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<document id="BDC3B1C18239CBB48B619C76DFC6E389" ID-DOI="10.5281/zenodo.12521148" ID-ISSN="1407-8619" ID-Zenodo-Dep="12521148" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719252154146" checkinUser="felipe" docAuthor="Rukmane, Anita" docDate="2019" docId="03CC878DFFD3DB40DC0B503EFEF6177D" docLanguage="en" docName="BalticJColeopterol.19.2.141-150.pdf" docOrigin="Baltic Journal of Coleopterology 19 (2)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.3:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="3" docTitle="Pachyrhynchus tetramaculatus Rukmane, 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="144" masterDocId="FFF5FFF5FFD1DB43DD245341FFCD152A" masterDocTitle="Four new species and two subspecies of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae: Pachyrhynchini) from Luzon Island, Philippines" masterLastPageNumber="150" masterPageNumber="141" pageNumber="143" updateTime="1719254198936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="4997483CA5B03B3BA5EC6A64E911418A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8B53548E7010C2BE690A9F583C088221">
<mods:title id="5DF5D94BF129AFA1B15AEF142571A48F">Four new species and two subspecies of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae: Pachyrhynchini) from Luzon Island, Philippines</mods:title>
</mods:titleInfo>
<mods:name id="6FA975F856FB3E12361A936893E6832A" type="personal">
<mods:role id="96E11C3F5ABCE7AD71C196312BB92349">
<mods:roleTerm id="CDE4DE56CF53041A62C462E7C7E9FD5D">Author</mods:roleTerm>
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<mods:namePart id="024F9C2469262546771BF3EDFA186656">Rukmane, Anita</mods:namePart>
</mods:name>
<mods:typeOfResource id="6234BBEA4D2571F8F55C8AF9B5A270DB">text</mods:typeOfResource>
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<mods:title id="57C0385941104B6C61E0198726413BB7">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="963782C95EF7C3AF3E4FB02AEF7EF77F">
<mods:date id="D49DC1F2598B04DE122FC6ED7CB31BF9">2019</mods:date>
<mods:detail id="56FF0C25088F3B8D40A3B58F830CA84D" type="pubDate">
<mods:number id="0627E23E72914D38F24A775BF228CB33">2019-12-31</mods:number>
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<mods:detail id="94C486F64C7872E7FE6ABD9BCAF0F1D4" type="volume">
<mods:number id="58B56301AFA3E41F2E440A29F401550B">19</mods:number>
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<mods:number id="827ADEA8A2F67FCCE680BB447EEBAFFD">2</mods:number>
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<mods:classification id="2780F2AF382F201FEBFC8BE4E9B0FB74">journal article</mods:classification>
<mods:identifier id="C79F23D5DE272B0F5ADA50A58A774B4B" type="DOI">10.5281/zenodo.12521148</mods:identifier>
<mods:identifier id="5F265FB3DBDF794F9415A0EF72C9408B" type="ISSN">1407-8619</mods:identifier>
<mods:identifier id="7751CD7054E32DC9442FE61E67C87086" type="Zenodo-Dep">12521148</mods:identifier>
</mods:mods>
<treatment id="03CC878DFFD3DB40DC0B503EFEF6177D" LSID="urn:lsid:plazi:treatment:03CC878DFFD3DB40DC0B503EFEF6177D" httpUri="http://treatment.plazi.org/id/03CC878DFFD3DB40DC0B503EFEF6177D" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143">
<subSubSection id="C37F6510FFD3DB41DC0B503EFD1416BD" box="[303,729,895,919]" pageId="2" pageNumber="143" type="nomenclature">
<paragraph id="8BDA369BFFD3DB41DC0B503EFD1416BD" blockId="2.[303,729,895,951]" box="[303,729,895,919]" pageId="2" pageNumber="143">
<heading id="D09281F7FFD3DB41DC0B503EFD1416BD" bold="true" box="[303,729,895,919]" fontSize="10" level="2" pageId="2" pageNumber="143" reason="4">
<emphasis id="B911EA89FFD3DB41DC0B503EFD1416BD" bold="true" box="[303,729,895,919]" pageId="2" pageNumber="143">
<taxonomicName id="4C654D18FFD3DB41DC0B503EFD4F16BD" authorityName="Rukmane" authorityYear="2019" box="[303,642,895,919]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov.">
<emphasis id="B911EA89FFD3DB41DC0B503EFD4F16BD" bold="true" box="[303,642,895,919]" italics="true" pageId="2" pageNumber="143">Pachyrhynchus tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F2FFD3DB41DFAC503EFD1416BD" box="[648,729,895,919]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD3DB41DC0B50DEFE2E169D" box="[303,483,927,951]" pageId="2" pageNumber="143" type="description">
<paragraph id="8BDA369BFFD3DB41DC0B50DEFE2E169D" blockId="2.[303,729,895,951]" box="[303,483,927,951]" pageId="2" pageNumber="143">
<figureCitation id="135E2A1EFFD3DB41DC0B50DEFEA8169D" box="[303,357,927,951]" captionStart="Fig" captionStartId="4.[303,344,1548,1572]" captionTargetBox="[334,1313,210,1406]" captionTargetId="figure-14@4.[323,1333,209,1459]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus of the following species: A – P. tetramaculatus sp. nov. male; B – P. tetramaculatus sp. nov. female; C – P.sagittatus sp. nov. male; D – P. kirklayroni sp. nov. male; P. congestus ssp. aedamlayroni subsp. nov. male; E - P.congestus ssp. aedamlayroni subsp. nov female" figureDoi="http://doi.org/10.5281/zenodo.12521150" httpUri="https://zenodo.org/record/12521150/files/figure.png" pageId="2" pageNumber="143">Fig. 1</figureCitation>
A-B, 4L-M.
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD3DB41DC0B509EFD84101D" pageId="2" pageNumber="143" type="materials_examined">
<paragraph id="8BDA369BFFD3DB41DC0B509EFD8411BD" blockId="2.[303,813,991,1175]" pageId="2" pageNumber="143">
<emphasis id="B911EA89FFD3DB41DC0B509EFE0116DD" bold="true" box="[303,460,991,1015]" pageId="2" pageNumber="143">Type material.</emphasis>
<materialsCitation id="3B0D3CC6FFD3DB41DCF6509EFCE5111D" collectingDate="2018-06" country="Philippines" location="Luzon" pageId="2" pageNumber="143" specimenCount="1" stateProvince="Nueva Vizcaya" typeStatus="holotype">
<typeStatus id="54DE8839FFD3DB41DCF6509EFDFF16DD" box="[466,562,991,1015]" pageId="2" pageNumber="143" type="holotype">Holotype</typeStatus>
. Male. ‘’
<collectingCountry id="F372760BFFD3DB41DFA5509EFCEC16DD" box="[641,801,991,1015]" name="Philippines" pageId="2" pageNumber="143">PHILIPPINES</collectingCountry>
/
<location id="8EBA6040FFD3DB41DC0B5741FEBB113D" LSID="urn:lsid:plazi:treatment:03CC878DFFD3DB40DC0B503EFEF6177D:8EBA6040FFD3DB41DC0B5741FEBB113D" box="[303,374,1024,1047]" country="Philippines" name="Luzon" pageId="2" pageNumber="143" stateProvince="Nueva Vizcaya">Luzon</location>
,
<collectingRegion id="49A1F879FFD3DB41DC5A5741FDD3113D" box="[382,542,1023,1047]" country="Philippines" name="Nueva Vizcaya" pageId="2" pageNumber="143">Nueva Vizcaya</collectingRegion>
,
<collectingRegion id="49A1F879FFD3DB41DF0C50BEFDB7113D" box="[552,634,1023,1047]" country="Philippines" name="Quezon" pageId="2" pageNumber="143">Quezon</collectingRegion>
/
<date id="FFDB105BFFD3DB41DFAF5741FD25113D" box="[651,744,1023,1047]" pageId="2" pageNumber="143" value="2018-06">
<collectingDate id="EF9FE9B3FFD3DB41DFAF5741FD25113D" box="[651,744,1023,1047]" pageId="2" pageNumber="143" value="2018-06">VI. 2018</collectingDate>
</date>
/ local collector leg.’’ (white rectangular label, printed)
</materialsCitation>
; ‘’
<typeStatus id="54DE8839FFD3DB41DC13577EFE03117D" box="[311,462,1087,1111]" pageId="2" pageNumber="143" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C654D18FFD3DB41DCC4577EFCE0117D" authorityName="Rukmane" authorityYear="2019" box="[480,813,1087,1111]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="tetramaculatus">
<emphasis id="B911EA89FFD3DB41DCC4577EFCE0117D" box="[480,813,1087,1111]" italics="true" pageId="2" pageNumber="143">Pachyrhynchus tetramaculatus</emphasis>
</taxonomicName>
/ Rukmane, 2019 / det. Rukmane A. 2019’’ (red rectangular label, printed).
</paragraph>
<paragraph id="8BDA369BFFD3DB41DC0B57FEFD84101D" blockId="2.[303,813,1215,1335]" pageId="2" pageNumber="143">
Female.
<typeStatus id="54DE8839FFD3DB41DCB35781FDCD11F2" box="[407,512,1216,1240]" pageId="2" pageNumber="143" type="paratype">Paratype</typeStatus>
. Same label as
<typeStatus id="54DE8839FFD3DB41DFE657FEFCE411FD" box="[706,809,1215,1239]" pageId="2" pageNumber="143" type="holotype">holotype</typeStatus>
. ‘’
<typeStatus id="54DE8839FFD3DB41DC12579EFE0811DD" box="[310,453,1247,1271]" pageId="2" pageNumber="143" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C654D18FFD3DB41DCF8579EFCE011DD" authorityName="Rukmane" authorityYear="2019" box="[476,813,1247,1271]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="tetramaculatus">
<emphasis id="B911EA89FFD3DB41DCF8579EFCE011DD" box="[476,813,1247,1271]" italics="true" pageId="2" pageNumber="143">Pachyrhynchus tetramaculatus</emphasis>
</taxonomicName>
/ Rukmane, 2019 / det. Rukmane A. 2019’’ (red rectangular label, printed).
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD3DB41DC0B561EFEAC10BD" pageId="2" pageNumber="143" type="distribution">
<paragraph id="8BDA369BFFD3DB41DC0B561EFEAC10BD" blockId="2.[303,813,1375,1431]" pageId="2" pageNumber="143">
<emphasis id="B911EA89FFD3DB41DC0B561EFE73105D" bold="true" box="[303,446,1375,1399]" pageId="2" pageNumber="143">Distribution:</emphasis>
Luzon Island,
<collectingRegion id="49A1F879FFD3DB41DF725621FE9010BD" country="Philippines" name="Nueva Vizcaya" pageId="2" pageNumber="143">Nueva Vizcaya Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD3DB40DC0B56FEFEF6177D" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143" type="description">
<paragraph id="8BDA369BFFD3DB41DC0B56FEFCE810DD" blockId="2.[303,814,1471,1527]" pageId="2" pageNumber="143">
<emphasis id="B911EA89FFD3DB41DC0B56FEFE7C10FD" bold="true" box="[303,433,1471,1495]" pageId="2" pageNumber="143">Description.</emphasis>
Dimensions (
<typeStatus id="54DE8839FFD3DB41DF1E56FEFD5B10FD" box="[570,662,1471,1495]" pageId="2" pageNumber="143" type="holotype">holotype</typeStatus>
): LB: 13.9; LE: 8.8; WE: 5.3; LP: 3.9; WP: 3.7; LR: 2.0; WR: 1.9.
</paragraph>
<paragraph id="8BDA369BFFD3DB41DC0B5561FD5213FD" blockId="2.[303,813,1567,1751]" pageId="2" pageNumber="143">
Integument black, strongly shiny, pronotum, head and legs shiny, underside with weaker luster; body with pale orange to yellow round recumbent scales; genae, coxa and femori with pale orange, yellow and shimmery green scales. Dorsal habitus as shown in
<figureCitation id="135E2A1EFFD3DB41DF0F55FEFDA813FD" box="[555,613,1727,1751]" captionStart="Fig" captionStartId="4.[303,344,1548,1572]" captionTargetBox="[334,1313,210,1406]" captionTargetId="figure-14@4.[323,1333,209,1459]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus of the following species: A – P. tetramaculatus sp. nov. male; B – P. tetramaculatus sp. nov. female; C – P.sagittatus sp. nov. male; D – P. kirklayroni sp. nov. male; P. congestus ssp. aedamlayroni subsp. nov. male; E - P.congestus ssp. aedamlayroni subsp. nov female" figureDoi="http://doi.org/10.5281/zenodo.12521150" httpUri="https://zenodo.org/record/12521150/files/figure.png" pageId="2" pageNumber="143">Fig. 1</figureCitation>
A-B.
</paragraph>
<paragraph id="8BDA369BFFD3DB41DE775381FA8817DD" blockId="2.[851,1361,191,760]" pageId="2" pageNumber="143">Rostrum slightly wider than long (LR/WR 1.05), slightly pubescent, bulging on apical part, with peak slightly before middle, ovate-triangular impression on basal half, weak longitudinal groove from middle to base of the forehead, two bulges on basal 1/3 and weak transverse impression on base; laterally with patch of round scaled on genae; few pubescence before antennal scrobes and long light hairs after antennal scrobes, with longer hairs near apex; antennomers mingled with long light hairs, scape covered with pubescence on basal part and few light hairs on apical part dorsally; pedicel and first antennomer sub equal in length, longer than wide, antennomers II-V subsphereical, sub equal in length. Head glabrous, finely punctured; eyes moderate, weakly prominent from the outline of the head. Forehead flat, more than two times as wide as eye width.</paragraph>
<paragraph id="8BDA369BFFD3DB41DE77505EFA88115D" blockId="2.[851,1361,799,1143]" pageId="2" pageNumber="143">Prothorax sub spherical, longer than wide (LP/ WP 1.05), widest at apical 1/2; weakly punctured; with the following markings of pale orange to yellow scales: 1) two roundish patches on disc laterally along apical margin; two ovate patches on basal part of the disc, redirected laterally; ovate patch on each latero-dorsal part; dorsal contour increased from moderate apex to peak at apical ½, then firmly rounded and decreased to just after the midline; then decreased strongly, with curve, sub basal margin slightly widened.</paragraph>
<paragraph id="8BDA369BFFD3DB41DE7757E1FB3A105D" blockId="2.[851,1361,1183,1399]" pageId="2" pageNumber="143">Legs stout; coxa patch of orange and shimmery green round scales; femur mingled with sparse pubescence and patch of scales on apical part along internal margin; tibiae incurved apically, covered with pubescence on all length and long light hairs on internal margin, with mucrones on all legs; tarsus with long, golden setae.</paragraph>
<paragraph id="8BDA369BFFD3DB41DE7756DEFA9C13FD" blockId="2.[851,1361,1439,1751]" pageId="2" pageNumber="143">Elytra sub ovate (LE/WE 1.66), with weakly pronounced intervals, nearly smooth; each elytron with the following scaly markings: 1) two ovate patches on basal ½; 2) four ovate spots on medial part; 3) two sutural patches – one sub medial, one near apex; 4) three ovate patches on apical part 5) triangular apical patch along apex; widest just in the middle; on dorsal dimension narrow at the base, gradually increases to middle, widest just in the middle, then roundish and</paragraph>
<paragraph id="8BDA369BFFD2DB40DF9E53D0FCE3158E" blockId="3.[698,814,145,164]" box="[698,814,145,164]" pageId="3" pageNumber="144">
<emphasis id="B911EA89FFD2DB40DF9E53D0FCE3158E" box="[698,814,145,164]" italics="true" pageId="3" pageNumber="144">Rukmane A.</emphasis>
</paragraph>
<paragraph id="8BDA369BFFD2DB40DDC753FEFD8E143D" blockId="3.[227,737,191,279]" pageId="3" pageNumber="144">gradually decreases to apex up to apical 1/3 where narrows more strongly in direction to apex; apex rounded, with weak pubescence.</paragraph>
<paragraph id="8BDA369BFFD2DB40DDC7527EFDF014BD" blockId="3.[227,737,319,407]" pageId="3" pageNumber="144">
Elytra wider than prothorax (WE/WP 1.43), more than twice as long as prothorax (LE/LP 2.26). Genitalia as shown in
<figureCitation id="135E2A1EFFD2DB40DCEE523EFDC814BD" box="[458,517,383,407]" captionStart="Fig" captionStartId="7.[227,267,1283,1307]" captionTargetBox="[239,1246,229,1199]" captionTargetId="figure-4@7.[234,1313,209,1209]" captionTargetPageId="7" captionText="Fig. 4. A, C, H, J, L – aedeagus in lateral view; B, D, I, K, M – aedeagus in ventral view; A, B – P.disargus sp. nov.; C, D – P.congestus ssp. aedamlayroni subsp. nov.; H, I – P.kirklayroni sp. nov.; J, K – P.sagittatus sp. nov.; L, M – P.tetramaculatus sp.nov.; E, F, G – female genitalia of P.phaleratus ssp. dannylayroni subsp. nov.; E – sternite VIII in ventral view; F – ovipositor in dorsal view; G – spermatheca. Scale bar 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.12521156" httpUri="https://zenodo.org/record/12521156/files/figure.png" pageId="3" pageNumber="144">Fig. 4</figureCitation>
L-M.
</paragraph>
<paragraph id="8BDA369BFFD2DB40DDC752FEFEF6177D" blockId="3.[227,738,447,599]" pageId="3" pageNumber="144">
<emphasis id="B911EA89FFD2DB40DDC752FEFEFA14FD" bold="true" box="[227,311,447,471]" pageId="3" pageNumber="144">Female.</emphasis>
Dimensions: LB: 15.3; LE: 10.3; WE: 7.4; LP: 4.7; WP: 4.3; LR: 2.1; WR: 2.4. Larger than male. Elytra more wide and more strongly rounded, as well as more strongly elongate apically.
</paragraph>
</subSubSection>
</treatment>
</document>
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