263 lines
37 KiB
XML
263 lines
37 KiB
XML
<document id="3371BE4BFE1B5E81F4CAD67B0058672D" ID-CLB-Dataset="28381" ID-DOI="10.11646/zootaxa.4565.3.8" ID-GBIF-Dataset="c16b340a-7325-43ea-9b33-7e0e7e25056f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2590345" ID-ZooBank="EF1B2384-F0B7-4ABD-8074-542849B7577B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552305307612" checkinUser="plazi" docAuthor="Neto, Jaime De Liege Gama, Ribeiro, José Moacir Ferreira & Passos, Mahedy Araujo Bastos" docDate="2019" docId="03CC87FB0F2B36447EB2FE78FC8EFEA7" docLanguage="en" docName="zootaxa.4565.3.8.pdf" docOrigin="Zootaxa 4565 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helicopsyche (Feropsyche) carajas Neto & Ribeiro & Passos 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="423" masterDocId="FFF5FF830F2936477E25FFA0FFC2FFBA" masterDocTitle="Two new species of Helicopsyche Siebold 1856 (Insecta: Trichoptera: Helicopsychidae) from Pará State, northern Brazil" masterLastPageNumber="426" masterPageNumber="420" pageNumber="422" updateTime="1698701578788" updateUser="ExternalLinkService">
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<mods:title id="459DA6D810DA8105E9EF6A03F422AD3B">Two new species of Helicopsyche Siebold 1856 (Insecta: Trichoptera: Helicopsychidae) from Pará State, northern Brazil</mods:title>
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<mods:name id="53C8729224A9DA2CDB3CECD77B2D9090" type="personal">
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<mods:namePart id="762204504DB5527A5A2C5762C46DDC6C">Neto, Jaime De Liege Gama</mods:namePart>
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<mods:namePart id="8A99E71F301CDD0A91CD3192CAC0DC7D">Ribeiro, José Moacir Ferreira</mods:namePart>
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<mods:name id="6702588E75D99295D3C32354C34EFEA1" type="personal">
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<mods:roleTerm id="5EA75C61ED8B70AC185E6CD3FA2AD1B7">Author</mods:roleTerm>
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<mods:namePart id="2287DFDEF3E9EDF767C47CDC3C2780D3">Passos, Mahedy Araujo Bastos</mods:namePart>
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<mods:title id="B8225B051D50B872675655B810D94D59">Zootaxa</mods:title>
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<mods:part id="4390399474E5B95779F7F8EFCF6865B7">
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<mods:date id="75AC218781E3CAFC638D7450288CCBC9">2019</mods:date>
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<mods:number id="4704534D789AD3CBF768A74A0593E5D4">2019-03-11</mods:number>
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<mods:number id="010D8E6819E7ADEC0A887E2217441A99">4565</mods:number>
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<treatment id="03CC87FB0F2B36447EB2FE78FC8EFEA7" ID-DOI="http://doi.org/10.5281/zenodo.5933441" ID-GBIF-Taxon="154367047" ID-Zenodo-Dep="5933441" LSID="urn:lsid:plazi:treatment:03CC87FB0F2B36447EB2FE78FC8EFEA7" httpUri="http://treatment.plazi.org/id/03CC87FB0F2B36447EB2FE78FC8EFEA7" lastPageId="3" lastPageNumber="423" pageId="2" pageNumber="422">
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<subSubSection id="C37F65660F2B36457EB2FE78FD61FE49" box="[151,675,472,499]" pageId="2" pageNumber="422" type="nomenclature">
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<paragraph id="8BDA36ED0F2B36457EB2FE78FD61FE49" blockId="2.[151,675,472,533]" box="[151,675,472,499]" pageId="2" pageNumber="422">
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<heading id="D09281810F2B36457EB2FE78FD61FE49" bold="true" box="[151,675,472,499]" fontSize="11" level="1" pageId="2" pageNumber="422" reason="1">
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<emphasis id="B911EAFF0F2B36457EB2FE78FD61FE49" bold="true" box="[151,675,472,499]" pageId="2" pageNumber="422">
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<taxonomicName id="4C654D6E0F2B36457EB2FE78FDFCFE48" ID-CoL="8P3HP" authority="Neto & Ribeiro & Passos, 2019" authorityName="Neto & Ribeiro & Passos" authorityYear="2019" box="[151,574,472,499]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Feropsyche">
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<emphasis id="B911EAFF0F2B36457EB2FE78FEFAFE48" bold="true" box="[151,312,472,498]" italics="true" pageId="2" pageNumber="422">Helicopsyche</emphasis>
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(
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<emphasis id="B911EAFF0F2B36457F6CFE78FE17FE48" bold="true" box="[329,469,472,498]" italics="true" pageId="2" pageNumber="422">Feropsyche</emphasis>
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)
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<emphasis id="B911EAFF0F2B36457FC3FE78FDFCFE48" bold="true" box="[486,574,472,498]" italics="true" pageId="2" pageNumber="422">carajas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257840F2B36457C60FE79FD61FE49" box="[581,675,473,499]" pageId="2" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37F65660F2B36457EB2FE5CFEEEFDAE" box="[151,300,508,533]" pageId="2" pageNumber="422" type="description">
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<paragraph id="8BDA36ED0F2B36457EB2FE5CFEEEFDAE" blockId="2.[151,675,472,533]" box="[151,300,508,533]" pageId="2" pageNumber="422">
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<figureCitation id="135E2A680F2B36457EB2FE5CFF20FDAE" box="[151,226,508,533]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Figs. 2</figureCitation>
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A–2G.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F65660F2B36457EB2FDE4FBF1FBEE" pageId="2" pageNumber="422" type="diagnosis">
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<paragraph id="8BDA36ED0F2B36457EB2FDE4FBF1FBEE" blockId="2.[151,1437,580,2009]" pageId="2" pageNumber="422">
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<emphasis id="B911EAFF0F2B36457EB2FDE4FEC8FDE7" bold="true" box="[151,266,580,605]" pageId="2" pageNumber="422">Diagnosis</emphasis>
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. The male of
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<taxonomicName id="4C654D6E0F2B36457F81FDE5FDD8FDE6" authority="Neto & Ribeiro & Passos, 2019" authorityName="Neto & Ribeiro & Passos" authorityYear="2019" box="[420,538,581,604]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="carajas" status="sp. nov.">
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<emphasis id="B911EAFF0F2B36457F81FDE5FDD8FDE6" box="[420,538,581,604]" italics="true" pageId="2" pageNumber="422">H. carajas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257840F2B36457C05FDE4FD9DFDE7" box="[544,607,580,605]" pageId="2" pageNumber="422" rank="species">
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<emphasis id="B911EAFF0F2B36457C05FDE4FD9DFDE7" bold="true" box="[544,607,580,605]" pageId="2" pageNumber="422">n. sp.</emphasis>
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</taxonomicNameLabel>
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is similar to those of
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<taxonomicName id="4C654D6E0F2B36457D6BFDE5FBA4FDE6" authority="Banks 1938" authorityName="Banks" authorityYear="1938" box="[846,1126,580,605]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="haitiensis">
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<emphasis id="B911EAFF0F2B36457D6BFDE5FC19FDE6" box="[846,987,580,604]" italics="true" pageId="2" pageNumber="422">H. haitiensis</emphasis>
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Banks 1938
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</taxonomicName>
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and
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<taxonomicName id="4C654D6E0F2B36457AB9FDE5FA5AFDE6" authority="Ross 1938" authorityName="Ross" authorityYear="1938" box="[1180,1432,580,604]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="limnella">
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<emphasis id="B911EAFF0F2B36457AB9FDE5FAD8FDE6" box="[1180,1306,580,604]" italics="true" pageId="2" pageNumber="422">H. limnella</emphasis>
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<bibRefCitation id="EFF44B1C0F2B36457B05FDE5FA5AFDE6" author="Ross, H. H." box="[1312,1432,580,604]" pageId="2" pageNumber="422" pagination="101 - 183" refId="ref3128" refString="Ross, H. H. (1938) Description of the Neartic caddisflies (Trichoptera), with special reference to Illinois species. Bulletin of the Illinois Natural History Survey, 21 (4), 101 - 183." type="journal article" year="1938">Ross 1938</bibRefCitation>
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</taxonomicName>
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, resembling each other mainly in the lateral view of the primary branch of each gonocoxite, with its posterior apex directed slightly ventrad, slender, and strongly pointed, and in the well-developed lateral lobes of the endotheca. In all three of these species, each gonocoxite has almost the same width basally and at midlength. However, in
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<taxonomicName id="4C654D6E0F2B36457BA4FD11FEC1FD56" authorityName="Banks" authorityYear="1938" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="haitiensis">
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<emphasis id="B911EAFF0F2B36457BA4FD11FEC1FD56" italics="true" pageId="2" pageNumber="422">H. haitiensis</emphasis>
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</taxonomicName>
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the dorsal margin of each gonocoxite is conspicuously concave in lateral view, but nearly straight in the other two species; in
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<taxonomicName id="4C654D6E0F2B36457FA3FD59FDCFFCAA" box="[390,525,760,784]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="haitensis">
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<emphasis id="B911EAFF0F2B36457FA3FD59FE58FCAA" box="[390,410,761,784]" italics="true" pageId="2" pageNumber="422">H</emphasis>
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.
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<emphasis id="B911EAFF0F2B36457F8CFD58FDCFFCAA" box="[425,525,760,784]" italics="true" pageId="2" pageNumber="422">haitensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C654D6E0F2B36457C61FD59FD00FCAA" authorityName="Ross" authorityYear="1938" box="[580,706,760,784]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="limnella">
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<emphasis id="B911EAFF0F2B36457C61FD59FD9AFCAA" box="[580,600,761,784]" italics="true" pageId="2" pageNumber="422">H</emphasis>
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.
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<emphasis id="B911EAFF0F2B36457C43FD58FD00FCAA" box="[614,706,760,784]" italics="true" pageId="2" pageNumber="422">limnella</emphasis>
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</taxonomicName>
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, each gonocoxite has a relatively smooth inner margin in ventral view while in the new species the inner margin of each gonocoxite has prominent setal chalazae about midlength; in
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<taxonomicName id="4C654D6E0F2B36457E96FCE1FEF0FCE2" authorityName="Ross" authorityYear="1938" box="[179,306,832,856]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="limnella">
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<emphasis id="B911EAFF0F2B36457E96FCE1FEF0FCE2" box="[179,306,832,856]" italics="true" pageId="2" pageNumber="422">H. limnella</emphasis>
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</taxonomicName>
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the gonocoxite as a whole is broader than that of the new species. In the new species the dorsal lobes of the endotheca are posteriorly protruding (but rounded in
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<taxonomicName id="4C654D6E0F2B36457D46FCC5FC35FCC6" authorityName="Banks" authorityYear="1938" box="[867,1015,868,892]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="haitiensis">
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<emphasis id="B911EAFF0F2B36457D46FCC5FC35FCC6" box="[867,1015,868,892]" italics="true" pageId="2" pageNumber="422">H. haitiensis</emphasis>
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</taxonomicName>
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and small in
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<taxonomicName id="4C654D6E0F2B36457A81FCC5FAE5FCC6" authorityName="Ross" authorityYear="1938" box="[1188,1319,868,892]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="limnella">
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<emphasis id="B911EAFF0F2B36457A81FCC5FAE5FCC6" box="[1188,1319,868,892]" italics="true" pageId="2" pageNumber="422">H. limnella</emphasis>
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</taxonomicName>
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;) and the phallotremal sclerite (= phallic sclerite of Johanson 2002) in ventral view it is bean-shaped and about midlength in endotheca (positioned anteriorly of the endotheca in
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<taxonomicName id="4C654D6E0F2B36457D2BFC0DFC60FC7E" authorityName="Banks" authorityYear="1938" box="[782,930,940,964]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="haitiensis">
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<emphasis id="B911EAFF0F2B36457D2BFC0DFC60FC7E" box="[782,930,940,964]" italics="true" pageId="2" pageNumber="422">H. haitiensis</emphasis>
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</taxonomicName>
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and circular in
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<taxonomicName id="4C654D6E0F2B36457A4DFC0DFB2EFC7E" authorityName="Ross" authorityYear="1938" box="[1128,1260,940,964]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="limnella">
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<emphasis id="B911EAFF0F2B36457A4DFC0DFB2EFC7E" box="[1128,1260,940,964]" italics="true" pageId="2" pageNumber="422">H. limnella</emphasis>
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</taxonomicName>
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). Additionally, segment X of the new species has 8 pairs of subequal megasetae distributed in 2 longitudinal rows distally (but in
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<taxonomicName id="4C654D6E0F2B36457EB2FC55FEEAFBB6" authorityName="Banks" authorityYear="1938" box="[151,296,1012,1036]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="haitiensis">
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<emphasis id="B911EAFF0F2B36457EB2FC55FEEAFBB6" box="[151,296,1012,1036]" italics="true" pageId="2" pageNumber="422">H. haitiensis</emphasis>
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</taxonomicName>
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it has about 23 short, stout megasetae in each of 2 irregular longitudinal rows starting laterally on anterior margin of tergite IX; and in
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<taxonomicName id="4C654D6E0F2B36457C16FBB9FD70FB8A" authorityName="Ross" authorityYear="1938" box="[563,690,1048,1072]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="limnella">
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<emphasis id="B911EAFF0F2B36457C16FBB9FD70FB8A" box="[563,690,1048,1072]" italics="true" pageId="2" pageNumber="422">H. limnella</emphasis>
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</taxonomicName>
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it has about 12 megasetae in each of 2 irregular longitudinal rows starting midway plus about 5 smaller megasetae in a transverse group at the apex).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F65660F2B36457EE2FBFFFD66F80F" pageId="2" pageNumber="422" type="description">
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<paragraph id="8BDA36ED0F2B36457EE2FBFFFBB3FAB2" blockId="2.[151,1437,580,2009]" pageId="2" pageNumber="422">
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<emphasis id="B911EAFF0F2B36457EE2FBFFFE93FBC2" bold="true" box="[199,337,1119,1144]" pageId="2" pageNumber="422">Description</emphasis>
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. Adult male. Head with cephalic warts oval and flat. Postantennal setal warts bean-shaped, almost as wide as scape. Antennal scape as long as eye diameter. Forewings brownish, each
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<quantity id="4C9D9B080F2B36457A68FB24FB61FB26" box="[1101,1187,1156,1181]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="422" unit="mm" value="3.5">3.5 mm</quantity>
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long (
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<typeStatus id="54DE884F0F2B36457ACFFB24FA91FB27" box="[1258,1363,1156,1181]" pageId="2" pageNumber="422" type="holotype">holotype</typeStatus>
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||
). Hind wings each
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||
<quantity id="4C9D9B080F2B36457F3FFB07FEB2FB7A" box="[282,368,1191,1216]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="422" unit="mm" value="2.5">2.5 mm</quantity>
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||
long (
|
||
<typeStatus id="54DE884F0F2B36457F92FB07FDDDFB7A" box="[439,543,1191,1216]" pageId="2" pageNumber="422" type="holotype">holotype</typeStatus>
|
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), with 22 hamuli on basal half of Costa; anal border with fringe of long setae, as long as hind wing width (
|
||
<figureCitation id="135E2A680F2B36457F91FB6CFDCCFB5E" box="[436,526,1228,1253]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2A</figureCitation>
|
||
). Apicomesal process of abdominal sternum VI about half as long as its segment, straight in lateral view, tubular, oriented posteroventrad, and apically rounded (
|
||
<figureCitation id="135E2A680F2B36457A29FB4FFBA6FAB2" box="[1036,1124,1263,1288]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA36ED0F2B36457EE2FAB4FE4DF82A" blockId="2.[151,1437,580,2009]" pageId="2" pageNumber="422">
|
||
<emphasis id="B911EAFF0F2B36457EE2FAB4FEBFFA97" bold="true" box="[199,381,1300,1325]" pageId="2" pageNumber="422">Male genitalia.</emphasis>
|
||
Segment IX, in lateral view, with anterior lobe on each side narrowly ellipsoid, oriented anteromesad; anterodorsal margin slightly convex; lateral apodeme on each side curved slightly dorsad posteriorly; tergal transverse apodeme absent (
|
||
<figureCitation id="135E2A680F2B36457C3FFAFCFDB1FACE" box="[538,627,1372,1397]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2C</figureCitation>
|
||
); in ventral view rectangular, anterior margin recessed, straight, without posterior process; sternal transverse apodeme present on each side (
|
||
<figureCitation id="135E2A680F2B36457D8DFADFFB81FA22" box="[936,1091,1407,1432]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Figs. 2C, 2D</figureCitation>
|
||
); in dorsal view with anterior margin deeply concave (
|
||
<figureCitation id="135E2A680F2B36457F8FFA04FDC0FA06" box="[426,514,1444,1469]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2E</figureCitation>
|
||
). Segment X directed posterad, in lateral view, dorsal margin nearly straight, apex truncate, ventral margin straight (
|
||
<figureCitation id="135E2A680F2B36457C32FA67FDB0FA5A" box="[535,626,1479,1504]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2C</figureCitation>
|
||
); in dorsal view, oblong, apex very shallowly notched, with 8 pairs of subequal megasetae in longitudinal rows near apex (
|
||
<figureCitation id="135E2A680F2B36457CC4FA4CFCFBF9BE" box="[737,825,1516,1541]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2E</figureCitation>
|
||
). Superior appendages clavate, oriented lateroposterad (
|
||
<figureCitation id="135E2A680F2B36457EBAF9AFFEF2F992" box="[159,304,1551,1576]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Figs. 2C, 2E</figureCitation>
|
||
). Gonocoxites each having primary branch, in lateral view, with basal half nearly as broad as median part and parallel-sided, posterior apex tapered and slender, curved slightly ventrad and strongly pointed, posterior margin unpigmented (
|
||
<figureCitation id="135E2A680F2B36457FAAF9F7FE2AF9CA" box="[399,488,1623,1648]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2C</figureCitation>
|
||
); in ventral view without incision between primary branch and basomesal lobe (
|
||
<figureCitation id="135E2A680F2B36457B4AF9F7FF7FF92E" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2D</figureCitation>
|
||
); in dorsal and ventral views elongate and strongly tapering apically, inner margin with 4 setal chalazae about midlength, external margin smooth (
|
||
<figureCitation id="135E2A680F2B36457C18F93FFD15F902" box="[573,727,1695,1720]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Figs. 2D, 2E</figureCitation>
|
||
); basomesal lobe, in lateral view, protruding beyond primary branch, with 7–9 apical megasetae (
|
||
<figureCitation id="135E2A680F2B36457C02F964FDBDF966" box="[551,639,1732,1757]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2C</figureCitation>
|
||
); in ventral view, subrectangular (
|
||
<figureCitation id="135E2A680F2B36457DDFF964FB96F966" box="[1018,1108,1732,1757]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2D</figureCitation>
|
||
). Basal plate, in ventral view, anteriorly tapered, with apex narrowly rounded (
|
||
<figureCitation id="135E2A680F2B36457CC0F947FC85F8BA" box="[741,839,1767,1792]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2D</figureCitation>
|
||
). Phallus, in lateral view, evenly bent ventrad at midlength, phallic shield (
|
||
<bibRefCitation id="EFF44B1C0F2B36457FE2F8ACFD97F89F" author="Morse, J. C." box="[455,597,1804,1829]" pageId="2" pageNumber="422" pagination="1 - 97" refId="ref3026" refString="Morse, J. C. (1975) A phylogeny and revision of the caddisfly genus Ceraclea (Trichoptera, Leptoceridae). Contributions of the American Entomological Institute, 11 (2), 1 - 97." type="journal article" year="1975">Morse 1975</bibRefCitation>
|
||
) evident at base in lateral and ventral views (
|
||
<figureCitation id="135E2A680F2B36457A51F8ACFAC9F89E" box="[1140,1291,1804,1829]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Figs. 2F, 2G</figureCitation>
|
||
); endotheca, lateral view, with protruding dorsal and ventral lobes posteriorly (
|
||
<figureCitation id="135E2A680F2B36457D87F88FFC3EF8F2" box="[930,1020,1839,1864]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Fig. 2F</figureCitation>
|
||
), in ventral view with lateral lobes prominent; sperm channel inflated basally (
|
||
<figureCitation id="135E2A680F2B36457CA4F8F4FCD0F8D6" box="[641,786,1876,1901]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Figs. 2F, 2G</figureCitation>
|
||
); phallotremal sclerite (
|
||
<bibRefCitation id="EFF44B1C0F2B36457A39F8F4FB77F8D7" author="Nielsen, A." box="[1052,1205,1876,1901]" pageId="2" pageNumber="422" pagination="1 - 159" refId="ref3088" refString="Nielsen, A. (1957) A comparative study of the genital segments and their appendages in male Trichoptera. Biologiske Skrifter udgivet af det Kongelige Danske Uidenskabernes Selskab, 8 (5), 1 - 159." type="journal article" year="1957">Nielsen 1957</bibRefCitation>
|
||
) conspicuous, beanshaped (
|
||
<figureCitation id="135E2A680F2B36457ED1F8D7FE41F82A" box="[244,387,1911,1936]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1993]" captionTargetBox="[158,1415,360,1906]" captionTargetId="figure@3.[150,1436,337,1950]" captionTargetPageId="3" captionText="FIGURE 2. Helicopsyche (Feropsyche) carajas sp. nov. 2A, male forewing (upper) and hind wing (lower). 2B, VI sternal process, left lateral. 2C–2G, male genitalia: 2C, left lateral; 2D, ventral; 2E, dorsal; 2F, phallus, left lateral; 2G, phallus, ventral." httpUri="https://zenodo.org/record/2590351/files/figure.png" pageId="2" pageNumber="422">Figs. 2F, 2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA36ED0F2B36457EE2F83CFD66F80F" blockId="2.[151,1437,580,2009]" box="[199,676,1948,1973]" pageId="2" pageNumber="422">
|
||
<emphasis id="B911EAFF0F2B36457EE2F83CFDDDF80F" bold="true" box="[199,543,1948,1973]" pageId="2" pageNumber="422">Female, pupa, larva, and egg</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F65660F2B36447EE2F860FEA2FF6E" lastPageId="3" lastPageNumber="423" pageId="2" pageNumber="422" type="materials_examined">
|
||
<paragraph id="8BDA36ED0F2B36447EE2F860FEA2FF6E" blockId="2.[151,1437,580,2009]" lastBlockId="3.[151,1437,151,285]" lastPageId="3" lastPageNumber="423" pageId="2" pageNumber="422">
|
||
<materialsCitation id="3B0D3CB00F2B36447EE2F860FEA2FF6E" ID-GBIF-Occurrence="2012823625" collectingDate="2018-08-07" collectingDateMax="2018-08-09" collectingDateMin="2018-08-07" collectionCode="MPEG" collectorName="Serra dos Carajas & J. L. Gama Neto & J. M. F. Ribeiro." country="Brazil" lastPageId="3" lastPageNumber="423" latitude="-6.04023" location="Parauapebas" longLatPrecision="1" longitude="-50.293938" pageId="2" pageNumber="422" specimenCount="1" stateProvince="Para" typeStatus="holotype">
|
||
<emphasis id="B911EAFF0F2B36457EE2F860FDC7F863" bold="true" box="[199,517,1984,2009]" pageId="2" pageNumber="422">
|
||
<typeStatus id="54DE884F0F2B36457EE2F860FEF6F863" box="[199,308,1984,2009]" pageId="2" pageNumber="422" type="holotype">Holotype</typeStatus>
|
||
male.
|
||
<collectingCountry id="F372767D0F2B36457FBCF860FDC7F863" box="[409,517,1984,2009]" name="Brazil" pageId="2" pageNumber="422">BRAZIL</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B911EAFF0F2B36457C3AF860FDA0F863" bold="true" box="[543,610,1984,2009]" pageId="2" pageNumber="422">
|
||
<collectingRegion id="49A1F80F0F2B36457C3AF860FD9FF863" box="[543,605,1984,2009]" country="Brazil" name="Para" pageId="2" pageNumber="422">Pará</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<location id="8EBA60360F2B36457C51F860FCC2F863" LSID="urn:lsid:plazi:treatment:03CC87FB0F2B36447EB2FE78FC8EFEA7:8EBA60360F2B36457C51F860FCC2F863" box="[628,768,1984,2009]" country="Brazil" latitude="-6.04023" longLatPrecision="1" longitude="-50.293938" name="Parauapebas" pageId="2" pageNumber="422" stateProvince="Para">Parauapebas</location>
|
||
municipality,
|
||
<collectorName id="2690533B0F2B36457D9CF860FB55F863" box="[953,1175,1984,2009]" pageId="2" pageNumber="422">Serra dos Carajás</collectorName>
|
||
, small order stream,
|
||
<geoCoordinate id="EE51502A0F2A36447EB2FF37FEF6FF0A" box="[151,308,151,176]" direction="south" orientation="latitude" pageId="3" pageNumber="423" precision="1" value="-6.04023">6°2’24.828”S</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE51502A0F2A36447F66FF37FDC6FF0A" box="[323,516,151,176]" direction="west" orientation="longitude" pageId="3" pageNumber="423" precision="1" value="-50.293938">50°17’38.184”W</geoCoordinate>
|
||
,
|
||
<collectingDate id="EF9FE9C50F2A36447C36FF37FD69FF0A" box="[531,683,151,176]" pageId="3" pageNumber="423" value="2018-08-07" valueMax="2018-08-09" valueMin="2018-08-07">
|
||
<date id="FFDB102D0F2A36447C36FF37FD69FF0A" box="[531,683,151,176]" pageId="3" pageNumber="423" value="2018-08-07" valueMax="2018-08-09" valueMin="2018-08-07">7–9.viii.2018</date>
|
||
</collectingDate>
|
||
.
|
||
<collectingMethod id="52244EFA0F2A36447C9FFF37FC05FF15" box="[698,967,151,176]" pageId="3" pageNumber="423">Pennsylvania light trap</collectingMethod>
|
||
, leg.
|
||
<collectorName id="2690533B0F2A36447A2EFF38FB06FF15" box="[1035,1220,152,176]" pageId="3" pageNumber="423">J.L. Gama Neto</collectorName>
|
||
&
|
||
<collectorName id="2690533B0F2A36447AD5FF38FA5FFF0A" box="[1264,1437,151,176]" pageId="3" pageNumber="423">J.M.F. Ribeiro.</collectorName>
|
||
(alcohol;
|
||
<collectionCode id="ED74AE280F2A36447F24FF1CFE96FF6E" box="[257,340,188,212]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="3" pageNumber="423">MPEG</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F65660F2A36447EE2FF7FFD14FF42" box="[199,726,223,248]" pageId="3" pageNumber="423" type="distribution">
|
||
<paragraph id="8BDA36ED0F2A36447EE2FF7FFD14FF42" blockId="3.[151,1437,151,285]" box="[199,726,223,248]" pageId="3" pageNumber="423">
|
||
<emphasis id="B911EAFF0F2A36447EE2FF7FFE9BFF42" bold="true" box="[199,345,223,248]" pageId="3" pageNumber="423">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F372767D0F2A36447F42FF7FFE69FF42" box="[359,427,223,248]" name="Brazil" pageId="3" pageNumber="423">Brazil</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A1F80F0F2A36447F9EFF40FE31FF42" box="[443,499,224,248]" country="Brazil" name="Para" pageId="3" pageNumber="423">Pará</collectingRegion>
|
||
).
|
||
<typeStatus id="54DE884F0F2A36447C20FF40FDFFFF42" box="[517,573,224,248]" pageId="3" pageNumber="423">Type</typeStatus>
|
||
locality only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F65660F2A36447EE2FEA4FC8EFEA7" box="[199,844,260,285]" pageId="3" pageNumber="423" type="etymology">
|
||
<paragraph id="8BDA36ED0F2A36447EE2FEA4FC8EFEA7" blockId="3.[151,1437,151,285]" box="[199,844,260,285]" pageId="3" pageNumber="423">
|
||
<emphasis id="B911EAFF0F2A36447EE2FEA4FE84FEA7" bold="true" box="[199,326,260,285]" pageId="3" pageNumber="423">Etymology</emphasis>
|
||
. The term “carajas” refers to the
|
||
<typeStatus id="54DE884F0F2A36447C9EFEA5FD29FEA7" box="[699,747,261,285]" pageId="3" pageNumber="423">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |