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<mods:title id="1B02BD8B698D3419CAD2F3C78C7D71D4">Revision of F. R. C. Reeds Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<mods:affiliation id="5259C19B1A1ED259850F0A77C47AAE8F">Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. &amp; Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA</mods:affiliation>
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<treatment id="03CB87846E441D1AFF3AC17A4BACF6CD" ID-DOI="http://doi.org/10.5281/zenodo.6816336" ID-GBIF-Taxon="197214852" ID-Zenodo-Dep="6816336" LSID="urn:lsid:plazi:treatment:03CB87846E441D1AFF3AC17A4BACF6CD" httpUri="http://treatment.plazi.org/id/03CB87846E441D1AFF3AC17A4BACF6CD" lastPageId="18" lastPageNumber="318" pageId="17" pageNumber="318">
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<paragraph id="8BDD36926E441D19FF3AC17A4BA1F018" blockId="17.[151,442,1338,1365]" box="[151,442,1338,1365]" pageId="17" pageNumber="318">
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<taxonomicName id="4C624D116E441D19FF3AC17A4BA1F018" ID-CoL="62FZQ" authority="Sheng, 1934" authorityName="Sheng" authorityYear="1934" box="[151,442,1338,1365]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E441D19FF3AC17A4BA1F018" bold="true" box="[151,442,1338,1365]" pageId="17" pageNumber="318">
<emphasis id="B916EA806E441D19FF3AC17A4B06F019" bold="true" box="[151,285,1339,1365]" italics="true" pageId="17" pageNumber="318">Birmanites</emphasis>
<bibRefCitation id="EFF34B636E441D19FE89C17B4BA1F018" author="Sheng, X. - F." box="[292,442,1338,1364]" pageId="17" pageNumber="318" pagination="1 - 19" refId="ref47171" refString="Sheng, X. - F. (1934) Lower Ordovician faunas of Chekiang. Palaeontologica Sinica New Series, 3, 1 - 19." type="journal article" year="1934">Sheng, 1934</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8BDD36926E441D19FF3AC13E4F6AF0DB" blockId="17.[151,1393,1407,1432]" box="[151,1393,1407,1432]" pageId="17" pageNumber="318">
<materialsCitation id="3B0A3CCF6E441D19FF3AC13E4F77F0DB" ID-GBIF-Occurrence="3861306315" box="[151,1388,1407,1432]" country="Myanmar" location="Hwe Mawng Beds'" pageId="17" pageNumber="318" specimenCount="1" stateProvince="Shan" typeStatus="holotype">
<emphasis id="B916EA806E441D19FF3AC13E4B05F0DB" bold="true" box="[151,286,1407,1431]" pageId="17" pageNumber="318">
<typeStatus id="54D988306E441D19FF3AC13E4AD7F0DB" box="[151,204,1407,1431]" pageId="17" pageNumber="318">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C624D116E441D19FE87C1C14862F0DB" authority="Reed, 1915" authorityName="Reed" authorityYear="1915" box="[298,633,1407,1431]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="B916EA806E441D19FE87C1C14BE7F0DB" box="[298,508,1407,1431]" italics="true" pageId="17" pageNumber="318">Ogygites birmanicus</emphasis>
<bibRefCitation id="EFF34B636E441D19FDAFC13E4862F0DB" author="Reed, F. R. C." box="[514,633,1407,1431]" pageId="17" pageNumber="318" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
</taxonomicName>
.
<location id="8EBD60496E441D19FD2AC13E4949F0DB" LSID="urn:lsid:plazi:treatment:03CB87846E441D1AFF3AC17A4BACF6CD:8EBD60496E441D19FD2AC13E4949F0DB" box="[647,850,1407,1432]" country="Myanmar" name="Hwe Mawng Beds'" pageId="17" pageNumber="318" stateProvince="Shan">Hwe Mawng Beds</location>
, northern
<collectingRegion id="49A6F8706E441D19FC14C13E4E3CF0DB" box="[953,1063,1407,1431]" country="Myanmar" name="Shan" pageId="17" pageNumber="318">Shan State</collectingRegion>
,
<collectingCountry id="F37576026E441D19FB9CC1C14E8EF0D4" box="[1073,1173,1408,1432]" name="Myanmar" pageId="17" pageNumber="318">Myanmar</collectingCountry>
, original designation
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.
</paragraph>
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<subSubSection id="C37865196E441D19FF3AC18449BBF33C" pageId="17" pageNumber="318" type="diagnosis">
<paragraph id="8BDD36926E441D19FF3AC18449BBF33C" blockId="17.[151,1437,1477,2008]" pageId="17" pageNumber="318">
<emphasis id="B916EA806E441D19FF3AC1844B09F093" bold="true" box="[151,274,1477,1503]" pageId="17" pageNumber="318">Diagnosis.</emphasis>
<bibRefCitation id="EFF34B636E441D19FEB5C1844B8FF0AC" author="Lu, Y. - H." box="[280,404,1477,1504]" pageId="17" pageNumber="318" pagination="1 - 453" refId="ref45640" refString="Lu, Y. - H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, New Series B, 11, 1 - 453." type="journal article" year="1975">Lus (1975</bibRefCitation>
, p. 3189) diagnosis is still largely appropriate. It should be modified to include the inverted U shape of the wide hypostomal fork, and to emphasise that the pygidial ring furrows are transverse, rather than chevronshaped. Despite the thin cuticle, the terrace ridges on the pygidial doublure are particularly strong. Only the best-preserved material shows a pointed junction of the facial sutures on the midline. In most flattened material the sutures appear to run nearly marginally over their anterior course.
</paragraph>
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<paragraph id="8BDD36926E441D1AFF6AC2384BACF6CD" blockId="17.[151,1437,1477,2008]" lastBlockId="18.[151,1437,150,897]" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318">
<emphasis id="B916EA806E441D19FF6AC2384B57F3DF" bold="true" box="[199,332,1657,1683]" pageId="17" pageNumber="318">Discussion.</emphasis>
<taxonomicName id="4C624D116E441D19FEFBC23B4842F3DF" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[342,601,1657,1683]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="B916EA806E441D19FEFBC23B4842F3DF" box="[342,601,1657,1683]" italics="true" pageId="17" pageNumber="318">Birmanites birmanicus</emphasis>
</taxonomicName>
was originally referred by
<bibRefCitation id="EFF34B636E441D19FC35C2384E3DF3D8" author="Reed, F. R. C." box="[920,1062,1657,1684]" pageId="17" pageNumber="318" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed (1915)</bibRefCitation>
to the genus
<taxonomicName id="4C624D116E441D19FB65C23B4B64F3FB" authority="Tromelin &amp; Lebesconte, 1876" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E441D19FB65C23B4F37F3D8" box="[1224,1324,1658,1684]" italics="true" pageId="17" pageNumber="318">Ogygites</emphasis>
<bibRefCitation id="EFF34B636E441D19FA98C2384B64F3FB" author="Tromelin, G. de &amp; Lebesconte, P." pageId="17" pageNumber="318" pagination="683 - 687" refId="ref47881" refString="Tromelin, G. de. &amp; Lebesconte, P. (1876) Presentation de fossiles paleozoiques du department d'Ille-et-Vilaine et note additionnelle sur la faune silurianne du l'Ouest de la France. Comptes Rendus 4 me session Association Francaise Advancement Science, Nantes, 1876, 683 - 687." type="journal article" year="1876">Tromelin &amp; Lebesconte, 1876</bibRefCitation>
</taxonomicName>
(type species
<taxonomicName id="4C624D116E441D19FD8AC2DF4965F3FB" authority="Brongniart" authorityName="Brongniart" box="[551,894,1693,1720]" class="Trilobita" family="Asaphidae" genus="Ogygia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="desmaresti">
<emphasis id="B916EA806E441D19FD8AC2DF48E0F3FB" box="[551,763,1693,1720]" italics="true" pageId="17" pageNumber="318">Ogygia desmaresti</emphasis>
Brongniart
</taxonomicName>
<emphasis id="B916EA806E441D19FC24C2DF4984F3FB" box="[905,927,1694,1719]" italics="true" pageId="17" pageNumber="318">in</emphasis>
Brongniart &amp; Desmarest 1822), and we are therefore obliged to discuss both genera here. The type material of the type species of
<taxonomicName id="4C624D116E441D19FBD5C2834EC7F390" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" box="[1144,1244,1730,1756]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E441D19FBD5C2834EC7F390" box="[1144,1244,1730,1756]" italics="true" pageId="17" pageNumber="318">Ogygites</emphasis>
</taxonomicName>
is both distorted and incomplete, and a very detailed discussion of it by
<bibRefCitation id="EFF34B636E441D19FCA5C2A449B1F3B3" author="Rabano, I." box="[776,938,1765,1791]" pageId="17" pageNumber="318" pagination="541 - 609" refId="ref46614" refString="Rabano, I. (1989) Trilobites del Ordovicico Medio del sector meridional de la Zona Centroiberica espanola. Part II Agnostina y Asaphida Publicaciones Especiales del Boletin Geologico y Minero, 100, 541 - 609." type="journal article" year="1989">Rabáno (1989</bibRefCitation>
. p. 7682) concluded with placing
<taxonomicName id="4C624D116E441D19FA94C2A74F86F24C" authority="Incertae" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" box="[1337,1437,1766,1792]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E441D19FA94C2A74F86F24C" box="[1337,1437,1766,1792]" italics="true" pageId="17" pageNumber="318">Ogygites</emphasis>
</taxonomicName>
Incertae subfamiliae” within
<taxonomicName id="4C624D116E441D19FE4CC3484840F26F" authorityName="Salter" authorityYear="1864" box="[481,603,1801,1827]" class="Trilobita" family="Asaphidae" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="family">Asaphidae</taxonomicName>
. She regarded the clarification of the identity of
<taxonomicName id="4C624D116E441D19FBC2C34B4EC8F268" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" box="[1135,1235,1802,1828]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E441D19FBC2C34B4EC8F268" box="[1135,1235,1802,1828]" italics="true" pageId="17" pageNumber="318">Ogygites</emphasis>
</taxonomicName>
as beyond resolution and recommended restricting its usage to the distorted
<typeStatus id="54D988306E441D19FC81C36C4989F20B" box="[812,914,1837,1863]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
.
<bibRefCitation id="EFF34B636E441D19FC36C36C4E59F20B" author="Rabano, I." box="[923,1090,1837,1863]" pageId="17" pageNumber="318" pagination="541 - 609" refId="ref46614" refString="Rabano, I. (1989) Trilobites del Ordovicico Medio del sector meridional de la Zona Centroiberica espanola. Part II Agnostina y Asaphida Publicaciones Especiales del Boletin Geologico y Minero, 100, 541 - 609." type="journal article" year="1989">Rabáno (1989)</bibRefCitation>
was unaware of a forthcoming revision of
<taxonomicName id="4C624D116E441D19FEBAC3134BAAF227" box="[279,433,1873,1900]" class="Trilobita" family="Asaphidae" genus="Ogygia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="desmaresti">
<emphasis id="B916EA806E441D19FEBAC3134BAAF227" box="[279,433,1873,1900]" italics="true" pageId="17" pageNumber="318">O. desmaresti</emphasis>
</taxonomicName>
from the type area of Angers by
<bibRefCitation id="EFF34B636E441D19FC8EC31049A9F227" author="Pillet, J." box="[803,946,1873,1899]" pageId="17" pageNumber="318" pagination="1 - 60" refId="ref46442" refString="Pillet, J. (1990) La faune des Ardoises d'Angers. Memoire de la Societe d'etudes scientifiques de l'Anjou, 7, 1 - 60." type="journal article" year="1990">Pillet (1990)</bibRefCitation>
, who attributed much additional material to the same species. Although it is distorted, much of it is articulated, and if Pillet is correct in attributing his material to
<taxonomicName id="4C624D116E441D19FF19C3DB4B54F2FF" box="[180,335,1945,1972]" class="Trilobita" family="Asaphidae" genus="Ogygia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="desmaresti">
<emphasis id="B916EA806E441D19FF19C3DB4B54F2FF" box="[180,335,1945,1972]" italics="true" pageId="17" pageNumber="318">O. desmaresti</emphasis>
</taxonomicName>
, as is likely, it is clear that the type species of
<taxonomicName id="4C624D116E441D19FCF4C3DB49A6F2F8" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" box="[857,957,1946,1972]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E441D19FCF4C3DB49A6F2F8" box="[857,957,1946,1972]" italics="true" pageId="17" pageNumber="318">Ogygites</emphasis>
</taxonomicName>
shares its principal features with both
<taxonomicName id="4C624D116E441D19FADDC3DB4AF5F29B" authorityName="Sheng" authorityYear="1934" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E441D19FADDC3DB4AF5F29B" italics="true" pageId="17" pageNumber="318">Birmanites</emphasis>
</taxonomicName>
and
<taxonomicName id="4C624D116E441D19FE88C3FC4BDCF29B" authorityName="Pribyl &amp; Vanek" authorityYear="1965" box="[293,455,1981,2007]" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E441D19FE88C3FC4BDCF29B" box="[293,455,1981,2007]" italics="true" pageId="17" pageNumber="318">Nobiliasaphus</emphasis>
</taxonomicName>
. The pygidium is not seen on the type material, but several specimens figured by Pillet (e.g. 1990, pl. 15) prove the presence of numerous backwardly directed chevron-shaped ridges/furrows on the long and narrow pygidial axis that are clearly present also on the type specimens of
<taxonomicName id="4C624D116E471D1AFC56C4FA4F75F599" authority="(Barrande)" baseAuthorityName="Barrande" box="[1019,1390,187,213]" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="B916EA806E471D1AFC56C4FA4EF4F599" box="[1019,1263,187,213]" italics="true" pageId="18" pageNumber="319">Nobiliasaphus nobilis</emphasis>
(Barrande)
</taxonomicName>
and
<emphasis id="B916EA806E471D1AFF3AC49E4BADF5B4" box="[151,438,223,249]" italics="true" pageId="18" pageNumber="319">Pamirotechites pamiricus</emphasis>
Balashova, but not on
<taxonomicName id="4C624D116E471D1AFD1FC49E49AAF5B5" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[690,945,223,249]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="B916EA806E471D1AFD1FC49E49AAF5B5" box="[690,945,223,249]" italics="true" pageId="18" pageNumber="319">Birmanites birmanicus</emphasis>
</taxonomicName>
or other oriental species of
<taxonomicName id="4C624D116E471D1AFB4EC49E4F45F5B4" authorityName="Sheng" authorityYear="1934" box="[1251,1374,223,248]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFB4EC49E4F45F5B4" box="[1251,1374,223,248]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
. This is such an unusual character that in our view it is likely a synapomorphy of species sharing it, and constitutes the defining character of
<taxonomicName id="4C624D116E471D1AFE26C5664836F40D" authorityName="Pribyl &amp; Vanek" authorityYear="1965" box="[395,557,295,321]" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFE26C5664836F40D" box="[395,557,295,321]" italics="true" pageId="18" pageNumber="319">Nobiliasaphus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C624D116E471D1AFD98C5664882F40D" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" box="[565,665,295,321]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFD98C5664882F40D" box="[565,665,295,321]" italics="true" pageId="18" pageNumber="319">Ogygites</emphasis>
</taxonomicName>
. For example,
<bibRefCitation id="EFF34B636E471D1AFCE9C56649F6F40D" author="Rabano, I." box="[836,1005,295,321]" pageId="18" pageNumber="319" pagination="541 - 609" refId="ref46614" refString="Rabano, I. (1989) Trilobites del Ordovicico Medio del sector meridional de la Zona Centroiberica espanola. Part II Agnostina y Asaphida Publicaciones Especiales del Boletin Geologico y Minero, 100, 541 - 609." type="journal article" year="1989">Rabáno (1989)</bibRefCitation>
illustrated several species from
<collectingCountry id="F37576026E471D1AFAF1C5664F87F40D" box="[1372,1436,295,321]" name="Spain" pageId="18" pageNumber="319">Spain</collectingCountry>
showing this feature, and
<bibRefCitation id="EFF34B636E471D1AFE14C50A48F1F429" author="Hammann, W. &amp; Leone, F." box="[441,746,331,357]" pageId="18" pageNumber="319" pagination="1 - 217" refId="ref44454" refString="Hammann, W. &amp; Leone, F. (1997) Trilobites from the post-Sardic (Upper Ordovician) sequence of southern Sardinia. Part 1. Beringeria, 20, 1 - 217." type="journal article" year="1997">Hammann &amp; Leone (1997)</bibRefCitation>
illustrated others from Sardinia. One of Rabános species,
<taxonomicName id="4C624D116E471D1AFA2EC50A4B16F4C5" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="hammanni">
<emphasis id="B916EA806E471D1AFA2EC50A4B16F4C5" italics="true" pageId="18" pageNumber="319">N. hammanni</emphasis>
</taxonomicName>
, has a posterior pygidial spine, which is not considered generically significant. However unsatisfactory its type material, it is likely that
<taxonomicName id="4C624D116E471D1AFE50C5D2487AF4E1" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" box="[509,609,403,429]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFE50C5D2487AF4E1" box="[509,609,403,429]" italics="true" pageId="18" pageNumber="319">Ogygites</emphasis>
</taxonomicName>
is the senior name for this clade. If
<taxonomicName id="4C624D116E471D1AFC43C5D24E72F4E0" authorityName="Sheng" authorityYear="1934" box="[1006,1129,403,428]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFC43C5D24E72F4E0" box="[1006,1129,403,428]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
is restricted to species with transverse pygidial ring furrows then some species may be better placed in
<taxonomicName id="4C624D116E471D1AFC43C5F64E72F49C" authorityName="Sheng" authorityYear="1934" box="[1006,1129,439,464]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFC43C5F64E72F49C" box="[1006,1129,439,464]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C624D116E471D1AFB5EC5F64F8EF49D" authorityName="Pribyl &amp; Vanek" authorityYear="1965" box="[1267,1429,439,465]" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFB5EC5F64F8EF49D" box="[1267,1429,439,465]" italics="true" pageId="18" pageNumber="319">Nobiliasaphus</emphasis>
</taxonomicName>
; for example,
<taxonomicName id="4C624D116E471D1AFE9DC59A48C9F4B9" authority="Hughes, 1979" authorityName="Hughes" authorityYear="1979" box="[304,722,475,501]" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="powysi">
<emphasis id="B916EA806E471D1AFE9DC59A4833F4B8" box="[304,552,475,501]" italics="true" pageId="18" pageNumber="319">Nobiliasaphus powysi</emphasis>
<bibRefCitation id="EFF34B636E471D1AFD9CC59A48C9F4B9" author="Hughes, C. P." box="[561,722,475,501]" pageId="18" pageNumber="319" pagination="109 - 181" refId="ref44696" refString="Hughes, C. P. (1979) The Ordovician trilobite faunas of the Builth-Llandrindod Inlier, central Wales. Part 3. Bulletin of the British Museum (Natural History) Geology Series, 32, 109 - 181." type="journal article" year="1979">Hughes, 1979</bibRefCitation>
</taxonomicName>
, from the Ordovician of the Builth Inlier,
<collectingCountry id="F37576026E471D1AFB1BC59A4EE6F4B9" box="[1206,1277,475,501]" name="United Kingdom" pageId="18" pageNumber="319">Wales</collectingCountry>
. The range of
<taxonomicName id="4C624D116E471D1AFF3AC5BE4B22F755" authorityName="Pribyl &amp; Vanek" authorityYear="1965" box="[151,313,511,537]" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFF3AC5BE4B22F755" box="[151,313,511,537]" italics="true" pageId="18" pageNumber="319">Nobiliasaphus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C624D116E471D1AFEECC5BE4BBEF755" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" box="[321,421,511,537]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFEECC5BE4BBEF755" box="[321,421,511,537]" italics="true" pageId="18" pageNumber="319">Ogygites</emphasis>
</taxonomicName>
does not extend as far to the east (present geography) over Ordovician Gondwana as does that of
<taxonomicName id="4C624D116E471D1AFF40C6624B73F770" authorityName="Sheng" authorityYear="1934" box="[237,360,547,572]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFF40C6624B73F770" box="[237,360,547,572]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
. Finally,
<taxonomicName id="4C624D116E471D1AFE7AC6624901F771" authority="Kielan, 1960" authorityName="Kielan" authorityYear="1960" box="[471,794,547,573]" class="Trilobita" family="Asaphidae" genus="Opsimasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFE7AC6624867F771" box="[471,636,547,573]" italics="true" pageId="18" pageNumber="319">Opsimasaphus</emphasis>
<bibRefCitation id="EFF34B636E471D1AFD2BC6624901F771" author="Kielan, Z." box="[646,794,547,573]" pageId="18" pageNumber="319" pagination="1 - 191" refId="ref44947" refString="Kielan, Z. (1960) Upper Ordovician trilobites from Poland and some related forms from Bohemia and Scandinavia. Acta Palaeontologica Polonica, 11, 1 - 191." type="journal article" year="1960">Kielan, 1960</bibRefCitation>
</taxonomicName>
(type species
<taxonomicName id="4C624D116E471D1AFC6EC6624F1AF771" authority="Kielan, 1960" authorityName="Kielan" authorityYear="1960" box="[963,1281,547,573]" class="Trilobita" family="Asaphidae" genus="Opsimasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="jaanussoni">
<emphasis id="B916EA806E471D1AFC6EC6624E7AF770" box="[963,1121,547,573]" italics="true" pageId="18" pageNumber="319">O. jaanussoni</emphasis>
<bibRefCitation id="EFF34B636E471D1AFBC1C6624F1AF771" author="Kielan, Z." box="[1132,1281,547,573]" pageId="18" pageNumber="319" pagination="1 - 191" refId="ref44947" refString="Kielan, Z. (1960) Upper Ordovician trilobites from Poland and some related forms from Bohemia and Scandinavia. Acta Palaeontologica Polonica, 11, 1 - 191." type="journal article" year="1960">Kielan, 1960</bibRefCitation>
</taxonomicName>
) differs from
<taxonomicName id="4C624D116E471D1AFF3AC6064B09F72C" authorityName="Sheng" authorityYear="1934" box="[151,274,583,608]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFF3AC6064B09F72C" box="[151,274,583,608]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
in having a relatively short (sag.) pygidium—the structure of the pygidial axis (with the exception of Chinese
<taxonomicName id="4C624D116E471D1AFF5AC62A4BFBF7C9" authorityName="Lu" authorityYear="1975" box="[247,480,619,645]" class="Trilobita" family="Asaphidae" genus="Opsimasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pseudodawanicus">
<emphasis id="B916EA806E471D1AFF5AC62A4BFBF7C9" box="[247,480,619,645]" italics="true" pageId="18" pageNumber="319">O. pseudodawanicus</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EFF34B636E471D1AFDBBC62A48BDF7C9" author="Turvey, S. T." box="[534,678,619,645]" pageId="18" pageNumber="319" pagination="347 - 399" refId="ref48057" refString="Turvey, S. T. (2007) Asaphoid trilobites from the Arenig-Llanvirn of the South China Plate. Palaeontology, 50, 347 - 399. https: // doi. org / 10.1111 / j. 1475 - 4983.2006.00641. x" type="journal article" year="2007">Turvey 2007</bibRefCitation>
) is similar to that of
<taxonomicName id="4C624D116E471D1AFC27C62A4E1EF7C8" authorityName="Sheng" authorityYear="1934" box="[906,1029,619,644]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFC27C62A4E1EF7C8" box="[906,1029,619,644]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C624D116E471D1AFB25C62A4F31F7C9" authorityName="Pribyl &amp; Vanek" authorityYear="1965" box="[1160,1322,619,645]" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFB25C62A4F31F7C9" box="[1160,1322,619,645]" italics="true" pageId="18" pageNumber="319">Nobiliasaphus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C624D116E471D1AFA9FC62A4F8DF7C9" authorityName="Tromelin &amp; Lebesconte" authorityYear="1876" box="[1330,1430,619,645]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFA9FC62A4F8DF7C9" box="[1330,1430,619,645]" italics="true" pageId="18" pageNumber="319">Ogygites</emphasis>
</taxonomicName>
, and the cephalic features seem to us identical to those of
<taxonomicName id="4C624D116E471D1AFC8AC6CE49B9F7E4" authorityName="Sheng" authorityYear="1934" box="[807,930,655,680]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFC8AC6CE49B9F7E4" box="[807,930,655,680]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF34B636E471D1AFC1CC6CE4E52F7E5" author="Kielan, Z." box="[945,1097,655,681]" pageId="18" pageNumber="319" pagination="1 - 191" refId="ref44947" refString="Kielan, Z. (1960) Upper Ordovician trilobites from Poland and some related forms from Bohemia and Scandinavia. Acta Palaeontologica Polonica, 11, 1 - 191." type="journal article" year="1960">Kielan (1960</bibRefCitation>
, p. 75-77) gave a detailed account of
<taxonomicName id="4C624D116E471D1AFF52C6F24BBFF781" authorityName="Kielan" authorityYear="1960" box="[255,420,691,717]" class="Trilobita" family="Asaphidae" genus="Opsimasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFF52C6F24BBFF781" box="[255,420,691,717]" italics="true" pageId="18" pageNumber="319">Opsimasaphus</emphasis>
</taxonomicName>
in relation to what would now be called
<taxonomicName id="4C624D116E471D1AFC2EC6F24F25F781" authority="Pribyl and Vanek, 1965" authorityName="Pribyl and Vanek" authorityYear="1965" box="[899,1342,690,717]" class="Trilobita" family="Asaphidae" genus="Nobiliasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFC2EC6F24E3EF781" box="[899,1061,691,717]" italics="true" pageId="18" pageNumber="319">Nobiliasaphus</emphasis>
<bibRefCitation id="EFF34B636E471D1AFB83C6F24F25F781" author="Pribyl, A. &amp; Vanek, J." box="[1070,1342,690,717]" pageId="18" pageNumber="319" pagination="277 - 282" refId="ref46547" refString="Pribyl, A. &amp; Vanek, J. (1965) Neue Trilobiten des bohmischen Ordoviziums. Vestnik Ustredniho Ustavu Geologickeho, 40, 277 - 282." type="journal article" year="1965">Přibyl and Vaněk, 1965</bibRefCitation>
</taxonomicName>
, but did not bring
<taxonomicName id="4C624D116E471D1AFEA5C6964B98F7BC" authorityName="Sheng" authorityYear="1934" box="[264,387,727,752]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFEA5C6964B98F7BC" box="[264,387,727,752]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
into the discussion, possibly because when she was writing this genus was still regarded as a dikelokephalinid.
<bibRefCitation id="EFF34B636E471D1AFED0C6BA4859F659" author="Chugaeva, M. N." box="[381,578,762,789]" pageId="18" pageNumber="319" pagination="1 - 138" refId="ref43103" refString="Chugaeva, M. N. (1958) Trilobites of the Ordovician of the Chu-Ili mountains. Ordovician of Kazakhstan 3. Trudy Geologicheskogo Instituta, 9, 1 - 138, 111 pls. [in Russian]" type="journal article" year="1958">Chugaeva (1958)</bibRefCitation>
described
<taxonomicName id="4C624D116E471D1AFD6EC6BA49ADF659" authorityName="Chugaeva" authorityYear="1958" box="[707,950,763,789]" class="Trilobita" family="Asaphidae" genus="Ogygites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="almatyensis">
<emphasis id="B916EA806E471D1AFD6EC6BA49ADF659" box="[707,950,763,789]" italics="true" pageId="18" pageNumber="319">Ogygites almatyensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F37576026E471D1AFBAFC6BA4E93F659" box="[1026,1160,763,789]" name="Kazakhstan" pageId="18" pageNumber="319">Kazakhstan</collectingCountry>
, which would be better placed in
<taxonomicName id="4C624D116E471D1AFEA8C75E4B9BF674" authorityName="Sheng" authorityYear="1934" box="[261,384,799,824]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFEA8C75E4B9BF674" box="[261,384,799,824]" italics="true" pageId="18" pageNumber="319">Birmanites</emphasis>
</taxonomicName>
on the criteria given here. It is possible that
<taxonomicName id="4C624D116E471D1AFCD5C75E4E06F675" authorityName="Kielan" authorityYear="1960" box="[888,1053,799,825]" class="Trilobita" family="Asaphidae" genus="Opsimasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFCD5C75E4E06F675" box="[888,1053,799,825]" italics="true" pageId="18" pageNumber="319">Opsimasaphus</emphasis>
</taxonomicName>
is a subjective junior synonym of
<emphasis id="B916EA806E471D1AFF3AC7024B02F610" box="[151,281,835,860]" italics="true" pageId="18" pageNumber="319">
<taxonomicName id="4C624D116E471D1AFF3AC7024B0EF610" authorityName="Sheng" authorityYear="1934" box="[151,277,835,860]" class="Trilobita" family="Asaphidae" genus="Birmanites" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">Birmanites</taxonomicName>
,
</emphasis>
as discussed by
<bibRefCitation id="EFF34B636E471D1AFE78C70248DAF611" author="Zhou, Z. - Y. &amp; Dean, W. T." box="[469,705,835,861]" pageId="18" pageNumber="319" pagination="743 - 786" refId="ref49452" refString="Zhou, Z. - Y. &amp; Dean, W. T. (1986) Ordovician trilobites from Chedao, Gansu Province, NW China. Palaeontology, 29, 743 - 786." type="journal article" year="1986">Zhou &amp; Dean (1986)</bibRefCitation>
and
<bibRefCitation id="EFF34B636E471D1AFD55C7024E09F611" author="Romano, M. &amp; Owen, A. W." box="[760,1042,835,861]" pageId="18" pageNumber="319" pagination="681 - 720" refId="ref46902" refString="Romano, M. &amp; Owen, A. W. (1993) Early Caradoc trilobites of eastern Ireland and their palaeogeographical significance. Palaeontology, 36, 681 - 720." type="journal article" year="1993">Romano &amp; Owen (1993)</bibRefCitation>
, while
<taxonomicName id="4C624D116E471D1AFBCEC7024F13F611" authorityName="Kielan" authorityYear="1960" box="[1123,1288,835,861]" class="Trilobita" family="Asaphidae" genus="Opsimasaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Asaphida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E471D1AFBCEC7024F13F611" box="[1123,1288,835,861]" italics="true" pageId="18" pageNumber="319">Opsimasaphus</emphasis>
</taxonomicName>
was retained by
<bibRefCitation id="EFF34B636E471D1AFF17C7264B4EF6CD" author="Turvey, S. T." box="[186,341,871,897]" pageId="18" pageNumber="319" pagination="347 - 399" refId="ref48057" refString="Turvey, S. T. (2007) Asaphoid trilobites from the Arenig-Llanvirn of the South China Plate. Palaeontology, 50, 347 - 399. https: // doi. org / 10.1111 / j. 1475 - 4983.2006.00641. x" type="journal article" year="2007">Turvey (2007</bibRefCitation>
, p. 366).
</paragraph>
</subSubSection>
</treatment>
</document>