273 lines
38 KiB
XML
273 lines
38 KiB
XML
<document id="E3399ADCA77B4AF875480A768097A330" ID-DOI="10.11646/zootaxa.5443.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="11064278" ID-ZooBank="A91A8D67-A81B-427D-8096-59E41326A8E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714028852870" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2024" docId="03CB87B7E94AFFE5F2CFFF0BFBFAFA14" docLanguage="en" docName="zootaxa.5443.4.2.pdf" docOrigin="Zootaxa 5443 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Octavius regius Janak 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="515" masterDocId="FFF2FFCFE95EFFF1F258FF9CFFC6FFD6" masterDocTitle="On Octavius from Maloti-Drakensberg and Golden Gate National Parks, South Africa (Coleoptera: Staphylinidae: Euaesthetinae)" masterLastPageNumber="522" masterPageNumber="495" pageNumber="515" updateTime="1714029516812" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3D8323AF43E625BBF2B9FCB908C10ACC">On Octavius from Maloti-Drakensberg and Golden Gate National Parks, South Africa (Coleoptera: Staphylinidae: Euaesthetinae)</mods:title>
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<mods:name id="922BF27BE2A92E9FAC2750FEA63A5143" type="personal">
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<mods:roleTerm id="C962C37EC8310FFD883D39F36C32E40A">Author</mods:roleTerm>
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<mods:namePart id="A31923E6FB628B2B87BBCD41891EA93B">Janák, Jiří</mods:namePart>
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<mods:title id="C12CED1BD89E6DE1E8DA5511F1191BCB">Zootaxa</mods:title>
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<mods:part id="8D06E259F593B9C1A902FD8607CF3B59">
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<mods:date id="B935FC18F684656EB56DB58AF8799BD7">2024</mods:date>
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<mods:number id="4961C7C9164EFA7450E024FF47252772">2024-04-24</mods:number>
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<mods:number id="593E6B8884A0C2F7EA7628FFD643131D">5443</mods:number>
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<mods:number id="930F858B40986434591A87B8118AD065">4</mods:number>
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<mods:url id="469A4B2C4CB7FB01669D265A3DA0CC5D">http://dx.doi.org/10.11646/zootaxa.5443.4.2</mods:url>
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<mods:classification id="16254E182F25CA86DCAFABC450F599B7">journal article</mods:classification>
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<mods:identifier id="89875311882259183D32675AB1E38034" type="DOI">10.11646/zootaxa.5443.4.2</mods:identifier>
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<mods:identifier id="5964013AE27EE284DD91062029FFF31B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CB87B7E94AFFE5F2CFFF0BFBFAFA14" ID-DOI="http://doi.org/10.5281/zenodo.11064387" ID-Zenodo-Dep="11064387" LSID="urn:lsid:plazi:treatment:03CB87B7E94AFFE5F2CFFF0BFBFAFA14" httpUri="http://treatment.plazi.org/id/03CB87B7E94AFFE5F2CFFF0BFBFAFA14" lastPageNumber="515" pageId="20" pageNumber="515">
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<subSubSection id="C378652AE94AFFE5F2CFFF0BFDD2FF67" box="[151,532,150,177]" pageId="20" pageNumber="515" type="nomenclature">
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<paragraph id="8BDD36A1E94AFFE5F2CFFF0BFDD2FF67" blockId="20.[151,532,150,214]" box="[151,532,150,177]" pageId="20" pageNumber="515">
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<heading id="D09581CDE94AFFE5F2CFFF0BFDD2FF67" bold="true" box="[151,532,150,177]" fontSize="11" level="1" pageId="20" pageNumber="515" reason="1">
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<emphasis id="B916EAB3E94AFFE5F2CFFF0BFDD2FF67" bold="true" box="[151,532,150,177]" pageId="20" pageNumber="515">
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<taxonomicName id="4C624D22E94AFFE5F2CFFF0BFE6FFF67" authority="Janak" authorityName="Janak" authorityYear="2024" box="[151,425,150,177]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="regius" status="sp. nov.">
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<emphasis id="B916EAB3E94AFFE5F2CFFF0BFE95FF67" bold="true" box="[151,339,151,177]" italics="true" pageId="20" pageNumber="515">Octavius regius</emphasis>
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Janák
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22557C8E94AFFE5F3EEFF0BFDD2FF67" box="[438,532,151,177]" pageId="20" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C378652AE94AFFE5F2CFFF27FE6AFF00" box="[151,428,187,214]" pageId="20" pageNumber="515" type="description">
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<paragraph id="8BDD36A1E94AFFE5F2CFFF27FE6AFF00" blockId="20.[151,532,150,214]" box="[151,428,187,214]" pageId="20" pageNumber="515">
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(
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<figureCitation id="13592A24E94AFFE5F2C7FF27FEF3FF03" box="[159,309,187,213]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Figs. 99–106</figureCitation>
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,
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<figureCitation id="13592A24E94AFFE5F319FF27FEABFF03" box="[321,365,187,213]" captionStart="FIGURES 113–121" captionStartId="23.[151,264,1358,1382]" captionTargetBox="[212,1375,191,1316]" captionTargetId="figure-15@23.[197,1390,181,1334]" captionTargetPageId="23" captionText="FIGURES 113–121. Octavius undulatus Janák, sp. nov.; 113–116, 118–119, holotype; 117, paratype. 113, habitus; 114, fore parts; 115, fore parts lateral; 116–117, aedeagus ventral; 118, male sternite VIII; 119, male sternite IX. Scale 0.5 mm: 113; scales 0.1 mm: 114–115; 116–117; 118–119; o: ommatidium. 120: Distribution of Octavius – subgroup A. 121: Distribution of Octavius – subgroup B." figureDoi="http://doi.org/10.5281/zenodo.11064300" httpUri="https://zenodo.org/record/11064300/files/figure.png" pageId="20" pageNumber="515">121</figureCitation>
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,
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<figureCitation id="13592A24E94AFFE5F321FF27FE62FF00" box="[377,420,187,214]" captionStart="FIGURES 122–127" captionStartId="24.[151,264,1647,1671]" captionTargetBox="[154,1434,181,1624]" captionTargetId="figure-29@24.[154,1434,181,1624]" captionTargetPageId="24" captionText="FIGURES 122–127. 122–123: Golden Gate. 124–125: Royal Natal: 124, Gudu forest, 125, Devilʼs Hoek. 126–127: Cathedral Peak: 126, Rainbow gorge, 127, Oqualweni Fern forest." figureDoi="http://doi.org/10.5281/zenodo.11064304" httpUri="https://zenodo.org/record/11064304/files/figure.png" pageId="20" pageNumber="515">125</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C378652AE94AFFE5F2CFFE9FFF25FE23" pageId="20" pageNumber="515" type="materials_examined">
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||
<paragraph id="8BDD36A1E94AFFE5F2CFFE9FFEF3FE97" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
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||
<materialsCitation id="3B0A3CFCE94AFFE5F2CFFE9FFEF7FE97" country="South Africa" county="Maloti-Drakensberg Park" location="Hoek" municipality="Royal Natal Nature Reserve" pageId="20" pageNumber="515" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
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||
<emphasis id="B916EAB3E94AFFE5F2CFFE9FFEFCFECB" bold="true" box="[151,314,259,285]" pageId="20" pageNumber="515">
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||
<typeStatus id="54D98803E94AFFE5F2CFFE9FFF14FECB" box="[151,210,259,285]" pageId="20" pageNumber="515">Type</typeStatus>
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locality.
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</emphasis>
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||
<collectingCountry id="F3757631E94AFFE5F31DFE9FFE1CFECB" box="[325,474,259,285]" name="South Africa" pageId="20" pageNumber="515">South Africa</collectingCountry>
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,
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<collectingRegion id="49A6F843E94AFFE5F3B2FE9FFCCDFECB" box="[490,779,259,285]" country="South Africa" name="KwaZulu-Natal" pageId="20" pageNumber="515">KwaZulu-Natal Province</collectingRegion>
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,
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<collectingCounty id="62BC4E2DE94AFFE5F142FE9FFBF8FECB" box="[794,1086,259,285]" pageId="20" pageNumber="515">Maloti-Drakensberg Park</collectingCounty>
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||
,
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||
<collectingMunicipality id="6BB9ACDBE94AFFE5F616FE9FFA5EFEC8" box="[1102,1432,259,286]" pageId="20" pageNumber="515">Royal Natal Nature Reserve</collectingMunicipality>
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||
, Devilʼs
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||
<location id="8EBD607AE94AFFE5F2A9FEBBFEF7FE97" LSID="urn:lsid:plazi:treatment:03CB87B7E94AFFE5F2CFFF0BFBFAFA14:8EBD607AE94AFFE5F2A9FEBBFEF7FE97" box="[241,305,295,321]" country="South Africa" county="Maloti-Drakensberg Park" municipality="Royal Natal Nature Reserve" name="Hoek" pageId="20" pageNumber="515" stateProvince="KwaZulu-Natal">Hoek</location>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BDD36A1E94AFFE5F29FFED7FF25FE23" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
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<emphasis id="B916EAB3E94AFFE5F29FFED7FEA8FEB3" bold="true" box="[199,366,331,357]" pageId="20" pageNumber="515">Type material</emphasis>
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||
(
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||
<specimenCount id="9D64FD28E94AFFE5F326FED7FDDEFEB3" box="[382,536,331,357]" count="13" pageId="20" pageNumber="515" type="generic">13 specimens</specimenCount>
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||
).
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||
<emphasis id="B916EAB3E94AFFE5F076FED7FD5DFEB3" bold="true" box="[558,667,331,357]" pageId="20" pageNumber="515">
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||
<typeStatus id="54D98803E94AFFE5F076FED7FD5DFEB3" box="[558,667,331,357]" pageId="20" pageNumber="515" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
<materialsCitation id="3B0A3CFCE94AFFE5F0FAFED0FBD6FE7B" collectingDate="2019-01-10" collectionCode="TMSA" collectorName="J. Janak" country="South Africa" determinerName="J. Janak" elevation="1620" location="Hoek" municipality="Maloti-Drakensberg Park" pageId="20" pageNumber="515" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
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||
<specimenCount id="9D64FD28E94AFFE5F0FAFED0FD7CFEB0" box="[674,698,332,358]" count="1" pageId="20" pageNumber="515" type="male">♂</specimenCount>
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||
: “
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||
<collectingCountry id="F3757631E94AFFE5F089FED7FCA2FEB3" box="[721,868,331,357]" name="South Africa" pageId="20" pageNumber="515">South Africa</collectingCountry>
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||
,
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||
<collectingRegion id="49A6F843E94AFFE5F128FED7FBE5FEB3" box="[880,1059,331,357]" country="South Africa" name="KwaZulu-Natal" pageId="20" pageNumber="515">KwaZulu-Natal</collectingRegion>
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||
,
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||
<collectingMunicipality id="6BB9ACDBE94AFFE5F676FED7FA88FEB3" box="[1070,1358,331,357]" pageId="20" pageNumber="515">Maloti-Drakensberg Park</collectingMunicipality>
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||
, Royal Natal NR, Devilʼs
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||
<location id="8EBD607AE94AFFE5F33FFEF3FE61FE5F" LSID="urn:lsid:plazi:treatment:03CB87B7E94AFFE5F2CFFF0BFBFAFA14:8EBD607AE94AFFE5F33FFEF3FE61FE5F" box="[359,423,367,393]" country="South Africa" municipality="Maloti-Drakensberg Park" name="Hoek" pageId="20" pageNumber="515" stateProvince="KwaZulu-Natal">Hoek</location>
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, 28°42.7ʼS, 28°55.4ʼE,
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||
<quantity id="4C9A9B44E94AFFE5F0E8FEF3FCC6FE5F" box="[688,768,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="20" pageNumber="515" unit="m" value="1620.0">
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<elevation id="004FD192E94AFFE5F0E8FEF3FCC6FE5F" box="[688,768,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="20" pageNumber="515" unit="m" value="1620.0">1620m</elevation>
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</quantity>
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,
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||
<date id="FFDC1061E94AFFE5F153FEF3FCB1FE5F" box="[779,887,367,393]" pageId="20" pageNumber="515" value="2019-01-10">
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||
<collectingDate id="EF98E989E94AFFE5F153FEF3FCB1FE5F" box="[779,887,367,393]" pageId="20" pageNumber="515" value="2019-01-10">10.i.2019</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="26975377E94AFFE5F1DAFEECFC1DFE5F" box="[898,987,367,394]" pageId="20" pageNumber="515">J. Janák</collectorName>
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||
lgt., Berlese extraction, leaf & log litter, sifting”, “
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||
<typeStatus id="54D98803E94AFFE5F35CFE0FFE6BFE7B" box="[260,429,403,429]" pageId="20" pageNumber="515" type="holotype">HOLOTYPUS</typeStatus>
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||
<taxonomicName id="4C624D22E94AFFE5F3EDFE08FDA0FE7B" authority="Janák, 2024" authorityName="Janák" authorityYear="2024" box="[437,614,404,430]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="regius" status="sp. nov.">
|
||
<emphasis id="B916EAB3E94AFFE5F3EDFE08FDA0FE7B" box="[437,614,404,430]" italics="true" pageId="20" pageNumber="515">Octavius regius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22557C8E94AFFE5F036FE08FD03FE78" box="[622,709,404,430]" pageId="20" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<determinerName id="7E01DA65E94AFFE5F095FE08FCE1FE7B" box="[717,807,403,430]" pageId="20" pageNumber="515">J. Janák</determinerName>
|
||
det. 2021” (
|
||
<collectionCode id="ED73AE64E94AFFE5F1ECFE0FFBCEFE7B" box="[948,1032,403,429]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="20" pageNumber="515">TMSA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0A3CFCE94AFFE5F643FE08FF19FE23" collectingDate="2019-01-10" collectionCode="TMSA, JJRC" collectorName="All" country="South Africa" determinerName="J. Janak" elevation="1620" location="Hoek" municipality="Maloti-Drakensberg Park" pageId="20" pageNumber="515" specimenCount="12" specimenCount-female="3" specimenCount-male="9" stateProvince="KwaZulu-Natal" typeStatus="paratype">
|
||
<typeStatus id="54D98803E94AFFE5F643FE08FB4AFE78" box="[1051,1164,404,430]" pageId="20" pageNumber="515" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D64FD28E94AFFE5F6C0FE0FFB03FE78" box="[1176,1221,403,430]" count="9" pageId="20" pageNumber="515" type="male">9 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D64FD28E94AFFE5F68AFE0FFB39FE78" box="[1234,1279,403,430]" count="3" pageId="20" pageNumber="515" type="female">3 ♀</specimenCount>
|
||
: same data as holotype (
|
||
<collectionCode id="ED73AE64E94AFFE5F350FE2BFE9CFE07" box="[264,346,439,465]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="20" pageNumber="515">TMSA</collectionCode>
|
||
,
|
||
<collectionCode id="ED73AE64E94AFFE5F33DFE2BFE63FE07" box="[357,421,439,465]" pageId="20" pageNumber="515">JJRC</collectionCode>
|
||
).
|
||
<collectorName id="26975377E94AFFE5F3EDFE2BFE1FFE07" box="[437,473,439,465]" pageId="20" pageNumber="515">All</collectorName>
|
||
<typeStatus id="54D98803E94AFFE5F3B8FE24FD8DFE04" box="[480,587,440,466]" pageId="20" pageNumber="515" type="paratype">paratypes</typeStatus>
|
||
with additional labels: “
|
||
<typeStatus id="54D98803E94AFFE5F102FE2BFC3DFE07" box="[858,1019,439,465]" pageId="20" pageNumber="515" type="paratype">PARATYPUS</typeStatus>
|
||
<taxonomicName id="4C624D22E94AFFE5F65AFE24FB77FE07" authority="Janák, 2024" authorityName="Janák" authorityYear="2024" box="[1026,1201,440,466]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="regius" status="sp. nov.">
|
||
<emphasis id="B916EAB3E94AFFE5F65AFE24FB77FE07" box="[1026,1201,440,466]" italics="true" pageId="20" pageNumber="515">Octavius regius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22557C8E94AFFE5F6EFFE24FAC8FE04" box="[1207,1294,440,466]" pageId="20" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<determinerName id="7E01DA65E94AFFE5F74CFE24FAABFE07" box="[1300,1389,439,466]" pageId="20" pageNumber="515">J. Janák</determinerName>
|
||
det. 2021”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378652AE94AFFE5F29FFE63FB88FBAB" pageId="20" pageNumber="515" type="description">
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFE63FC6EFDEB" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
<emphasis id="B916EAB3E94AFFE5F29FFE63FE9FFDCF" bold="true" box="[199,345,511,537]" pageId="20" pageNumber="515">Description.</emphasis>
|
||
Body length
|
||
<quantity id="4C9A9B44E94AFFE5F3ABFE63FDBDFDCC" box="[499,635,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="20" pageNumber="515" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
|
||
(M
|
||
<quantity id="4C9A9B44E94AFFE5F0F5FE63FCD2FDCC" box="[685,788,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="515" unit="mm" value="1.2">1.20 mm</quantity>
|
||
, HT
|
||
<quantity id="4C9A9B44E94AFFE5F115FE63FC6EFDCF" box="[845,936,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="515" unit="mm" value="1.1">1.1 mm</quantity>
|
||
), forebody length
|
||
<quantity id="4C9A9B44E94AFFE5F62FFE63FB39FDCC" box="[1143,1279,511,538]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="20" pageNumber="515" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
|
||
(M
|
||
<quantity id="4C9A9B44E94AFFE5F769FE63FA5EFDCC" box="[1329,1432,511,538]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="20" pageNumber="515" unit="mm" value="0.51">0.51 mm</quantity>
|
||
, HT
|
||
<quantity id="4C9A9B44E94AFFE5F29BFDBFFEDBFDEB" box="[195,285,547,574]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="515" unit="mm" value="0.5">0.5 mm</quantity>
|
||
). Microphalmous, apterous, rusty, slightly shiny (
|
||
<figureCitation id="13592A24E94AFFE5F11FFDBFFC5AFDEB" box="[839,924,547,573]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Fig. 99</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFDDBFDBDFD1B" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
Head (
|
||
<figureCitation id="13592A24E94AFFE5F349FDDBFE74FDB7" box="[273,434,583,609]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Figs. 100–101</figureCitation>
|
||
) slightly broader than pronotum (R 1.01–1.09, M 1.05, HT 1.04), eyes very small and flat, with a few small ommatidia (
|
||
<figureCitation id="13592A24E94AFFE5F3BDFDF7FD4CFD53" box="[485,650,619,645]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Figs. 101–102</figureCitation>
|
||
), sides of head rounded, posterior angles moderately angular, median impression on disc absent, surface with very fine reticulation consisting of irregular polygonal fields and very fine punctures raising from center of each field.
|
||
</paragraph>
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFD44FB2FFCEF" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
Pronotum (
|
||
<figureCitation id="13592A24E94AFFE5F310FD4BFE2BFD27" box="[328,493,727,753]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Figs. 100–101</figureCitation>
|
||
) markedly longer than broad (R 1.10–1.21, M 1.16, HT 1.15), strongly narrowed posteriorly; anterior angles rounded, disc regularly convex, disc and base without impressions, posterior angles shortly rounded; surface finely irregularly reticulate and very finely punctate, similarly as on head.
|
||
</paragraph>
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFCDFFAA2FC73" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
Elytra (
|
||
<figureCitation id="13592A24E94AFFE5F346FCDFFE02FC8B" box="[286,452,835,861]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Figs. 100–101</figureCitation>
|
||
) roundly trapezoid, markedly broader than long (R 1.12–1.23, M 1.18, HT 1.20), as broad as or broader than head (R 1.01–1.12, M 1.05, HT 1.08), without sharp longitudinal ridge laterally (
|
||
<figureCitation id="13592A24E94AFFE5F776FCFBFA49FC57" box="[1326,1423,871,897]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Fig. 101</figureCitation>
|
||
), surface with very fine reticulation consisting of irregular polygonal fields and sparsely, very finely punctured.
|
||
</paragraph>
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFC33FE56FC38" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">Abdomen with two paratergites, with reticulation consisting of polygonal, slightly transverse fields, very finely and sparsely punctate.</paragraph>
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFC6BFB88FBAB" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
Male. Sternite VIII deeply roundly emarginated in posterior quarter (
|
||
<figureCitation id="13592A24E94AFFE5F1B5FC6BFB95FBC4" box="[1005,1107,1015,1042]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Fig. 105</figureCitation>
|
||
), sternite IX as in
|
||
<figureCitation id="13592A24E94AFFE5F76CFC6BFA5EFBC7" box="[1332,1432,1015,1041]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Fig. 106</figureCitation>
|
||
. Aedeagus symmetrical, relatively large (length
|
||
<quantity id="4C9A9B44E94AFFE5F0E3FB87FCA5FBE0" box="[699,867,1051,1078]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="20" pageNumber="515" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.20–0.23 mm</quantity>
|
||
, M
|
||
<quantity id="4C9A9B44E94AFFE5F1CDFB87FC39FBE0" box="[917,1023,1051,1078]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="515" unit="mm" value="0.22">0.22 mm</quantity>
|
||
, HT
|
||
<quantity id="4C9A9B44E94AFFE5F665FB87FB6EFBE3" box="[1085,1192,1051,1077]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="20" pageNumber="515" unit="mm" value="0.21">0.21 mm</quantity>
|
||
), apical part pointed, internal sac with apical part covered by curved teeth, in basal part with small scales; parameres wide, markedly longer than median lobe, with 7–8 subapical laterally oriented setae (
|
||
<figureCitation id="13592A24E94AFFE5F1C4FBFFFB84FBAB" box="[924,1090,1123,1149]" captionStart="FIGURES 99–112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99–112. 99–106: Octavius regius Janák, sp. nov.; 99–103, 105–106, holotype; 104, paratype. 107–112: Octavius subulatus Janák, sp. nov.; 107–110, 112–113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103–104, 110–111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100–101, 108–109; 103–104; 110–111; 105–106, 112–113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="20" pageNumber="515">Figs. 103–104</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378652AE94AFFE5F29FFB1BFB9AFB3F" pageId="20" pageNumber="515" type="diagnosis">
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFB1BFB9AFB3F" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
<emphasis id="B916EAB3E94AFFE5F29FFB1BFE0AFB77" bold="true" box="[199,460,1159,1185]" pageId="20" pageNumber="515">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C624D22E94AFFE5F389FB14FD01FB77" authority="Janak" authorityName="Janak" authorityYear="2024" box="[465,711,1159,1186]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="regius" status="sp. nov.">
|
||
<emphasis id="B916EAB3E94AFFE5F389FB14FDB9FB77" box="[465,639,1160,1186]" italics="true" pageId="20" pageNumber="515">Octavius regius</emphasis>
|
||
Janák
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B916EAB3E94AFFE5F089FB1BFCEEFB77" bold="true" box="[721,808,1159,1185]" pageId="20" pageNumber="515">
|
||
<taxonomicNameLabel id="A22557C8E94AFFE5F089FB1BFCEEFB77" box="[721,808,1159,1185]" pageId="20" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is to be arranged in the key of Southern African
|
||
<taxonomicName id="4C624D22E94AFFE5F761FB14FEEFFB13" authority="(Puthz 2006)" baseAuthorityName="Puthz" baseAuthorityYear="2006" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916EAB3E94AFFE5F761FB14FA5BFB74" box="[1337,1437,1160,1186]" italics="true" pageId="20" pageNumber="515">Octavius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF34B50E94AFFE5F2C7FB37FEE7FB13" author="Puthz, V." box="[159,289,1195,1221]" pageId="20" pageNumber="515" pagination="1 - 28" refId="ref16421" refString="Puthz, V. (2006) New southern African species and distribution records of the genus Octavius Fauvel (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 43, 1 - 28." type="journal article" year="2006">Puthz 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
at the couplet 83 (84) near
|
||
<taxonomicName id="4C624D22E94AFFE5F006FB30FCA0FB13" authority="Puthz, 1986" authorityName="Puthz" authorityYear="1986" box="[606,870,1195,1222]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="bacillus">
|
||
<emphasis id="B916EAB3E94AFFE5F006FB30FD1FFB13" box="[606,729,1195,1222]" italics="true" pageId="20" pageNumber="515">O. bacillus</emphasis>
|
||
<bibRefCitation id="EFF34B50E94AFFE5F0B8FB37FCA0FB13" author="Puthz, V." box="[736,870,1195,1221]" pageId="20" pageNumber="515" pagination="185 - 201" refId="ref16376" refString="Puthz, V. (1986) Neue Arten der Gattung Octavius Fauvel, 1872, aus Sudafrika nebst einer Bestimmungstabelle der in der Athiopis lebenden Gattungsvertreter (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 34, 185 - 201." type="journal article" year="1986">Puthz, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
but differs markedly by emarginated male sternite VIII, by median lobe much broader and parameres markedly longer than median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378652AE94AFFE5F29FFB6FFD47FAE4" pageId="20" pageNumber="515" type="etymology">
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFB6FFD47FAE4" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
<emphasis id="B916EAB3E94AFFE5F29FFB6FFE63FADB" bold="true" box="[199,421,1267,1293]" pageId="20" pageNumber="515">Derivatio nominis.</emphasis>
|
||
The name of this species is derived from the latin word regius (royal), which refers to the
|
||
<typeStatus id="54D98803E94AFFE5F2CFFA84FF01FAE4" box="[151,199,1304,1330]" pageId="20" pageNumber="515">type</typeStatus>
|
||
locality – Royal Natal Nature Reserve.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378652AE94AFFE5F29FFAA7FBFAFA14" pageId="20" pageNumber="515" type="distribution">
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFAA7FB2FFAAF" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
<emphasis id="B916EAB3E94AFFE5F29FFAA7FEA7FA83" bold="true" box="[199,353,1339,1365]" pageId="20" pageNumber="515">Distribution.</emphasis>
|
||
<taxonomicName id="4C624D22E94AFFE5F332FAA0FDACFA83" authority="Janak" authorityName="Janak" authorityYear="2024" box="[362,618,1339,1366]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="regius" status="sp. nov.">
|
||
<emphasis id="B916EAB3E94AFFE5F332FAA0FDDBFA83" box="[362,541,1340,1366]" italics="true" pageId="20" pageNumber="515">Octavius regius</emphasis>
|
||
Janák
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B916EAB3E94AFFE5F020FAA7FD12FA83" bold="true" box="[632,724,1339,1365]" pageId="20" pageNumber="515">
|
||
<taxonomicNameLabel id="A22557C8E94AFFE5F020FAA7FD12FA83" box="[632,724,1339,1365]" pageId="20" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is currently recorded only from the Devilʼs Hoek area in the Royal Natal Nature Reserve, Drakensberg Mts.,
|
||
<collectingRegion id="49A6F843E94AFFE5F0EEFAC3FC14FAAF" box="[694,978,1375,1401]" country="South Africa" name="KwaZulu-Natal" pageId="20" pageNumber="515">KwaZulu-Natal province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3757631E94AFFE5F185FAC3FBAAFAAF" box="[989,1132,1375,1401]" name="South Africa" pageId="20" pageNumber="515">South Africa</collectingCountry>
|
||
(
|
||
<figureCitation id="13592A24E94AFFE5F622FAC3FB1AFAAF" box="[1146,1244,1375,1401]" captionStart="FIGURES 113–121" captionStartId="23.[151,264,1358,1382]" captionTargetBox="[212,1375,191,1316]" captionTargetId="figure-15@23.[197,1390,181,1334]" captionTargetPageId="23" captionText="FIGURES 113–121. Octavius undulatus Janák, sp. nov.; 113–116, 118–119, holotype; 117, paratype. 113, habitus; 114, fore parts; 115, fore parts lateral; 116–117, aedeagus ventral; 118, male sternite VIII; 119, male sternite IX. Scale 0.5 mm: 113; scales 0.1 mm: 114–115; 116–117; 118–119; o: ommatidium. 120: Distribution of Octavius – subgroup A. 121: Distribution of Octavius – subgroup B." figureDoi="http://doi.org/10.5281/zenodo.11064300" httpUri="https://zenodo.org/record/11064300/files/figure.png" pageId="20" pageNumber="515">Fig. 121</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD36A1E94AFFE5F29FFA1FFBFAFA14" blockId="20.[151,1437,259,1474]" pageId="20" pageNumber="515">
|
||
<emphasis id="B916EAB3E94AFFE5F29FFA1FFE8DFA4B" bold="true" box="[199,331,1411,1437]" pageId="20" pageNumber="515">Bionomics.</emphasis>
|
||
All specimens were collected in siftings of litter in an indigenous forest patch at the elevation of about
|
||
<quantity id="4C9A9B44E94AFFE5F284FA3BFEF7FA14" box="[220,305,1447,1474]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="20" pageNumber="515" unit="m" value="1600.0">1600 m</quantity>
|
||
a.s.l. with abundance
|
||
<specimenCount id="9D64FD28E94AFFE5F073FA3BFD6CFA17" box="[555,682,1447,1473]" count="1" pageId="20" pageNumber="515" type="generic">1 specimen</specimenCount>
|
||
per kg of sifted material (
|
||
<figureCitation id="13592A24E94AFFE5F195FA3BFBE9FA14" box="[973,1071,1447,1474]" captionStart="FIGURES 122–127" captionStartId="24.[151,264,1647,1671]" captionTargetBox="[154,1434,181,1624]" captionTargetId="figure-29@24.[154,1434,181,1624]" captionTargetPageId="24" captionText="FIGURES 122–127. 122–123: Golden Gate. 124–125: Royal Natal: 124, Gudu forest, 125, Devilʼs Hoek. 126–127: Cathedral Peak: 126, Rainbow gorge, 127, Oqualweni Fern forest." figureDoi="http://doi.org/10.5281/zenodo.11064304" httpUri="https://zenodo.org/record/11064304/files/figure.png" pageId="20" pageNumber="515">Fig. 125</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |