" pageNumber="355">Coll. Brit. Mus.</collectorName>
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1922-482. // Hollyford.
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<date id="FFC9529CFFE5C91DFA27421DFE8DF9E5" box="[280,368,1586,1610]" pageId="14" pageNumber="355" value="14-02-23">23.2.14</date>
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. [in
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<collectingMunicipality id="6BACEE26FFE5C91DFA94421DFDF9F9E5" box="[427,516,1586,1610]" pageId="14" pageNumber="355">Broun’s</collectingMunicipality>
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hand] //
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<taxonomicName id="4C770FDFFFE5C91DF947421EFC83F9E5" authorityName="Broun" authorityYear="1921" box="[632,894,1585,1610]" class="Insecta" family="Curculionidae" genus="Etheophanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="optandus">Etheophanus optandus</taxonomicName>
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. [in
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<collectingCounty id="62A90CD0FFE5C91DF887421DFBECF9E5" box="[952,1041,1586,1610]" pageId="14" pageNumber="355">Broun’s</collectingCounty>
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hand] //
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<typeStatus id="54CCCAFEFFE5C91DFF45421DFAF0F9E5" box="[1146,1293,1586,1610]" pageId="14" pageNumber="355" type="holotype">HOLOTYPE</typeStatus>
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[printed]
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<specimenCount id="9D71BFD5FFE5C91DFEB8421DFA61F9E6" box="[1415,1436,1586,1609]" pageId="14" pageNumber="355" type="female">♀</specimenCount>
|
||
<taxonomicName id="4C770FDFFFE5C91DFBA8427AFDDBF9C1" authority="Broun, 1923" authorityName="Broun" authorityYear="1921" box="[151,550,1621,1646]" class="Insecta" family="Curculionidae" genus="Etheophanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="optandus">
|
||
Etheophanus optandus
|
||
<bibRefCitation id="EFE609ADFFE5C91DFAA44279FDDBF9C1" author="Broun, T." box="[411,550,1622,1646]" pageId="14" pageNumber="355" pagination="667 - 708" refId="ref10401" refString="Broun, T. (1923) Descriptions of new genera and species of Coleoptera. Part VIII. Bulletin of the New Zealand Institute, 1, 667 - 708." type="journal article" year="1923">Broun, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
det.
|
||
<determinerName id="7E149898FFE5C91DF9614279FD40F9C1" box="[606,701,1622,1646]" pageId="14" pageNumber="355">R. Craw</determinerName>
|
||
, 1987 [in Craw’s hand on orange dayglow label]”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC8745CFFE5C91DFBF84256FCD3F975" blockId="14.[151,1437,1189,1754]" pageId="14" pageNumber="355">
|
||
<emphasis id="B903A84EFFE5C91DFBF84256FD06F93D" bold="true" box="[199,763,1657,1682]" pageId="14" pageNumber="355">Dissected material (NZAC). South Island. FD.</emphasis>
|
||
Lake Gunn, Eglington Val.,
|
||
<date id="FFC9529CFFE5C91DFF724255FB15F93D" box="[1101,1256,1657,1682]" pageId="14" pageNumber="355" value="1966-10-30">30 Oct. 1966</date>
|
||
, J.I. Townsend, litter under
|
||
<taxonomicName id="4C770FDFFFE5C91DFA2042B2FE5DF91A" box="[287,416,1693,1717]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="355" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B903A84EFFE5C91DFA2042B2FE5DF91A" box="[287,416,1693,1717]" italics="true" pageId="14" pageNumber="355">Nothofagus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<specimenCount id="9D71BFD5FFE5C91DFA8C42B1FDECF919" box="[435,529,1693,1718]" pageId="14" pageNumber="355" type="male">2 males</specimenCount>
|
||
).
|
||
<materialsCitation id="3B1F7E01FFE5C91DF91842B2FAFDF919" ID-GBIF-Occurrence="2147525089" box="[551,1280,1693,1718]" collectingDate="1968-11-12" collectionCode="WD" collectorName="Fox Glacier & J. I. Townsend" pageId="14" pageNumber="355" specimenCount="1" specimenCount-male="1">
|
||
<emphasis id="B903A84EFFE5C91DF91842B2FDA3F919" bold="true" box="[551,606,1693,1718]" pageId="14" pageNumber="355">
|
||
<collectionCode id="ED66EC99FFE5C91DF91842B2FDA7F919" box="[551,602,1693,1718]" pageId="14" pageNumber="355">WD</collectionCode>
|
||
.
|
||
</emphasis>
|
||
<collectorName id="2682118AFFE5C91DF95842B1FD0CF919" box="[615,753,1693,1718]" pageId="14" pageNumber="355">Fox Glacier</collectorName>
|
||
,
|
||
<date id="FFC9529CFFE5C91DF9C042B1FC62F919" box="[767,927,1693,1718]" pageId="14" pageNumber="355" value="1968-11-12">
|
||
<collectingDate id="EF8DAB74FFE5C91DF9C042B1FC62F919" box="[767,927,1693,1718]" pageId="14" pageNumber="355" value="1968-11-12">12 Nov. 1968</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2682118AFFE5C91DF89242B1FBB2F919" box="[941,1103,1693,1718]" pageId="14" pageNumber="355">J.I. Townsend</collectorName>
|
||
, litter (
|
||
<specimenCount id="9D71BFD5FFE5C91DFF9942B1FB05F919" box="[1190,1272,1693,1718]" pageId="14" pageNumber="355" type="male">1 male</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
; Fox Glacier, Minnehaha Tr.,
|
||
<date id="FFC9529CFFE5C91DFA7642EDFE2DF975" box="[329,464,1729,1754]" pageId="14" pageNumber="355" value="1965-02-08">8 Feb. 1965</date>
|
||
, N.A. Walker, litter (
|
||
<specimenCount id="9D71BFD5FFE5C91DF98242EDFCDCF975" box="[701,801,1729,1754]" pageId="14" pageNumber="355" type="female">1 female</specimenCount>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |