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<document id="17A0B477A3363B15F7D179FB77241C86" ID-DOI="10.11646/zootaxa.4747.2.6" ID-GBIF-Dataset="cefd46f6-f601-4ccf-bfbf-91b41c58937f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3694896" ID-ZooBank="96716FDE-BA1F-4903-8998-33A7780E0A9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583221284863" checkinUser="plazi" docAuthor="Král, David, Jákl, Stanislav, Lu, Yuanyuan &amp; Bai, Ming" docDate="2020" docId="03CC879B8A65FFADFF5BFCE5F34FF80D" docLanguage="en" docName="zootaxa.4747.2.6.pdf.imf" docOrigin="Zootaxa 4747 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratophyus amdoensis Kral &amp; Jakl 2020, new species" docType="treatment" docVersion="7" lastPageNumber="341" masterDocId="FFF5FFE38A64FFAFFFCCFFA0F011FFE8" masterDocTitle="The genus Ceratophyus Fischer von Waldheim, 1824 (Coleoptera: Geotrupidae) in China, along with description of a new species" masterLastPageNumber="349" masterPageNumber="339" pageNumber="340" updateTime="1698724118592" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="4AA9CD20658E4F659C1DAB1C648E4244" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4AA9BB99FD2A8099459772759C9BEB40">The genus Ceratophyus Fischer von Waldheim, 1824 (Coleoptera: Geotrupidae) in China, along with description of a new species</mods:title>
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<mods:name id="16B19C8C9B6D9160B2D7EC8950D7C9CE" type="personal">
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<mods:namePart id="1015D1CB3C7206C2B3D647449F1EA881">Král, David</mods:namePart>
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<mods:namePart id="B59D5E86A3D2BE8B92B6D59E2E813208">Jákl, Stanislav</mods:namePart>
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<mods:namePart id="347BB1CDC2370CCDEC99A62D5535D092">Lu, Yuanyuan</mods:namePart>
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<mods:namePart id="AF1ED73CA69C9F2D663D6E698F56A362">Bai, Ming</mods:namePart>
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<mods:title id="E217E5A454066989944678554440C2D8">Zootaxa</mods:title>
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<mods:part id="3DEB812A6F2994E6C2A4343EC33590F7">
<mods:date id="3A62CA3E6345B77F4549C16ECF8A8BEB">2020</mods:date>
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<mods:number id="5200C7B2BA7E446ADDCA1345152A13FE">2020-03-03</mods:number>
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<mods:number id="2E6FD3D60F86051F9EFBA9343848290D">4747</mods:number>
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<mods:classification id="21AFD7788C9E22EEA60C379E5BCBAB54">journal article</mods:classification>
<mods:identifier id="90B7EC8B8956C60317D70E1035966DE3" type="DOI">10.11646/zootaxa.4747.2.6</mods:identifier>
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<mods:identifier id="DCEA85E39243F632DB0D6B9189E79912" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CC879B8A65FFADFF5BFCE5F34FF80D" ID-DOI="http://doi.org/10.5281/zenodo.3704194" ID-GBIF-Taxon="162381238" ID-Zenodo-Dep="3704194" LSID="urn:lsid:plazi:treatment:03CC879B8A65FFADFF5BFCE5F34FF80D" httpUri="http://treatment.plazi.org/id/03CC879B8A65FFADFF5BFCE5F34FF80D" lastPageId="2" lastPageNumber="341" pageId="1" pageNumber="340">
<subSubSection id="C37F65068A65FFAEFF5BFCE5F3D0FCB6" box="[151,961,836,864]" pageId="1" pageNumber="340" type="nomenclature">
<paragraph id="8BDA368D8A65FFAEFF5BFCE5F3D0FCB6" blockId="1.[151,961,836,900]" box="[151,961,836,864]" pageId="1" pageNumber="340">
<emphasis id="B911EA9F8A65FFAEFF5BFCE5F2E5FCB7" bold="true" box="[151,756,836,864]" pageId="1" pageNumber="340">
<taxonomicName id="4C654D0E8A65FFAEFF5BFCE5F246FCB7" ID-CoL="SP6G" authority="Kral &amp; Jakl" authorityName="Kral &amp; Jakl" authorityYear="2020" box="[151,599,836,864]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="amdoensis" status="sp. nov.">
<emphasis id="B911EA9F8A65FFAEFF5BFCE5F1A6FC88" bold="true" box="[151,439,837,864]" italics="true" pageId="1" pageNumber="340">Ceratophyus amdoensis</emphasis>
Král &amp; Jákl
</taxonomicName>
,
<taxonomicNameLabel id="A22257E48A65FFAEFDA8FCE5F2E5FCB7" box="[612,756,837,863]" pageId="1" pageNumber="340" rank="species">new species</taxonomicNameLabel>
</emphasis>
[安多角粪金龟]
</paragraph>
</subSubSection>
<subSubSection id="C37F65068A65FFAEFF5BFCC9F103FC6C" box="[151,274,873,900]" pageId="1" pageNumber="340" type="description">
<paragraph id="8BDA368D8A65FFAEFF5BFCC9F103FC6C" blockId="1.[151,961,836,900]" box="[151,274,873,900]" pageId="1" pageNumber="340">
(
<figureCitation id="135E2A088A65FFAEFF53FCC9F118FC6C" box="[159,265,873,900]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–12" captionStartId-0="2.[151,264,1671,1696]" captionStartId-1="3.[151,264,1811,1836]" captionTargetBox-0="[177,1411,244,1633]" captionTargetBox-1="[184,1353,211,1675]" captionTargetId-0="figure@2.[164,1423,234,1644]" captionTargetId-1="figure@3.[155,1429,178,1788]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–3. Ceratophyus amdoensis Král &amp; Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale." captionText-1="FIGURES 4–12. Aedeagus of Ceratophyus Fischer von Waldheim, 1824 species. 4–6, Ceratophyus amdoensis Král &amp; Jákl, new species, holotype; 7–9, C. dauricus (Jekel, 1866) (“Transbaikalia”, NMPC); 10–12, C. kabaki Nikolajev, 2007 (Sichuan, Yading, NMPC). 4, 7, 10, dorsal view; 5, 8, 11, left lateral view; 6, 9, 12, tips, ventral view. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.3694898" figureDoi-1="http://doi.org/10.5281/zenodo.3694900" httpUri-0="https://zenodo.org/record/3694898/files/figure.png" httpUri-1="https://zenodo.org/record/3694900/files/figure.png" pageId="1" pageNumber="340">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F65068A65FFAEFF5BFC11F53BFBDF" pageId="1" pageNumber="340" type="materials_examined">
<paragraph id="8BDA368D8A65FFAEFF5BFC11F4A2FC23" blockId="1.[151,1437,945,2015]" box="[151,1203,945,971]" pageId="1" pageNumber="340">
<heading id="D09281E18A65FFAEFF5BFC11F4A2FC23" bold="true" box="[151,1203,945,971]" fontSize="11" level="1" pageId="1" pageNumber="340" reason="1">
<materialsCitation id="3B0D3CD08A65FFAEFF5BFC11F4A2FC23" ID-GBIF-Occurrence="2573218802" box="[151,1203,945,971]" country="China" elevation="2300" latitude="38.366665" location="Gansu" longLatPrecision="1175" longitude="101.333336" pageId="1" pageNumber="340" specimenCount="1" stateProvince="Gansu" typeStatus="holotype">
<emphasis id="B911EA9F8A65FFAEFF5BFC11F4A2FC23" bold="true" box="[151,1203,945,971]" pageId="1" pageNumber="340">
<typeStatus id="54DE882F8A65FFAEFF5BFC11F0C5FC23" box="[151,212,945,971]" pageId="1" pageNumber="340">Type</typeStatus>
locality.
<collectingCountry id="F372761D8A65FFAEFE88FC11F183FC23" box="[324,402,945,971]" name="China" pageId="1" pageNumber="340">China</collectingCountry>
,
<collectingRegion id="49A1F86F8A65FFAEFE53FC11F1E3FC23" box="[415,498,945,971]" country="China" name="Gansu" pageId="1" pageNumber="340">Gansu</collectingRegion>
,
<quantity id="4C9D9B688A65FFAEFDCCFC11F25CFC23" box="[512,589,945,971]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="340" unit="km" value="50.0">50 km</quantity>
SE of
<collectingRegion id="49A1F86F8A65FFAEFD53FC11F301FC23" box="[671,784,945,971]" country="China" name="Shandan" pageId="1" pageNumber="340">Shandan</collectingRegion>
,
<geoCoordinate id="EE51504A8A65FFAEFCD1FC11F392FC23" box="[797,899,945,971]" degrees="38" direction="north" minutes="22" orientation="latitude" pageId="1" pageNumber="340" precision="925" value="38.366665">38°22’N</geoCoordinate>
<geoCoordinate id="EE51504A8A65FFAEFC46FC11F3ECFC23" box="[906,1021,945,971]" degrees="101" direction="east" minutes="20" orientation="longitude" pageId="1" pageNumber="340" precision="925" value="101.333336">101°20’E</geoCoordinate>
,
<elevation id="0048D1BE8A65FFAEFBC7FC11F4A2FC23" box="[1035,1203,945,971]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="340" unit="m" value="2300.0">
<quantity id="4C9D9B688A65FFAEFBC7FC11F474FC23" box="[1035,1125,945,971]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="340" unit="m" value="2300.0">2300 m</quantity>
a. s. l.
</elevation>
</emphasis>
</materialsCitation>
</heading>
</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BFC75F53BFBDF" blockId="1.[151,1437,945,2015]" pageId="1" pageNumber="340">
<emphasis id="B911EA9F8A65FFAEFF0BFC75F167FC07" bold="true" box="[199,374,980,1007]" pageId="1" pageNumber="340">Type material.</emphasis>
<materialsCitation id="3B0D3CD08A65FFAEFEB2FC74F368FBFB" ID-GBIF-Occurrence="2573218845" collectionCode="NMPC" country="China" elevation="2300" latitude="38.366665" location="Gansu" longLatPrecision="1175" longitude="101.333336" pageId="1" pageNumber="340" specimenCount="1" specimenCount-male="1" stateProvince="Gansu" typeStatus="holotype">
<emphasis id="B911EA9F8A65FFAEFEB2FC74F23CFC07" bold="true" box="[382,557,980,1007]" pageId="1" pageNumber="340">
<collectingCountry id="F372761D8A65FFAEFEB2FC74F1D8FC07" box="[382,457,980,1007]" name="China" pageId="1" pageNumber="340">China</collectingCountry>
,
<collectingRegion id="49A1F86F8A65FFAEFE1AFC74F239FC07" box="[470,552,980,1007]" country="China" name="Gansu" pageId="1" pageNumber="340">Gansu</collectingRegion>
:
</emphasis>
<typeStatus id="54DE882F8A65FFAEFDFAFC75F2B2FC07" box="[566,675,981,1007]" pageId="1" pageNumber="340" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D63FD048A65FFAEFD62FC75F2D2FC07" box="[686,707,981,1007]" pageId="1" pageNumber="340" type="male">♂</specimenCount>
(
<collectionCode id="ED74AE488A65FFAEFD1EFC75F336FC07" box="[722,807,981,1007]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="340">NMPC</collectionCode>
), “CHINA–GANSU |
<quantity id="4C9D9B688A65FFAEFBE9FC74F474FC07" box="[1061,1125,980,1007]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="340" unit="km" value="50.0">50km</quantity>
SE SHANDAN |
<geoCoordinate id="EE51504A8A65FFAEFAF8FC75F586FC07" box="[1332,1431,981,1007]" degrees="38" direction="north" minutes="22" orientation="latitude" pageId="1" pageNumber="340" precision="925" value="38.366665">38°22’N</geoCoordinate>
,
<geoCoordinate id="EE51504A8A65FFAEFF5BFC59F113FBFB" box="[151,258,1017,1043]" degrees="101" direction="east" minutes="20" orientation="longitude" pageId="1" pageNumber="340" precision="925" value="101.333336">101°20’E</geoCoordinate>
|| 15-25-05.2005 H=
<quantity id="4C9D9B688A65FFAEFE26FC59F228FBFB" box="[490,569,1017,1043]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="340" unit="m" value="2300.0">
<elevation id="0048D1BE8A65FFAEFE26FC59F228FBFB" box="[490,569,1017,1043]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="340" unit="m" value="2300.0">2300m</elevation>
</quantity>
| leg/Coll Siniaev [printed]”
</materialsCitation>
.
<materialsCitation id="3B0D3CD08A65FFAEFC48FC59F589FBFB" ID-GBIF-Occurrence="2573218811" box="[900,1432,1017,1043]" collectionCode="NMPC" country="China" elevation="2300" latitude="38.366665" location="Gansu" longLatPrecision="1175" longitude="101.333336" pageId="1" pageNumber="340" specimenCount="1" specimenCount-male="1" stateProvince="Gansu" typeStatus="paratype">
<typeStatus id="54DE882F8A65FFAEFC48FC59F3E4FBFB" box="[900,1013,1017,1043]" pageId="1" pageNumber="340" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D63FD048A65FFAEFC33FC59F438FBFB" box="[1023,1065,1017,1043]" pageId="1" pageNumber="340" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED74AE488A65FFAEFBFBFC59F49DFBFB" box="[1079,1164,1017,1043]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="340">NMPC</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B0D3CD08A65FFAEFF5BFBBCF537FBDF" ID-GBIF-Occurrence="2573218805" box="[151,1318,1052,1079]" collectingDate="1994-08-03" collectingDateMax="1994-08-07" collectingDateMin="1994-08-03" collectionCode="SJCP" country="China" elevation="3350" location="Qinghai" pageId="1" pageNumber="340" specimenCount="1" specimenCount-male="1" stateProvince="Qinghai" typeStatus="paratype">
<emphasis id="B911EA9F8A65FFAEFF5BFBBCF110FBDF" bold="true" box="[151,257,1052,1079]" pageId="1" pageNumber="340">
<collectingRegion id="49A1F86F8A65FFAEFF5BFBBCF0EDFBDF" box="[151,252,1052,1079]" country="China" name="Qinghai" pageId="1" pageNumber="340">Qinghai</collectingRegion>
:
</emphasis>
<specimenCount id="9D63FD048A65FFAEFEC4FBBDF123FBDF" box="[264,306,1053,1079]" pageId="1" pageNumber="340" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED74AE488A65FFAEFE8DFBBDF192FBDF" box="[321,387,1053,1079]" pageId="1" pageNumber="340">SJCP</collectionCode>
), “CHINA-Qinghai, Nan Shan | Heimahe,
<date id="FFDB104D8A65FFAEFCADFBBDF412FBDF" box="[865,1027,1053,1079]" pageId="1" pageNumber="340" value="1994-08-03" valueMax="1994-08-07" valueMin="1994-08-03">
<collectingDate id="EF9FE9A58A65FFAEFCADFBBDF412FBDF" box="[865,1027,1053,1079]" pageId="1" pageNumber="340" value="1994-08-03" valueMax="1994-08-07" valueMin="1994-08-03">3.-7.VIII.1994</collectingDate>
</date>
|
<quantity id="4C9D9B688A65FFAEFBDBFBBCF4BFFBDF" box="[1047,1198,1052,1079]" metricMagnitude="3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="1" pageNumber="340" unit="m" value="3350.0" valueMax="3500.0" valueMin="3200.0">
<elevation id="0048D1BE8A65FFAEFBDBFBBCF4BFFBDF" box="[1047,1198,1052,1079]" metricMagnitude="3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="1" pageNumber="340" unit="m" value="3350.0" valueMax="3500.0" valueMin="3200.0">3200-3500 m</elevation>
</quantity>
[printed]”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F65068A65FFADFF0BFBE0F113FF3F" lastPageId="2" lastPageNumber="341" pageId="1" pageNumber="340" type="description">
<paragraph id="8BDA368D8A65FFAEFF0BFBE0F303FB97" blockId="1.[151,1437,945,2015]" pageId="1" pageNumber="340">
<emphasis id="B911EA9F8A65FFAEFF0BFBE0F1CCFBB3" bold="true" box="[199,477,1088,1115]" pageId="1" pageNumber="340">
Description of
<typeStatus id="54DE882F8A65FFAEFEBAFBE1F1CCFBB3" box="[374,477,1089,1115]" pageId="1" pageNumber="340" type="holotype">holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D63FD048A65FFAEFE21FBE1F215FBB3" box="[493,516,1089,1115]" pageId="1" pageNumber="340" type="male">♂</specimenCount>
). Weakly convex, black, dorsal surface shining; macrosetation of lateral margin of head and pronotum and ventral surface blackish (
<figureCitation id="135E2A088A65FFAEFD73FBC5F311FB97" box="[703,768,1125,1151]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1671,1696]" captionTargetBox="[177,1411,244,1633]" captionTargetId="figure@2.[164,1423,234,1644]" captionTargetPageId="2" captionText="FIGURES 1–3. Ceratophyus amdoensis Král &amp; Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3694898" httpUri="https://zenodo.org/record/3694898/files/figure.png" pageId="1" pageNumber="340">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BFB29F147FA93" blockId="1.[151,1437,945,2015]" pageId="1" pageNumber="340">
Head (
<figureCitation id="135E2A088A65FFAEFEDEFB29F16FFB4B" box="[274,382,1161,1187]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1671,1696]" captionTargetBox="[177,1411,244,1633]" captionTargetId="figure@2.[164,1423,234,1644]" captionTargetPageId="2" captionText="FIGURES 1–3. Ceratophyus amdoensis Král &amp; Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3694898" httpUri="https://zenodo.org/record/3694898/files/figure.png" pageId="1" pageNumber="340">Figs. 1, 3</figureCitation>
). Mandibles considerably protruding anteriorly, with broadly rounded, bisinuate external outline, smooth. Labrum transverse, broader than long, anterior edge weakly emarginate, surface distinctly punctate. Clypeus triangularly elongated anteriorly and slightly upward, with long, acute triangular apical process. Clypeal horn including its base finely, sparsely punctate. Oblique keels above eyes distinct, reaching to level of anterior outline of eyes; genae elongated in distinct tooth anterolaterally. Clypeus shallowly, sparsely but distinctly punctate, each puncture very fine, separated by about ten puncture diameters. Frons, vertex and occiput except area of clypeal horn impunctate.
</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BFA25F51CF9E3" blockId="1.[151,1437,945,2015]" pageId="1" pageNumber="340">
Pronotum (
<figureCitation id="135E2A088A65FFAEFE8BFA25F1A4FA77" box="[327,437,1413,1439]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1671,1696]" captionTargetBox="[177,1411,244,1633]" captionTargetId="figure@2.[164,1423,234,1644]" captionTargetPageId="2" captionText="FIGURES 1–3. Ceratophyus amdoensis Král &amp; Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3694898" httpUri="https://zenodo.org/record/3694898/files/figure.png" pageId="1" pageNumber="340">Figs. 1, 3</figureCitation>
) distinctly transverse, broadest at midlength, entirely bordered; lateral margins broadly widened; anterior angles broadly rounded; lateral fovea deeply, coarsely and densely punctate; midline extremely shallow to absent. Punctation of pronotum coarse and dense, disc distinctly doubly punctate. Pronotal horn well developed, straight, narrowly and weakly upturned in lateral aspect, without subapical tooth ventrally.
</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BF9B5F2B5F9C7" blockId="1.[151,1437,945,2015]" box="[199,676,1557,1583]" pageId="1" pageNumber="340">Scutellum cordate, transverse, impunctate.</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BF999F318F957" blockId="1.[151,1437,945,2015]" pageId="1" pageNumber="340">
Elytra (
<figureCitation id="135E2A088A65FFAEFED7F999F197F9BB" box="[283,390,1593,1619]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1671,1696]" captionTargetBox="[177,1411,244,1633]" captionTargetId="figure@2.[164,1423,234,1644]" captionTargetPageId="2" captionText="FIGURES 1–3. Ceratophyus amdoensis Král &amp; Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3694898" httpUri="https://zenodo.org/record/3694898/files/figure.png" pageId="1" pageNumber="340">Figs. 1, 3</figureCitation>
) distinctly longer than wide, maximum width posterior of middle; surface moderately shiny; between suture and humerus with 7 striae, all of them well impressed along entire length of elytron, stria 5 distinctly shortened before base; intervals convex, almost impunctate; humeral umbone distinct; base of epipleura in area of humeral umbone distinctly narrowed to apex of elytron.
</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BF969F17BF90B" blockId="1.[151,1437,945,2015]" box="[199,362,1737,1763]" pageId="1" pageNumber="340">Macropterous.</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BF94DF31FF8A7" blockId="1.[151,1437,945,2015]" pageId="1" pageNumber="340">
Legs (
<figureCitation id="135E2A088A65FFAEFEC0F94DF166F8EF" box="[268,375,1773,1799]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1671,1696]" captionTargetBox="[177,1411,244,1633]" captionTargetId="figure@2.[164,1423,234,1644]" captionTargetPageId="2" captionText="FIGURES 1–3. Ceratophyus amdoensis Král &amp; Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3694898" httpUri="https://zenodo.org/record/3694898/files/figure.png" pageId="1" pageNumber="340">Figs. 1–2</figureCitation>
). Femora unarmed, glabrous, impunctate, with two macrosetaceous transverse carinae. Protibia with 8 external teeth gradually diminishing proximally, apical tooth distinctly bifid, ventromedial edge unarmed; meso- and metatibiae with 3 transverse external carinae.
</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BF8F9F230F87F" blockId="1.[151,1437,945,2015]" pageId="1" pageNumber="340">
<emphasis id="B911EA9F8A65FFAEFF0BF8F9F163F89B" box="[199,370,1881,1907]" italics="true" pageId="1" pageNumber="340">Ventral surface</emphasis>
(
<figureCitation id="135E2A088A65FFAEFE4FF8F9F1D8F89B" box="[387,457,1881,1907]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1671,1696]" captionTargetBox="[177,1411,244,1633]" captionTargetId="figure@2.[164,1423,234,1644]" captionTargetPageId="2" captionText="FIGURES 1–3. Ceratophyus amdoensis Král &amp; Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3694898" httpUri="https://zenodo.org/record/3694898/files/figure.png" pageId="1" pageNumber="340">Fig. 2</figureCitation>
). Abdominal ventrites scabrous, coarsely and densely punctate, with short, recumbent, greyish to reddish colored setation.
</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BF801F3B9F853" blockId="1.[151,1437,945,2015]" box="[199,936,1953,1979]" pageId="1" pageNumber="340">
<emphasis id="B911EA9F8A65FFAEFF0BF801F162F853" box="[199,371,1953,1979]" italics="true" pageId="1" pageNumber="340">Male genitalia.</emphasis>
Parameres of characteristic shape, as in
<figureCitation id="135E2A088A65FFAEFCF4F801F3B5F853" box="[824,932,1953,1979]" captionStart="FIGURES 4–12" captionStartId="3.[151,264,1811,1836]" captionTargetBox="[184,1353,211,1675]" captionTargetId="figure@3.[155,1429,178,1788]" captionTargetPageId="3" captionText="FIGURES 4–12. Aedeagus of Ceratophyus Fischer von Waldheim, 1824 species. 4–6, Ceratophyus amdoensis Král &amp; Jákl, new species, holotype; 7–9, C. dauricus (Jekel, 1866) (“Transbaikalia”, NMPC); 10–12, C. kabaki Nikolajev, 2007 (Sichuan, Yading, NMPC). 4, 7, 10, dorsal view; 5, 8, 11, left lateral view; 6, 9, 12, tips, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3694900" httpUri="https://zenodo.org/record/3694900/files/figure.png" pageId="1" pageNumber="340">Figs. 4–6</figureCitation>
.
</paragraph>
<paragraph id="8BDA368D8A65FFAEFF0BF864F296F837" blockId="1.[151,1437,945,2015]" box="[199,647,1988,2015]" pageId="1" pageNumber="340">
<emphasis id="B911EA9F8A65FFAEFF0BF864F1A6F837" bold="true" box="[199,439,1988,2015]" pageId="1" pageNumber="340">Sexual dimorphism.</emphasis>
Female unknown.
</paragraph>
<paragraph id="8BDA368D8A66FFADFF0BFF39F113FF3F" blockId="2.[151,1436,152,215]" pageId="2" pageNumber="341">
<emphasis id="B911EA9F8A66FFADFF0BFF39F16FFF5B" bold="true" box="[199,382,153,179]" pageId="2" pageNumber="341">Measurements.</emphasis>
Total body length:
<typeStatus id="54DE882F8A66FFADFDADFF39F2DDFF5B" box="[609,716,153,179]" pageId="2" pageNumber="341" type="holotype">Holotype</typeStatus>
, 23.0 mm;
<typeStatus id="54DE882F8A66FFADFC9DFF39F3A0FF5B" box="[849,945,153,179]" pageId="2" pageNumber="341" type="paratype">paratype</typeStatus>
(Shandan),
<quantity id="4C9D9B688A66FFADFBF2FF38F4B7FF5B" box="[1086,1190,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="2" pageNumber="341" unit="mm" value="18.5">18.5 mm</quantity>
,
<typeStatus id="54DE882F8A66FFADFB78FF39F505FF5B" box="[1204,1300,153,179]" pageId="2" pageNumber="341" type="paratype">paratype</typeStatus>
(Heimahe),
<quantity id="4C9D9B688A66FFADFF5BFF1CF0EFFF3F" box="[151,254,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="2" pageNumber="341" unit="mm" value="21.5">21.5 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF1A66058A66FFADFF5BF927F1D0F92C" ID-DOI="http://doi.org/10.5281/zenodo.3694898" ID-Zenodo-Dep="3694898" httpUri="https://zenodo.org/record/3694898/files/figure.png" pageId="2" pageNumber="341" startId="2.[151,264,1671,1696]" targetBox="[177,1411,244,1633]" targetPageId="2">
<paragraph id="8BDA368D8A66FFADFF5BF927F1D0F92C" blockId="2.[151,1436,1671,1732]" pageId="2" pageNumber="341">
<emphasis id="B911EA9F8A66FFADFF5BF927F12AF948" bold="true" box="[151,315,1671,1696]" pageId="2" pageNumber="341">FIGURES 1–3.</emphasis>
<taxonomicName id="4C654D0E8A66FFADFE8EF928F2A7F948" authority="Kral &amp; Jakl" authorityName="Kral &amp; Jakl" authorityYear="2020" box="[322,694,1671,1696]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="amdoensis" status="sp. nov.">
<emphasis id="B911EA9F8A66FFADFE8EF928F224F948" box="[322,565,1672,1696]" italics="true" pageId="2" pageNumber="341">Ceratophyus amdoensis</emphasis>
Král &amp; Jákl
</taxonomicName>
,
<taxonomicNameLabel id="A22257E48A66FFADFD0DF929F32FF948" box="[705,830,1671,1696]" pageId="2" pageNumber="341" rank="species">new species</taxonomicNameLabel>
, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale.
</paragraph>
</caption>
<subSubSection id="C37F65068A66FFADFF0BF952F244F8D8" pageId="2" pageNumber="341" type="discussion">
<paragraph id="8BDA368D8A66FFADFF0BF952F244F8D8" blockId="2.[151,1436,1778,2021]" pageId="2" pageNumber="341">
<emphasis id="B911EA9F8A66FFADFF0BF952F15CF8E5" bold="true" box="[199,333,1778,1805]" pageId="2" pageNumber="341">Variability.</emphasis>
In small
<typeStatus id="54DE882F8A66FFADFE7BF953F206F8E5" box="[439,535,1779,1805]" pageId="2" pageNumber="341" type="paratype">paratype</typeStatus>
from Shandan head horn short, more or less straight, and pronotal horn less developed, much shorter than in
<typeStatus id="54DE882F8A66FFADFE20F8B6F243F8D8" box="[492,594,1814,1840]" pageId="2" pageNumber="341" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F65068A66FFADFF0BF89AF320F8BC" box="[199,817,1850,1877]" pageId="2" pageNumber="341" type="diagnosis">
<paragraph id="8BDA368D8A66FFADFF0BF89AF320F8BC" blockId="2.[151,1436,1778,2021]" box="[199,817,1850,1877]" pageId="2" pageNumber="341">
<emphasis id="B911EA9F8A66FFADFF0BF89AF1DFF8BD" bold="true" box="[199,462,1850,1877]" pageId="2" pageNumber="341">Differential diagnosis.</emphasis>
Refer to the species key below.
</paragraph>
</subSubSection>
<subSubSection id="C37F65068A66FFADFF0BF8FFF0F3F828" pageId="2" pageNumber="341" type="etymology">
<paragraph id="8BDA368D8A66FFADFF0BF8FFF0F3F828" blockId="2.[151,1436,1778,2021]" pageId="2" pageNumber="341">
<emphasis id="B911EA9F8A66FFADFF0BF8FFF15FF891" bold="true" box="[199,334,1887,1913]" pageId="2" pageNumber="341">Etymology.</emphasis>
Toponymic; an adjective derived from Amdo, the name of one of three traditional regions of
<collectingRegion id="49A1F86F8A66FFADFAAEF8FEF58DF890" box="[1378,1436,1886,1912]" country="China" name="Xizang" pageId="2" pageNumber="341">Tibet</collectingRegion>
(nowadays approximately consisting of the
<collectingRegion id="49A1F86F8A66FFADFD4DF822F2CAF874" box="[641,731,1922,1948]" country="China" name="Qinghai" pageId="2" pageNumber="341">Qinghai</collectingRegion>
and western
<collectingRegion id="49A1F86F8A66FFADFCBCF823F3A9F875" box="[880,952,1923,1949]" country="China" name="Gansu" pageId="2" pageNumber="341">Gansu</collectingRegion>
Provinces, where the new species was collected.
</paragraph>
</subSubSection>
<subSubSection id="C37F65068A66FFADFF0BF86AF34FF80D" box="[199,862,1994,2021]" pageId="2" pageNumber="341" type="distribution">
<paragraph id="8BDA368D8A66FFADFF0BF86AF34FF80D" blockId="2.[151,1436,1778,2021]" box="[199,862,1994,2021]" pageId="2" pageNumber="341">
<emphasis id="B911EA9F8A66FFADFF0BF86AF170F80D" bold="true" box="[199,353,1994,2021]" pageId="2" pageNumber="341">Distribution.</emphasis>
<collectingCountry id="F372761D8A66FFADFEABF86AF1BBF80C" box="[359,426,1994,2020]" name="China" pageId="2" pageNumber="341">China</collectingCountry>
(
<collectingRegion id="49A1F86F8A66FFADFE75F86BF214F80D" box="[441,517,1995,2021]" country="China" name="Gansu" pageId="2" pageNumber="341">Gansu</collectingRegion>
,
<collectingRegion id="49A1F86F8A66FFADFDDCF86AF27EF80D" box="[528,623,1994,2021]" country="China" name="Qinghai" pageId="2" pageNumber="341">Qinghai</collectingRegion>
); see map on
<figureCitation id="135E2A088A66FFADFCCBF86AF34BF80D" box="[775,858,1994,2021]" captionStart="FIGURE 19" captionStartId="7.[151,250,1344,1369]" captionTargetBox="[151,1436,410,1318]" captionTargetId="figure@7.[151,1436,410,1318]" captionTargetPageId="7" captionText="FIGURE 19. Sketch map of China with marked distribution of Ceratophyus Fischer von Waldheim, 1824 species." figureDoi="http://doi.org/10.5281/zenodo.3694906" httpUri="https://zenodo.org/record/3694906/files/figure.png" pageId="2" pageNumber="341">Fig. 19</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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