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<mods:title id="03E6E58AC027F510C1F3FA5F2858A679">Leech Collections from Washington State, with the Description of Two New Species of Placobdella (Annelida: Glossiphoniidae)</mods:title>
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<taxonomicName id="4C632888B578FFC7D8A0FE340BA9139F" authority="(Verrill, 1872) Castle, 1900" authorityName="Castle" authorityYear="1900" baseAuthorityName="Verrill" baseAuthorityYear="1872" box="[415,977,455,481]" class="Clitellata" family="Glossiphoniidae" genus="Glossiphonia" kingdom="Animalia" order="Rhynchobdellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="elegans">
<emphasis id="B9178F19B578FFC7D8A0FE340AF4139F" box="[415,652,455,481]" italics="true" pageId="8" pageNumber="9">Glossiphonia elegans</emphasis>
(Verrill, 1872) Castle, 1900
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<figureCitation id="13584F8EB578FFC7DBBFFE0F0A881068" box="[640,752,508,534]" captionStart="FIGURES 1419" captionStartId="8.[144,242,1701,1722]" captionTargetBox="[144,1248,937,1674]" captionTargetId="figure-234@8.[144,1248,937,1674]" captionTargetPageId="8" captionText="FIGURES 1419. Glossiphoniids found in Washington in this study; dorsal and ventral views, left and right respectively. 14. Glossiphonia elegans. 15. Helobdella elongata. 16. H. modesta. 17. Helobdella papillata. 18. Placobdella montifera. 19. Theromyzon cf. rude. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5359194" httpUri="https://zenodo.org/record/5359194/files/figure.png" pageId="8" pageNumber="9">Figure 14</figureCitation>
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The presence of three pairs of eyespots, dorsal papillation and pigmentation pattern, two paramedial longitudinal stripes in the ventral surface, two annuli between gonopores, and a relatively large size correspond with the description of
<taxonomicName id="4C632888B578FFC7DA21FD8F0C7410E8" authorityName="Castle" authorityYear="1900" baseAuthorityName="Verrill" baseAuthorityYear="1872" box="[798,1036,636,662]" class="Clitellata" family="Glossiphoniidae" genus="Glossiphonia" kingdom="Animalia" order="Rhynchobdellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="elegans">
<emphasis id="B9178F19B578FFC7DA21FD8F0C7410E8" box="[798,1036,636,662]" italics="true" pageId="8" pageNumber="9">Glossiphonia elegans</emphasis>
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. North American and European forms have very similar morphological traits, to the extent that previous taxonomists (i.e.,
<bibRefCitation id="EFF22EFAB578FFC7D82EFD3409CA109F" author="Klemm, D. J." box="[273,434,711,737]" pageId="8" pageNumber="9" refId="ref5983" refString="Klemm, D. J. 1982. The leeches (Annelida: Hirudinea) of North America. Cincinnati, OH: Aquatic Biology Section, Environmental Monitoring and Support Laboratory. Office of Research and Development, U. S. Environmental Protection Agency." type="book" year="1982">Klemm, 1982</bibRefCitation>
;
<bibRefCitation id="EFF22EFAB578FFC7D881FD340A21109C" author="Sawyer, R. T." box="[446,601,711,738]" pageId="8" pageNumber="9" refId="ref6623" refString="Sawyer, R. T. 1986. Leech biology and behavior. Oxford: Clarendon Press." type="book" year="1986">Sawyer, 1986</bibRefCitation>
) considered them to be the same species:
<taxonomicName id="4C632888B578FFC7DD72FD34099E1179" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Clitellata" family="Glossiphoniidae" genus="Glossiphonia" kingdom="Animalia" order="Rhynchobdellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="complanata">
<emphasis id="B9178F19B578FFC7DD72FD34096E1178" italics="true" pageId="8" pageNumber="9">Glossiphonia complanata</emphasis>
(Linnaeus, 1758)
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. However,
<bibRefCitation id="EFF22EFAB578FFC7DB5AFD1F0B311179" author="Siddall, M. E. &amp; R. B. Budinoff &amp; E. Borda" box="[613,841,748,775]" pageId="8" pageNumber="9" pagination="105 - 112" refId="ref6781" refString="Siddall, M. E., R. B. Budinoff, and E. Borda. 2005. Phylogenetic evaluation of the systematics and biogeography of the leech family Glossiphoniidae. Invertebrate Systematics 19: 105 - 112." type="journal article" year="2005">Siddall et al. (2005)</bibRefCitation>
, using phylogenetic criteria, found that they represent different evolutionary lineages and resurrected Verrills 1872 name
<taxonomicName id="4C632888B578FFC7DDADFCE20941112F" authorityName="Castle" authorityYear="1900" baseAuthorityName="Verrill" baseAuthorityYear="1872" class="Clitellata" family="Glossiphoniidae" genus="Glossiphonia" kingdom="Animalia" order="Rhynchobdellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="elegans">
<emphasis id="B9178F19B578FFC7DDADFCE20941112F" italics="true" pageId="8" pageNumber="9">Glossiphonia elegans</emphasis>
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for the North American species (see also
<bibRefCitation id="EFF22EFAB578FFC7DA1DFCC40C23112F" author="Madill, J. &amp; P. Hovingh" box="[802,1115,823,849]" pageId="8" pageNumber="9" pagination="1 - 21" refId="ref6094" refString="Madill, J., and P. Hovingh. 2007. Freshwater leech (Annelida: Hirudinida) distribution in the Canadian province of Newfoundland and Labrador and adjacent regions: check-list, new records, new pigmentation forms, and Pleistocene refugia. Zootaxa 1657: 1 - 21." type="journal article" year="2007">Madill and Hovingh, 2007</bibRefCitation>
). Found on the underside of submerged rocks and wood.
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