treatments-xml/data/03/CC/87/03CC87B17643FF8B9293FA48FC5DD1F1.xml

<document id="767196334CB93F91CDA8437D7D3A9827" ID-DOI="10.5281/zenodo.275727" ID-GBIF-Dataset="1e79f840-6ced-48b5-ab20-94992c6f0a88" ID-ISSN="1175-5326" ID-Zenodo-Dep="275727" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460160971924" checkinUser="plazi" docAuthor="West, Rick C. &amp; Nunn, Steven C." docDate="2010" docId="03CC87B17643FF8B9293FA48FC5DD1F1" docLanguage="en" docName="zt02362p043.pdf" docOrigin="Zootaxa 2362" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lyrognathus robustus Smith 1988" docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="FFF5FFC9765DFFA89204FFFAFF92D456" masterDocTitle="A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae" masterLastPageNumber="43" masterPageNumber="1" pageNumber="31" updateTime="1698235793967" updateUser="plazi">
<mods:mods id="358A4A82A7DCAA2617F38D9926995A6C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0FC7DBD45C8BBF39FC336F768E397A31">
<mods:title id="EAEE4B7E3ECA0D5DA8FA8D93191F1850">A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae</mods:title>
</mods:titleInfo>
<mods:name id="87529B507330547C22747CBEDB449699" type="personal">
<mods:role id="AA868A380529F90F82E965FE786CEE86">
<mods:roleTerm id="E495A20DE34046FB241C4D0F37590268">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0BC54E5331CF1D3B70FC1C539449B240">West, Rick C.</mods:namePart>
</mods:name>
<mods:name id="6BD01C0BC7C7DDA6D4734575771DA3CC" type="personal">
<mods:role id="6F1972DDB479C6935735BA22B0B25B85">
<mods:roleTerm id="39521031E0892E1A03E13C978A227A9B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="44864D12376E712E15D78BF2001AB423">Nunn, Steven C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="929BBFD54A3CCAE84C92BC76BB6651FF">text</mods:typeOfResource>
<mods:relatedItem id="6E961AF2078BB2A07A6F2030F763F04A" type="host">
<mods:titleInfo id="0A5A428FF7B6D1C48B2A80BC4E056C3C">
<mods:title id="7E9A729267D21C7CCB790F11D08AE1CF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D435F750110D1A9330935A86EB20F9A7">
<mods:date id="C7406B2D155EF831BB8ED48A5107B705">2010</mods:date>
<mods:detail id="E56BA3B49DAB885BD9D5E2B795F96BB4" type="volume">
<mods:number id="8283790500D8FD4ED12E453E28E4764A">2362</mods:number>
</mods:detail>
<mods:extent id="BF4A5AA51E5DBD1CDB0CA3CE1B3FF52F" unit="page">
<mods:start id="87E335018259632F901E67787F82A730">1</mods:start>
<mods:end id="00798339FC48B18DAA6BA43C308FC89E">43</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4FFD890A8CBFEF08F154B9278314DA48">journal article</mods:classification>
<mods:identifier id="1A0F98D6F1E2F1B858597A48B92DE5BA" type="DOI">10.5281/zenodo.275727</mods:identifier>
<mods:identifier id="426FFA38A57ADF57F67E2BDB0FD60098" type="GBIF-Dataset">1e79f840-6ced-48b5-ab20-94992c6f0a88</mods:identifier>
<mods:identifier id="C58C7D9CD2D37F7D819E526AE4F08714" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8B1409BBA5CDD5B766185FCBC3EBC3AF" type="Zenodo-Dep">275727</mods:identifier>
</mods:mods>
<treatment id="03CC87B17643FF8B9293FA48FC5DD1F1" ID-DOI="http://doi.org/10.5281/zenodo.6210158" ID-GBIF-Taxon="127654813" ID-Zenodo-Dep="6210158" LSID="urn:lsid:plazi:treatment:03CC87B17643FF8B9293FA48FC5DD1F1" httpUri="http://treatment.plazi.org/id/03CC87B17643FF8B9293FA48FC5DD1F1" lastPageId="35" lastPageNumber="36" pageId="30" pageNumber="31">
<subSubSection id="C37F652C7643FFB69293FA48FEE6D1B9" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BDA36A77643FFB69293FA48FDA6D19A" blockId="30.[151,564,1458,1519]" box="[151,564,1458,1484]" pageId="30" pageNumber="31">
<heading id="D09281CB7643FFB69293FA48FDA6D19A" bold="true" box="[151,564,1458,1484]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<taxonomicName id="4C654D247643FFB69293FA48FDA6D19A" ID-CoL="3WTQQ" authority="Smith 1988" authorityName="Smith" authorityYear="1988" box="[151,564,1458,1484]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB57643FFB69293FA48FDA6D19A" bold="true" box="[151,564,1458,1484]" pageId="30" pageNumber="31">
<emphasis id="B911EAB57643FFB69293FA48FE32D19A" bold="true" box="[151,416,1458,1484]" italics="true" pageId="30" pageNumber="31">Lyrognathus robustus</emphasis>
<bibRefCitation id="EFF44B567643FFB693A3FA48FDA6D19A" author="Smith" box="[423,564,1458,1484]" pageId="30" pageNumber="43" refString="Smith, A. M. (1988) Lyrognathus robustus, a new species of theraphosid spider from Malaysia. British Tarantula Society Journal, 4 (2), 15 - 19." type="journal article" year="1988">Smith 1988</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDA36A77643FFB69293FA2FFEE6D1B9" blockId="30.[151,564,1458,1519]" box="[151,372,1493,1519]" pageId="30" pageNumber="31">
(
<figureCitation id="135E2A227643FFB6929BFA2FFEA6D1B9" box="[159,308,1493,1519]" captionStart-0="FIGURE 91" captionStart-1="FIGURES 92 – 101" captionStart-2="FIGURES 102 – 112" captionStartId-0="31.[151,255,1152,1176]" captionStartId-1="32.[151,270,1604,1628]" captionStartId-2="33.[151,271,1848,1872]" captionTargetBox-0="[151,1435,343,1130]" captionTargetBox-1="[160,1422,253,1586]" captionTargetBox-2="[151,1426,375,1821]" captionTargetId-0="figure@31.[151,1436,343,1130]" captionTargetId-1="figure@32.[151,1436,253,1601]" captionTargetId-2="figure@33.[151,1436,354,1833]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionText-0="FIGURE 91. Lyrognathus robustus, adult female habitus, dorsolateral view. Photo: R. C. West." captionText-1="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." captionText-2="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri-0="https://zenodo.org/record/275747/files/figure.png" httpUri-1="https://zenodo.org/record/275748/files/figure.png" httpUri-2="https://zenodo.org/record/275749/files/figure.png" pageId="30" pageNumber="31">Figs 91–112</figureCitation>
, 
<figureCitation id="135E2A227643FFB69344FA2FFEFED1B9" box="[320,364,1493,1519]" captionStart="FIGURE 114" captionStartId="38.[151,255,1297,1321]" captionTargetBox="[151,1436,194,1276]" captionTargetId="figure@38.[151,1436,194,1276]" captionTargetPageId="38" captionText="FIGURE 114. Records of Lyrognathus species in Southeast Asia. C. Senescall" httpUri="https://zenodo.org/record/275751/files/figure.png" pageId="30" pageNumber="31">114</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F652C7643FFB69293F9DAFCB5D362" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8BDA36A77643FFB69293F9DAFCCFD261" blockId="30.[151,1033,1568,1623]" box="[151,861,1568,1591]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB7511897643FFB69293F9DAFCCFD261" box="[151,861,1568,1591]" pageId="30" pageNumber="31">
<taxonomicName id="4C654D247643FFB69293F9DAFDBAD261" ID-CoL="3WTQQ" authority="Smith 1988: 15" authorityName="Smith" authorityPageNumber="15" authorityYear="1988" box="[151,552,1568,1591]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB57643FFB69293F9DAFEECD261" box="[151,382,1568,1591]" italics="true" pageId="30" pageNumber="31">Lyrognathus robustus</emphasis>
<treatmentCitation id="0AC410B67643FFB69380F9DAFDBAD261" author="Smith" box="[388,552,1568,1591]" page="15" pageId="30" pageNumber="31" year="1988">
<bibRefCitation id="EFF44B567643FFB69380F9DAFD91D261" author="Smith" box="[388,515,1568,1591]" pageId="30" pageNumber="43" refString="Smith, A. M. (1988) Lyrognathus robustus, a new species of theraphosid spider from Malaysia. British Tarantula Society Journal, 4 (2), 15 - 19." type="journal article" year="1988">Smith 1988</bibRefCitation>
: 15
</treatmentCitation>
</taxonomicName>
–19, figs1–9; 
<bibRefCitation id="EFF44B567643FFB690BFF9DAFCCBD261" author="Platnick" box="[699,857,1568,1591]" pageId="30" pageNumber="43" refString="Platnick, N. I. (2009) The world spider catalog, version 9.5. American Museum of Natural History, available from http: // research. amnh. org / entomology / spiders / catalog / index. html (accessed 30 June 2009)" type="journal article" year="2009">Platnick, 2009</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDA36A77643FFB69293F9BAFB9BD201" blockId="30.[151,1033,1568,1623]" box="[151,1033,1600,1623]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB7511897643FFB69293F9BAFB9BD201" box="[151,1033,1600,1623]" pageId="30" pageNumber="31">
<taxonomicName id="4C654D247643FFB69293F9BAFD96D201" ID-CoL="3WTQN" authority="West 1991: 615" authorityName="West" authorityPageNumber="615" authorityYear="1991" box="[151,516,1600,1623]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="liewi">
<emphasis id="B911EAB57643FFB69293F9BAFEC5D201" box="[151,343,1600,1623]" italics="true" pageId="30" pageNumber="31">Lyrognathus liewi</emphasis>
<treatmentCitation id="0AC410B67643FFB69359F9BAFD96D201" author="West" box="[349,516,1600,1623]" page="615" pageId="30" pageNumber="31" year="1991">
<bibRefCitation id="EFF44B567643FFB69359F9BAFE43D201" author="West" box="[349,465,1600,1623]" pageId="30" pageNumber="43" refString="West, R. C. (1991) A new theraphosid spider from the Cameron Highlands, Malaysia. Memoirs of the Queensland Museum, 30, 615 - 619." type="journal article" year="1991">West 1991</bibRefCitation>
: 615
</treatmentCitation>
</taxonomicName>
–619, figs 1–11. Schmidt &amp; von 
<treatmentCitation id="0AC410B67643FFB69164F9BAFB97D201" author="Wirth" box="[864,1029,1600,1623]" page="10" pageId="30" pageNumber="31" year="1992">
<bibRefCitation id="EFF44B567643FFB69164F9BAFC4FD201" author="Wirth" box="[864,989,1600,1623]" pageId="30" pageNumber="43" refString="Schmidt, G. &amp; von Wirth, V. (1992) Neue veroffentlichungen uber vogelspinnen. Arachnologischer Anzeiger, 3 (4), 10." type="journal article" year="1992">Wirth 1992</bibRefCitation>
: 10
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDA36A77643FFB69293F97FFCB5D362" blockId="30.[151,1437,1669,1994]" pageId="30" pageNumber="31">
<emphasis id="B911EAB57643FFB69293F97FFF64D2C9" bold="true" box="[151,246,1669,1695]" pageId="30" pageNumber="31">
<typeStatus id="54DE88057643FFB69293F97FFF78D2C9" box="[151,234,1669,1695]" pageId="30" pageNumber="31">Types</typeStatus>
:
</emphasis>
<taxonomicName id="4C654D247643FFB69307F97FFD4CD2C9" ID-CoL="3WTQQ" authority="Smith 1988" authorityName="Smith" authorityYear="1988" box="[259,734,1669,1695]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB57643FFB69307F97FFDA0D2C8" box="[259,562,1669,1694]" italics="true" pageId="30" pageNumber="31">Lyrognathus robustus</emphasis>
<bibRefCitation id="EFF44B567643FFB6903AF97FFD4CD2C9" author="Smith" box="[574,734,1669,1695]" pageId="30" pageNumber="43" refString="Smith, A. M. (1988) Lyrognathus robustus, a new species of theraphosid spider from Malaysia. British Tarantula Society Journal, 4 (2), 15 - 19." type="journal article" year="1988">Smith 1988</bibRefCitation>
</taxonomicName>
, female 
<typeStatus id="54DE88057643FFB6915AF97FFC44D2C9" box="[862,982,1669,1695]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
, 1988.2.6, 
<collectingCountry id="F37276377643FFB69679F97FFB6AD2C9" box="[1149,1272,1669,1695]" name="Cameroon" pageId="30" pageNumber="31">Cameron</collectingCountry>
Highlands, 
<collectingCountry id="F37276377643FFB69293F950FE9BD292" box="[151,265,1706,1732]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
, (unknown collector and date), NHM.–images (R. Raven, pers. comm.)–examined; 
<emphasis id="B911EAB57643FFB69701F951FECCD2BF" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="4C654D247643FFB69701F951FECBD2BF" ID-CoL="3WTQN" authority="West 1991" authorityName="West" authorityYear="1991" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="liewi">
Lyrognathus liewi 
<bibRefCitation id="EFF44B567643FFB692DDF92AFECBD2BF" author="West" box="[217,345,1744,1769]" pageId="30" pageNumber="43" refString="West, R. C. (1991) A new theraphosid spider from the Cameron Highlands, Malaysia. Memoirs of the Queensland Museum, 30, 615 - 619." type="journal article" year="1991">West 1991</bibRefCitation>
</taxonomicName>
,
</emphasis>
female 
<typeStatus id="54DE88057643FFB693C4F92AFDBFD2BC" box="[448,557,1744,1770]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D63FD2E7643FFB6903EF92AFD30D2BC" box="[570,674,1744,1770]" pageId="30" pageNumber="31" type="female">2 female</specimenCount>
<typeStatus id="54DE88057643FFB690AFF92AFCB0D2BC" box="[683,802,1744,1770]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
, 
<accessionNumber id="9436AB447643FFB6912BF92AFC1CD2BC" box="[815,910,1744,1770]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S15460" pageId="30" pageNumber="31" type="EnaNcbi">S15460</accessionNumber>
, 
<accessionNumber id="9436AB447643FFB6919FF92AFC65D2BC" box="[923,1015,1744,1770]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S15461" pageId="30" pageNumber="31" type="EnaNcbi">S15461</accessionNumber>
and 
<accessionNumber id="9436AB447643FFB69630F92AFB01D2BC" box="[1076,1171,1744,1770]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S15462" pageId="30" pageNumber="31" type="EnaNcbi">S15462</accessionNumber>
, respectively, 
<quantity id="4C9D9B427643FFB69744F92AFA0ED2BC" box="[1344,1436,1744,1770]" metricMagnitude="4" metricUnit="m" metricValue="3.8624256" pageId="30" pageNumber="31" unit="mi" value="24.0">24 mile</quantity>
road marker, 
<collectingCountry id="F37276377643FFB69333F90FFE36D359" box="[311,420,1781,1807]" name="Cameroon" pageId="30" pageNumber="31">Cameron</collectingCountry>
Highlands (
<geoCoordinate id="EE5150607643FFB6903CF90FFD05D359" box="[568,663,1781,1807]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="925" value="4.6833334">4°41' N</geoCoordinate>
, 
<geoCoordinate id="EE5150607643FFB690A0F90FFC8FD359" box="[676,797,1781,1807]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="101.86667">101°52' E</geoCoordinate>
), Pahang, West 
<collectingCountry id="F37276377643FFB691DAF90FFBDCD359" box="[990,1102,1781,1807]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
, col. K. C. Liew, 
<date id="FFDB10677643FFB69726F90FFF46D362" pageId="30" pageNumber="31" value="1986-10" valueMax="1986-11-31" valueMin="1986-10-00">Oct.–Nov. 1986</date>
, QM.–images (R. Raven, pers. comm.)–examined.
</paragraph>
</subSubSection>
<subSubSection id="C37F652C7643FFB692C2F8C5FA0ED39C" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BDA36A77643FFB692C2F8C5FA0ED39C" blockId="30.[151,1437,1669,1994]" pageId="30" pageNumber="31">
<emphasis id="B911EAB57643FFB692C2F8C5FE01D30F" bold="true" box="[198,403,1855,1881]" pageId="30" pageNumber="31">Other material:</emphasis>
<specimenCount id="9D63FD2E7643FFB693A6F8BAFD8CD30C" box="[418,542,1856,1882]" pageId="30" pageNumber="31" type="female">3 females</specimenCount>
, 
<specimenCount id="9D63FD2E7643FFB69035F8BAFD19D30C" box="[561,651,1856,1882]" pageId="30" pageNumber="31" type="male">1 male</specimenCount>
, S74339, S74341, S74342 and S74340, respectively, 
<collectingCountry id="F37276377643FFB69734F8BAFA09D30C" box="[1328,1435,1856,1882]" name="Cameroon" pageId="30" pageNumber="31">Cameron</collectingCountry>
Highlands, Pahang, West 
<collectingCountry id="F37276377643FFB693CEF89FFDA8D329" box="[458,570,1893,1919]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
, QM–examined; 
<specimenCount id="9D63FD2E7643FFB6910DF89FFCCFD329" box="[777,861,1893,1919]" pageId="30" pageNumber="31" type="male">1 male</specimenCount>
, 
<accessionNumber id="9436AB447643FFB6916DF89FFC55D329" box="[873,967,1893,1919]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88073" pageId="30" pageNumber="31" type="EnaNcbi">S88073</accessionNumber>
, Maxwell Hill (= Bukit Larut, 4°52΄ N, 100°48΄ E), Perak, West 
<collectingCountry id="F37276377643FFB693BEF870FDBBD3F2" box="[442,553,1930,1956]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
, QM–examined; 
<specimenCount id="9D63FD2E7643FFB690FAF870FCE0D3F2" box="[766,882,1930,1956]" pageId="30" pageNumber="31" type="female">2 females</specimenCount>
, 
<collectionCode id="ED74AE627643FFB6917BF870FC2AD3F2" box="[895,952,1930,1956]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="30" pageNumber="31">ZRC</collectionCode>
<collectionCode id="ED74AE627643FFB691C4F870FB91D3F2" box="[960,1027,1930,1956]" pageId="30" pageNumber="31">ARA</collectionCode>
.747, Taiping (
<geoCoordinate id="EE5150607643FFB696A8F870FA9BD3F2" box="[1196,1289,1930,1956]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="925" value="4.85">4°51' N</geoCoordinate>
, 
<geoCoordinate id="EE5150607643FFB69712F870FA1CD3F2" box="[1302,1422,1930,1956]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="100.8">100°48' E</geoCoordinate>
), Perak, West 
<collectingCountry id="F37276377643FFB6932DF84AFE05D39C" box="[297,407,1968,1994]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
, col. H. C. Abraham, 
<date id="FFDB10677643FFB69093F84AFCB8D39C" box="[663,810,1968,1994]" pageId="30" pageNumber="31" value="1925-11-01">1 Nov. 1925</date>
, 
<collectionCode id="ED74AE627643FFB69132F84AFC1FD39C" box="[822,909,1968,1994]" httpUri="http://grbio.org/cool/phd9-bbbi" name="Raffles Museum of Biodiversity Research" pageId="30" pageNumber="31">RMBR</collectionCode>
.–images (D. Court, pers. comm.)–examined.
</paragraph>
</subSubSection>
<subSubSection id="C37F652C7642FFB792C2FF6DFEECD574" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BDA36A77642FFB792C2FF6DFEECD574" blockId="31.[151,1436,151,290]" pageId="31" pageNumber="32">
<emphasis id="B911EAB57642FFB792C2FF6DFEDAD4E7" bold="true" box="[198,328,151,177]" pageId="31" pageNumber="32">Diagnosis:</emphasis>
Differs from other 
<taxonomicName id="4C654D247642FFB79040FF62FD45D4E7" box="[580,727,152,177]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB57642FFB79040FF62FD45D4E7" box="[580,727,152,177]" italics="true" pageId="31" pageNumber="32">Lyrognathus</emphasis>
</taxonomicName>
species in the basally incrassate metatarsi IV that tapers distally (
<figureCitation id="135E2A227642FFB79305FF47FECED481" box="[257,348,189,215]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Fig. 99</figureCitation>
). Also differs from all other species in having uniquely divided scopula on metatarsi IV, a third claw on tarsi IV (
<figureCitation id="135E2A227642FFB793B6FF18FD98D4AA" box="[434,522,226,252]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Fig. 98</figureCitation>
and see: Character discussion) and bilobular spermathecae without apical swelling (
<figureCitation id="135E2A227642FFB7930EFEF2FEE2D574" box="[266,368,264,290]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Fig. 101</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1A662F7642FFB79293FB7AFB03D0C1" box="[151,1169,1152,1176]" httpUri="https://zenodo.org/record/275747/files/figure.png" pageId="31" pageNumber="32" targetBox="[151,1435,343,1130]" targetPageId="31">
<paragraph id="8BDA36A77642FFB79293FB7AFB03D0C1" blockId="31.[151,1169,1152,1176]" box="[151,1169,1152,1176]" pageId="31" pageNumber="32">
<emphasis id="B911EAB57642FFB79293FB7AFEB4D0CE" bold="true" box="[151,294,1152,1176]" pageId="31" pageNumber="32">FIGURE 91.</emphasis>
<taxonomicName id="4C654D247642FFB79329FB7AFD86D0C1" box="[301,532,1152,1175]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB57642FFB79329FB7AFD86D0C1" box="[301,532,1152,1175]" italics="true" pageId="31" pageNumber="32">Lyrognathus robustus</emphasis>
</taxonomicName>
, adult female habitus, dorsolateral view. Photo: R. C. West.
</paragraph>
</caption>
<subSubSection id="C37F652C7642FF8A92C0FB10FB11D3E4" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32" type="description">
<paragraph id="8BDA36A77642FFB792C0FB10FB0FD153" blockId="31.[151,1437,1258,2032]" box="[196,1181,1258,1285]" pageId="31" pageNumber="32">
<emphasis id="B911EAB57642FFB792C0FB10FECCD152" bold="true" box="[196,350,1258,1284]" pageId="31" pageNumber="32">Description:</emphasis>
female QM 
<accessionNumber id="9436AB447642FFB793F7FB11FDDFD153" box="[499,589,1259,1285]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S15641" pageId="31" pageNumber="32" type="EnaNcbi">S15641</accessionNumber>
(unless otherwise stated) with body length: 38.14.
</paragraph>
<paragraph id="8BDA36A77642FFB792C0FAEAFC2ED17C" blockId="31.[151,1437,1258,2032]" box="[196,956,1296,1322]" pageId="31" pageNumber="32">Color (in alcohol): reddish brown with darker femora on all legs.</paragraph>
<paragraph id="8BDA36A77642FFB792C0FACCFA0ED106" blockId="31.[151,1437,1258,2032]" box="[196,1436,1334,1360]" pageId="31" pageNumber="32">Carapace: length 16.7, width 12.8 (width across anterior edge 9.6). Fovea 2.3, procurved, deep, very wide.</paragraph>
<paragraph id="8BDA36A77642FFB792C0FAA1FDEAD1CC" blockId="31.[151,1437,1258,2032]" pageId="31" pageNumber="32">
Eyes (
<figureCitation id="135E2A227642FFB7930BFAA1FEF8D123" box="[271,362,1371,1397]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Fig. 93</figureCitation>
): ocular tubercle; length 1.85, width 2.65. Anterior row transverse. Posterior row slightly recurved. Sizes: AME, ALE, PLE, PME.
</paragraph>
<paragraph id="8BDA36A77642FFB792C0FA5CFAF3D203" blockId="31.[151,1437,1258,2032]" pageId="31" pageNumber="32">
Chelicerae (
<figureCitation id="135E2A227642FFB7935CFA5CFE79D196" box="[344,491,1446,1472]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Figs 95, 96</figureCitation>
): length 6.05, width at base of each lobe 4.25. Ectal lyrate region a series of strikers (&gt;80), up to 7 disordered horizontal rows (
<figureCitation id="135E2A227642FFB790FAFA31FCCBD1B3" box="[766,857,1483,1509]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Fig. 96</figureCitation>
). Strongest and longest strikers on lowest rows. Each striker needle-form with some filiform ends. Teeth: 14 (with&gt;2 small teeth proximally) and&gt;60 basomesal teeth. Intercheliceral pegs (&gt;7 larger,
<emphasis id="B911EAB57642FFB790D7F9ECFD75D279" box="[723,743,1558,1583]" italics="true" pageId="31" pageNumber="32">&gt;</emphasis>
20 smaller) in tight cluster on basodorsal surface (
<figureCitation id="135E2A227642FFB79732F9ECFA1CD266" box="[1334,1422,1558,1584]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Fig. 95</figureCitation>
). On retrolateral surface basomedially, with small cluster of short strong spines (&gt;3) point distad (
<figureCitation id="135E2A227642FFB796F8F9C1FAC6D203" box="[1276,1364,1595,1621]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Fig. 96</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A77642FFB792C0F99AFCB3D30C" blockId="31.[151,1437,1258,2032]" pageId="31" pageNumber="32">
Maxillae (
<figureCitation id="135E2A227642FFB79338F99AFE54D22C" box="[316,454,1632,1658]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Figs 94, 97</figureCitation>
): prolaterally plano-convex. Anterior lobe well pronounced. Many cuspules (&gt;320) on inner basoventral surface. Diagonally of anterior lobe, a basal projection with long stout setae. 
<taxonomicName id="4C654D247642FFB79721F97CFAC9D2F6" box="[1317,1371,1670,1696]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="42" phylum="Arthropoda" rank="species" species="lyra">Lyra</taxonomicName>
(
<figureCitation id="135E2A227642FFB79768F97CFF2BD293" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="31" pageNumber="32">Fig. 97</figureCitation>
): many bacilliform rods (&gt;280) form dense, ovoid patch on medial prolateral surface, lowest row with up to&gt;40 bacillae, longest rods finish center of lowest row. Rods distally paddle-shaped with long shafts, some with distal blades. Truncate posteriorly. At widest point, lyrate patch up to 10 rows deep with smallest rods dorsally. Immediately above suture, 30 small spines on anterior margin. Labium: length 2.55, width 3.67. Many small cuspules (&gt;600) along anterior 1/3 surface.
</paragraph>
<paragraph id="8BDA36A77642FF8892C0F89CFD14D481" blockId="31.[151,1437,1258,2032]" lastBlockId="32.[151,1436,152,215]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">Sternum: length 7.65, width 6.9. Slightly wider posteriorly than anteriorly, with even cover of short setae, interspersed with longer darker setae. Lateral and posterior points acuminate with series of short stout setae cover posterior points. Sigilla: 3 pairs; posterior pair, large, 3.2 lengths apart, 1.85 of their length from margin. Median pair 1/2 size of posterior, similar shape, border margin. Anterior pair smallest, somewhat obscured, border margin. Labio-sternal sigilla largest, separated by their own diameter. Some strong spiniform setae around posterior edges, border cox. III/IV.</paragraph>
<caption id="DF1A662F767DFF889293F9BEFE87D34D" httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="32" pageNumber="33" targetBox="[160,1422,253,1586]" targetPageId="32">
<paragraph id="8BDA36A7767DFF889293F9BEFE87D34D" blockId="32.[151,1437,1604,1819]" pageId="32" pageNumber="33">
<emphasis id="B911EAB5767DFF889293F9BEFEFFD20A" bold="true" box="[151,365,1604,1628]" pageId="32" pageNumber="33">FIGURES 92–101.</emphasis>
<taxonomicName id="4C654D24767DFF889370F9BEFDA4D20D" box="[372,566,1604,1627]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="liewi">
<emphasis id="B911EAB5767DFF889370F9BEFDA4D20D" box="[372,566,1604,1627]" italics="true" pageId="32" pageNumber="33">Lyrognathus liewi</emphasis>
</taxonomicName>
(jun. syn. 
<taxonomicName id="4C654D24767DFF8890A8F9BEFCB7D20D" box="[684,805,1604,1627]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767DFF8890A8F9BEFCB7D20D" box="[684,805,1604,1627]" italics="true" pageId="32" pageNumber="33">L. robustus</emphasis>
</taxonomicName>
), paratype female QM S19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing 
<taxonomicName id="4C654D24767DFF88909EF95EFD5AD2ED" box="[666,712,1700,1723]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10mm for 92, 98, 99; 5mm for 100; 2mm for 93–97; 1mm for 101. Illustrations: S. C. Nunn.
</paragraph>
</caption>
<paragraph id="8BDA36A7767DFF8892C0F8BEFA0ED3A2" blockId="32.[151,1436,1860,2036]" pageId="32" pageNumber="33">
Leg setation: retrolateral surfaces of pat., tib. and met. IV with distinct thick short brushes of penicillate setae. Legs: formula; (length only) IV, I, II, III. Leg 
<collectingCountry id="F3727637767DFF889101F890FCBED3D2" box="[773,812,1898,1924]" name="Russia" pageId="32" pageNumber="33">RF</collectingCountry>
~82.67. Leg lengths (fem., pat., tib., met., tar., total): 
<typeStatus id="54DE8805767DFF889293F875FF6ED3FF" box="[151,252,1935,1961]" pageId="32" pageNumber="33" type="paratype">paratype</typeStatus>
<accessionNumber id="9436AB44767DFF889306F875FEF2D3FF" box="[258,352,1935,1961]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S15460" pageId="32" pageNumber="33" type="EnaNcbi">S15460</accessionNumber>
: palp: 9.08, 5.76, 6.49, 0.00, 5.94, 27.18. I: 14.02, 9.02, 9.9, 8.42, 4.95, 46.31. II: 11.72, 7.87, 7.98, 7.7, 4.57, 39.84. III: 10.78, 6.99, 6.77, 8.91, 4.4, 37.85. IV: 14.69, 8.8, 12.21, 15.4, 5.5, 56.6. 
<accessionNumber id="9436AB44767DFF88973DF84EFA0BD398" box="[1337,1433,1972,1998]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S15460" pageId="32" pageNumber="33" type="EnaNcbi">S15460</accessionNumber>
: palp: 9.15, 5.65, 5.85, 0.00, 5.9, 26.55. I: 11.95, 7.8, 9.3, 7.35, 5.35, 41.75. II: 10.2, 6.85, 6.45, 6.7, 5.0, 35.2.
</paragraph>
<paragraph id="8BDA36A7767CFF899293FF62FEE9D4AA" blockId="33.[151,1436,152,327]" pageId="33" pageNumber="34">
III: 8.95, 5.85, 5.6, 7.65, 5.2, 33.25. IV: 13.25, 7.1, 11.25, 12.9, 6.0, 50.5. Tib. IV incrassate. Met. IV basally incrassate, tapers distally. Tar. IV with transverse suture at distal 1/3 of tarsus, most obvious ventrally and laterally (
<figureCitation id="135E2A22767CFF899303FF18FEFCD4AA" box="[263,366,226,252]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="33" pageNumber="34">Fig. 100</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A7767CFF8992C2FEF2FB95D511" blockId="33.[151,1436,152,327]" pageId="33" pageNumber="34">
Scopula: on met./tar. I–IV. Undivided on met./tar. I–III. Met. and tar. IV divided by band of extremely strong spiniform setae with scopula only on apical 1/3 of met. IV (
<figureCitation id="135E2A22767CFF8991A6FED7FC68D511" box="[930,1018,301,327]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="33" pageNumber="34">Fig. 98</figureCitation>
).
</paragraph>
<caption id="DF1A662F767CFF899293F8C2FA12D3B9" httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="33" pageNumber="34" targetBox="[151,1426,375,1821]" targetPageId="33">
<paragraph id="8BDA36A7767CFF899293F8C2FA12D3B9" blockId="33.[151,1437,1848,2031]" pageId="33" pageNumber="34">
<emphasis id="B911EAB5767CFF899293F8C2FEE8D306" bold="true" box="[151,378,1848,1872]" pageId="33" pageNumber="34">FIGURES 102–112.</emphasis>
<taxonomicName id="4C654D24767CFF899386F8C2FDFED319" box="[386,620,1848,1871]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767CFF899386F8C2FDFED319" box="[386,620,1848,1871]" italics="true" pageId="33" pageNumber="34">Lyrognathus robustus</emphasis>
</taxonomicName>
male QM S74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing 
<taxonomicName id="4C654D24767CFF89976EF882FA0AD3D9" box="[1386,1432,1912,1935]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20mm for 102; 10mm for 108; 5mm for 109; 2mm for 104, 107, 110, 111; 1mm for 103, 105, 106; 0.5mm for 112. Photo &amp; illustrations: S. C. Nunn.
</paragraph>
</caption>
<paragraph id="8BDA36A7767FFF8A92C0FF62FC20D5E1" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Coxae: many short black thorns prolatero-dorsally, no thorns retrolaterally I–IV; easily seen dorsally. I longest, 
<emphasis id="B911EAB5767FFF8A9304FF47FEB6D480" box="[256,292,189,214]" italics="true" pageId="34" pageNumber="35">ca.</emphasis>
1.8 times length of II. IV widest, 1.3 times longer than III, basally rectangular with rounded corners; with ventral thorns prolaterally on I–IV. I–III ventrally with many long thick blunt setae, pallid, some short black setae intermixed. IV with mixture of long thick blunt setae, pallid intermixed with shorter thin pallid setae. Ventral surface of I–IV gently sloping anteriorly. Retrolateral setation: I–III with median narrow light brush, IV completely covered in light setae. I–IV retrolaterally lack ventral ledge, ventrally with uniform setation. Ventral measurements: I length 7.2, width 3.1; II 6.1, 3.1; III 5.8, 2.9; IV 6.7, 4.4. Trochantera (ventral): I length 4.0, width 3.8; II 3.4, 3.1; III 2.8, 2.7; IV 4.0, 3.9.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FE38FB43D624" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Trichobothria: tarsi: on all, basal filiform field slightly wider than clavate field, merges evenly. Clavates on tar. I only in distal 1/2, long filiforms only in basal 1/2, shorter filiforms intermixed with clavates distally. Clavate extent on II–IV 
<emphasis id="B911EAB5767FFF8A93B3FDF7FE46D670" box="[439,468,525,550]" italics="true" pageId="34" pageNumber="35">cf.</emphasis>
I, only in distal 1/2. Shorter filiforms for length. Short epitrichobothrial field on I shorter than clavates, increasing in height proximally. Tarsal organ visible (coniform, high domed), positioned immediately behind prolateral paired tarsal claw, but obscured by border setae, on legs I–IV.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FD87FD73D6B4" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Spines: met. I with 1 DV, met. II with 1 DV, 1 DPV, 1 DRV, met. III with 2 DV, 1 DPV, 1 DRV, 1 DD and met. IV with 2 DV, 1 DPV, 1 DRV, 1 DD, 1 DL. Disordered cluster of small, short spines cover entire ventral region of met. IV outside scopulate area (
<figureCitation id="135E2A22767FFF8A9078FD32FD46D6B4" box="[636,724,712,738]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="34" pageNumber="35">Fig. 98</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FD17FB1BD751" blockId="34.[151,1437,152,1970]" box="[196,1161,749,775]" pageId="34" pageNumber="35">
Claws: unarmed paired claws on all legs. Reduced third claw on leg IV (
<figureCitation id="135E2A22767FFF8A9612FD17FBEED751" box="[1046,1148,749,775]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="34" pageNumber="35">Fig. 100</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FCE8FDA2D794" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Genitalia (
<figureCitation id="135E2A22767FFF8A9344FCE8FE3AD77A" box="[320,424,786,812]" captionStart="FIGURES 92 – 101" captionStartId="32.[151,270,1604,1628]" captionTargetBox="[160,1422,253,1586]" captionTargetId="figure@32.[151,1436,253,1601]" captionTargetPageId="32" captionText="FIGURES 92 – 101. Lyrognathus liewi (jun. syn. L. robustus), paratype female QM S 19651.15461. 92. Habitus, dorsal view. 93. Eyes, dorsal view. 94. Sternum, labium and right maxillae, ventral view. 95. Intercheliceral pegs, prolateral view. 96. Right chelicerae, lower retrolateral view showing strikers and arrow pointing to retrolateral basomedial cheliceral spines. 97. Right maxillae showing lyra, prolateral view. 98. Right leg IV, ventral view. 99. Right leg IV, retrolateral view. 100. Right tar. IV, prolateral view showing transverse suture and reduced third claw. 101. Spermathecae, dorsal view. Scale bar = 10 mm for 92, 98, 99; 5 mm for 100; 2 mm for 93 – 97; 1 mm for 101. Illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275748/files/figure.png" pageId="34" pageNumber="35">Fig. 101</figureCitation>
): spermathecae typically bilobular, dividing basally. Lobes not apically swollen, with lateral lobes 2/3 length of medials. Medials very long. Heavily sclerotized distally, proximally reduces to weak sclerotization at bases of all lobes. N.B.: 
<bibRefCitation id="EFF44B56767FFF8A90C0FCA7FCC6D721" author="West" box="[708,852,861,887]" pageId="34" pageNumber="43" refString="West, R. C. (1991) A new theraphosid spider from the Cameron Highlands, Malaysia. Memoirs of the Queensland Museum, 30, 615 - 619." type="journal article" year="1991">West (1991)</bibRefCitation>
noted extreme variation in 
<typeStatus id="54DE8805767FFF8A969FFCA7FB4BD721" box="[1179,1241,861,887]" pageId="34" pageNumber="35">types</typeStatus>
held at the QM. In one example (
<typeStatus id="54DE8805767FFF8A936CFC78FE31D7CA" box="[360,419,898,924]" pageId="34" pageNumber="35">type</typeStatus>
: 
<accessionNumber id="9436AB44767FFF8A93B4FC78FD80D7CA" box="[432,530,898,924]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S15641" pageId="34" pageNumber="35" type="EnaNcbi">S15641</accessionNumber>
), the spermathecae were trilobular, in another (
<typeStatus id="54DE8805767FFF8A965BFC78FB08D7CA" box="[1119,1178,898,924]" pageId="34" pageNumber="35">type</typeStatus>
: 
<accessionNumber id="9436AB44767FFF8A96A3FC78FA9BD7CA" box="[1191,1289,898,924]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S15460" pageId="34" pageNumber="35" type="EnaNcbi">S15460</accessionNumber>
), a bud was present on one of the medial lobes.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FC37FBB9D7B1" blockId="34.[151,1437,152,1970]" box="[196,1067,973,999]" pageId="34" pageNumber="35">
Male QM S74340 (
<figureCitation id="135E2A22767FFF8A93ACFC37FD9FD7B1" box="[424,525,973,999]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 102</figureCitation>
, as in female except) with body length: 28.54.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FC08FC9FD05A" blockId="34.[151,1437,152,1970]" box="[196,781,1010,1036]" pageId="34" pageNumber="35">Color (in alcohol): dark brown in natural lighting.</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FBE2FEC9D001" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Carapace (
<figureCitation id="135E2A22767FFF8A934DFBE2FE2BD064" box="[329,441,1048,1074]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 102</figureCitation>
): length 11.2, width 10.1 (width across anterior edge 6.3). Fovea 1.8, distinctly procurved, deep.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FB98FDEAD0F4" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Eyes (
<figureCitation id="135E2A22767FFF8A9314FB98FEE9D02A" box="[272,379,1122,1148]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 103</figureCitation>
): ocular tubercle; length 1.2, width 2.1. Anterior row transverse. Posterior row slightly recurved. Sizes: AME, ALE, PLE, PME.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FB57FB47D0BA" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Chelicerae: length 6.7, width at base of each lobe 4.8. Teeth: 12 with&gt;40 small basomesal teeth. Intercheliceral pegs (&gt;2 larger,&gt;13 smaller) in tight cluster on basodorsal surface (
<figureCitation id="135E2A22767FFF8A9664FB28FB5AD0BA" box="[1120,1224,1234,1260]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 105</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FB02FF66D1D4" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Maxillae (
<figureCitation id="135E2A22767FFF8A9345FB02FE63D144" box="[321,497,1272,1298]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Figs 104, 107</figureCitation>
): anterior lobe well pronounced, many cuspules (&gt;280) on inner basoventral surface. 
<taxonomicName id="4C654D24767FFF8A92FEFAE7FEA2D161" box="[250,304,1309,1335]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="42" phylum="Arthropoda" rank="species" species="lyra">Lyra</taxonomicName>
(
<figureCitation id="135E2A22767FFF8A9344FAE7FE3AD161" box="[320,424,1309,1335]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 107</figureCitation>
): immediately above suture no elongated spines (only typical hair-like setae) medially to anterior margin. Labium (
<figureCitation id="135E2A22767FFF8A93EDFAB8FDC0D10A" box="[489,594,1346,1372]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 104</figureCitation>
): length 1.4, width 2.3. Many small cuspules (&gt;500) along anterior 1/4 surface.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FA77FF73D1A4" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Sternum (
<figureCitation id="135E2A22767FFF8A933EFA77FE37D1F1" box="[314,421,1421,1447]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 104</figureCitation>
): length 5.4, width 4.2. Clearly wider posteriorly than anteriorly, with even cover of short setae, interspersed with many longer darker setae. Differs from females in that no spiniform setae around edges.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0FA07FE86D34A" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Legs: formula; (length) IV, I, II, III; (width) III, IV, I, II. Leg 
<collectingCountry id="F3727637767FFF8A9194FA07FC24D241" box="[912,950,1533,1559]" name="Russia" pageId="34" pageNumber="35">RF</collectingCountry>
~93.45. Leg lengths (fem., pat., tib., met., tar., total): palp: 10.1, 5.15, 9.05, 0.00, 2.9, 27.2. I: 12.5, 6.75, 10.6, 9.55, 6.43, 45.83. II: 11.2, 5.9, 8.94, 8.95, 5.6, 40.59. III: 9.05, 4.65, 6.7, 8.85, 5.53, 34.78. IV: 12.6, 6.15, 10.4, 14.55, 5.34, 49.09. Leg mid-widths (fem., pat., tib., met., tar., total.): palp: 2.15, 2.03, 2.02, 0.00, 2.4, 8.6. I: 2.45, 2.35, 2.2, 1.95, 1.63, 10.58. II: 2.44, 2.35, 2.15, 1.87, 1.5, 10.31. III: 3.9, 2.9, 2.05, 1.83, 1.75, 12.43. IV: 2.43, 2.85, 2.3, 1.82, 1.72, 11.12. Coxae ventral measurements: palp–length 4.99, width 2.16; I–5.21, 2.66; II–4.94, 2.66; III–4.04, 2.67; IV– 4.52, 2.70. Trochantera ventral: palp–length 1.79, width 1.97; I–2.16, 2.29; II–2.05, 2.17; III–2.13, 2.41; IV– 2.16, 2.45.
</paragraph>
<paragraph id="8BDA36A7767FFF8A92C0F8D2FB11D3E4" blockId="34.[151,1437,152,1970]" pageId="34" pageNumber="35">
Genitalia. Palp (
<figureCitation id="135E2A22767FFF8A9382F8D2FDA2D314" box="[390,560,1832,1858]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Figs 109–112</figureCitation>
): tib. with dorsal arrangement of distinct long pallid sensory setae 2–1–2 (proximally, medially and distally), distal pair with medial bend dorsally, as in typical metatarsal distal paired sensory setae (
<figureCitation id="135E2A22767FFF8A934DF888FE21D3DA" box="[329,435,1906,1932]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 109</figureCitation>
); embolus tip tapers distally with no flaring (
<figureCitation id="135E2A22767FFF8A91D3F888FBD0D3DA" box="[983,1090,1906,1932]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 110</figureCitation>
). Twisted through 90° at tip. Relative length of tib. about twice length of embolus. No longitudinal keel (
<figureCitation id="135E2A22767FFF8A9614F862FBE7D3E4" box="[1040,1141,1944,1970]" captionStart="FIGURES 102 – 112" captionStartId="33.[151,271,1848,1872]" captionTargetBox="[151,1426,375,1821]" captionTargetId="figure@33.[151,1436,354,1833]" captionTargetPageId="33" captionText="FIGURES 102 – 112. Lyrognathus robustus male QM S 74340. 102, Habitus, dorsal view. 103. Eyes, dorsal view. 104. Sternum, labium and right maxillae, ventral view. 105. Right intercheliceral pegs, prolateral view. 106. Right lower retrolateral cheliceral surface showing strikers and retrolaeral basomedial spines. 107. Right maxillae showing lyra, prolateral view. 108. Right leg IV, ventral view. 109. Right palpal tibia, prolateral view. 110. Right bulb, retrolateral view. 111. Right bulb, prolateral view. 112. Left embolus, transverse cross section. Scale bar = 20 mm for 102; 10 mm for 108; 5 mm for 109; 2 mm for 104, 107, 110, 111; 1 mm for 103, 105, 106; 0.5 mm for 112. Photo &amp; illustrations: S. C. Nunn." httpUri="https://zenodo.org/record/275749/files/figure.png" pageId="34" pageNumber="35">Fig. 112</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F652C767EFF8B92C0FF6DFCEFD58A" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BDA36A7767EFF8B92C0FF6DFCEFD58A" blockId="35.[151,1437,151,1447]" pageId="35" pageNumber="36">
<emphasis id="B911EAB5767EFF8B92C0FF6DFDFCD4E7" bold="true" box="[196,622,151,177]" pageId="35" pageNumber="36">Distribution and natural history</emphasis>
(
<figureCitation id="135E2A22767EFF8B9084FF62FD78D4E4" box="[640,746,152,178]" captionStart="FIGURE 114" captionStartId="38.[151,255,1297,1321]" captionTargetBox="[151,1436,194,1276]" captionTargetId="figure@38.[151,1436,194,1276]" captionTargetPageId="38" captionText="FIGURE 114. Records of Lyrognathus species in Southeast Asia. C. Senescall" httpUri="https://zenodo.org/record/275751/files/figure.png" pageId="35" pageNumber="36">Fig. 114</figureCitation>
): 
<taxonomicName id="4C654D24767EFF8B9101FF62FC02D4E7" box="[773,912,152,177]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B9101FF62FC02D4E7" box="[773,912,152,177]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
is found among the wet montane forested regions of 
<collectingCountry id="F3727637767EFF8B931DFF47FE14D481" box="[281,390,189,215]" name="Cameroon" pageId="35" pageNumber="36">Cameron</collectingCountry>
Highlands, Pahang State, in the main Central range and from Baling, Kedah State and Bukit Larut and Gerik, Perak State in the Eastern range of West 
<collectingCountry id="F3727637767EFF8B917BFF18FC7DD4AA" box="[895,1007,226,252]" name="Malaysia" pageId="35" pageNumber="36">Malaysia</collectingCountry>
. 
<taxonomicName id="4C654D24767EFF8B91FDFF18FBE9D4AD" box="[1017,1147,226,251]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B91FDFF18FBE9D4AD" box="[1017,1147,226,251]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
construct simple tubular burrows in clay-like soil with either a plain opening or a slight flare-mouth that utilizes surrounding organic debris or vegetation around the mouth on sloped or nearly vertical embankments. Adult female burrows were J-shaped and terminatined in an enlarged chamber about 
<quantity id="4C9D9B42767EFF8B914CFEA8FC2DD53A" box="[840,959,338,364]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" metricValueMax="3.0" metricValueMin="1.8" pageId="35" pageNumber="36" unit="cm" value="24.0" valueMax="30.0" valueMin="18.0">18–30 cm</quantity>
deep. An ultimate male was found in a burrow in January (S. Hogg, pers. comm.). A “nervous” spider, 
<taxonomicName id="4C654D24767EFF8B91AAFE82FBAAD5C7" box="[942,1080,376,401]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B91AAFE82FBAAD5C7" box="[942,1080,376,401]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
will readily bite if provoked (West, pers. obs.). The local native Orang Asli people know these tarantulas as “Tei Foo” (translates as “Earth Tiger”) and still believe the bite to be fatal (
<bibRefCitation id="EFF44B56767EFF8B9092FE38FCE2D58A" author="West" box="[662,880,450,476]" pageId="35" pageNumber="43" refString="West, R. C. (1991) A new theraphosid spider from the Cameron Highlands, Malaysia. Memoirs of the Queensland Museum, 30, 615 - 619." type="journal article" year="1991" yearSuffix="p">West 1991, p. 616</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F652C767EFF8B92C0FE1DFC5DD1F1" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BDA36A7767EFF8B92C0FE1DFC58D0BA" blockId="35.[151,1437,151,1447]" pageId="35" pageNumber="36">
<emphasis id="B911EAB5767EFF8B92C0FE1DFEADD657" bold="true" box="[196,319,487,513]" pageId="35" pageNumber="36">Remarks:</emphasis>
Von 
<bibRefCitation id="EFF44B56767EFF8B9378FE12FDC4D654" author="Wirth" box="[380,598,488,514]" pageId="35" pageNumber="43" refString="von Wirth, V. (2000) Uber die tatsachliche Identitat der im Vorartikel als Lyrognathus robustus Smith, 1988 bezeichneten Vogelspinnenart (Araneae, Theraphosidae, Selenocosmiinae). Arthropoda, 3 (8), 17 - 19." type="journal article" year="2000" yearSuffix="p">Wirth (2000, p.17)</bibRefCitation>
discussed some issues between 
<taxonomicName id="4C654D24767EFF8B91CBFE12FBC0D657" box="[975,1106,488,513]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B91CBFE12FBC0D657" box="[975,1106,488,513]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C654D24767EFF8B968DFE12FAD2D657" box="[1161,1344,488,513]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B968DFE12FAD2D657" box="[1161,1344,488,513]" italics="true" pageId="35" pageNumber="36">Coremiocnemis</emphasis>
</taxonomicName>
species within 
<collectingCountry id="F3727637767EFF8B92EEFDF7FECBD671" box="[234,345,525,551]" name="Malaysia" pageId="35" pageNumber="36">Malaysia</collectingCountry>
, compared to 
<taxonomicName id="4C654D24767EFF8B9005FDF7FD06D670" box="[513,660,525,550]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B9005FDF7FD06D670" box="[513,660,525,550]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
of 
<collectingCountry id="F3727637767EFF8B90B8FDF7FD6ED671" box="[700,764,525,551]" name="India" pageId="35" pageNumber="36">India</collectingCountry>
. He assumed it might be best to transfer 
<taxonomicName id="4C654D24767EFF8B96E6FDF7FAF4D670" box="[1250,1382,525,550]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B96E6FDF7FAF4D670" box="[1250,1382,525,550]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
into 
<taxonomicName id="4C654D24767EFF8B9293FDC8FEC0D61D" box="[151,338,562,587]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B9293FDC8FEC0D61D" box="[151,338,562,587]" italics="true" pageId="35" pageNumber="36">Coremiocnemis</emphasis>
</taxonomicName>
based on plesiomorphic metatarsal scopula division on leg IV that 
<taxonomicName id="4C654D24767EFF8B9680FDC8FA9ED61D" box="[1156,1292,562,587]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B9680FDC8FA9ED61D" box="[1156,1292,562,587]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
shares with 
<emphasis id="B911EAB5767EFF8B9293FDA2FEF0D627" box="[151,354,600,625]" italics="true" pageId="35" pageNumber="36">
<taxonomicName id="4C654D24767EFF8B9293FDA2FECFD627" box="[151,349,600,625]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Coremiocnemis</taxonomicName>
,
</emphasis>
as well as presence of intercheliceral pegs and bilobular spermathceae, not known in 
<taxonomicName id="4C654D24767EFF8B9293FD87FEA2D6C0" box="[151,304,637,662]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B9293FD87FEA2D6C0" box="[151,304,637,662]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
prior to Smith’s 
<taxonomicName id="4C654D24767EFF8B9000FD87FD1DD6C0" box="[516,655,637,662]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B9000FD87FD1DD6C0" box="[516,655,637,662]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
description. He also considered zoogeographic ranges for both groups. However, with the inclusion of three new 
<taxonomicName id="4C654D24767EFF8B910BFD58FC38D6ED" box="[783,938,674,699]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B910BFD58FC38D6ED" box="[783,938,674,699]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
species that display traits diagnostic of 
<taxonomicName id="4C654D24767EFF8B9293FD32FEB9D6B7" box="[151,299,712,737]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B9293FD32FEB9D6B7" box="[151,299,712,737]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
(undivided scopula on metatarsi IV), but share with 
<taxonomicName id="4C654D24767EFF8B91A2FD32FBB9D6B7" box="[934,1067,712,737]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B91A2FD32FBB9D6B7" box="[934,1067,712,737]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
the presence of intercheliceral pegs and bilobular spermathecae and further, are found within 
<collectingCountry id="F3727637767EFF8B9186FD17FC6AD751" box="[898,1016,749,775]" name="Indonesia" pageId="35" pageNumber="36">Indonesia</collectingCountry>
, the previously considered dubious placement of this species into 
<taxonomicName id="4C654D24767EFF8B93FDFCE8FD19D77D" box="[505,651,786,811]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B93FDFCE8FD19D77D" box="[505,651,786,811]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
is no longer questionable. Further, clavate trichobothria on tarsi of 
<taxonomicName id="4C654D24767EFF8B9293FCC2FE8BD707" box="[151,281,824,849]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B9293FCC2FE8BD707" box="[151,281,824,849]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
are like that in all other 
<taxonomicName id="4C654D24767EFF8B9030FCC2FD54D707" box="[564,710,824,849]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B9030FCC2FD54D707" box="[564,710,824,849]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
and not those of any Malaysian 
<taxonomicName id="4C654D24767EFF8B963BFCC2FB64D707" box="[1087,1270,824,849]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B963BFCC2FB64D707" box="[1087,1270,824,849]" italics="true" pageId="35" pageNumber="36">Coremiocnemis</emphasis>
</taxonomicName>
species. Male embolus morphology does not conform to 
<taxonomicName id="4C654D24767EFF8B9083FCA7FCACD720" box="[647,830,861,886]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B9083FCA7FCACD720" box="[647,830,861,886]" italics="true" pageId="35" pageNumber="36">Coremiocnemis</emphasis>
</taxonomicName>
either, lacking a distal “kiss curl” (West &amp; Nunn, in press). There is at most likely a close relationship shared between 
<taxonomicName id="4C654D24767EFF8B919AFC78FBA2D7CD" box="[926,1072,898,923]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B919AFC78FBA2D7CD" box="[926,1072,898,923]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C654D24767EFF8B966DFC78FAB2D7CD" box="[1129,1312,898,923]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B966DFC78FAB2D7CD" box="[1129,1312,898,923]" italics="true" pageId="35" pageNumber="36">Coremiocnemis</emphasis>
</taxonomicName>
by way of several shared derived characters (see: retrolateral cheliceral spines and spermathecae in Character discussion), however, the presence of penicillate retrolateral setal brushes on tibiae IV in combination with incrassate tibiae IV in females and ventral projection of the male embolus is seen only in 
<taxonomicName id="4C654D24767EFF8B96FFFC08FA07D05D" box="[1275,1429,1010,1035]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B96FFFC08FA07D05D" box="[1275,1429,1010,1035]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C654D24767EFF8B9293FBE2FEB3D067" box="[151,289,1048,1073]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B9293FBE2FEB3D067" box="[151,289,1048,1073]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
shows some traits that differ from other members of the genus and, in that regard, might be considered the most extreme representative of 
<taxonomicName id="4C654D24767EFF8B90BEFBC7FCDED000" box="[698,844,1085,1110]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B90BEFBC7FCDED000" box="[698,844,1085,1110]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
, but cannot be considered separate to the group as it clearly does not fit 
<taxonomicName id="4C654D24767EFF8B939DFB99FDC1D02A" box="[409,595,1123,1148]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B939DFB99FDC1D02A" box="[409,595,1123,1148]" italics="true" pageId="35" pageNumber="36">Coremiocnemis</emphasis>
</taxonomicName>
or any other selenocosmiine genus as suggested by von 
<bibRefCitation id="EFF44B56767EFF8B96FDFB98FA0BD02A" author="Wirth" box="[1273,1433,1122,1148]" pageId="35" pageNumber="43" refString="von Wirth, V. (2000) Uber die tatsachliche Identitat der im Vorartikel als Lyrognathus robustus Smith, 1988 bezeichneten Vogelspinnenart (Araneae, Theraphosidae, Selenocosmiinae). Arthropoda, 3 (8), 17 - 19." type="journal article" year="2000">Wirth (2000)</bibRefCitation>
. The inclusion of 
<taxonomicName id="4C654D24767EFF8B935AFB72FE4AD0F7" box="[350,472,1160,1185]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
<emphasis id="B911EAB5767EFF8B935AFB72FE4AD0F7" box="[350,472,1160,1185]" italics="true" pageId="35" pageNumber="36">L. achilles</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE767EFF8B93DBFB7DFDAED0F7" box="[479,572,1159,1185]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B911EAB5767EFF8B93DBFB7DFDAED0F7" bold="true" box="[479,572,1159,1185]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C654D24767EFF8B904FFB72FD21D0F7" box="[587,691,1160,1185]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B911EAB5767EFF8B904FFB72FD21D0F7" box="[587,691,1160,1185]" italics="true" pageId="35" pageNumber="36">L. fuscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE767EFF8B90BEFB7DFC85D0F7" box="[698,791,1159,1185]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B911EAB5767EFF8B90BEFB7DFC85D0F7" bold="true" box="[698,791,1159,1185]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C654D24767EFF8B9154FB72FC44D0F7" box="[848,982,1160,1185]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB5767EFF8B9154FB72FC44D0F7" box="[848,982,1160,1185]" italics="true" pageId="35" pageNumber="36">L. lessunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE767EFF8B91D9FB7DFBA8D0F7" box="[989,1082,1159,1185]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B911EAB5767EFF8B91D9FB7DFBA8D0F7" bold="true" box="[989,1082,1159,1185]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
, herein, shows that traits such as intercheliceral pegs, bilobular spermathecae, retrolateral basomedial cheliceral spines and zoogeographic ranges outside of 
<collectingCountry id="F3727637767EFF8B9362FB28FE30D0BA" box="[358,418,1234,1260]" name="India" pageId="35" pageNumber="36">India</collectingCountry>
clearly fit parsimoniously within 
<taxonomicName id="4C654D24767EFF8B9134FB29FC50D0BA" box="[816,962,1235,1260]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB5767EFF8B9134FB29FC50D0BA" box="[816,962,1235,1260]" italics="true" pageId="35" pageNumber="36">Lyrognathus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDA36A7767EFF8B92C0FB02FC5DD1F1" blockId="35.[151,1437,151,1447]" pageId="35" pageNumber="36">
Even though 
<taxonomicName id="4C654D24767EFF8B9363FB02FE51D147" box="[359,451,1272,1297]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="liewi">
<emphasis id="B911EAB5767EFF8B9363FB02FE51D147" box="[359,451,1272,1297]" italics="true" pageId="35" pageNumber="36">L. liewi</emphasis>
</taxonomicName>
was placed as a junior synonym of 
<taxonomicName id="4C654D24767EFF8B9178FB02FB96D147" box="[892,1028,1272,1297]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B9178FB02FB96D147" box="[892,1028,1272,1297]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
, a subsequent search through the senior author’s collection yielded a male, now deposited in the QM. A second 
<taxonomicName id="4C654D24767EFF8B9646FAE7FB55D160" box="[1090,1223,1309,1334]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B9646FAE7FB55D160" box="[1090,1223,1309,1334]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
male was located near the summit of Bukit Larut, Perak, West 
<collectingCountry id="F3727637767EFF8B90ACFAB8FC81D10A" box="[680,787,1346,1372]" name="Malaysia" pageId="35" pageNumber="36">Malaysia</collectingCountry>
by Mr Stephen Hogg, and then donated to the QM. So far no other males from 
<taxonomicName id="4C654D24767EFF8B93BCFA92FDAFD1D7" box="[440,573,1384,1409]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5767EFF8B93BCFA92FDAFD1D7" box="[440,573,1384,1409]" italics="true" pageId="35" pageNumber="36">L. robustus</emphasis>
</taxonomicName>
have ever been located, in these specimens, informative characters exist that are not present in females, hence the need to define the male here.
</paragraph>
</subSubSection>
</treatment>
</document>