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<document id="CC386C8D4AD51DEE3DFD77CCCF40FDF9" ID-DOI="10.11646/zootaxa.4750.1.3" ID-GBIF-Dataset="295158ed-c627-49b9-adab-9c0c9f46ee7c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3702831" ID-ZooBank="04D225FD-F6C9-4503-9536-BCD723194C14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583849190549" checkinUser="plazi" docAuthor="Kovačić, Marcelo, Sadeghi, Reza &amp; Esmaeili, Hamid Reza" docDate="2020" docId="03CC87F7FFAFA204AAE8A2783CBBFB0C" docLanguage="en" docName="zootaxa.4750.1.3.pdf" docOrigin="Zootaxa 4750 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Silhouettea in Miller 1988" docType="key" docVersion="7" lastPageNumber="55" masterDocId="FFF5FF8FFFA9A202AA7FA3513927FFB3" masterDocTitle="New species of Silhouettea (Teleostei: Gobiidae) from Qeshm Island, Iran and the DNA barcoding of the Persian Gulf and Oman Sea gobies" masterLastPageNumber="66" masterPageNumber="49" pageNumber="55" updateTime="1698726614545" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A4EB6B9A0669164DD163946EC003210C">New species of Silhouettea (Teleostei: Gobiidae) from Qeshm Island, Iran and the DNA barcoding of the Persian Gulf and Oman Sea gobies</mods:title>
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<treatment id="03CC87F7FFAFA204AAE8A2783CBBFB0C" ID-DOI="http://doi.org/10.5281/zenodo.3706317" ID-GBIF-Taxon="162481308" ID-Zenodo-Dep="3706317" LSID="urn:lsid:plazi:treatment:03CC87F7FFAFA204AAE8A2783CBBFB0C" httpUri="http://treatment.plazi.org/id/03CC87F7FFAFA204AAE8A2783CBBFB0C" lastPageNumber="55" pageId="6" pageNumber="55">
<subSubSection id="C37F656AFFAFA204AAE8A2783D1DFEDB" pageId="6" pageNumber="55" type="nomenclature">
<paragraph id="8BDA36E1FFAFA204AAE8A2783B1AFEF7" blockId="6.[151,1082,297,360]" box="[151,573,297,324]" pageId="6" pageNumber="55">
<heading id="D092818DFFAFA204AAE8A2783B1AFEF7" bold="true" box="[151,573,297,324]" fontSize="11" level="1" pageId="6" pageNumber="55" reason="1">
<emphasis id="B911EAF3FFAFA204AAE8A2783B1AFEF7" bold="true" box="[151,573,297,324]" pageId="6" pageNumber="55">
Key to valid species of
<taxonomicName id="4C654D62FFAFA204ABCBA2783B1AFEF7" ID-CoL="7GV6" ID-ENA="1365714" authorityName="in Miller" authorityYear="1988" box="[436,573,297,324]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B911EAF3FFAFA204ABCBA2783B1AFEF7" bold="true" box="[436,573,297,324]" italics="true" pageId="6" pageNumber="55">Silhouettea</emphasis>
</taxonomicName>
</emphasis>
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</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A21C3D1DFEDB" blockId="6.[151,1082,297,360]" box="[151,1082,333,360]" pageId="6" pageNumber="55">
(modified from
<bibRefCitation id="EFF44B10FFAFA204AB35A21C38C5FED4" author="Miller, P. J." box="[330,482,333,359]" pageId="6" pageNumber="55" pagination="241 - 273" refId="ref12805" refString="Miller, P. J. (1988) Studies on Silhouettea Smith 1959 and an account of Ebomegobius Herre 1946 (Pisces: Gobiidae). Senckenbergiana Biologica, 68 (4 / 6), 241 - 273." type="journal article" year="1988">Miller (1988)</bibRefCitation>
to accommodate
<taxonomicName id="4C654D62FFAFA204A8D7A21C3A6DFED4" authorityName="Randall" authorityYear="2008" box="[680,842,333,359]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="capitlineata">
<emphasis id="B911EAF3FFAFA204A8D7A21C3A6DFED4" box="[680,842,333,359]" italics="true" pageId="6" pageNumber="55">S. capitlineata</emphasis>
</taxonomicName>
and the new species)
</paragraph>
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<subSubSection id="C37F656AFFAFA204AAE8A2C33CBBFB0C" pageId="6" pageNumber="55" type="key">
<keyStep id="30912E44FFAFA204AAE8A2C33CBBFE4C" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A2C33CBBFE77" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A2C33CBBFE77" pageId="6" pageNumber="55">1a First dorsal fin spines IIIV, not extending rearwards beyond anterior end of the second dorsal fin when depressed; body with lateral midline blotches more or less evident; estuarine or marine............................................... 2</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A29D3CBBFE4C" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A29D3CBBFE4C" pageId="6" pageNumber="55">
1b First dorsal fin spines IIIV, extending rearwards beyond middle of the second dorsal fin when depressed; body with four narrow vertical dark striae; freshwater, Lake Sibaya,
<collectingCountry id="F3727671FFAFA204A8BFA2BB3A18FDB3" box="[704,831,490,512]" name="Mozambique" pageId="6" pageNumber="55">Mozambique</collectingCountry>
...........................................
<taxonomicName id="4C654D62FFAFA204AF34A2BB3CBBFE4C" ID-CoL="4XDK5" authorityName="Farquharson" authorityYear="1970" box="[1355,1436,490,511]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="sibayi">
<emphasis id="B911EAF3FFAFA204AF34A2BB3CBBFE4C" box="[1355,1436,490,511]" italics="true" pageId="6" pageNumber="55">S. sibayi</emphasis>
</taxonomicName>
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</paragraph>
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<keyStep id="30912E44FFAFA204AAE8A1563CBBFD8A" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A1563CBBFDAF" blockId="6.[151,1437,402,1216]" box="[151,1436,519,541]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A1563CBBFDAF" box="[151,1436,519,541]" pageId="6" pageNumber="55">2a Head length 31.435.5% of standard length, head width 20.524.6% of standard length.............................. 3</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A1753CBBFD8A" blockId="6.[151,1437,402,1216]" box="[151,1436,548,570]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A1753CBBFD8A" box="[151,1436,548,570]" pageId="6" pageNumber="55">2b Head length 20.130.0% of standard length, head width 11.615.5% of standard length.............................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E44FFAFA204AAE8A1133CBBFDC7" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A1133CBAFDE4" blockId="6.[151,1437,402,1216]" box="[151,1437,578,600]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A1133CBAFDE4" box="[151,1437,578,600]" pageId="6" pageNumber="55">
3a Anal fin I/12, opercle and predorsal area with scales,
<collectingCountry id="F3727671FFAFA204A89DA1133AA3FDEB" box="[738,900,578,600]" name="Marshall Islands" pageId="6" pageNumber="55">Marshall Islands</collectingCountry>
, Pacific Ocean.....................
<taxonomicName id="4C654D62FFAFA204AF6DA1133CBAFDE4" ID-CoL="6YQHG" authorityName="Randall" authorityYear="2008" box="[1298,1437,578,599]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="capitlineata">
<emphasis id="B911EAF3FFAFA204AF6DA1133CBAFDE4" box="[1298,1437,578,599]" italics="true" pageId="6" pageNumber="55">S. capitlineata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A10E3CBBFDC7" blockId="6.[151,1437,402,1216]" box="[151,1436,607,629]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A10E3CBBFDC7" box="[151,1436,607,629]" pageId="6" pageNumber="55">
3b Anal fin I/13, opercle and predorsal area naked,
<collectingCountry id="F3727671FFAFA204A8CDA10E3A1EFDC6" box="[690,825,607,629]" name="Qeshm Island" pageId="6" pageNumber="55">Qeshm Island</collectingCountry>
, Indian Ocean...............
<taxonomicName id="4C654D62FFAFA204AEFEA10E3C6DFDC7" authority="Kovačić &amp; Sadeghi &amp; Esmaeili, 2020" authorityName="Kovačić &amp; Sadeghi &amp; Esmaeili" authorityYear="2020" box="[1153,1354,607,628]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="ghazalae" status="sp. nov.">
<emphasis id="B911EAF3FFAFA204AEFEA10E3C6DFDC7" box="[1153,1354,607,628]" italics="true" pageId="6" pageNumber="55">Silhouettea ghazalae</emphasis>
</taxonomicName>
<emphasis id="B911EAF3FFAFA204AF2FA10E3CBBFDC7" bold="true" box="[1360,1436,607,628]" pageId="6" pageNumber="55">
<taxonomicNameLabel id="A2225788FFAFA204AF2FA10E3CBBFDC7" box="[1360,1436,607,628]" pageId="6" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<keyStep id="30912E44FFAFA204AAE8A12D3CBBFCB4" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A12D3CBAFD7E" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A12D3CBAFD7E" pageId="6" pageNumber="55">
4a Lateral midline with 9-10 short vertical dark marks, each within oblong or rhomboidal dark blotch; body scale pocket beneath scale with numerous fine melanophores in crescentic dark mark, the Northern Red Sea, introduced to Mediterranean..................................................................................................
<taxonomicName id="4C654D62FFAFA204AF4DA1E63CBAFD7E" ID-CoL="4XDJT" baseAuthorityName="Chabanaud" baseAuthorityYear="1933" box="[1330,1437,695,717]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="aegyptia">
<emphasis id="B911EAF3FFAFA204AF4DA1E63CBAFD7E" box="[1330,1437,695,717]" italics="true" pageId="6" pageNumber="55">S. aegyptia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A1853CBBFCB4" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A1853CBBFCB4" pageId="6" pageNumber="55">
4b Lateral midline with at least four to six oblong to triangular blotches more or less evident but lacking distinct vertical central mark (up to ten in
<taxonomicName id="4C654D62FFAFA204ABEBA1A338FAFCB4" ID-CoL="325V5" authorityName="Takagi" authorityYear="1957" box="[404,477,754,775]" class="Actinopterygii" family="Gobiidae" genus="Ctenogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="dotui">
<emphasis id="B911EAF3FFAFA204ABEBA1A338FAFCB4" box="[404,477,754,775]" italics="true" pageId="6" pageNumber="55">S. dotui</emphasis>
</taxonomicName>
); body scale pocket with crescentic mark beneath scale formed by fewer larger melanophores... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E44FFAFA204AAE8A05E3CBBFCF1" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A05E3CBBFC96" blockId="6.[151,1437,402,1216]" box="[151,1436,783,805]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A05E3CBBFC96" box="[151,1436,783,805]" pageId="6" pageNumber="55">5a Anal fin I/1314...................................................................................... 6</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A07D3CBBFCF1" blockId="6.[151,1437,402,1216]" box="[151,1436,812,834]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A07D3CBBFCF1" box="[151,1436,812,834]" pageId="6" pageNumber="55">5b Anal fin I/1112...................................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E44FFAFA204AAE8A0183CBAFC04" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A0183CBBFCCE" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A0183CBBFCCE" pageId="6" pageNumber="55">
6a Nape lacking conspicuous white spot; the first dorsal fin with anterior dark spot but no obvious posterior spot; pectoral fin 1415; Indian Ocean and Red Sea...............................................................
<taxonomicName id="4C654D62FFAFA204AF57A0363CBBFCCE" ID-CoL="4XDK3" authorityName="Smith" authorityYear="1959" box="[1320,1436,871,893]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="insinuans">
<emphasis id="B911EAF3FFAFA204AF57A0363CBBFCCE" box="[1320,1436,871,893]" italics="true" pageId="6" pageNumber="55">S. insinuans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A0D53CBAFC04" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A0D53CBAFC04" pageId="6" pageNumber="55">
6b Nape with conspicuous white spot; the first dorsal fin with posterior dark spot but no obvious anterior spot; pectoral fin 16;
<collectingCountry id="F3727671FFAFA204AA9DA0F33869FC0B" box="[226,334,930,952]" name="Philippines" pageId="6" pageNumber="55">Philippines</collectingCountry>
.............................................................................
<taxonomicName id="4C654D62FFAFA204AE8DA0F33CBAFC04" ID-CoL="325W6" authorityName="Herre" authorityYear="1934" box="[1266,1437,930,951]" class="Actinopterygii" family="Gobiidae" genus="Ctenogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="nuchipunctatus">
<emphasis id="B911EAF3FFAFA204AE8DA0F33CBAFC04" box="[1266,1437,930,951]" italics="true" pageId="6" pageNumber="55">S. nuchipunctatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E44FFAFA204AAE8A0EE3CBBFC42" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A0EE3CBBFC67" blockId="6.[151,1437,402,1216]" box="[151,1436,959,981]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A0EE3CBBFC67" box="[151,1436,959,981]" pageId="6" pageNumber="55">
7a Anal fin I/12;
<collectingCountry id="F3727671FFAFA204AB14A0EE3885FC67" box="[363,418,959,980]" name="Japan" pageId="6" pageNumber="55">Japan</collectingCountry>
and
<collectingCountry id="F3727671FFAFA204ABAFA0EE3B26FC66" box="[464,513,959,981]" name="India" pageId="6" pageNumber="55">India</collectingCountry>
........................................................................... 8
</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A08D3CBBFC42" blockId="6.[151,1437,402,1216]" box="[151,1436,988,1010]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A08D3CBBFC42" box="[151,1436,988,1010]" pageId="6" pageNumber="55">
7b Anal fin I/11; northern
<collectingCountry id="F3727671FFAFA204ABBFA08D3B3FFC41" box="[448,536,988,1010]" name="Australia" pageId="6" pageNumber="55">Australia</collectingCountry>
......................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E44FFAFA204AAE8A0AB3CBBFBFA" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A0AB3CBBFB9F" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A0AB3CBBFB9F" pageId="6" pageNumber="55">
8a Body densely pigmented with coarse reticulation of numerous crescentic scale pocket marks; up to ten small not very evident lateral midline markings small, median fins less than body height; breast scaled;
<collectingCountry id="F3727671FFAFA204A996A7463D07FB9F" box="[1001,1056,1047,1068]" name="Japan" pageId="6" pageNumber="55">Japan</collectingCountry>
.........................
<taxonomicName id="4C654D62FFAFA204AF2CA7463CBBFB9F" ID-CoL="325V5" authorityName="Takagi" authorityYear="1957" box="[1363,1436,1047,1068]" class="Actinopterygii" family="Gobiidae" genus="Ctenogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="dotui">
<emphasis id="B911EAF3FFAFA204AF2CA7463CBBFB9F" box="[1363,1436,1047,1068]" italics="true" pageId="6" pageNumber="55">S. dotui</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A7653CBBFBFA" blockId="6.[151,1437,402,1216]" box="[151,1436,1076,1098]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A7653CBBFBFA" box="[151,1436,1076,1098]" pageId="6" pageNumber="55">
8b Body with six oval blotches along lateral midline; median fins higher than body, breast naked;
<collectingCountry id="F3727671FFAFA204AE0FA7653D86FBF9" box="[1136,1185,1076,1098]" name="India" pageId="6" pageNumber="55">India</collectingCountry>
.............
<taxonomicName id="4C654D62FFAFA204AF37A7643CBBFBFA" ID-CoL="4XDJZ" authorityName="Rao" authorityYear="1971" box="[1352,1436,1076,1098]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="indica">
<emphasis id="B911EAF3FFAFA204AF37A7643CBBFBFA" box="[1352,1436,1076,1098]" italics="true" pageId="6" pageNumber="55">S. indica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E44FFAFA204AAE8A7033CBBFB0C" pageId="6" pageNumber="55">
<paragraph id="8BDA36E1FFAFA204AAE8A7033CBBFB37" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A7033CBBFB37" pageId="6" pageNumber="55">
9a Axilla pigmented; lateral blotches small; nape with transverse band of fine melanophores narrower than pupil; branchiostegal membrane pale................................................................................
<taxonomicName id="4C654D62FFAFA204AF47A73E3CBBFB37" ID-CoL="4XDJX" authorityName="Larson &amp; Miller" authorityYear="1986" box="[1336,1436,1135,1156]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="evanida">
<emphasis id="B911EAF3FFAFA204AF47A73E3CBBFB37" box="[1336,1436,1135,1156]" italics="true" pageId="6" pageNumber="55">S. evanida</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDA36E1FFAFA204AAE8A7DD3CBBFB0C" blockId="6.[151,1437,402,1216]" pageId="6" pageNumber="55">
<keyLead id="309495D4FFAFA204AAE8A7DD3CBBFB0C" pageId="6" pageNumber="55">
9b Axilla pale, lateral blotches large; nape with transverse band of fine melanophores wider than pupil; branchiostegal membrane densely pigmented..............................................................................
<taxonomicName id="4C654D62FFAFA204AF38A7FB3CBBFB0C" ID-CoL="4XDJY" authorityName="Larson &amp; Miller" authorityYear="1986" box="[1351,1436,1194,1215]" class="Actinopterygii" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="hoesei">
<emphasis id="B911EAF3FFAFA204AF38A7FB3CBBFB0C" box="[1351,1436,1194,1215]" italics="true" pageId="6" pageNumber="55">S. hoesei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>