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<document id="940B3765BC9FCC7AEA9C4D198C764697" ID-DOI="10.11646/zootaxa.3904.4.9" ID-GBIF-Dataset="b62ba0f5-0801-48e1-9287-1397b6777463" ID-ISSN="1175-5326" ID-Zenodo-Dep="245329" ID-ZooBank="9BCD9549-508A-4710-A4B1-4887FE48A68E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460893235200" checkinUser="plazi" docAuthor="Hlaváč, Peter &amp; Brachat, Volker" docDate="2015" docId="03CB879B6637C85B139AFD301953F8BE" docLanguage="en" docName="zt03904p599.pdf" docOrigin="Zootaxa 3904 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyphobythus anatolicus Hlavac &amp; Brachat, sp. nov." docType="treatment" docVersion="8" lastPageNumber="597" masterDocId="FFF2FFE36636C85A130DFFA41D03FFF7" masterDocTitle="New species of the genus Glyphobythus Raffray, 1904 (Coleoptera: Staphylinidae: Pselaphinae) from Turkey" masterLastPageNumber="599" masterPageNumber="596" pageNumber="597" updateTime="1698359757808" updateUser="plazi">
<mods:mods id="B245BD7DB6B17DB4524463C61B218304" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B2887B31D47B2EC5C3A3731B60FE4125">
<mods:title id="512E2382625DFBF865199BC57F83332E">New species of the genus Glyphobythus Raffray, 1904 (Coleoptera: Staphylinidae: Pselaphinae) from Turkey</mods:title>
</mods:titleInfo>
<mods:name id="FF051BC733ACD23783FCA94A60210154" type="personal">
<mods:role id="45DF4BD29F6A15841FF8F9D72DCD12A6">
<mods:roleTerm id="04901DECBA11EF8DA763CAA84EA8D1C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="37ADF4476548D066736097EBF9235589">Hlaváč, Peter</mods:namePart>
</mods:name>
<mods:name id="B1749DE95D9029B4D3E0871EC35C7838" type="personal">
<mods:role id="38A6E13BA51E72BBA4E9E342F689917F">
<mods:roleTerm id="033906A58A86121877B28A7E03A73A83">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="67386A0110EEE0315373CCED31DBBCE7">Brachat, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="E92D7DBBAEDC938DF308A4C782211037">text</mods:typeOfResource>
<mods:relatedItem id="84E956E42FE98EA32DAF6016CF3D936C" type="host">
<mods:titleInfo id="2A8DCDA004DB34E0EA96CCAD6419024F">
<mods:title id="EF2679A674FD22DD97F8B70C9DF480F7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5D3B56C3F2B064149FF81568B30DF9B3">
<mods:date id="A724A9894C900688BE85880E61BC7409">2015</mods:date>
<mods:detail id="3A08D0B493A53B356CEBF0C21F167DA8" type="volume">
<mods:number id="55D60BD656768831D0E7C2E257913877">3904</mods:number>
</mods:detail>
<mods:detail id="DB99E455400A6FD9BE58566B774C2319" type="issue">
<mods:number id="78F9D1E94112567C63E6F827BAE73B8D">4</mods:number>
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<mods:start id="262ED5C6FAB4677736A68372505CDA93">596</mods:start>
<mods:end id="4BEE46C35C09612B8B4C75CEF77F2486">599</mods:end>
</mods:extent>
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<mods:classification id="112DBE302E289D14D829B12D3AD99CB2">journal article</mods:classification>
<mods:identifier id="07015B94C695E71C15975C202B6358E7" type="DOI">10.11646/zootaxa.3904.4.9</mods:identifier>
<mods:identifier id="C20D933B3F75B7C2EA22593AA02CF948" type="GBIF-Dataset">b62ba0f5-0801-48e1-9287-1397b6777463</mods:identifier>
<mods:identifier id="9743C98D093CF69406CCC9771DADA469" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="289DEC723DD76830FE18A756E2118B93" type="Zenodo-Dep">245329</mods:identifier>
<mods:identifier id="CFC29A2D34EB11A849764F4AC477DB14" type="ZooBank">9BCD9549-508A-4710-A4B1-4887FE48A68E</mods:identifier>
</mods:mods>
<treatment id="03CB879B6637C85B139AFD301953F8BE" ID-DOI="http://doi.org/10.5281/zenodo.6103158" ID-GBIF-Taxon="119587128" ID-Zenodo-Dep="6103158" LSID="urn:lsid:plazi:treatment:03CB879B6637C85B139AFD301953F8BE" httpUri="http://treatment.plazi.org/id/03CB879B6637C85B139AFD301953F8BE" lastPageNumber="597" pageId="1" pageNumber="597">
<subSubSection id="C37865066637C85B139AFD301C10FD38" pageId="1" pageNumber="597" type="nomenclature">
<paragraph id="8BDD368D6637C85B139AFD301E10FD58" blockId="1.[151,787,660,719]" box="[151,787,660,687]" pageId="1" pageNumber="597">
<heading id="D09581E16637C85B139AFD301E10FD58" bold="true" box="[151,787,660,687]" fontSize="11" level="1" pageId="1" pageNumber="597" reason="1">
<emphasis id="B916EA9F6637C85B139AFD301E10FD58" bold="true" box="[151,787,660,687]" pageId="1" pageNumber="597">
<taxonomicName id="4C624D0E6637C85B139AFD301FADFD58" ID-CoL="3GGLT" authority="Hlavac &amp; Brachat" authorityName="Hlavac &amp; Brachat" box="[151,686,660,687]" class="Insecta" family="Staphylinidae" genus="Glyphobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="B916EA9F6637C85B139AFD301CC0FD59" bold="true" box="[151,451,660,686]" italics="true" pageId="1" pageNumber="597">Glyphobythus anatolicus</emphasis>
Hlaváč &amp; Brachat
</taxonomicName>
<taxonomicNameLabel id="A22557E46637C85B11BBFD311E10FD58" box="[694,787,661,687]" pageId="1" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD368D6637C85B1390FD1C1C10FD38" blockId="1.[151,787,660,719]" box="[157,275,696,719]" pageId="1" pageNumber="597">
(
<figureCitation id="13592A086637C85B13A8FD1C1C09FD38" box="[165,266,696,719]" captionStart-0="FIGURES 1 – 4. 1" captionStart-1="FIGURE 5" captionStartId-0="2.[151,269,1931,1954]" captionStartId-1="3.[151,255,1198,1221]" captionTargetBox-0="[174,1343,202,1870]" captionTargetBox-1="[151,1436,157,1173]" captionTargetId-0="figure@2.[172,1396,194,1878]" captionTargetId-1="figure@3.[151,1436,157,1173]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 4. 1, Glyphobythus anatolicus sp. nov., male holotype, dorsal habitus (scale = 0.5 mm); 2, female, paratype, dorsal habitus (scale = 0.5 mm); 3, aedeagus, dorsal view (scale = 0.2 mm); 4, aedeagus, lateral view (scale = 0.2 mm)." captionText-1="FIGURE 5. Macrohabitat of the type locality of Glyphobythus anatolicus sp. nov." httpUri-0="https://zenodo.org/record/245330/files/figure.png" httpUri-1="https://zenodo.org/record/245331/files/figure.png" pageId="1" pageNumber="597">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37865066637C85B139AFD5919FBFCAB" pageId="1" pageNumber="597" type="etymology">
<paragraph id="8BDD368D6637C85B139AFD5919FBFCAB" blockId="1.[151,1436,765,1865]" pageId="1" pageNumber="597">
<emphasis id="B916EA9F6637C85B139AFD591C18FCE3" bold="true" box="[151,283,765,788]" pageId="1" pageNumber="597">Etymology:</emphasis>
Named after
<collectingCountry id="F375761D6637C85B12A1FD591F09FCE3" box="[428,522,765,788]" name="Turkey" pageId="1" pageNumber="597">Anatolia</collectingCountry>
, a historic name for
<collectingCountry id="F375761D6637C85B11E8FD591E31FCE3" box="[741,818,765,788]" name="Turkey" pageId="1" pageNumber="597">Turkey</collectingCountry>
, the country where the new species has been discovered.
<emphasis id="B916EA9F6637C85B139AFC861C55FCCE" bold="true" box="[151,342,802,825]" pageId="1" pageNumber="597">Material studied</emphasis>
:
<typeStatus id="54D9882F6637C85B126AFC861CF4FCCE" box="[359,503,802,825]" pageId="1" pageNumber="597" type="holotype">HOLOTYPE</typeStatus>
,
<specimenCount id="9D64FD046637C85B1109FC861F50FCCE" box="[516,595,802,825]" pageId="1" pageNumber="597" type="male">1 male</specimenCount>
:
<geoCoordinate id="EE56504A6637C85B1152FC861FBFFCCE" box="[607,700,800,825]" direction="north" orientation="latitude" pageId="1" pageNumber="597" precision="925" value="41.216667">N 41o13'</geoCoordinate>
20
<geoCoordinate id="EE56504A6637C85B11ECFC861E3DFCCE" box="[737,830,800,825]" direction="east" orientation="longitude" pageId="1" pageNumber="597" precision="925" value="31.883333">E 31o53'</geoCoordinate>
6,
<collectingCountry id="F375761D6637C85B1057FC861E7FFCCE" box="[858,892,802,825]" name="Chile" pageId="1" pageNumber="597">TR</collectingCountry>
Zonguldak
<date id="FFDC104D6637C85B170FFC861962FCCE" box="[1026,1121,802,825]" pageId="1" pageNumber="597" value="2014-05-03">3.5.2014</date>
, Devrek.,
<quantity id="4C9A9B686637C85B17D9FC86181EFCCE" box="[1236,1309,802,825]" metricMagnitude="2" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="597" unit="m" value="590.0">590 m</quantity>
, Brachat &amp; Meybohm (NMPC).
<typeStatus id="54D9882F6637C85B127BFCE11F0FFCAB" box="[374,524,837,860]" pageId="1" pageNumber="597" type="paratype">PARATYPES</typeStatus>
,
<specimenCount id="9D64FD046637C85B111BFCE11F7BFCAB" box="[534,632,837,860]" pageId="1" pageNumber="597" type="male">32 males</specimenCount>
,
<specimenCount id="9D64FD046637C85B118EFCE11FF9FCAB" box="[643,762,837,860]" pageId="1" pageNumber="597" type="female">11 females</specimenCount>
: same data as
<typeStatus id="54D9882F6637C85B109CFCE11EEDFCAB" box="[913,1006,837,860]" pageId="1" pageNumber="597" type="holotype">holotype</typeStatus>
(OUMNH, PHC, VBC).
</paragraph>
</subSubSection>
<subSubSection id="C37865066637C85B13CAFCCC1C0DF9C3" pageId="1" pageNumber="597" type="description">
<paragraph id="8BDD368D6637C85B13CAFCCC1953FA18" blockId="1.[151,1436,765,1865]" pageId="1" pageNumber="597">
<emphasis id="B916EA9F6637C85B13CAFCCC1C51FC88" bold="true" box="[199,338,872,895]" pageId="1" pageNumber="597">Description.</emphasis>
Male: Body (
<figureCitation id="13592A086637C85B12E5FCCC1F25FC88" box="[488,550,872,895]" captionStart="FIGURES 1 – 4. 1" captionStartId="2.[151,269,1931,1954]" captionTargetBox="[174,1343,202,1870]" captionTargetId="figure@2.[172,1396,194,1878]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1, Glyphobythus anatolicus sp. nov., male holotype, dorsal habitus (scale = 0.5 mm); 2, female, paratype, dorsal habitus (scale = 0.5 mm); 3, aedeagus, dorsal view (scale = 0.2 mm); 4, aedeagus, lateral view (scale = 0.2 mm)." httpUri="https://zenodo.org/record/245330/files/figure.png" pageId="1" pageNumber="597">Fig. 1</figureCitation>
) glossy, dark reddish-brown, legs, antennae and maxillary palpi yellowish-brown, length
<quantity id="4C9A9B686637C85B13E9FC2E1C86FC56" box="[228,389,906,929]" metricMagnitude="-3" metricUnit="m" metricValue="1.125" metricValueMax="1.15" metricValueMin="1.1" pageId="1" pageNumber="597" unit="mm" value="1.125" valueMax="1.15" valueMin="1.1">1.10–1.15 mm</quantity>
, maximum width of elytra about
<quantity id="4C9A9B686637C85B11F3FC2E1E67FC56" box="[766,868,906,929]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="597" unit="mm" value="0.55">0.55 mm</quantity>
. Head and pronotum with sparse, long and uneven pubescence, pubescence on elytra and abdominal tergites even and denser. Head slightly wider than long in both sexes, triangular, vertex about twice as wide as frons measured across supra-antennal prominent tubercles, rostrum with a large deep sulcus reaching vertexal foveae which are located in line of anterior margin of eyes, distance between vertexal foveae slightly wider than distance from fovea to lateral margin of head, vertex with median keel reaching posterior margin of vertexal foveae. Maxillary palpi long, slightly shorter than antennae, palpomere II long, pipe-shaped, palpomere III short, both granulate on dorsal surface, terminal palpomere subtriangular, with well-defined basal setose fovea,
<quantity id="4C9A9B686637C85B13EDFBD91C7FFB63" box="[224,380,1149,1172]" metricMagnitude="-4" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="1" pageNumber="597" unit="mm" value="0.205" valueMax="0.21" valueMin="0.2">0.20–0.21 mm</quantity>
long. Antennae short, scape very long, 3.75 times as long as wide, slightly expanded from base to apex, three times as long as pedicel, antennomere III elongate, slightly pedunculate, 1.4 times as long as IV, antennomeres IV–VIII subequal in size, spherical, IX and X transverse, IX twice as wide as long, X 2.4 times as wide as long, terminal antennomere 1.8 as long as wide, pointed, relative length of antennomeres: 15.0 / 5.0 / 3.5 / 2.5 / 3.0 / 2.0 / 2.0 / 2.0 / 2.0 / 2.5 / 14.0. Pronotum slightly wider than long and about 1.2 times as long as head, widest in anterior third, lateral foveae well-defined, connected by shallow, well-defined antebasal sulcus, median antebasal fovea absent. Elytra about as long as wide, about 1.8 times as long as pronotum, with two large deep basal setose foveae and well defined entire sutural stria on each elytron. Visible part of abdomen slightly narrower and much shorter than elytra, length of visible abdominal tergites 0.3–0.4 of elytral length, first visible tergite (IV) slightly longer than second (V). Aedeagus (
<figureCitation id="13592A086637C85B121CFA111C79FA3B" box="[273,378,1461,1484]" captionStart="FIGURES 1 – 4. 1" captionStartId="2.[151,269,1931,1954]" captionTargetBox="[174,1343,202,1870]" captionTargetId="figure@2.[172,1396,194,1878]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1, Glyphobythus anatolicus sp. nov., male holotype, dorsal habitus (scale = 0.5 mm); 2, female, paratype, dorsal habitus (scale = 0.5 mm); 3, aedeagus, dorsal view (scale = 0.2 mm); 4, aedeagus, lateral view (scale = 0.2 mm)." httpUri="https://zenodo.org/record/245330/files/figure.png" pageId="1" pageNumber="597">Figs. 3, 4</figureCitation>
) elongate,
<quantity id="4C9A9B686637C85B12FBFA111F55FA3B" box="[502,598,1461,1484]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="597" unit="mm" value="0.45">0.45 mm</quantity>
long, median lobe almost symmetrical, its apex bifurcate, parameres curved outwards at apices, with two preapical setae, internal sac bearing four accessory sclerites.
</paragraph>
<paragraph id="8BDD368D6637C85B13CAFA5E1C0DF9C3" blockId="1.[151,1436,765,1865]" pageId="1" pageNumber="597">
<emphasis id="B916EA9F6637C85B13CAFA5E1CA6F9E6" bold="true" box="[199,421,1530,1553]" pageId="1" pageNumber="597">Sexual dimorphism</emphasis>
: Females (
<figureCitation id="13592A086637C85B1111FA5E1F5EF9E6" box="[540,605,1530,1553]" captionStart="FIGURES 1 – 4. 1" captionStartId="2.[151,269,1931,1954]" captionTargetBox="[174,1343,202,1870]" captionTargetId="figure@2.[172,1396,194,1878]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1, Glyphobythus anatolicus sp. nov., male holotype, dorsal habitus (scale = 0.5 mm); 2, female, paratype, dorsal habitus (scale = 0.5 mm); 3, aedeagus, dorsal view (scale = 0.2 mm); 4, aedeagus, lateral view (scale = 0.2 mm)." httpUri="https://zenodo.org/record/245330/files/figure.png" pageId="1" pageNumber="597">Fig. 2</figureCitation>
) have narrower head and much smaller eyes, there is no difference in size of both sexes.
</paragraph>
</subSubSection>
<subSubSection id="C37865066637C85B13CAF9E41E1CF8D0" pageId="1" pageNumber="597" type="biology_ecology">
<paragraph id="8BDD368D6637C85B13CAF9E419B3F96B" blockId="1.[151,1436,765,1865]" pageId="1" pageNumber="597">
<emphasis id="B916EA9F6637C85B13CAF9E41C21F9A0" bold="true" box="[199,290,1600,1623]" pageId="1" pageNumber="597">Biology.</emphasis>
Females of
<taxonomicName id="4C624D0E6637C85B12A0F9E41F42F9A0" box="[429,577,1600,1623]" class="Insecta" family="Staphylinidae" genus="Glyphobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9F6637C85B12A0F9E41F42F9A0" box="[429,577,1600,1623]" italics="true" pageId="1" pageNumber="597">Glyphobythus</emphasis>
</taxonomicName>
have been exclusively found in caves while males can also be found outside, living in leaf litter or under deeply embedded stones (
<bibRefCitation id="EFF34B7C6637C85B11D5F9C61E68F98E" author="Jeannel" box="[728,875,1634,1657]" pageId="1" pageNumber="597" refString="Jeannel, R. (1950) Faune de France, 53. Coleopteres Pselaphides. Librarie de la Faculte des Sciences, Paris, i - iii, 421." type="journal article" year="1950">Jeannel 1950</bibRefCitation>
). All specimens of
<taxonomicName id="4C624D0E6637C85B1731F9C619CAF98E" box="[1084,1225,1634,1657]" class="Insecta" family="Staphylinidae" genus="Glyphobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="anatolicus">
<emphasis id="B916EA9F6637C85B1731F9C619CAF98E" box="[1084,1225,1634,1657]" italics="true" pageId="1" pageNumber="597">G. anatolicus</emphasis>
</taxonomicName>
were collected in a small area at the base of a rocky vertical limestone face (
<figureCitation id="13592A086637C85B11F9F9211E30F96B" box="[756,819,1669,1692]" captionStart="FIGURE 5" captionStartId="3.[151,255,1198,1221]" captionTargetBox="[151,1436,157,1173]" captionTargetId="figure@3.[151,1436,157,1173]" captionTargetPageId="3" captionText="FIGURE 5. Macrohabitat of the type locality of Glyphobythus anatolicus sp. nov." httpUri="https://zenodo.org/record/245331/files/figure.png" pageId="1" pageNumber="597">Fig. 5</figureCitation>
) by sifting mainly
<taxonomicName id="4C624D0E6637C85B10F2F9211941F96B" box="[1023,1090,1669,1692]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="597" phylum="Tracheophyta" rank="genus">
<emphasis id="B916EA9F6637C85B10F2F9211941F96B" box="[1023,1090,1669,1692]" italics="true" pageId="1" pageNumber="597">Fagus</emphasis>
</taxonomicName>
leaf litter.
</paragraph>
<paragraph id="8BDD368D6637C85B13CAF90C1E1CF8D0" blockId="1.[151,1436,765,1865]" pageId="1" pageNumber="597">
<emphasis id="B916EA9F6637C85B13CAF90C1CC2F948" bold="true" box="[199,449,1704,1727]" pageId="1" pageNumber="597">Differential diagnosis.</emphasis>
The new species from
<collectingCountry id="F375761D6637C85B11B4F90C1E06F948" box="[697,773,1704,1727]" name="Turkey" pageId="1" pageNumber="597">Turkey</collectingCountry>
is assigned to the genus
<taxonomicName id="4C624D0E6637C85B1702F90C19A0F948" box="[1039,1187,1704,1727]" class="Insecta" family="Staphylinidae" genus="Glyphobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9F6637C85B1702F90C19A0F948" box="[1039,1187,1704,1727]" italics="true" pageId="1" pageNumber="597">Glyphobythus</emphasis>
</taxonomicName>
basing on the presence of a well-defined dorsal basal fovea on the palpomere IV in males. The new species differs from all other known
<taxonomicName id="4C624D0E6637C85B139AF9491C28F8F3" box="[151,299,1773,1796]" class="Insecta" family="Staphylinidae" genus="Glyphobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9F6637C85B139AF9491C28F8F3" box="[151,299,1773,1796]" italics="true" pageId="1" pageNumber="597">Glyphobythus</emphasis>
</taxonomicName>
in having this fovea located close to the basal margin of palpomere, far from the apex, in unmodified fore tibiae in both sexes, and in a different shape of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C37865066637C85B13CAF8961953F8BE" box="[199,1104,1842,1865]" pageId="1" pageNumber="597" type="distribution">
<paragraph id="8BDD368D6637C85B13CAF8961953F8BE" blockId="1.[151,1436,765,1865]" box="[199,1104,1842,1865]" pageId="1" pageNumber="597">
<emphasis id="B916EA9F6637C85B13CAF8961C5AF8BE" bold="true" box="[199,345,1842,1865]" pageId="1" pageNumber="597">Distribution.</emphasis>
Known only from the
<typeStatus id="54D9882F6637C85B1141F8961F79F8BE" box="[588,634,1842,1865]" pageId="1" pageNumber="597">type</typeStatus>
locality, the province Zonguldak of
<collectingCountry id="F375761D6637C85B10F3F896194FF8BE" box="[1022,1100,1842,1865]" name="Turkey" pageId="1" pageNumber="597">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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