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<document id="A2EC494B5CDD2FC8A06B8B7CBF9DAB24" ID-DOI="10.11646/zootaxa.5443.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="11064278" ID-ZooBank="A91A8D67-A81B-427D-8096-59E41326A8E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714028852870" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2024" docId="03CB87B7E956FFFBF2CFFC86FF14FF2F" docLanguage="en" docName="zootaxa.5443.4.2.pdf" docOrigin="Zootaxa 5443 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Octavius kambergensis Janak 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="505" masterDocId="FFF2FFCFE95EFFF1F258FF9CFFC6FFD6" masterDocTitle="On Octavius from Maloti-Drakensberg and Golden Gate National Parks, South Africa (Coleoptera: Staphylinidae: Euaesthetinae)" masterLastPageNumber="522" masterPageNumber="495" pageNumber="503" updateTime="1714029516812" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C54BA88A558B900502666C6B3CF135B4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="84E1EC5DF5DF9E6474B5908FC7E18A5C">
<mods:title id="FE8F533A21420257602F5E59427AFB42">On Octavius from Maloti-Drakensberg and Golden Gate National Parks, South Africa (Coleoptera: Staphylinidae: Euaesthetinae)</mods:title>
</mods:titleInfo>
<mods:name id="6B3F7C6D314CAF58825CBEB7FF32ED40" type="personal">
<mods:role id="40DEE9BB5ADCED57F628D905A471EA26">
<mods:roleTerm id="B33221BE0CAA6B1192C1E2D2E78FC284">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FCAD556D5155025528DD537F96569DA2">Janák, Jiří</mods:namePart>
</mods:name>
<mods:typeOfResource id="F9A48098CB0D0D852CE8F5277656418D">text</mods:typeOfResource>
<mods:relatedItem id="CA8B7A39B3560CDFEAE35D5C59A27069" type="host">
<mods:titleInfo id="64134EF36593E15176F1519668C6F36B">
<mods:title id="2E518D215F4D11E52BCC7C740A08CF2D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1652DF334D616815AEBBA21FFE482840">
<mods:date id="BFB26E65B191AE861EB2337144069227">2024</mods:date>
<mods:detail id="D62C0CEEE82384FC4AA9B8BB1178DE24" type="pubDate">
<mods:number id="D92E1439D9820B9389730E37BEA7028C">2024-04-24</mods:number>
</mods:detail>
<mods:detail id="1DEA38D570159E7A1386D592C8F55E74" type="volume">
<mods:number id="2E705315FEB5FACB8D88D11A319C70FF">5443</mods:number>
</mods:detail>
<mods:detail id="1EC240A72089EC3EC894167458F2169D" type="issue">
<mods:number id="EB04908D3F1B12C59B8327EB208C81F2">4</mods:number>
</mods:detail>
<mods:extent id="741EA03BD7F7665A1EBA0CB78E8C7154" unit="page">
<mods:start id="3A7F2DCDCF32DE8F823471DA64AF94F8">495</mods:start>
<mods:end id="88D8289D988F0A9B2BDDB6601DE8B549">522</mods:end>
</mods:extent>
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<mods:location id="5B00B69CC53A7605CA6C872C8DA8EC0E">
<mods:url id="EF4CFA42904A6D1A8F50579FEC0E2E46">http://dx.doi.org/10.11646/zootaxa.5443.4.2</mods:url>
</mods:location>
<mods:classification id="9B1A21F9586C13D8D06695BF72A977E0">journal article</mods:classification>
<mods:identifier id="2D53AF4B9189C8FBE879F1A6CD8D25CA" type="DOI">10.11646/zootaxa.5443.4.2</mods:identifier>
<mods:identifier id="E422B14A9D347E4B4BDB716C8CD46DEE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BD25E2E3253006686C8953CD72E07012" type="Zenodo-Dep">11064278</mods:identifier>
<mods:identifier id="B1DE3B23077157A337ACF536F3E50DBE" type="ZooBank">A91A8D67-A81B-427D-8096-59E41326A8E3</mods:identifier>
</mods:mods>
<treatment id="03CB87B7E956FFFBF2CFFC86FF14FF2F" ID-DOI="http://doi.org/10.5281/zenodo.11064356" ID-Zenodo-Dep="11064356" LSID="urn:lsid:plazi:treatment:03CB87B7E956FFFBF2CFFC86FF14FF2F" httpUri="http://treatment.plazi.org/id/03CB87B7E956FFFBF2CFFC86FF14FF2F" lastPageId="10" lastPageNumber="505" pageId="8" pageNumber="503">
<subSubSection id="C378652AE956FFF9F2CFFC86FDA9FCE2" box="[151,623,793,820]" pageId="8" pageNumber="503" type="nomenclature">
<paragraph id="8BDD36A1E956FFF9F2CFFC86FDA9FCE2" blockId="8.[151,623,793,856]" box="[151,623,793,820]" pageId="8" pageNumber="503">
<heading id="D09581CDE956FFF9F2CFFC86FDA9FCE2" bold="true" box="[151,623,793,820]" fontSize="11" level="1" pageId="8" pageNumber="503" reason="1">
<emphasis id="B916EAB3E956FFF9F2CFFC86FDA9FCE2" bold="true" box="[151,623,793,820]" pageId="8" pageNumber="503">
<taxonomicName id="4C624D22E956FFF9F2CFFC86FDC2FCE2" authority="Janak" authorityName="Janak" authorityYear="2024" box="[151,516,793,820]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="kambergensis" status="sp. nov.">
<emphasis id="B916EAB3E956FFF9F2CFFC86FE68FCE2" bold="true" box="[151,430,793,820]" italics="true" pageId="8" pageNumber="503">Octavius kambergensis</emphasis>
Janák
</taxonomicName>
,
<taxonomicNameLabel id="A22557C8E956FFF9F049FC86FDA9FCE2" box="[529,623,794,820]" pageId="8" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C378652AE956FFF9F2CFFCA2FE4DFC8E" box="[151,395,830,856]" pageId="8" pageNumber="503" type="description">
<paragraph id="8BDD36A1E956FFF9F2CFFCA2FE4DFC8E" blockId="8.[151,623,793,856]" box="[151,395,830,856]" pageId="8" pageNumber="503">
<figureCitation id="13592A24E956FFF9F2CFFCA2FED8FC8E" box="[151,286,830,856]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="8" pageNumber="503">Figs. 39–43</figureCitation>
,
<figureCitation id="13592A24E956FFF9F371FCA2FE93FC8E" box="[297,341,830,856]" captionStart="FIGURES 113–121" captionStartId="23.[151,264,1358,1382]" captionTargetBox="[212,1375,191,1316]" captionTargetId="figure-15@23.[197,1390,181,1334]" captionTargetPageId="23" captionText="FIGURES 113–121. Octavius undulatus Janák, sp. nov.; 113–116, 118–119, holotype; 117, paratype. 113, habitus; 114, fore parts; 115, fore parts lateral; 116–117, aedeagus ventral; 118, male sternite VIII; 119, male sternite IX. Scale 0.5 mm: 113; scales 0.1 mm: 114–115; 116–117; 118–119; o: ommatidium. 120: Distribution of Octavius – subgroup A. 121: Distribution of Octavius – subgroup B." figureDoi="http://doi.org/10.5281/zenodo.11064300" httpUri="https://zenodo.org/record/11064300/files/figure.png" pageId="8" pageNumber="503">120</figureCitation>
,
<figureCitation id="13592A24E956FFF9F339FCA2FE4DFC8E" box="[353,395,830,856]" captionStart="FIGURES 128–133" captionStartId="25.[151,264,1651,1675]" captionTargetBox="[154,1434,182,1627]" captionTargetId="figure-15@25.[154,1434,181,1627]" captionTargetPageId="25" captionText="FIGURES 128–133. 128: Cathedral Peak, Masongwane river banks. 129: Injisuthi. 130: Monkʼs Cowl, Hlatikulu forest. 131: Lotheni. 132: Kamberg, locality 1. 133: Kamberg locality 2." figureDoi="http://doi.org/10.5281/zenodo.11064306" httpUri="https://zenodo.org/record/11064306/files/figure.png" pageId="8" pageNumber="503">133</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C378652AE956FFF9F2CFFC1AFDB6FB82" pageId="8" pageNumber="503" type="materials_examined">
<paragraph id="8BDD36A1E956FFF9F2CFFC1AFAA2FC76" blockId="8.[151,1437,901,2008]" box="[151,1380,901,928]" pageId="8" pageNumber="503">
<materialsCitation id="3B0A3CFCE956FFF9F2CFFC1AFAA6FC76" box="[151,1376,901,928]" country="South Africa" location="Kamberg National Reserve" municipality="Maloti-Drakensberg Park" pageId="8" pageNumber="503" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<emphasis id="B916EAB3E956FFF9F2CFFC1AFEF0FC49" bold="true" box="[151,310,901,928]" pageId="8" pageNumber="503">
<typeStatus id="54D98803E956FFF9F2CFFC1AFF14FC76" box="[151,210,902,928]" pageId="8" pageNumber="503">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3757631E956FFF9F364FC1AFE0BFC76" box="[316,461,902,928]" name="South Africa" pageId="8" pageNumber="503">South Africa</collectingCountry>
,
<collectingRegion id="49A6F843E956FFF9F381FC1AFD33FC76" box="[473,757,902,928]" country="South Africa" name="KwaZulu-Natal" pageId="8" pageNumber="503">KwaZulu-Natal Province</collectingRegion>
,
<collectingMunicipality id="6BB9ACDBE956FFF9F159FC1AFBE6FC76" box="[769,1056,902,928]" pageId="8" pageNumber="503">Maloti-Drakensberg Park</collectingMunicipality>
,
<location id="8EBD607AE956FFF9F674FC1AFAA6FC76" LSID="urn:lsid:plazi:treatment:03CB87B7E956FFFBF2CFFC86FF14FF2F:8EBD607AE956FFF9F674FC1AFAA6FC76" box="[1068,1376,902,928]" country="South Africa" municipality="Maloti-Drakensberg Park" name="Kamberg National Reserve" pageId="8" pageNumber="503" stateProvince="KwaZulu-Natal">Kamberg National Reserve</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDD36A1E956FFF9F29FFC36FDB6FB82" blockId="8.[151,1437,901,2008]" pageId="8" pageNumber="503">
<emphasis id="B916EAB3E956FFF9F29FFC36FEB3FC15" bold="true" box="[199,373,937,964]" pageId="8" pageNumber="503">Type material</emphasis>
(
<specimenCount id="9D64FD28E956FFF9F3D4FC36FDE8FC12" box="[396,558,938,964]" count="70" pageId="8" pageNumber="503" type="generic">70 specimens</specimenCount>
).
<emphasis id="B916EAB3E956FFF9F014FC36FD7FFC12" bold="true" box="[588,697,938,964]" pageId="8" pageNumber="503">
<typeStatus id="54D98803E956FFF9F014FC36FD7FFC12" box="[588,697,938,964]" pageId="8" pageNumber="503" type="holotype">Holotype</typeStatus>
</emphasis>
<materialsCitation id="3B0A3CFCE956FFF9F090FC36FADBFBDA" collectingDate="2019-01-12" collectionCode="TMSA" collectorName="J. Janak" country="South Africa" determinerName="J. Janak" elevation="40" location="Maloti-Drakensberg Park" pageId="8" pageNumber="503" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<specimenCount id="9D64FD28E956FFF9F090FC36FD26FC12" box="[712,736,938,964]" count="1" pageId="8" pageNumber="503" type="male">♂</specimenCount>
: “
<collectingCountry id="F3757631E956FFF9F0A5FC36FC5EFC12" box="[765,920,938,964]" name="South Africa" pageId="8" pageNumber="503">South Africa</collectingCountry>
,
<collectingRegion id="49A6F843E956FFF9F1F3FC36FB98FC12" box="[939,1118,938,964]" country="South Africa" name="KwaZulu-Natal" pageId="8" pageNumber="503">KwaZulu-Natal</collectingRegion>
,
<location id="8EBD607AE956FFF9F629FC36FA5EFC12" LSID="urn:lsid:plazi:treatment:03CB87B7E956FFFBF2CFFC86FF14FF2F:8EBD607AE956FFF9F629FC36FA5EFC12" box="[1137,1432,938,964]" country="South Africa" name="Maloti-Drakensberg Park" pageId="8" pageNumber="503" stateProvince="KwaZulu-Natal">Maloti-Drakensberg Park</location>
, Kamberg NR, ind. forest patches 29°21.85ʼS 29°37.1ʼE, 1820–
<quantity id="4C9A9B44E956FFF9F13BFC52FC51FC3E" box="[867,919,974,1000]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="503" unit="m" value="40.0">
<elevation id="004FD192E956FFF9F13BFC52FC51FC3E" box="[867,919,974,1000]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="503" unit="m" value="40.0">40m</elevation>
</quantity>
,
<date id="FFDC1061E956FFF9F1FBFC52FBC9FC3E" box="[931,1039,974,1000]" pageId="8" pageNumber="503" value="2019-01-12">
<collectingDate id="EF98E989E956FFF9F1FBFC52FBC9FC3E" box="[931,1039,974,1000]" pageId="8" pageNumber="503" value="2019-01-12">12.i.2019</collectingDate>
</date>
,
<collectorName id="26975377E956FFF9F643FC52FBB3FC3E" box="[1051,1141,974,1000]" pageId="8" pageNumber="503">J. Janák</collectorName>
lgt.”, “Berlese extraction, leaf &amp; log litter, sifting”, “
<typeStatus id="54D98803E956FFF9F398FC6EFDAFFBDA" box="[448,617,1010,1036]" pageId="8" pageNumber="503" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C624D22E956FFF9F029FC6EFCB3FBDA" authority="Janák, 2024" authorityName="Janák" authorityYear="2024" box="[625,885,1010,1036]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="kambergensis" status="sp. nov.">
<emphasis id="B916EAB3E956FFF9F029FC6EFCB3FBDA" box="[625,885,1010,1036]" italics="true" pageId="8" pageNumber="503">Octavius kambergensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557C8E956FFF9F125FC6EFC12FBDA" box="[893,980,1010,1036]" pageId="8" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="7E01DA65E956FFF9F183FC6EFBF2FBDA" box="[987,1076,1010,1036]" pageId="8" pageNumber="503">J. Janák</determinerName>
det. 2021” (
<collectionCode id="ED73AE64E956FFF9F698FC6EFAD2FBDA" box="[1216,1300,1010,1036]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="503">TMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B0A3CFCE956FFF9F770FC6EFDAAFB82" collectingDate="2019-01-12" collectionCode="TMSA, JJRC" collectorName="All" country="South Africa" determinerName="J. Janak" elevation="40" location="Maloti-Drakensberg Park" pageId="8" pageNumber="503" specimenCount="69" specimenCount-female="35" specimenCount-male="34" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<typeStatus id="54D98803E956FFF9F770FC6EFA5FFBDA" box="[1320,1433,1010,1036]" pageId="8" pageNumber="503" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D64FD28E956FFF9F2CFFB8AFF14FBE6" box="[151,210,1046,1072]" count="34" pageId="8" pageNumber="503" type="male">34 ♂</specimenCount>
,
<specimenCount id="9D64FD28E956FFF9F2B8FB8AFEDDFBE6" box="[224,283,1046,1073]" count="35" pageId="8" pageNumber="503" type="female">35 ♀</specimenCount>
: same data as holotype (
<collectionCode id="ED73AE64E956FFF9F06BFB8AFD43FBE6" box="[563,645,1046,1072]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="503">TMSA</collectionCode>
,
<collectionCode id="ED73AE64E956FFF9F0C9FB8AFD17FBE6" box="[657,721,1046,1072]" pageId="8" pageNumber="503">JJRC</collectionCode>
).
<collectorName id="26975377E956FFF9F0BBFB8AFCC1FBE6" box="[739,775,1046,1072]" pageId="8" pageNumber="503">All</collectorName>
<typeStatus id="54D98803E956FFF9F157FB8AFCBCFBE6" box="[783,890,1046,1072]" pageId="8" pageNumber="503" type="paratype">paratypes</typeStatus>
with additional labels: “
<typeStatus id="54D98803E956FFF9F6C8FB8AFAF7FBE6" box="[1168,1329,1046,1072]" pageId="8" pageNumber="503" type="paratype">PARATYPUS</typeStatus>
<taxonomicName id="4C624D22E956FFF9F761FB8AFEF6FB82" authority="Janák, 2024" authorityName="Janák" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="kambergensis" status="sp. nov.">
<emphasis id="B916EAB3E956FFF9F761FB8AFEF6FB82" italics="true" pageId="8" pageNumber="503">Octavius kambergensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557C8E956FFF9F36FFBA6FE48FB82" box="[311,398,1082,1108]" pageId="8" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="7E01DA65E956FFF9F3CCFBA6FE2BFB82" box="[404,493,1082,1108]" pageId="8" pageNumber="503">J. Janák</determinerName>
det. 2021”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C378652AE956FFF9F29FFBC1FD64F8F2" pageId="8" pageNumber="503" type="description">
<paragraph id="8BDD36A1E956FFF9F29FFBC1FE0FFB16" blockId="8.[151,1437,901,2008]" pageId="8" pageNumber="503">
<emphasis id="B916EAB3E956FFF9F29FFBC1FE9FFBA1" bold="true" box="[199,345,1117,1143]" pageId="8" pageNumber="503">Description.</emphasis>
Body length
<quantity id="4C9A9B44E956FFF9F3AEFBC2FDB8FBAE" box="[502,638,1118,1144]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="8" pageNumber="503" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.6–1.8 mm</quantity>
(M
<quantity id="4C9A9B44E956FFF9F0EAFBC2FCCAFBAE" box="[690,780,1118,1144]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="503" unit="mm" value="1.6">1.6 mm</quantity>
, HT
<quantity id="4C9A9B44E956FFF9F11FFBC2FC64FBAE" box="[839,930,1118,1144]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="503" unit="mm" value="1.7">1.7 mm</quantity>
), forebody length
<quantity id="4C9A9B44E956FFF9F62CFBC2FB3BFBAE" box="[1140,1277,1118,1144]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="8" pageNumber="503" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
(M
<quantity id="4C9A9B44E956FFF9F768FBC2FA5EFBAE" box="[1328,1432,1118,1145]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="503" unit="mm" value="0.75">0.75 mm</quantity>
, HT
<quantity id="4C9A9B44E956FFF9F29FFB1EFEE0FB4A" box="[199,294,1154,1180]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="503" unit="mm" value="0.8">0.8 mm</quantity>
). Macrophthalmous, apterous, light rusty, head dull, pronotum slightly shiny, elytra and abdomen moderately shiny (
<figureCitation id="13592A24E956FFF9F330FB3AFE7BFB16" box="[360,445,1190,1216]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="8" pageNumber="503">Fig. 39</figureCitation>
).
</paragraph>
<paragraph id="8BDD36A1E956FFF9F29FFB56FD2AFA86" blockId="8.[151,1437,901,2008]" pageId="8" pageNumber="503">
Head (
<figureCitation id="13592A24E956FFF9F349FB56FEA4FB32" box="[273,354,1226,1252]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="8" pageNumber="503">Fig. 39</figureCitation>
) slightly narrower than pronotum (R 0.88–0.94, M 0.91, HT 0.90), eyes relatively small, temples mostly more than twice as long as eyes (R 1.97–2.93, M 2.37, HT 2.11), sides of head distinctly roundly widened behind, posterior angles slightly angular, median impression on disc absent, lateral parts of head finely granulose, median part densely and moderately finely reticulate.
</paragraph>
<paragraph id="8BDD36A1E956FFF9F29FFAC6FBBCFA36" blockId="8.[151,1437,901,2008]" pageId="8" pageNumber="503">
Pronotum (
<figureCitation id="13592A24E956FFF9F31BFAC6FE55FAA2" box="[323,403,1370,1396]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="8" pageNumber="503">Fig. 39</figureCitation>
) slightly broader than long (R 1.09–1.18, M 1.13, HT 1.18), moderately narrowed posteriorly; anterior angles slightly angular, dorsal impressions deep, transverse impression deep, lateral impressions deep, delimited by very narrow longitudinal ridge laterally; lateral parts beside lateral impressions moderately densely and finely granulose, remainder of surface very densely and moderately finely reticulate.
</paragraph>
<paragraph id="8BDD36A1E956FFF9F29FFA76FEFEF99A" blockId="8.[151,1437,901,2008]" pageId="8" pageNumber="503">
Elytra (
<figureCitation id="13592A24E956FFF9F343FA76FEAAF9D2" box="[283,364,1514,1540]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="8" pageNumber="503">Fig. 39</figureCitation>
) subquadrate, much broader than long (R 1.36–1.50, M 1.43, HT 1.38), with sharp longitudinal ridge laterally; between latter and suture irregularly sparsely granulate, and shallowly, moderately reticulate, sparsely shortly setose.
</paragraph>
<paragraph id="8BDD36A1E956FFF9F29FF9CAFDB8F942" blockId="8.[151,1437,901,2008]" pageId="8" pageNumber="503">Abdomen subparallel, slightly widened to segment V, with one paratergite and microsculpture consisting of triangular or rhomboid fields, finely setose.</paragraph>
<paragraph id="8BDD36A1E956FFF9F29FF902FD64F8F2" blockId="8.[151,1437,901,2008]" pageId="8" pageNumber="503">
Male. Sternite VIII triangularly emarginated in posterior one-fifth (
<figureCitation id="13592A24E956FFF9F1ECF902FBC1F96E" box="[948,1031,1694,1720]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="8" pageNumber="503">Fig. 42</figureCitation>
), sternite IX as in
<figureCitation id="13592A24E956FFF9F68BF902FAE3F96E" box="[1235,1317,1694,1720]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="8" pageNumber="503">Fig. 43</figureCitation>
. Aedeagus slightly asymmetrical, moderately long and narrow (length
<quantity id="4C9A9B44E956FFF9F16FF95EFC1AF90A" box="[823,988,1730,1757]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="8" pageNumber="503" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
, M
<quantity id="4C9A9B44E956FFF9F652F95EFBB6F90A" box="[1034,1136,1730,1756]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="503" unit="mm" value="0.46">0.46 mm</quantity>
, HT
<quantity id="4C9A9B44E956FFF9F6F1F95EFAD4F90A" box="[1193,1298,1730,1757]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="503" unit="mm" value="0.45">0.45 mm</quantity>
), apical part pointed, internal structure small, tooth shaped, strongly sclerotised; parameres slightly shorter than median lobe, with about 4–5 apicolateral setae (
<figureCitation id="13592A24E956FFF9F04FF896FD50F8F2" box="[535,662,1802,1828]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="8" pageNumber="503">Fig. 40–41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C378652AE956FFF9F29FF8B1FC6FF80E" pageId="8" pageNumber="503" type="diagnosis">
<paragraph id="8BDD36A1E956FFF9F29FF8B1FC6FF80E" blockId="8.[151,1437,901,2008]" pageId="8" pageNumber="503">
<emphasis id="B916EAB3E956FFF9F29FF8B1FE16F891" bold="true" box="[199,464,1837,1863]" pageId="8" pageNumber="503">Differential diagnosis.</emphasis>
<taxonomicName id="4C624D22E956FFF9F380F8B2FCEFF89E" authority="Janak" authorityName="Janak" authorityYear="2024" box="[472,809,1838,1864]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="kambergensis" status="sp. nov.">
<emphasis id="B916EAB3E956FFF9F380F8B2FD18F89E" box="[472,734,1838,1864]" italics="true" pageId="8" pageNumber="503">Octavius kambergensis</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E956FFF9F16EF8B2FC57F89E" bold="true" box="[822,913,1838,1864]" pageId="8" pageNumber="503">
<taxonomicNameLabel id="A22557C8E956FFF9F16EF8B2FC57F89E" box="[822,913,1838,1864]" pageId="8" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs among species with small eyes, with temples more than twice as long as eyes, with the head distinctly widened posteriorly, with large body and the disc of the head without a median impression. In the most recent key of South African
<taxonomicName id="4C624D22E956FFF9F61AF8EAFA87F846" authority="(Janak 2014)" baseAuthorityName="Janak" baseAuthorityYear="2014" box="[1090,1345,1910,1936]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B916EAB3E956FFF9F61AF8EAFB60F846" box="[1090,1190,1910,1936]" italics="true" pageId="8" pageNumber="503">Octavius</emphasis>
(
<bibRefCitation id="EFF34B50E956FFF9F6EDF8EAFAFFF846" author="Janak, J." box="[1205,1337,1910,1936]" pageId="8" pageNumber="503" pagination="195 - 231" refId="ref16165" refString="Janak, J. (2014) New species and subspecies of Octavius from South Africa, with a key and additional distribution records (Coleoptera: Staphylinidae: Euaesthetinae). Acta Entomologica Musei Nationalis Pragae, 54 (1), 195 - 231." type="journal article" year="2014">Janák 2014</bibRefCitation>
)
</taxonomicName>
is to be arranged at the couplet 103 (106) near
<taxonomicName id="4C624D22E956FFF9F00CF806FC55F862" authority="Puthz, 2006" authorityName="Puthz" authorityYear="2006" box="[596,915,1946,1972]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="extralobatus">
<emphasis id="B916EAB3E956FFF9F00CF806FCC5F862" box="[596,771,1946,1972]" italics="true" pageId="8" pageNumber="503">O. extralobatus</emphasis>
<bibRefCitation id="EFF34B50E956FFF9F153F806FC55F862" author="Puthz, V." box="[779,915,1946,1972]" pageId="8" pageNumber="503" pagination="1 - 28" refId="ref16421" refString="Puthz, V. (2006) New southern African species and distribution records of the genus Octavius Fauvel (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 43, 1 - 28." type="journal article" year="2006">Puthz, 2006</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C624D22E956FFF9F194F806FB31F862" authority="Puthz, 2006" authorityName="Puthz" authorityYear="2006" box="[972,1271,1946,1972]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="pugionifer">
<emphasis id="B916EAB3E956FFF9F194F806FBA2F862" box="[972,1124,1946,1972]" italics="true" pageId="8" pageNumber="503">O. pugionifer</emphasis>
<bibRefCitation id="EFF34B50E956FFF9F635F806FB31F862" author="Puthz, V." box="[1133,1271,1946,1972]" pageId="8" pageNumber="503" pagination="1 - 28" refId="ref16421" refString="Puthz, V. (2006) New southern African species and distribution records of the genus Octavius Fauvel (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 43, 1 - 28." type="journal article" year="2006">Puthz, 2006</bibRefCitation>
</taxonomicName>
, but differs by different shape of aedeagus with small tooth shaped internal structure.
</paragraph>
</subSubSection>
<caption id="DF1D6629E957FFF8F2CFF963FBD1F889" ID-DOI="http://doi.org/10.5281/zenodo.11064286" ID-Zenodo-Dep="11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="9" pageNumber="504" startId="9.[151,264,1791,1815]" targetBox="[185,1403,194,1748]" targetPageId="9" targetType="figure">
<paragraph id="8BDD36A1E957FFF8F2CFF963FBD1F889" blockId="9.[151,1436,1791,1887]" pageId="9" pageNumber="504">
<emphasis id="B916EAB3E957FFF8F2CFF963FE9EF8C1" bold="true" box="[151,344,1791,1815]" pageId="9" pageNumber="504">FIGURES 39–48.</emphasis>
39–43:
<taxonomicName id="4C624D22E957FFF8F3E9F963FD20F8C1" authority="Janak" authorityName="Janak" authorityYear="2024" box="[433,742,1791,1815]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="504" phylum="Arthropoda" rank="species" species="kambergensis" status="sp. nov.">
<emphasis id="B916EAB3E957FFF8F3E9F963FD59F8C1" box="[433,671,1791,1815]" italics="true" pageId="9" pageNumber="504">Octavius kambergensis</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E957FFF8F0ACF963FC8EF8C1" bold="true" box="[756,840,1791,1815]" pageId="9" pageNumber="504">
<taxonomicNameLabel id="A22557C8E957FFF8F0ACF963FC8EF8C1" box="[756,840,1791,1815]" pageId="9" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 39–40, 42–43, holotype; 41, paratype. 44–48:
<taxonomicName id="4C624D22E957FFF8F71AF963FE87F8ED" authority="Janak" authorityName="Janak" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="504" phylum="Arthropoda" rank="species" species="lothenicus" status="sp. nov.">
<emphasis id="B916EAB3E957FFF8F71AF963FF39F8ED" italics="true" pageId="9" pageNumber="504">Octavius lothenicus</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E957FFF8F314F8BFFE5AF8ED" bold="true" box="[332,412,1827,1851]" pageId="9" pageNumber="504">
<taxonomicNameLabel id="A22557C8E957FFF8F314F8BFFE5AF8ED" box="[332,412,1827,1851]" pageId="9" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures.
</paragraph>
</caption>
<subSubSection id="C378652AE957FFF8F29FF811FB1BF87E" box="[199,1245,1933,1960]" pageId="9" pageNumber="504" type="etymology">
<paragraph id="8BDD36A1E957FFF8F29FF811FB1BF87E" blockId="9.[151,1436,1933,2032]" box="[199,1245,1933,1960]" pageId="9" pageNumber="504">
<emphasis id="B916EAB3E957FFF8F29FF811FE62F871" bold="true" box="[199,420,1933,1959]" pageId="9" pageNumber="504">Derivatio nominis.</emphasis>
This species is named after the
<typeStatus id="54D98803E957FFF8F152F812FCFCF87E" box="[778,826,1934,1960]" pageId="9" pageNumber="504">type</typeStatus>
locality – Kamberg Natural Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C378652AE957FFFBF29FF82DFF14FF2F" lastPageId="10" lastPageNumber="505" pageId="9" pageNumber="504" type="distribution">
<paragraph id="8BDD36A1E957FFF8F29FF82DFBE6F839" blockId="9.[151,1436,1933,2032]" pageId="9" pageNumber="504">
<emphasis id="B916EAB3E957FFF8F29FF82DFEA7F81D" bold="true" box="[199,353,1969,1995]" pageId="9" pageNumber="504">Distribution.</emphasis>
<taxonomicName id="4C624D22E957FFF8F333F82EFD79F81D" authority="Janak" authorityName="Janak" authorityYear="2024" box="[363,703,1969,1996]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="504" phylum="Arthropoda" rank="species" species="kambergensis" status="sp. nov.">
<emphasis id="B916EAB3E957FFF8F333F82EFDB4F81D" box="[363,626,1969,1996]" italics="true" pageId="9" pageNumber="504">Octavius kambergensis</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E957FFF8F095F82DFCECF81D" bold="true" box="[717,810,1969,1995]" pageId="9" pageNumber="504">
<taxonomicNameLabel id="A22557C8E957FFF8F095F82DFCECF81D" box="[717,810,1969,1995]" pageId="9" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently recorded only from the Kamberg Natural Reserve in Drakensberg Mts.,
<collectingRegion id="49A6F843E957FFF8F3B3F849FCCEF839" box="[491,776,2005,2031]" country="South Africa" name="KwaZulu-Natal" pageId="9" pageNumber="504">KwaZulu-Natal Province</collectingRegion>
,
<collectingCountry id="F3757631E957FFF8F14CF849FC64F839" box="[788,930,2005,2031]" name="South Africa" pageId="9" pageNumber="504">South Africa</collectingCountry>
(
<figureCitation id="13592A24E957FFF8F1E9F849FBD5F839" box="[945,1043,2005,2031]" captionStart="FIGURES 113–121" captionStartId="23.[151,264,1358,1382]" captionTargetBox="[212,1375,191,1316]" captionTargetId="figure-15@23.[197,1390,181,1334]" captionTargetPageId="23" captionText="FIGURES 113–121. Octavius undulatus Janák, sp. nov.; 113–116, 118–119, holotype; 117, paratype. 113, habitus; 114, fore parts; 115, fore parts lateral; 116–117, aedeagus ventral; 118, male sternite VIII; 119, male sternite IX. Scale 0.5 mm: 113; scales 0.1 mm: 114–115; 116–117; 118–119; o: ommatidium. 120: Distribution of Octavius – subgroup A. 121: Distribution of Octavius – subgroup B." figureDoi="http://doi.org/10.5281/zenodo.11064300" httpUri="https://zenodo.org/record/11064300/files/figure.png" pageId="9" pageNumber="504">Fig. 120</figureCitation>
).
</paragraph>
<paragraph id="8BDD36A1E954FFFBF29FFF0AFF14FF2F" blockId="10.[151,1437,150,249]" pageId="10" pageNumber="505">
<emphasis id="B916EAB3E954FFFBF29FFF0AFE8DFF66" bold="true" box="[199,331,150,176]" pageId="10" pageNumber="505">Bionomics.</emphasis>
All specimens were collected in siftings of forest litter in forest patches of an indigenous forest at the elevation of about
<quantity id="4C9A9B44E954FFFBF3EEFF27FD95FF03" box="[438,595,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.83" metricValueMax="1.8399999999999999" metricValueMin="1.8199999999999998" pageId="10" pageNumber="505" unit="m" value="1830.0" valueMax="1840.0" valueMin="1820.0">1820–1840 m</quantity>
a.s.l. with the abundance of 5.3 specimens per kg of sifted material (
<figureCitation id="13592A24E954FFFBF737FF27FF03FF2F" captionStart="FIGURES 128–133" captionStartId="25.[151,264,1651,1675]" captionTargetBox="[154,1434,182,1627]" captionTargetId="figure-15@25.[154,1434,181,1627]" captionTargetPageId="25" captionText="FIGURES 128–133. 128: Cathedral Peak, Masongwane river banks. 129: Injisuthi. 130: Monkʼs Cowl, Hlatikulu forest. 131: Lotheni. 132: Kamberg, locality 1. 133: Kamberg locality 2." figureDoi="http://doi.org/10.5281/zenodo.11064306" httpUri="https://zenodo.org/record/11064306/files/figure.png" pageId="10" pageNumber="505">Fig. 133</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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