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<mods:title id="50ACAD65AB88E6BC145570D5FBB0BEB6">Revision and Phylogeny of the Genus Phausis (Coleoptera: Lampyridae) with the Description of Three New Species</mods:title>
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<subSubSection id="C36F18AEFFBCFFD77DB2557D5675F9F4" box="[151,564,1597,1624]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BCA4B25FFBCFFD77DB2557D5675F9F4" blockId="18.[151,564,1597,1660]" box="[151,564,1597,1624]" pageId="18" pageNumber="19">
<heading id="D082FC49FFBCFFD77DB2557D5675F9F4" bold="true" box="[151,564,1597,1624]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C7530A6FFBCFFD77DB2557D5675F9F4" ID-CoL="4FXJH" authority="LeConte, 1878" authorityName="LeConte" authorityYear="1878" box="[151,564,1597,1624]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBCFFD77DB2557D5675F9F4" bold="true" box="[151,564,1597,1624]" pageId="18" pageNumber="19">
<emphasis id="B9019737FFBCFFD77DB2557D5539F9F4" bold="true" box="[151,376,1597,1624]" italics="true" pageId="18" pageNumber="19">Phausis inaccensa</emphasis>
<bibRefCitation id="EFE436D4FFBCFFD77C5A557D5675F9F4" author="LeConte, J. L." box="[383,564,1597,1623]" pageId="18" pageNumber="19" pagination="593 - 669" refId="ref39939" refString="LeConte, J. L. (1878) Descriptions of new species. In: Hubbard, H. G. &amp; Schwarz, E. A., The Coleoptera of Florida and Michigan. Proceedings of the American Philosophical Society, 17 (101), pp. 593 - 669. https: // doi. org / 10.5962 / bhl. title. 60253" type="journal article" year="1878">LeConte, 1878</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C36F18AEFFBCFFD77DB255215583F9DF" box="[151,450,1633,1660]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BCA4B25FFBCFFD77DB255215583F9DF" blockId="18.[151,564,1597,1660]" box="[151,450,1633,1660]" pageId="18" pageNumber="19">
(
<figureCitation id="134E57A0FFBCFFD77DBA552154A9F9DF" box="[159,232,1633,1660]" captionStart="FIGURE 8" captionStartId="21.[152,255,1112,1136]" captionTargetBox="[204,1383,183,1082]" captionTargetId="figure-17@21.[200,1387,179,1086]" captionTargetPageId="21" captionText="FIGURE 8. Detail of North American distribution of Phausis inaccensa." figureDoi="http://doi.org/10.5281/zenodo.11348292" httpUri="https://zenodo.org/record/11348292/files/figure.png" pageId="18" pageNumber="19">Figs 8</figureCitation>
; 
<figureCitation id="134E57A0FFBCFFD77DD055225569F9DF" box="[245,296,1634,1660]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="18" pageNumber="19">17D</figureCitation>
; 
<figureCitation id="134E57A0FFBCFFD77C115522553CF9DF" box="[308,381,1634,1660]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (A–Jj) (from left to right: dorsal, lateral, ventral view). (A–C) Phausis branhami sp. nov.; (D–F) Phausis californica; (G–I) Phausis dorothae; (J–L) Phausis inaccensa; (M–O) Phausis luminosa; (P–R) Phausis marina; (S–U) Phausis milleri sp. nov.; (V–X) Phausis nigra; (Y–Aa) Phausis reticulata; (Bb–Dd) Phausis rhombica; (Ee–Gg) Phausis riversi; (Hh–Jj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="18" pageNumber="19">22J–L</figureCitation>
; 
<figureCitation id="134E57A0FFBCFFD77CAD552255FBF9DF" box="[392,442,1634,1660]" captionStart="FIGURE 24" captionStartId="48.[152,255,1943,1967]" captionTargetBox="[234,1354,181,1919]" captionTargetId="figure-28@48.[234,1354,181,1919]" captionTargetPageId="48" captionText="FIGURE 24. Type labels of Phausis (A–L). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina; (G) Phausis milleri sp. nov.; (H) Phausis nigra; (I) Phausis reticulata; (J) Phausis rhombica; (K) Phausis riversi; (L) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348327" httpUri="https://zenodo.org/record/11348327/files/figure.png" pageId="18" pageNumber="19">24D</figureCitation>
)
</paragraph>
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<subSubSection id="C36F18AEFFBCFFD77DB255E65687F958" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8BCA4B25FFBCFFD77DB255E6565DF91D" blockId="18.[151,710,1702,1787]" box="[151,540,1702,1726]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB656C0BFFBCFFD77DB255E6565DF91D" box="[151,540,1702,1726]" pageId="18" pageNumber="19">
<treatmentCitation id="0AD46D34FFBCFFD77DB255E6565DF91D" author="LeConte, J. L." box="[151,540,1702,1726]" page="611" pageId="18" pageNumber="19" year="1878">
<taxonomicName id="4C7530A6FFBCFFD77DB255E6565DF91D" ID-CoL="4FXJH" authority="LeConte 1878: 611" authorityName="LeConte" authorityPageNumber="611" authorityYear="1878" box="[151,540,1702,1726]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBCFFD77DB255E65512F91D" box="[151,339,1702,1726]" italics="true" pageId="18" pageNumber="19">Phausis inaccensa</emphasis>
<bibRefCitation id="EFE436D4FFBCFFD77C7C55E6565DF91D" author="LeConte, J. L." box="[345,540,1702,1726]" pageId="18" pageNumber="19" pagination="593 - 669" refId="ref39939" refString="LeConte, J. L. (1878) Descriptions of new species. In: Hubbard, H. G. &amp; Schwarz, E. A., The Coleoptera of Florida and Michigan. Proceedings of the American Philosophical Society, 17 (101), pp. 593 - 669. https: // doi. org / 10.5962 / bhl. title. 60253" type="journal article" year="1878">LeConte 1878: 611</bibRefCitation>
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</paragraph>
<paragraph id="8BCA4B25FFBCFFD77DB2558556EEF97E" blockId="18.[151,710,1702,1787]" box="[151,687,1732,1757]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB656C0BFFBCFFD77DB2558556EEF97E" box="[151,687,1732,1757]" pageId="18" pageNumber="19">
<taxonomicName id="4C7530A6FFBCFFD77DB2558555AEF97E" authority="(LeConte)" baseAuthorityName="LeConte" baseAuthorityYear="1885" box="[151,495,1733,1757]" class="Insecta" family="Lampyridae" genus="Lamprohiza" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBCFFD77DB25585553DF97F" box="[151,380,1733,1757]" italics="true" pageId="18" pageNumber="19">Lamprohiza inaccensa</emphasis>
(LeConte)
</taxonomicName>
: 
<treatmentCitation id="0AD46D34FFBCFFD77CDD558556EEF97E" author="LeConte, J. L." box="[504,687,1732,1757]" page="20" pageId="18" pageNumber="19" year="1885">
<bibRefCitation id="EFE436D4FFBCFFD77CDD558556EEF97E" author="LeConte, J. L." box="[504,687,1732,1757]" pageId="18" pageNumber="19" pagination="1 - 32" refId="ref40128" refString="LeConte, J. L. (1885) Short studies of North American Coleoptera. Transactions of the Entomological Society of America, 12 (2), 1 - 32. https: // doi. org / 10.2307 / 25076447" type="journal article" year="1885">LeConte 1885: 20</bibRefCitation>
</treatmentCitation>
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<paragraph id="8BCA4B25FFBCFFD77DB255A35687F958" blockId="18.[151,710,1702,1787]" box="[151,710,1763,1787]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB656C0BFFBCFFD77DB255A35687F958" box="[151,710,1763,1787]" pageId="18" pageNumber="19">
<taxonomicName id="4C7530A6FFBCFFD77DB255A355AEF958" authority="(LeConte)" baseAuthorityName="LeConte" baseAuthorityYear="1885" box="[151,495,1763,1787]" class="Insecta" family="Lampyridae" genus="Lamprohiza" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBCFFD77DB255A3553DF958" box="[151,380,1763,1787]" italics="true" pageId="18" pageNumber="19">Lamprohiza inaccensa</emphasis>
(LeConte)
</taxonomicName>
: 
<treatmentCitation id="0AD46D34FFBCFFD77CDD55A35687F958" author="Blatchley, W. S." box="[504,710,1763,1787]" page="823" pageId="18" pageNumber="19" year="1910">
<bibRefCitation id="EFE436D4FFBCFFD77CDD55A35687F958" author="Blatchley, W. S." box="[504,710,1763,1787]" pageId="18" pageNumber="19" pagination="1 - 1386" refId="ref38804" refString="Blatchley, W. S. (1910) The Coleoptera or Beetles of Indiana. Indiana Department of Geology and Natural Resources, No. 1, 1 - 1386. https: // doi. org / 10.5962 / bhl. title. 79314" type="book chapter" year="1910">Blatchley 1910: 823</bibRefCitation>
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="C36F18AEFFBCFFD67DB2546954A7F832" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BCA4B25FFBCFFD67DB25469559EFE72" blockId="18.[151,1437,1833,2040]" lastBlockId="19.[151,1437,151,2010]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Diagnosis. 
<emphasis id="B9019737FFBCFFD77C31546A5510F8E7" bold="true" box="[276,337,1834,1860]" pageId="18" pageNumber="19">Male</emphasis>
. 
<taxonomicName id="4C7530A6FFBCFFD77C785469566DF8E0" authorityName="LeConte" authorityYear="1878" box="[349,556,1833,1859]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBCFFD77C785469566DF8E0" box="[349,556,1833,1859]" italics="true" pageId="18" pageNumber="19">Phausis inaccensa</emphasis>
</taxonomicName>
can be distinguished from its congeners by the following characters: mandibles short (apices of mandibles not crossing); pronotum broadly transverse and subrhombus, with hind angles finely denticulate (
<figureCitation id="134E57A0FFBCFFD77C07543155CAF82F" box="[290,395,1905,1932]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="18" pageNumber="19">Fig. 17D</figureCitation>
); elytra reticulate to punctate but not as in 
<emphasis id="B9019737FFBCFFD77E5454315552F80C" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C7530A6FFBCFFD77E5454315000F828" authorityName="LeConte" authorityYear="1852" baseAuthorityName="Say" baseAuthorityYear="1825" box="[881,1089,1905,1931]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reticulata">Phausis reticulata</taxonomicName>
, 
<taxonomicName id="4C7530A6FFBCFFD7796854315157F828" authorityName="Fender" authorityYear="1966" box="[1101,1302,1905,1931]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="luminosa">Phausis luminosa</taxonomicName>
, or 
<taxonomicName id="4C7530A6FFBCFFD7786154315552F80C" authorityName="Hodson &amp; Branham" authorityYear="2024" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="christineae">Phausis christineae</taxonomicName>
</emphasis>
; elytral humeral angle (shoulder) sinuate (
<figureCitation id="134E57A0FFBCFFD77FC654D55708F813" box="[739,841,1941,1968]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="18" pageNumber="19">Fig. 17D</figureCitation>
); abdominal segment VII elongate, with apical margin emarginated; basal piece of aedeagus longer than wide, segments contiguous, and unpigmented; parameres with both outer and inner lateral margins convex. 
<taxonomicName id="4C7530A6FFBCFFD77FB3549D571AF854" authorityName="Fender" authorityYear="1962" box="[662,859,2013,2039]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dorothae">
<emphasis id="B9019737FFBCFFD77FB3549D571AF854" box="[662,859,2013,2039]" italics="true" pageId="18" pageNumber="19">Phausis dorothae</emphasis>
</taxonomicName>
also has short mandibles, but can be distinguished from 
<taxonomicName id="4C7530A6FFBDFFD67DF053D75520FF13" authorityName="LeConte" authorityYear="1878" box="[213,353,151,176]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBDFFD67DF053D75520FF13" box="[213,353,151,176]" italics="true" pageId="19" pageNumber="20">P. inaccensa</emphasis>
</taxonomicName>
by the absence of anterior vitreous areas (pronotal spots) and the body being dark brown versus yellowish-brown. 
<taxonomicName id="4C7530A6FFBDFFD67C4753FB5670FF77" authorityName="LeConte" authorityYear="1878" box="[354,561,187,213]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBDFFD67C4753FB5670FF77" box="[354,561,187,213]" italics="true" pageId="19" pageNumber="20">Phausis inaccensa</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C7530A6FFBDFFD67FDB53FB578BFF76" authorityName="Hodson &amp; Branham" authorityYear="2024" box="[766,970,187,213]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="branhami">
<emphasis id="B9019737FFBDFFD67FDB53FB578BFF76" box="[766,970,187,213]" italics="true" pageId="19" pageNumber="20">Phausis branhami</emphasis>
</taxonomicName>
but can be distinguished by the following characters: anterior vitreous pronotal spots are broadly falcate (
<figureCitation id="134E57A0FFBDFFD67E6A539F57F2FF5A" box="[847,947,223,249]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="19" pageNumber="20">Fig. 17D</figureCitation>
) (anterior vitreous areas (pronotal spots) are triangular with rounded edges in 
<taxonomicName id="4C7530A6FFBDFFD67F2E52435696FEBE" authorityName="Hodson &amp; Branham" authorityYear="2024" box="[523,727,259,285]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="branhami">
<emphasis id="B9019737FFBDFFD67F2E52435696FEBE" box="[523,727,259,285]" italics="true" pageId="19" pageNumber="20">Phausis branhami</emphasis>
</taxonomicName>
); the basal piece (dorsal view) (
<figureCitation id="134E57A0FFBDFFD67965524350DCFEBE" box="[1088,1181,259,285]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (A–Jj) (from left to right: dorsal, lateral, ventral view). (A–C) Phausis branhami sp. nov.; (D–F) Phausis californica; (G–I) Phausis dorothae; (J–L) Phausis inaccensa; (M–O) Phausis luminosa; (P–R) Phausis marina; (S–U) Phausis milleri sp. nov.; (V–X) Phausis nigra; (Y–Aa) Phausis reticulata; (Bb–Dd) Phausis rhombica; (Ee–Gg) Phausis riversi; (Hh–Jj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="19" pageNumber="20">Fig. 22J</figureCitation>
) is quadrate, segments contiguous, and unpigmented (basal piece is c-shaped, segments widely divergent, and moderately sclerotized (light brown) in 
<taxonomicName id="4C7530A6FFBDFFD67C36520B55A5FEC6" authorityName="Hodson &amp; Branham" authorityYear="2024" box="[275,484,331,357]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="branhami">
<emphasis id="B9019737FFBDFFD67C36520B55A5FEC6" box="[275,484,331,357]" italics="true" pageId="19" pageNumber="20">Phausis branhami</emphasis>
</taxonomicName>
); the ventral opening (
<figureCitation id="134E57A0FFBDFFD67FD7520B5718FEC6" box="[754,857,331,357]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (A–Jj) (from left to right: dorsal, lateral, ventral view). (A–C) Phausis branhami sp. nov.; (D–F) Phausis californica; (G–I) Phausis dorothae; (J–L) Phausis inaccensa; (M–O) Phausis luminosa; (P–R) Phausis marina; (S–U) Phausis milleri sp. nov.; (V–X) Phausis nigra; (Y–Aa) Phausis reticulata; (Bb–Dd) Phausis rhombica; (Ee–Gg) Phausis riversi; (Hh–Jj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="19" pageNumber="20">Fig. 22L</figureCitation>
) on the median lobe is greater than apical 1/2 of median lobe (ventral opening comprises of apical1/4 of median lobe in 
<taxonomicName id="4C7530A6FFBDFFD67E8E522F5037FE2A" authorityName="Hodson &amp; Branham" authorityYear="2024" box="[939,1142,367,393]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="branhami">
<emphasis id="B9019737FFBDFFD67E8E522F5037FE2A" box="[939,1142,367,393]" italics="true" pageId="19" pageNumber="20">Phausis branhami</emphasis>
</taxonomicName>
). This species is similar in coloration to 
<taxonomicName id="4C7530A6FFBDFFD67C0952D35597FE0E" authorityName="Hodson &amp; Branham" authorityYear="2024" box="[300,470,403,429]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B9019737FFBDFFD67C0952D35597FE0E" box="[300,470,403,429]" italics="true" pageId="19" pageNumber="20">Phausis milleri</emphasis>
</taxonomicName>
, but can be distinguished from it by the outer margin of the mandible is convex versus straight as in 
<taxonomicName id="4C7530A6FFBDFFD67C0852F75599FE72" authorityName="Hodson &amp; Branham" authorityYear="2024" box="[301,472,439,465]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B9019737FFBDFFD67C0852F75599FE72" box="[301,472,439,465]" italics="true" pageId="19" pageNumber="20">Phausis milleri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCA4B25FFBDFFD67DE2529B56EEFD0A" blockId="19.[151,1437,151,2010]" pageId="19" pageNumber="20">
Redescription. 
<emphasis id="B9019737FFBDFFD67C50529B55F3FE56" bold="true" box="[373,434,475,501]" pageId="19" pageNumber="20">Male</emphasis>
(
<figureCitation id="134E57A0FFBDFFD67CE3529B567CFE56" box="[454,573,475,501]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="19" pageNumber="20">Figs. 17D</figureCitation>
; 
<figureCitation id="134E57A0FFBDFFD67F68529B56D9FE56" box="[589,664,475,501]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (A–Jj) (from left to right: dorsal, lateral, ventral view). (A–C) Phausis branhami sp. nov.; (D–F) Phausis californica; (G–I) Phausis dorothae; (J–L) Phausis inaccensa; (M–O) Phausis luminosa; (P–R) Phausis marina; (S–U) Phausis milleri sp. nov.; (V–X) Phausis nigra; (Y–Aa) Phausis reticulata; (Bb–Dd) Phausis rhombica; (Ee–Gg) Phausis riversi; (Hh–Jj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="19" pageNumber="20">22J–L</figureCitation>
). Length 5.0– 
<quantity id="4C8DE6C0FFBDFFD67E19529A57D8FE56" box="[828,921,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="20" unit="mm" value="6.5">6.5 mm</quantity>
. Width 
<quantity id="4C8DE6C0FFBDFFD67EDD529A50C6FE56" box="[1016,1159,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="19" pageNumber="20" unit="mm" value="1.85" valueMax="2.5" valueMin="1.2">1.2–2.5 mm</quantity>
. 
<emphasis id="B9019737FFBDFFD679B3529B509DFE56" box="[1174,1244,475,501]" italics="true" pageId="19" pageNumber="20">Color.</emphasis>
Body yellowish brown; eyes black; mouthparts yellowish-brown, except tips of mandibles brunneous to ferrous; antennae yellowish brown; central pronotal disc yellowish-brown basally, dark brown apically; thorax overall yellowish-brown; legs yellowish-brown; lateral margins of elytra paler than elytral disc; abdomen brown with lateral margins of abdominal segments VII and VIII flavus; photic organs absent on abdominal segments VI, VII and VIII; pubescence grayish. Anterior vitreous areas (pronotal spots) present.
</paragraph>
<paragraph id="8BCA4B25FFBDFFD67DE251F35756FC22" blockId="19.[151,1437,151,2010]" pageId="19" pageNumber="20">
<emphasis id="B9019737FFBDFFD67DE251F35545FD6E" box="[199,260,691,717]" italics="true" pageId="19" pageNumber="20">Head</emphasis>
small and transverse (frontal view). Vertex longitudinally concave with fine punctures dorsally. Eyes large and approximate, partially obstructing labium postventrally. Mandibles short (apices of mandibles not crossing frontally), robust, and begins tapering at half length of mandible; incisor region short (less than half the total length of mandible); outer margin convex versus straight as in other taxa (
<taxonomicName id="4C7530A6FFBDFFD67EAA505F57BDFC9B" authorityName="Fender" authorityYear="1966" box="[911,1020,799,824]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="marina">
<emphasis id="B9019737FFBDFFD67EAA505F57BDFC9B" box="[911,1020,799,824]" italics="true" pageId="19" pageNumber="20">P. marina</emphasis>
</taxonomicName>
, 
<emphasis id="B9019737FFBDFFD6792E505F50A3FC9A" box="[1035,1250,799,825]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C7530A6FFBDFFD6792E505F5027FC9B" authorityName="Hopping, Det. Ralph Hopping" authorityYear="1937" box="[1035,1126,799,824]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigra">P. nigra</taxonomicName>
, 
<taxonomicName id="4C7530A6FFBDFFD67957505F50A3FC9A" box="[1138,1250,799,825]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="skelleyi">P. skelleyi</taxonomicName>
</emphasis>
, and 
<taxonomicName id="4C7530A6FFBDFFD67804505F51CAFC9A" authorityName="Hodson &amp; Branham" authorityYear="2024" box="[1313,1419,799,825]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B9019737FFBDFFD67804505F5173FC9B" box="[1313,1330,799,824]" italics="true" pageId="19" pageNumber="20">P</emphasis>
. 
<emphasis id="B9019737FFBDFFD67865505F51CAFC9A" box="[1344,1419,799,825]" italics="true" pageId="19" pageNumber="20">milleri</emphasis>
</taxonomicName>
); basal angle approximately less than 90 degrees dorsally. Maxillary palps with four palpomeres, last palpomere securiform; Labial palps reduced, with three palpomeres.
</paragraph>
<paragraph id="8BCA4B25FFBDFFD67DE250CB50B7FB96" blockId="19.[151,1437,151,2010]" pageId="19" pageNumber="20">
<emphasis id="B9019737FFBDFFD67DE250CB5570FC06" box="[199,305,907,933]" italics="true" pageId="19" pageNumber="20">Antennae</emphasis>
filiform, long, extending to posterior end of metathorax. Scape 2.0×longer than pedicel. Pedicel approximately 1/3 the length of antennomere 3. Antennomere 3 to 6 subequal in length. Antennomeres 7 to 10 shorter than previous antennomeres. Antennomere 11 longer and ovate at apex. Flagellomeres feebly compressed, but not as compressed as in 
<taxonomicName id="4C7530A6FFBDFFD67CF350B7567BFBB2" authorityName="Laporte" authorityYear="1833" box="[470,570,1015,1041]" class="Insecta" family="Lampyridae" genus="Lucidota" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBDFFD67CF350B7567BFBB2" box="[470,570,1015,1041]" italics="true" pageId="19" pageNumber="20">Lucidota</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C7530A6FFBDFFD67F4450B7568DFBB2" box="[609,716,1015,1041]" class="Insecta" family="Lampyridae" genus="Ellychnia" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBDFFD67F4450B7568DFBB2" box="[609,716,1015,1041]" italics="true" pageId="19" pageNumber="20">Ellychnia</emphasis>
</taxonomicName>
. When viewed laterally, flagellomeres are wider than in ventral view. Sensorium present on the distal end of antennomere 11, semi-transparent, small, and subulate.
</paragraph>
<paragraph id="8BCA4B25FFBDFFD67DE2577F5750FAAF" blockId="19.[151,1437,151,2010]" pageId="19" pageNumber="20">
<emphasis id="B9019737FFBDFFD67DE2577F5577FBFB" box="[199,310,1087,1112]" italics="true" pageId="19" pageNumber="20">Pronotum</emphasis>
broadly transverse especially basally, subrhombus (
<figureCitation id="134E57A0FFBDFFD67E53577E579FFBFA" box="[886,990,1086,1113]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="19" pageNumber="20">Fig. 17D</figureCitation>
); median carina present, extending from base to almost the apex of pronotum; anterior margin sometimes feebly emarginated; lateral margins arcuate and diverging; all margins reflexed, sides becoming explanate toward the hind angles; basal margin broadly convex; hind angles finely denticulate. Central pronotal disc present; quadrate; shiny and overall impunctate, with fine sparse punctures basally. Anterior vitreous areas (spots) present, large (less than one width of spot between windows), broadly falcate, appearing almost triangular, impunctate.
</paragraph>
<paragraph id="8BCA4B25FFBDFFD67DE256565618FAF7" blockId="19.[151,1437,151,2010]" pageId="19" pageNumber="20">
<taxonomicName id="4C7530A6FFBDFFD67DE256565574FA93" box="[199,309,1302,1328]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBDFFD67DE256565574FA93" box="[199,309,1302,1328]" italics="true" pageId="19" pageNumber="20">Scutellum</emphasis>
</taxonomicName>
spatulate, sides curving inward and posterior apex truncate-rounded with longitudinal groove (
<figureCitation id="134E57A0FFBDFFD6784A5656548AFAF7" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="19" pageNumber="20">Fig. 17D</figureCitation>
); surface dull and finely punctured.
</paragraph>
<paragraph id="8BCA4B25FFBDFFD67DE2561D574AFA47" blockId="19.[151,1437,151,2010]" pageId="19" pageNumber="20">
<emphasis id="B9019737FFBDFFD67DE2561D554CFAD4" box="[199,269,1373,1399]" italics="true" pageId="19" pageNumber="20">Elytra</emphasis>
well developed; rugose basally, becoming reticulate medially to apically; humeral angle (shoulder) sinuate (
<figureCitation id="134E57A0FFBDFFD67DDD56C15523FA38" box="[248,354,1409,1435]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="19" pageNumber="20">Fig. 17D</figureCitation>
); lateral margins widely explanate, with fine reticulation Elytral costa absent medially on each elytron. Longitudinal elevated lines present and transverse lines present, transverse lines somewhat irregular in the interstitial spaces giving elytra a reticulated appearance.
</paragraph>
<paragraph id="8BCA4B25FFBDFFD67DE256AE5125F9D1" blockId="19.[151,1437,151,2010]" pageId="19" pageNumber="20">
<emphasis id="B9019737FFBDFFD67DE256AE5552F9A4" box="[199,275,1518,1543]" italics="true" pageId="19" pageNumber="20">Venter.</emphasis>
Thorax transverse, rhomboidal. Metathorax with posterior apex notched; medial line present; surface shiny, overall impunctate, but sparsely-finely punctate laterally. Tergal apicolateral angles strongly lobed. Photic organs absent on abdominal segments VI, VII and VIII. Abdominal segment VII elongate (longer than wide), apical margin emarginated. Abdominal segment VIII same as abdominal segment VII except apical margin rounded.
</paragraph>
<paragraph id="8BCA4B25FFBDFFD67DE2553C54A7F832" blockId="19.[151,1437,151,2010]" pageId="19" pageNumber="20">
<emphasis id="B9019737FFBDFFD67DE2553C5539F935" box="[199,376,1660,1686]" italics="true" pageId="19" pageNumber="20">Male Genitalia.</emphasis>
Basal piece (dorsal view; 
<figureCitation id="134E57A0FFBDFFD67FBD553C56B3F934" box="[664,754,1660,1687]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (A–Jj) (from left to right: dorsal, lateral, ventral view). (A–C) Phausis branhami sp. nov.; (D–F) Phausis californica; (G–I) Phausis dorothae; (J–L) Phausis inaccensa; (M–O) Phausis luminosa; (P–R) Phausis marina; (S–U) Phausis milleri sp. nov.; (V–X) Phausis nigra; (Y–Aa) Phausis reticulata; (Bb–Dd) Phausis rhombica; (Ee–Gg) Phausis riversi; (Hh–Jj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="19" pageNumber="20">Fig. 22J</figureCitation>
) longer than wide, quadrate, contiguous (segments touching), and unsclerotized (translucent); median lobe (dorsal view) approximates tips of parameres, elongate and rounded (greater than 45 degrees) at apex; parallel to parameres; sensilla absent; dorsal inner-basal margins convex; dorsal opening absent; longitudinal dorsal carina absent; lateral line absent; ventral opening present (
<figureCitation id="134E57A0FFBDFFD6799F55A85161F8A1" box="[1210,1312,1768,1794]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (A–Jj) (from left to right: dorsal, lateral, ventral view). (A–C) Phausis branhami sp. nov.; (D–F) Phausis californica; (G–I) Phausis dorothae; (J–L) Phausis inaccensa; (M–O) Phausis luminosa; (P–R) Phausis marina; (S–U) Phausis milleri sp. nov.; (V–X) Phausis nigra; (Y–Aa) Phausis reticulata; (Bb–Dd) Phausis rhombica; (Ee–Gg) Phausis riversi; (Hh–Jj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="19" pageNumber="20">Fig. 22L</figureCitation>
), elliptical, comprising of more than apical half of medial lobe. Parameres (dorsal view) elongate, slender, sub-parallel; inner lateral margins convex; outer lateral margins broadly convex; apex roundly acute (lateral view; 
<figureCitation id="134E57A0FFBDFFD6799854705165F8E9" box="[1213,1316,1840,1866]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (A–Jj) (from left to right: dorsal, lateral, ventral view). (A–C) Phausis branhami sp. nov.; (D–F) Phausis californica; (G–I) Phausis dorothae; (J–L) Phausis inaccensa; (M–O) Phausis luminosa; (P–R) Phausis marina; (S–U) Phausis milleri sp. nov.; (V–X) Phausis nigra; (Y–Aa) Phausis reticulata; (Bb–Dd) Phausis rhombica; (Ee–Gg) Phausis riversi; (Hh–Jj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="19" pageNumber="20">Fig. 22K</figureCitation>
); proximal flanges (extensions of parameres that curve or extend outward from inner margin) absent (dorsal view); sensilla absent.
</paragraph>
</subSubSection>
<subSubSection id="C36F18AEFFBDFFD17DE254DB559FF85A" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="8BCA4B25FFBDFFD17DE254DB5571FF77" blockId="19.[151,1437,151,2010]" lastBlockId="20.[151,1437,150,2041]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="B9019737FFBDFFD67DE254DB5564F816" bold="true" box="[199,293,1947,1973]" pageId="19" pageNumber="20">Female.</emphasis>
Length 
<quantity id="4C8DE6C0FFBDFFD67CA754DB55A7F815" box="[386,486,1947,1974]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" pageId="19" pageNumber="20" unit="mm" value="6.05">6.05 mm</quantity>
(average). Apterous and appear physogastric with 8 true abdominal segments. 
<emphasis id="B9019737FFBDFFD6787254DB51DCF816" box="[1367,1437,1947,1973]" italics="true" pageId="19" pageNumber="20">Color.</emphasis>
Body pale-cream to yellow, parts transparent; thorax with a dorsal-central, dark irregular shaped spot; eyes black; antennomeres pale-yellow to light brown, with 3 to 5 segments expanding apically.Anterior vitreous areas (pronotal spots) absent.
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE2539E56DEFEBF" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
<emphasis id="B9019737FFBAFFD17DE2539E5545FF5B" box="[199,260,222,248]" italics="true" pageId="20" pageNumber="21">Head</emphasis>
Eyes compound, small and separated, not partially obstructing labium post-ventrally as it is in males. Smaller in diameter than 
<taxonomicName id="4C7530A6FFBAFFD17C975243567BFEBF" authorityName="LeConte" authorityYear="1852" baseAuthorityName="Say" baseAuthorityYear="1825" box="[434,570,258,284]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="B9019737FFBAFFD17C975243567BFEBF" box="[434,570,258,284]" italics="true" pageId="20" pageNumber="21">P. reticulata</emphasis>
</taxonomicName>
females.
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE252665669FEC7" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
<emphasis id="B9019737FFBAFFD17DE252665570FEE3" box="[199,305,294,320]" italics="true" pageId="20" pageNumber="21">Antennae</emphasis>
with 3 to 5 antennomeres instead of 11 as in males; clavate, membranous, with no distinct segmentation, short, does not reach to metathorax.
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE2522F576FFE50" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
<emphasis id="B9019737FFBAFFD17DE2522F5577FE2B" box="[199,310,367,392]" italics="true" pageId="20" pageNumber="21">Pronotum</emphasis>
transverse, rhombus; sides less arcuate and less strongly convergent before the rounded anterior margin compared to 
<taxonomicName id="4C7530A6FFBAFFD17CA552D256D0FE0F" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[384,657,402,428]" class="Insecta" family="Lampyridae" genus="Lamprohiza" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="splendidula">
<emphasis id="B9019737FFBAFFD17CA552D256D0FE0F" box="[384,657,402,428]" italics="true" pageId="20" pageNumber="21">Lamprohiza splendidula</emphasis>
</taxonomicName>
; all margins reflexed, very strongly so anteriorly, becoming flattened toward the hind angles; basal margin weakly undulate; hind angles rounded; anterior margin emarginated. Central pronotal disc absent. Anterior vitreous areas (spots) absent.
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE252BD5522FDB4" blockId="20.[151,1437,150,2041]" box="[199,355,509,535]" pageId="20" pageNumber="21">
<emphasis id="B9019737FFBAFFD17DE252BD554CFDB4" box="[199,269,509,535]" italics="true" pageId="20" pageNumber="21">Elytra</emphasis>
absent.
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE251625501FD04" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
<emphasis id="B9019737FFBAFFD17DE251625556FD98" box="[199,279,546,571]" italics="true" pageId="20" pageNumber="21">Venter.</emphasis>
Abdomen with 8 visible true abdominal segments. Tergal apicolateral angles strongly lobed as in males. Abdominal segments VII bearing paired photic organs, consisting of a luminous spot along the lateral margins of each segment (
<bibRefCitation id="EFE436D4FFBAFFD17CF3512956A1FD20" author="Faust, L. F. &amp; Forrest, T. G." box="[470,736,617,643]" pageId="20" pageNumber="21" pagination="177 - 189" refId="ref39141" refString="Faust, L. F. &amp; Forrest, T. G. (2017) Bringing light to the lives of the shadow ghosts, Phausis inaccensa (Coleoptera: Lampyridae). American Entomologist, 63 (3), 177 - 189. https: // doi. org / 10.1093 / ae / tmx 027" type="journal article" year="2017">Faust &amp; Forrest 2017</bibRefCitation>
). The two photic organs can only be seen if the female is emitting light.
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE251F15512FCF9" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
<emphasis id="B9019737FFBAFFD17DE251F15556FD68" bold="true" box="[199,279,689,715]" pageId="20" pageNumber="21">Larva.</emphasis>
1 
<superScript id="7C00E66DFFBAFFD17C0A51F1557BFD1C" attach="left" box="[303,314,689,703]" fontSize="6" pageId="20" pageNumber="21">st</superScript>
Instar. Measures 
<quantity id="4C8DE6C0FFBAFFD17F2C51F05631FD68" box="[521,624,688,715]" metricMagnitude="-3" metricUnit="m" metricValue="1.57" pageId="20" pageNumber="21" unit="mm" value="1.57">1.57 mm</quantity>
long and 
<quantity id="4C8DE6C0FFBAFFD17FC251F15701FD68" box="[743,832,689,715]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="20" pageNumber="21" unit="mm" value="0.7">0.7 mm</quantity>
wide. Pronotum wider than long (0.61 x 
<quantity id="4C8DE6C0FFBAFFD1780051F051D1FD68" box="[1317,1424,688,715]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="20" pageNumber="21" unit="mm" value="0.35">0.35 mm</quantity>
). 
<emphasis id="B9019737FFBAFFD17DB25194549CFD4D" box="[151,221,724,750]" italics="true" pageId="20" pageNumber="21">Color.</emphasis>
Initially pale white overall but turns greyish tan with a pale dorsal-central carina after 24 hours; eyes dark; antennae translucent white; legs pale yellow; thorax and abdominal segments pale yellow ventrally. 
<emphasis id="B9019737FFBAFFD1780E51B951DCFCB0" box="[1323,1437,761,787]" italics="true" pageId="20" pageNumber="21">Antennae.</emphasis>
Present. 
<emphasis id="B9019737FFBAFFD17DD2505D5502FC95" box="[247,323,797,822]" italics="true" pageId="20" pageNumber="21">Venter.</emphasis>
Has 8 true abdominal segments, plus a pygopodia. Tergal apical lateral angles strongly lobed (
<bibRefCitation id="EFE436D4FFBAFFD1787B505C5506FCF9" author="Faust, L. F. &amp; Forrest, T. G." pageId="20" pageNumber="21" pagination="177 - 189" refId="ref39141" refString="Faust, L. F. &amp; Forrest, T. G. (2017) Bringing light to the lives of the shadow ghosts, Phausis inaccensa (Coleoptera: Lampyridae). American Entomologist, 63 (3), 177 - 189. https: // doi. org / 10.1093 / ae / tmx 027" type="journal article" year="2017">Faust &amp; Forrest 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE25024571FFC01" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
<emphasis id="B9019737FFBAFFD17DE250245549FCDD" bold="true" box="[199,264,868,894]" pageId="20" pageNumber="21">Eggs.</emphasis>
Round. 
<quantity id="4C8DE6C0FFBAFFD17C4A50245588FCDC" box="[367,457,868,895]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="mm" value="0.5">0.5 mm</quantity>
in diameter. 
<emphasis id="B9019737FFBAFFD17F43502456EDFCDD" box="[614,684,868,894]" italics="true" pageId="20" pageNumber="21">Color.</emphasis>
Pale yellow. Clutch totals are approximately 20– 
<specimenCount id="9D7380ACFFBAFFD179CB5024510AFCDC" box="[1262,1355,868,895]" count="32" pageId="20" pageNumber="21" type="egg">32 eggs</specimenCount>
. Hatch approximately 34 days after oviposition (
<bibRefCitation id="EFE436D4FFBAFFD17F4150C85716FC01" author="Faust, L. F. &amp; Forrest, T. G." box="[612,855,904,930]" pageId="20" pageNumber="21" pagination="177 - 189" refId="ref39141" refString="Faust, L. F. &amp; Forrest, T. G. (2017) Bringing light to the lives of the shadow ghosts, Phausis inaccensa (Coleoptera: Lampyridae). American Entomologist, 63 (3), 177 - 189. https: // doi. org / 10.1093 / ae / tmx 027" type="journal article" year="2017">Faust &amp; Forrest 2017</bibRefCitation>
)
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE250EC5587FC65" blockId="20.[151,1437,150,2041]" box="[199,454,940,966]" pageId="20" pageNumber="21">Etymology. Unknown.</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE2508F55A7FB92" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
<materialsCitation id="3B1D4178FFBAFFD17DE2508F55A7FB92" collectionCode="MCZC" country="United States of America" location="MICHIGAN" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Michigan" typeStatus="lectotype">
Specimens Examined. 
<emphasis id="B9019737FFBAFFD17CF35090563AFC4A" box="[470,635,975,1001]" italics="true" pageId="20" pageNumber="21">Type Material</emphasis>
. 
<typeStatus id="54CEF587FFBAFFD17FB550905741FC49" box="[656,768,976,1002]" pageId="20" pageNumber="21" type="lectotype">Lectotype</typeStatus>
(designated here; 
<figureCitation id="134E57A0FFBAFFD17EC0508F500EFC4A" box="[997,1103,975,1001]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (A–F). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="20" pageNumber="21">Figs.17D</figureCitation>
; 
<figureCitation id="134E57A0FFBAFFD17946509050D4FC49" box="[1123,1173,976,1002]" captionStart="FIGURE 24" captionStartId="48.[152,255,1943,1967]" captionTargetBox="[234,1354,181,1919]" captionTargetId="figure-28@48.[234,1354,181,1919]" captionTargetPageId="48" captionText="FIGURE 24. Type labels of Phausis (A–L). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina; (G) Phausis milleri sp. nov.; (H) Phausis nigra; (I) Phausis reticulata; (J) Phausis rhombica; (K) Phausis riversi; (L) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348327" httpUri="https://zenodo.org/record/11348327/files/figure.png" pageId="20" pageNumber="21">24D</figureCitation>
) 
<emphasis id="B9019737FFBAFFD1798E508F51DDFC4A" bold="true" box="[1195,1436,975,1001]" pageId="20" pageNumber="21">
<collectingCountry id="F3620BB5FFBAFFD1798E508F50A6FC4A" box="[1195,1255,975,1001]" name="United States of America" pageId="20" pageNumber="21">USA</collectingCountry>
: 
<collectingRegion id="49B185C7FFBAFFD179DF508F51D6FC4A" box="[1274,1431,975,1001]" country="United States of America" name="Michigan" pageId="20" pageNumber="21">MICHIGAN</collectingRegion>
:
</emphasis>
“MARQUETTE Co., 37, Mich., 28, Type No 2798, 
<collectionCode id="ED64D3E0FFBAFFD17E2550B4570EFBAD" box="[768,847,1012,1038]" pageId="20" pageNumber="21">MCZC</collectionCode>
Image Database,” 
<specimenCount id="9D7380ACFFBAFFD1791450B4501EFBAD" box="[1073,1119,1012,1038]" count="1" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
[originally a syntype from LeConte Collection] (
<collectionCode id="ED64D3E0FFBAFFD17CAE5757559FFB92" box="[395,478,1047,1073]" pageId="20" pageNumber="21">MCZC</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE2577B564FFAF2" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
<emphasis id="B9019737FFBAFFD17DE2577B553CFBF6" box="[199,381,1083,1109]" italics="true" pageId="20" pageNumber="21">Other material.</emphasis>
<materialsCitation id="3B1D4178FFBAFFD17CAD577B5704FBF6" box="[392,837,1083,1109]" country="United States of America" location="Co" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Oklahoma">
<emphasis id="B9019737FFBAFFD17CAD577B56CFFBF6" bold="true" box="[392,654,1083,1109]" pageId="20" pageNumber="21">
<collectingCountry id="F3620BB5FFBAFFD17CAD577B5584FBF6" box="[392,453,1083,1109]" name="United States of America" pageId="20" pageNumber="21">USA</collectingCountry>
: 
<collectingRegion id="49B185C7FFBAFFD17CF0577B56C8FBF6" box="[469,649,1083,1109]" country="United States of America" name="Oklahoma" pageId="20" pageNumber="21">OKLAHOMA</collectingRegion>
:
</emphasis>
LATIMER 
<location id="8EAA1DFEFFBAFFD17E3B577B5704FBF6" LSID="urn:lsid:plazi:treatment:03DCFA33FFBCFFD17DB2557D559FF85A:8EAA1DFEFFBAFFD17E3B577B5704FBF6" box="[798,837,1083,1109]" country="United States of America" name="Co" pageId="20" pageNumber="21" stateProvince="Oklahoma">Co</location>
</materialsCitation>
., 
<materialsCitation id="3B1D4178FFBAFFD17E7C577B55F5FBDA" collectingDate="1983-04" collectionCode="FSCA, V" country="United States of America" pageId="20" pageNumber="21" specimenCount="3" specimenCount-male="3" stateProvince="Oklahoma">
<date id="FFCB6DE5FFBAFFD17E7C577B57F6FBF6" box="[857,951,1083,1109]" pageId="20" pageNumber="21" value="1983-04">
<collectingDate id="EF8F940DFFBAFFD17E7C577B57F6FBF6" box="[857,951,1083,1109]" pageId="20" pageNumber="21" value="1983-04">IV.1983</collectingDate>
</date>
, Coll. K. Stephan, Det. K. Stephan, 
<specimenCount id="9D7380ACFFBAFFD1784B577B51DDFBF5" box="[1390,1436,1083,1110]" count="3" pageId="20" pageNumber="21" type="male">3 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17DBA571F54ACFBDA" box="[159,237,1119,1145]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except 
<date id="FFCB6DE5FFBAFFD17CBB571F55F5FBDA" box="[414,436,1119,1145]" pageId="20" pageNumber="21" value="1983-05">
<collectionCode id="ED64D3E0FFBAFFD17CBB571F55F5FBDA" box="[414,436,1119,1145]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="21" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17C91571F5719FBDA" box="[436,856,1119,1146]" collectionCode="FSCA, V" country="United States of America" pageId="20" pageNumber="21" specimenCount="6" specimenCount-male="6" stateProvince="Oklahoma">
.1983, 
<specimenCount id="9D7380ACFFBAFFD17F24571F566EFBD9" box="[513,559,1119,1146]" count="6" pageId="20" pageNumber="21" type="male">6 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17F67571F56D1FBDA" box="[578,656,1119,1145]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except 
<date id="FFCB6DE5FFBAFFD17E67571F5719FBDA" box="[834,856,1119,1145]" pageId="20" pageNumber="21" value="1984-05">
<collectionCode id="ED64D3E0FFBAFFD17E67571F5719FBDA" box="[834,856,1119,1145]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="21" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17E7D571F5156FBD9" box="[856,1303,1119,1146]" collectionCode="FSCA" country="United States of America" pageId="20" pageNumber="21" specimenCount="15" specimenCount-male="15" stateProvince="Oklahoma">
.1984, 
<specimenCount id="9D7380ACFFBAFFD17E80571F5051FBDA" box="[933,1040,1119,1146]" count="15" pageId="20" pageNumber="21" type="male">15 males</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17907571F5031FBDA" box="[1058,1136,1119,1145]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17804571F5591FB3E" collectingDate="1985-04" collectionCode="FSCA, V" country="United States of America" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Oklahoma">
<date id="FFCB6DE5FFBAFFD17804571F513EFBD9" box="[1313,1407,1119,1146]" pageId="20" pageNumber="21" value="1985-04">
<collectingDate id="EF8F940DFFBAFFD17804571F513EFBD9" box="[1313,1407,1119,1146]" pageId="20" pageNumber="21" value="1985-04">IV.1985</collectingDate>
</date>
, 
<specimenCount id="9D7380ACFFBAFFD178AB571F54EDFB3D" count="1" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17D9B57C3554DFB3E" box="[190,268,1155,1181]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except 
<date id="FFCB6DE5FFBAFFD17C9F57C35591FB3E" box="[442,464,1155,1181]" pageId="20" pageNumber="21" value="1986-05">
<collectionCode id="ED64D3E0FFBAFFD17C9F57C35591FB3E" box="[442,464,1155,1181]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="21" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17CF557C35725FB3D" box="[464,868,1155,1182]" collectionCode="FSCA" country="United States of America" pageId="20" pageNumber="21" specimenCount="10" specimenCount-male="10" stateProvince="Oklahoma">
.1986, 
<specimenCount id="9D7380ACFFBAFFD17F3957C35618FB3D" box="[540,601,1155,1182]" count="10" pageId="20" pageNumber="21" type="male">10 ♂</specimenCount>
, (
<collectionCode id="ED64D3E0FFBAFFD17F5457C356FEFB3E" box="[625,703,1155,1181]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17E4857C3514AFB3D" box="[877,1291,1155,1182]" collectingDate="1986-04" collectionCode="FSCA" country="United States of America" pageId="20" pageNumber="21" specimenCount="9" specimenCount-male="9" stateProvince="Oklahoma">
<date id="FFCB6DE5FFBAFFD17E4857C3578AFB3E" box="[877,971,1155,1181]" pageId="20" pageNumber="21" value="1986-04">
<collectingDate id="EF8F940DFFBAFFD17E4857C3578AFB3E" box="[877,971,1155,1181]" pageId="20" pageNumber="21" value="1986-04">IV.1986</collectingDate>
</date>
, 
<specimenCount id="9D7380ACFFBAFFD17EFC57C35047FB3D" box="[985,1030,1155,1182]" count="9" pageId="20" pageNumber="21" type="male">9 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD1793D57C35027FB3E" box="[1048,1126,1155,1181]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD1783157C35593FB62" collectingDate="1987-04" collectionCode="FCSA, V" country="United States of America" pageId="20" pageNumber="21" specimenCount="18" specimenCount-male="18" stateProvince="Oklahoma">
<date id="FFCB6DE5FFBAFFD1783157C35130FB3E" box="[1300,1393,1155,1181]" pageId="20" pageNumber="21" value="1987-04">
<collectingDate id="EF8F940DFFBAFFD1783157C35130FB3E" box="[1300,1393,1155,1181]" pageId="20" pageNumber="21" value="1987-04">IV.1987</collectingDate>
</date>
, 
<specimenCount id="9D7380ACFFBAFFD178A557C354EDFB61" count="18" pageId="20" pageNumber="21" type="male">18 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17D9A57E7554CFB62" box="[191,269,1191,1217]" pageId="20" pageNumber="21">FCSA</collectionCode>
); same except 
<date id="FFCB6DE5FFBAFFD17C9957E75593FB62" box="[444,466,1191,1217]" pageId="20" pageNumber="21" value="1987-05">
<collectionCode id="ED64D3E0FFBAFFD17C9957E75593FB62" box="[444,466,1191,1217]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="21" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17CF757E75713FB61" box="[466,850,1191,1218]" collectionCode="FCSA" country="United States of America" pageId="20" pageNumber="21" specimenCount="8" specimenCount-male="8" stateProvince="Oklahoma">
.1987, 
<specimenCount id="9D7380ACFFBAFFD17F3B57E7560DFB61" box="[542,588,1191,1218]" count="8" pageId="20" pageNumber="21" type="male">8 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17F7B57E756EDFB62" box="[606,684,1191,1217]" pageId="20" pageNumber="21">FCSA</collectionCode>
); same except
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17E7957E7514BFB61" box="[860,1290,1191,1218]" collectingDate="1988-04" collectionCode="FSCA" country="United States of America" pageId="20" pageNumber="21" specimenCount="27" specimenCount-male="27" stateProvince="Oklahoma">
<date id="FFCB6DE5FFBAFFD17E7957E757F8FB62" box="[860,953,1191,1217]" pageId="20" pageNumber="21" value="1988-04">
<collectingDate id="EF8F940DFFBAFFD17E7957E757F8FB62" box="[860,953,1191,1217]" pageId="20" pageNumber="21" value="1988-04">IV.1988</collectingDate>
</date>
, 
<specimenCount id="9D7380ACFFBAFFD17EED57E75045FB61" box="[968,1028,1191,1218]" count="27" pageId="20" pageNumber="21" type="male">27 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD1793357E75025FB62" box="[1046,1124,1191,1217]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD1783157E75594FB46" collectingDate="1989-04" collectionCode="FSCA, V" country="United States of America" pageId="20" pageNumber="21" specimenCount="18" specimenCount-male="18" stateProvince="Oklahoma">
<date id="FFCB6DE5FFBAFFD1783157E75133FB62" box="[1300,1394,1191,1217]" pageId="20" pageNumber="21" value="1989-04">
<collectingDate id="EF8F940DFFBAFFD1783157E75133FB62" box="[1300,1394,1191,1217]" pageId="20" pageNumber="21" value="1989-04">IV.1989</collectingDate>
</date>
, 
<specimenCount id="9D7380ACFFBAFFD178A557E754EDFB45" count="18" pageId="20" pageNumber="21" type="male">18 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17D9A578B554CFB46" box="[191,269,1227,1253]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except 
<date id="FFCB6DE5FFBAFFD17C9A578B5594FB46" box="[447,469,1227,1253]" pageId="20" pageNumber="21" value="1989-05">
<collectionCode id="ED64D3E0FFBAFFD17C9A578B5594FB46" box="[447,469,1227,1253]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="21" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17CF0578B55DCFAAA" collectionCode="FSCA, R" country="United States of America" pageId="20" pageNumber="21" specimenCount="2" specimenCount-male="2" stateProvince="Oklahoma">
.1989, 
<specimenCount id="9D7380ACFFBAFFD17F07578B5611FB45" box="[546,592,1227,1254]" count="2" pageId="20" pageNumber="21" type="male">2 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17F46578B56F0FB46" box="[611,689,1227,1253]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
); same except 
<quantity id="4C8DE6C0FFBAFFD17E41578B57D1FB45" box="[868,912,1227,1254]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="20" pageNumber="21" unit="mi" value="5.0">5mi</quantity>
<collectorName id="26802EF3FFBAFFD17EBF578C506EFB46" box="[922,1071,1227,1254]" pageId="20" pageNumber="21">W. Red Oak</collectorName>
, 10.IV.85, forest litter, 
<collectorName id="26802EF3FFBAFFD17862578B54B7FAAA" pageId="20" pageNumber="21">Coll. J. Pakaluk</collectorName>
, #262, 
<collectorName id="26802EF3FFBAFFD17C6957B0553CFAA9" box="[332,381,1264,1290]" pageId="20" pageNumber="21">Det.</collectorName>
<collectionCode id="ED64D3E0FFBAFFD17CA057AF55DCFAAA" box="[389,413,1263,1289]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="21">R</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B1D4178FFBAFFD17C8757AF5056FAAA" box="[418,1047,1263,1290]" collectionCode="SEMC" collectorName="S. Miller" country="United States of America" location="Red Oak" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Oklahoma">
<collectorName id="26802EF3FFBAFFD17C8757AF5648FAAA" box="[418,521,1263,1289]" pageId="20" pageNumber="21">S. Miller</collectorName>
, 
<specimenCount id="9D7380ACFFBAFFD17F3057AF5600FAA9" box="[533,577,1263,1290]" count="1" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBAFFD17F7757AF56E7FAAA" box="[594,678,1263,1289]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="20" pageNumber="21" type="Museum">SEMC</collectionCode>
); same except 
<quantity id="4C8DE6C0FFBAFFD17E7757AF573FFAA9" box="[850,894,1263,1290]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="20" pageNumber="21" unit="mi" value="5.0">5mi</quantity>
W. 
<location id="8EAA1DFEFFBAFFD17E8B57AF5056FAAA" LSID="urn:lsid:plazi:treatment:03DCFA33FFBCFFD17DB2557D559FF85A:8EAA1DFEFFBAFFD17E8B57AF5056FAAA" box="[942,1047,1263,1289]" country="United States of America" name="Red Oak" pageId="20" pageNumber="21" stateProvince="Oklahoma">Red Oak</location>
</materialsCitation>
, 
<date id="FFCB6DE5FFBAFFD1790157AF50CCFAAA" box="[1060,1165,1263,1289]" pageId="20" pageNumber="21" value="1977-05-07">
7.V
<materialsCitation id="3B1D4178FFBAFFD1796857AF50CCFAAA" box="[1101,1165,1263,1289]" collectionCode="FSCA" country="United States of America" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Oklahoma">.1977</materialsCitation>
</date>
, Coll. K. Stephan, Det. K. Stephan, 
<specimenCount id="9D7380ACFFBAFFD17C0256535513FA8D" box="[295,338,1299,1326]" count="1" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
(FSCA). 
<materialsCitation id="3B1D4178FFBAFFD17CE1565357BCFA8E" box="[452,1021,1299,1325]" country="United States of America" location="Tannersville" municipality="Co." pageId="20" pageNumber="21" specimenCount="1" stateProvince="Pennsylvania">
<emphasis id="B9019737FFBAFFD17CE1565356E8FA8E" bold="true" box="[452,681,1299,1325]" pageId="20" pageNumber="21">
<collectingRegion id="49B185C7FFBAFFD17CE1565356E5FA8E" box="[452,676,1299,1325]" country="United States of America" name="Pennsylvania" pageId="20" pageNumber="21">PENNSYLVANIA</collectingRegion>
:
</emphasis>
MONROE 
<collectingMunicipality id="6BAED15FFFBAFFD17E1056535721FA8E" box="[821,864,1299,1325]" pageId="20" pageNumber="21">Co.</collectingMunicipality>
, 
<location id="8EAA1DFEFFBAFFD17E49565357BCFA8E" LSID="urn:lsid:plazi:treatment:03DCFA33FFBCFFD17DB2557D559FF85A:8EAA1DFEFFBAFFD17E49565357BCFA8E" box="[876,1021,1299,1325]" country="United States of America" municipality="Co." name="Tannersville" pageId="20" pageNumber="21" stateProvince="Pennsylvania">Tannersville</location>
</materialsCitation>
, 
<date id="FFCB6DE5FFBAFFD1792C565350CFFA8E" box="[1033,1166,1299,1325]" pageId="20" pageNumber="21" value="1948-06-14">
14.VI
<materialsCitation id="3B1D4178FFBAFFD17969565350CFFA8E" box="[1100,1166,1299,1325]" collectionCode="AMNH" collectorName="Coll. J. W. Green, J. W. &amp; Green Collection" country="United States of America" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Pennsylvania">.1948</materialsCitation>
</date>
, 
<collectorName id="26802EF3FFBAFFD179BE5653551EFAF2" pageId="20" pageNumber="21">Coll. J.W. Green, J.W. Green Collection</collectorName>
, 
<specimenCount id="9D7380ACFFBAFFD17C4F567755D4FAF1" box="[362,405,1335,1362]" count="1" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
(AMNH).
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE2561B55CFFA1E" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
Distribution. Widely distributed throughout eastern 
<collectingCountry id="F3620BB5FFBAFFD17E20561B57DBFAD6" box="[773,922,1371,1397]" name="United States of America" pageId="20" pageNumber="21">United States</collectingCountry>
in 
<collectingRegion id="49B185C7FFBAFFD17E9E561B5053FAD6" box="[955,1042,1371,1397]" country="United States of America" name="Indiana" pageId="20" pageNumber="21">Indiana</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17939561B50CBFAD6" box="[1052,1162,1371,1397]" country="United States of America" name="Michigan" pageId="20" pageNumber="21">Michigan</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD179B1561B514DFAD6" box="[1172,1292,1371,1397]" country="United States of America" name="Oklahoma" pageId="20" pageNumber="21">Oklahoma</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17833561B51D9FAD6" box="[1302,1432,1371,1397]" country="United States of America" name="Mississippi" pageId="20" pageNumber="21">Mississippi</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17DB2563F54BCFA3A" box="[151,253,1407,1433]" country="United States of America" name="Missouri" pageId="20" pageNumber="21">Missouri</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17C35563F55C8FA3A" box="[272,393,1407,1433]" country="United States of America" name="Minnesota" pageId="20" pageNumber="21">Minnesota</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17CBF563F5643FA3A" box="[410,514,1407,1433]" country="United States of America" name="Alabama" pageId="20" pageNumber="21">Alabama</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17F36563F563FFA3A" box="[531,638,1407,1433]" country="United States of America" name="Arkansas" pageId="20" pageNumber="21">Arkansas</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17FB556C05747FA39" box="[656,774,1408,1434]" country="United States of America" name="Tennessee" pageId="20" pageNumber="21">Tennessee</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17E3C563F5737FA3A" box="[793,886,1407,1433]" country="United States of America" name="Georgia" pageId="20" pageNumber="21">Georgia</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17EAD563F507BFA3A" box="[904,1082,1407,1433]" country="United States of America" name="North Carolina" pageId="20" pageNumber="21">North Carolina</collectingRegion>
, 
<collectingRegion id="49B185C7FFBAFFD17968563F50FBFA3A" box="[1101,1210,1407,1433]" country="United States of America" name="Maryland" pageId="20" pageNumber="21">Maryland</collectingRegion>
(Faust 2017), and 
<collectingRegion id="49B185C7FFBAFFD17DB256E3556DFA1E" box="[151,300,1443,1469]" country="United States of America" name="Pennsylvania" pageId="20" pageNumber="21">Pennsylvania</collectingRegion>
(
<figureCitation id="134E57A0FFBAFFD17C1956E355C1FA1E" box="[316,384,1443,1469]" captionStart="FIGURE 8" captionStartId="21.[152,255,1112,1136]" captionTargetBox="[204,1383,183,1082]" captionTargetId="figure-17@21.[200,1387,179,1086]" captionTargetPageId="21" captionText="FIGURE 8. Detail of North American distribution of Phausis inaccensa." figureDoi="http://doi.org/10.5281/zenodo.11348292" httpUri="https://zenodo.org/record/11348292/files/figure.png" pageId="20" pageNumber="21">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE256875638FA42" blockId="20.[151,1437,150,2041]" box="[199,633,1479,1505]" pageId="20" pageNumber="21">Phenology. Occurs April through May.</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE256AA509CF930" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
Remarks. Originally described as 
<taxonomicName id="4C7530A6FFBAFFD17F6456AA5750F9A7" authorityName="LeConte" authorityYear="1878" box="[577,785,1514,1540]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBAFFD17F6456AA5750F9A7" box="[577,785,1514,1540]" italics="true" pageId="20" pageNumber="21">Phausis inaccensa</emphasis>
</taxonomicName>
by LeConte in 1878 and 1881 it was placed into the genus 
<taxonomicName id="4C7530A6FFBAFFD17DB2554E555CF98B" baseAuthorityName="Olivier" baseAuthorityYear="1911" box="[151,285,1550,1576]" class="Insecta" family="Lampyridae" genus="Lamprohiza" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBAFFD17DB2554E555CF98B" box="[151,285,1550,1576]" italics="true" pageId="20" pageNumber="21">Lamprohiza</emphasis>
</taxonomicName>
first by LeConte in 1885, then by Blatchley in 1910. 
<taxonomicName id="4C7530A6FFBAFFD17E52554E5006F98B" authorityName="LeConte" authorityYear="1878" box="[887,1095,1550,1576]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBAFFD17E52554E5006F98B" box="[887,1095,1550,1576]" italics="true" pageId="20" pageNumber="21">Phausis inaccensa</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C7530A6FFBAFFD179FE554E5120F98B" baseAuthorityName="Olivier" baseAuthorityYear="1911" box="[1243,1377,1550,1576]" class="Insecta" family="Lampyridae" genus="Lamprohiza" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBAFFD179FE554E5120F98B" box="[1243,1377,1550,1576]" italics="true" pageId="20" pageNumber="21">Lamprohiza</emphasis>
</taxonomicName>
(like many other species in 
<taxonomicName id="4C7530A6FFBAFFD17CAE557255A5F9EF" authorityName="LeConte" authorityYear="1852" box="[395,484,1586,1612]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBAFFD17CAE557255A5F9EF" box="[395,484,1586,1612]" italics="true" pageId="20" pageNumber="21">Phausis</emphasis>
</taxonomicName>
) by having longer antennae, the presence of apical sensorium on the last antennomere, and having no tibial spurs. This species has since been returned to 
<taxonomicName id="4C7530A6FFBAFFD17EA555155798F9CC" authorityName="LeConte" authorityYear="1852" box="[896,985,1621,1647]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBAFFD17EA555155798F9CC" box="[896,985,1621,1647]" italics="true" pageId="20" pageNumber="21">Phausis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE436D4FFBAFFD17ECD55155039F9D3" author="Olivier, E." box="[1000,1144,1621,1648]" pageId="20" pageNumber="21" pagination="41 - 42" refId="ref41053" refString="Olivier, E. (1907) Coleoptera Family. Lampyridae. Wytsman Genera Insectorum, 53, 41 - 42." type="journal article" year="1907">Olivier 1907</bibRefCitation>
&amp; 
<bibRefCitation id="EFE436D4FFBAFFD179B05516508EF9D3" author="Olivier, E." box="[1173,1231,1622,1648]" pageId="20" pageNumber="21" pagination="31 - 68" refId="ref41075" refString="Olivier, E. (1910) Family Lampyridae. In: Schenkling, S. (Ed.), Coleopterum Catalogus. Pars 9. Wytsman Junk, Berlin, pp. 31 - 68." type="book chapter" year="1910">1910</bibRefCitation>
, 
<bibRefCitation id="EFE436D4FFBAFFD179FE551651DDF9CC" author="McDermott, F. A." box="[1243,1436,1621,1648]" pageId="20" pageNumber="21" pagination="1 - 72" refId="ref40774" refString="McDermott, F. A. (1964) The taxonomy of the Lampyridae (Coleoptera). Transactions of the American Entomological Society, 90, 1 - 72." type="journal article" year="1964">McDermott 1964</bibRefCitation>
&amp; 
<bibRefCitation id="EFE436D4FFBAFFD17D88553954A6F930" author="McDermott, F. A." box="[173,231,1657,1683]" pageId="20" pageNumber="21" pagination="1 - 149" refId="ref40805" refString="McDermott, F. A. (1966) Lampyridae. In: Steel, W. O. (Ed.) Coleopterum Catalogus Supplementa, Second edition. The Hague, Junk, pp. 1 - 149." type="book chapter" year="1966">1966</bibRefCitation>
, 
<bibRefCitation id="EFE436D4FFBAFFD17DD6553955C5F930" author="Fender, K. M." box="[243,388,1657,1683]" pageId="20" pageNumber="21" pagination="83 - 95" refId="ref39317" refString="Fender, K. M. (1966) The genus Phausis in America North of Mexico (Coleoptera-Lampyridae). Northwest Science, 40 (3), 83 - 95." type="journal article" year="1966">Fender 1966</bibRefCitation>
). This species is not mentioned in 
<bibRefCitation id="EFE436D4FFBAFFD17E23553957E6F930" author="Hopping, R." box="[774,935,1657,1683]" pageId="20" pageNumber="21" pagination="89 - 90" refId="ref39674" refString="Hopping, R. (1937) New Coleoptera from Western Canada VI. The Canadian Entomologist, 69, 89 - 90. https: // doi. org / 10.4039 / Ent 6989 - 4" type="journal article" year="1937">Hopping 1937</bibRefCitation>
as cited by Fender in 1966.
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE255DD5658F82D" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
A 
<materialsCitation id="3B1D4178FFBAFFD17DC555DD5160F978" collectorName="Marquette &amp; One" location="LeConte" municipality="In" pageId="20" pageNumber="21" specimenCount="2" specimenCount-male="2" stateProvince="Michigan" typeStatus="holotype">
<typeStatus id="54CEF587FFBAFFD17DC555DD5503F914" box="[224,322,1693,1719]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
has not been designated for this species. 
<collectingMunicipality id="6BAED15FFFBAFFD17E2955DE5762F91B" box="[780,803,1694,1720]" pageId="20" pageNumber="21">In</collectingMunicipality>
the original description 
<location id="8EAA1DFEFFBAFFD1791655DD50D2F914" LSID="urn:lsid:plazi:treatment:03DCFA33FFBCFFD17DB2557D559FF85A:8EAA1DFEFFBAFFD1791655DD50D2F914" box="[1075,1171,1693,1719]" municipality="In" name="LeConte" pageId="20" pageNumber="21" stateProvince="Michigan">LeConte</location>
recorded and described 
<specimenCount id="9D7380ACFFBAFFD17DB255815548F978" box="[151,265,1729,1755]" count="2" pageId="20" pageNumber="21" type="male">two males</specimenCount>
from 
<collectorName id="26802EF3FFBAFFD17C6B55815585F978" box="[334,452,1729,1755]" pageId="20" pageNumber="21">Marquette</collectorName>
, 
<collectingRegion id="49B185C7FFBAFFD17CEA5581567BF978" box="[463,570,1729,1755]" country="United States of America" name="Michigan" pageId="20" pageNumber="21">Michigan</collectingRegion>
(
<bibRefCitation id="EFE436D4FFBAFFD17F6D558156ACF978" author="LeConte, J. L." box="[584,749,1729,1755]" pageId="20" pageNumber="21" pagination="593 - 669" refId="ref39939" refString="LeConte, J. L. (1878) Descriptions of new species. In: Hubbard, H. G. &amp; Schwarz, E. A., The Coleoptera of Florida and Michigan. Proceedings of the American Philosophical Society, 17 (101), pp. 593 - 669. https: // doi. org / 10.5962 / bhl. title. 60253" type="journal article" year="1878">LeConte 1878</bibRefCitation>
). 
<collectorName id="26802EF3FFBAFFD17E2455815771F978" box="[769,816,1729,1755]" pageId="20" pageNumber="21">One</collectorName>
of these specimens has been designated as a
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD1780D55815794F881" collectionCode="MCZ" collectorName="In" location="Museum of Comparative Zoological Collection" municipality="LeConte Collection" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Michigan" typeStatus="syntype">
<typeStatus id="54CEF587FFBAFFD1780D5581513EF978" box="[1320,1407,1729,1755]" pageId="20" pageNumber="21" type="syntype">syntype</typeStatus>
in the 
<collectingMunicipality id="6BAED15FFFBAFFD17D9A55A555DBF95D" box="[191,410,1764,1791]" pageId="20" pageNumber="21">LeConte Collection</collectingMunicipality>
at the 
<location id="8EAA1DFEFFBAFFD17CC555A557AEF95D" LSID="urn:lsid:plazi:treatment:03DCFA33FFBCFFD17DB2557D559FF85A:8EAA1DFEFFBAFFD17CC555A557AEF95D" box="[480,1007,1764,1791]" municipality="LeConte Collection" name="Museum of Comparative Zoological Collection" pageId="20" pageNumber="21" stateProvince="Michigan">Museum of Comparative Zoological Collection</location>
(
<collectionCode id="ED64D3E0FFBAFFD17EDE55A5507BF95C" box="[1019,1082,1765,1791]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="20" pageNumber="21" type="Museum">MCZ</collectionCode>
web 2010). 
<collectorName id="26802EF3FFBAFFD179E455A55099F95C" box="[1217,1240,1765,1791]" pageId="20" pageNumber="21">In</collectorName>
order to maintain taxonomic stability and clarification of type locality, I here designate the
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17EFB5449562AF8C9" collectionCode="MCZC" location="Museum of Comparative Zoology Collection" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Michigan" typeStatus="syntype">
<typeStatus id="54CEF587FFBAFFD17EFB54495074F880" box="[990,1077,1801,1827]" pageId="20" pageNumber="21" type="syntype">syntype</typeStatus>
specimen from the 
<location id="8EAA1DFEFFBAFFD1783F54495648F8E5" LSID="urn:lsid:plazi:treatment:03DCFA33FFBCFFD17DB2557D559FF85A:8EAA1DFEFFBAFFD1783F54495648F8E5" name="Museum of Comparative Zoology Collection" pageId="20" pageNumber="21" stateProvince="Michigan">Museum of Comparative Zoology Collection</location>
with locality label data: “Marquette, 37, Mich.; 28; 
<taxonomicName id="4C7530A6FFBAFFD17968546C515DF8E5" authorityName="LeConte" authorityYear="1878" box="[1101,1308,1836,1862]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="inaccensa">
<emphasis id="B9019737FFBAFFD17968546C515DF8E5" box="[1101,1308,1836,1862]" italics="true" pageId="20" pageNumber="21">Phausis inaccensa</emphasis>
</taxonomicName>
LeC.; 
<emphasis id="B9019737FFBAFFD1784D546D51DDF8E5" box="[1384,1436,1837,1862]" italics="true" pageId="20" pageNumber="21">Type</emphasis>
No 2798; 
<collectionCode id="ED64D3E0FFBAFFD17C2C54105519F8C9" box="[265,344,1872,1898]" pageId="20" pageNumber="21">MCZC</collectionCode>
Image Database” as the
</materialsCitation>
<materialsCitation id="3B1D4178FFBAFFD17F5154105654F82D" collectionCode="ICZN" location="International Code of Zoological Nomenclature" municipality="Article" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Michigan" typeStatus="lectotype">
<typeStatus id="54CEF587FFBAFFD17F515410569AF8C9" box="[628,731,1872,1898]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
specimen according to 
<collectingMunicipality id="6BAED15FFFBAFFD17EC254105074F8C9" box="[999,1077,1872,1898]" pageId="20" pageNumber="21">Article</collectingMunicipality>
74 of the 
<location id="8EAA1DFEFFBAFFD1798E541055F6F82E" LSID="urn:lsid:plazi:treatment:03DCFA33FFBCFFD17DB2557D559FF85A:8EAA1DFEFFBAFFD1798E541055F6F82E" municipality="Article" name="International Code of Zoological Nomenclature" pageId="20" pageNumber="21" stateProvince="Michigan">International Code of Zoological Nomenclature</location>
(
<collectionCode id="ED64D3E0FFBAFFD17CE05434564CF82D" box="[453,525,1908,1934]" pageId="20" pageNumber="21">ICZN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE254D755B2F812" blockId="20.[151,1437,150,2041]" box="[199,499,1943,1970]" pageId="20" pageNumber="21">Variation. None identified.</paragraph>
<paragraph id="8BCA4B25FFBAFFD17DE254FB559FF85A" blockId="20.[151,1437,150,2041]" pageId="20" pageNumber="21">
Biology. Can be found in mixed-forest and dry oak ridgetops, moist vegetation near streams and rivers (
<bibRefCitation id="EFE436D4FFBAFFD1787B54FB5504F85A" author="Faust, L. F. &amp; Forrest, T. G." pageId="20" pageNumber="21" pagination="177 - 189" refId="ref39141" refString="Faust, L. F. &amp; Forrest, T. G. (2017) Bringing light to the lives of the shadow ghosts, Phausis inaccensa (Coleoptera: Lampyridae). American Entomologist, 63 (3), 177 - 189. https: // doi. org / 10.1093 / ae / tmx 027" type="journal article" year="2017">Faust &amp; Forrest 2017</bibRefCitation>
; Faust 2017).
</paragraph>
</subSubSection>
</treatment>
</document>