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<document id="C5796828A03F256F61D021D4B2C81F05" ID-DOI="10.11646/zootaxa.3980.4.9" ID-GBIF-Dataset="8a28a9c7-80a9-4b8b-9726-a6b0fe238039" ID-ISSN="1175-5326" ID-Zenodo-Dep="245332" ID-ZooBank="B91A84A7-42C3-49F1-83CC-958E77E55A22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461244609838" checkinUser="plazi" docAuthor="Aristizábal-García, Hernán, Floriano, Carla Fernanda Burguez, Moreira, Felipe Ferraz Figueiredo &amp; Bispo, Pitágoras Da Conceição" docDate="2015" docId="03CA614CA7009C1B438562E9FC7E2A9F" docLanguage="en" docName="zt03980p596.pdf" docOrigin="Zootaxa 3980 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microvelia" docType="treatment" docVersion="8" lastPageNumber="596" masterDocId="FFF31934A7049C1E43126333FFF22B32" masterDocTitle="New species and notes on Microveliinae from northern South America (Hemiptera: Heteroptera: Veliidae)" masterLastPageNumber="596" masterPageNumber="591" pageNumber="595" updateTime="1698605998038" updateUser="plazi">
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<mods:title id="03B4C54085038C7B938E22889340BC33">New species and notes on Microveliinae from northern South America (Hemiptera: Heteroptera: Veliidae)</mods:title>
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<mods:namePart id="87E102D9EE5FBEA45295299B6C888E14">Aristizábal-García, Hernán</mods:namePart>
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<treatment id="03CA614CA7009C1B438562E9FC7E2A9F" ID-DOI="http://doi.org/10.5281/zenodo.6102952" ID-GBIF-Taxon="119634468" ID-Zenodo-Dep="6102952" LSID="urn:lsid:plazi:treatment:03CA614CA7009C1B438562E9FC7E2A9F" httpUri="http://treatment.plazi.org/id/03CA614CA7009C1B438562E9FC7E2A9F" lastPageId="5" lastPageNumber="596" pageId="4" pageNumber="595">
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<paragraph id="8BDCD05AA7009C1A438562E9FD7B2AC1" blockId="4.[151,649,474,499]" box="[151,649,474,499]" pageId="4" pageNumber="595">
<heading id="D0946736A7009C1A438562E9FD7B2AC1" bold="true" box="[151,649,474,499]" fontSize="10" level="2" pageId="4" pageNumber="595" reason="2">
<emphasis id="B9170C48A7009C1A438562E9FD7B2AC1" bold="true" box="[151,649,474,499]" pageId="4" pageNumber="595">
Two misplaced
<taxonomicName id="4C63ABD9A7009C1A425D62E9FE3E2AC1" ID-CoL="5SLM" box="[335,460,474,499]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A425D62E9FE3E2AC1" bold="true" box="[335,460,474,499]" italics="true" pageId="4" pageNumber="595">Microvelia</emphasis>
</taxonomicName>
from
<collectingCountry id="F37490CAA7009C1A410662E9FD7B2AC1" box="[532,649,474,499]" name="Colombia" pageId="4" pageNumber="595">Colombia</collectingCountry>
</emphasis>
</heading>
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<subSubSection id="C37983D1A7009C1A43856111FBFF2F01" pageId="4" pageNumber="595" type="description">
<paragraph id="8BDCD05AA7009C1A43856111FEF62869" blockId="4.[151,1437,546,1075]" pageId="4" pageNumber="595">
Representatives of the subfamily
<taxonomicName id="4C63ABD9A7009C1A41036111FD9F2909" box="[529,621,546,571]" class="Insecta" family="Veliidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="subFamily" subFamily="Veliinae">Veliinae</taxonomicName>
(
<taxonomicName id="4C63ABD9A7009C1A416E6110FCF52909" box="[636,775,547,571]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
:
<taxonomicName id="4C63ABD9A7009C1A40036111FC9D2909" box="[785,879,546,571]" class="Insecta" family="Veliidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="family">Veliidae</taxonomicName>
) are characterized by the absence of ocelli, tarsal formula 3-3-3, and absence of a swimming fan on middle tarsus (
<bibRefCitation id="EFF2ADABA7009C1A40796175FBE9296D" author="Andersen" box="[875,1051,582,607]" pageId="4" pageNumber="595" refString="Andersen, N. M. (1982) The semiaquatic bugs (Hemiptera, Gerromorpha) phylogeny, adaptation, biogeography and classification. Entomonograph, 3, 1 - 455." type="journal article" year="1982">Andersen 1982</bibRefCitation>
). Six genera of the subfamily have been recorded from South
<collectingCountry id="F37490CAA7009C1A42CB6159FDCB29B1" box="[473,569,618,643]" name="United States of America" pageId="4" pageNumber="595">America</collectingCountry>
so far:
<emphasis id="B9170C48A7009C1A418D6158FCF229B1" box="[671,768,619,643]" italics="true" pageId="4" pageNumber="595">Veloidea</emphasis>
Gould, 1934,
<taxonomicName id="4C63ABD9A7009C1A40A06158FAEA29B1" authority="Hungerford, 1929" authorityName="Hungerford" authorityYear="1929" box="[946,1304,618,643]" class="Insecta" family="Veliidae" genus="Stridulivelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A40A06158FBCE29B1" box="[946,1084,619,643]" italics="true" pageId="4" pageNumber="595">Stridulivelia</emphasis>
Hungerford, 1929
</taxonomicName>
,
<taxonomicName id="4C63ABD9A7009C1A46386158FE1A2995" authority="Polhemus &amp; Polhemus, 1993" authorityName="Polhemus &amp; Polhemus" authorityYear="1993" class="Insecta" family="Veliidae" genus="Platyvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A46386158FA6E29B1" box="[1322,1436,619,643]" italics="true" pageId="4" pageNumber="595">Platyvelia</emphasis>
Polhemus &amp; Polhemus, 1993
</taxonomicName>
,
<taxonomicName id="4C63ABD9A7009C1A42E761BDFC392995" authority="Polhemus &amp; Polhemus, 1993" authorityName="Polhemus &amp; Polhemus" authorityYear="1993" box="[501,971,654,679]" class="Insecta" family="Veliidae" genus="Steinovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A42E761BDFD832994" box="[501,625,654,678]" italics="true" pageId="4" pageNumber="595">Steinovelia</emphasis>
Polhemus &amp; Polhemus, 1993
</taxonomicName>
,
<emphasis id="B9170C48A7009C1A40CA61BDFBCB2994" box="[984,1081,654,678]" italics="true" pageId="4" pageNumber="595">Oiovelia</emphasis>
Drake &amp; Maldonado-Capriles, 1952, and
<taxonomicName id="4C63ABD9A7009C1A42066180FDC229F9" authority="Breddin, 1898" authorityName="Breddin" authorityYear="1898" box="[276,560,690,715]" class="Insecta" family="Veliidae" genus="Paravelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A42066180FE7029F9" box="[276,386,691,715]" italics="true" pageId="4" pageNumber="595">Paravelia</emphasis>
Breddin, 1898
</taxonomicName>
(
<bibRefCitation id="EFF2ADABA7009C1A41516181FCD829F9" author="Moreira" box="[579,810,690,715]" pageId="4" pageNumber="595" refString="Moreira, F. F. F., Barbosa, J. F., Ribeiro, J. R. I. &amp; Alecrim, V. P. (2011) Checklist and distribution of semiaquatic and aquatic Heteroptera (Gerromorpha and Nepomorpha) occurring in Brazil. Zootaxa, 2958, 1 - 74." type="journal article" year="2011">
Moreira
<emphasis id="B9170C48A7009C1A41B86180FD1729F9" box="[682,741,691,715]" italics="true" pageId="4" pageNumber="595">et al.</emphasis>
2011
</bibRefCitation>
; Rodrigues
<emphasis id="B9170C48A7009C1A40A56180FC0129F9" box="[951,1011,691,715]" italics="true" pageId="4" pageNumber="595">et al.</emphasis>
2014). Fifty-nine species have been described in
<taxonomicName id="4C63ABD9A7009C1A423461E5FE6629DC" box="[294,404,726,750]" class="Insecta" family="Veliidae" genus="Paravelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A423461E5FE6629DC" box="[294,404,726,750]" italics="true" pageId="4" pageNumber="595">Paravelia</emphasis>
</taxonomicName>
, two of which are
<taxonomicName id="4C63ABD9A7009C1A417461E4FC5129DD" authority="Padilla-Gil, 2013" authorityName="Padilla-Gil" authorityYear="2013" box="[614,931,726,751]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="B9170C48A7009C1A417461E4FD2929DC" box="[614,731,726,750]" italics="true" pageId="4" pageNumber="595">P. acantha</emphasis>
<bibRefCitation id="EFF2ADABA7009C1A41F161E5FC5129DD" author="Padilla-Gil" box="[739,931,726,751]" pageId="4" pageNumber="595" refString="Padilla-Gil, D. N. (2013) Two new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3693 (4), 491 - 502. http: // dx. doi. org / 10.11646 / zootaxa. 3693.4.5" type="journal article" year="2013">Padilla-Gil, 2013</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C63ABD9A7009C1A40C861E4FAF829DD" authority="Padilla-Gil, 2013" authorityName="Padilla-Gil" authorityYear="2013" box="[986,1290,726,751]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="fanera">
<emphasis id="B9170C48A7009C1A40C861E4FBCC29DD" box="[986,1086,727,751]" italics="true" pageId="4" pageNumber="595">P. fanera</emphasis>
<bibRefCitation id="EFF2ADABA7009C1A475761E5FAF829DD" author="Padilla-Gil" box="[1093,1290,726,751]" pageId="4" pageNumber="595" refString="Padilla-Gil, D. N. (2013) Two new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3693 (4), 491 - 502. http: // dx. doi. org / 10.11646 / zootaxa. 3693.4.5" type="journal article" year="2013">Padilla-Gil, 2013</bibRefCitation>
</taxonomicName>
. Members of the subfamily
<taxonomicName id="4C63ABD9A7009C1A422861C9FE282821" box="[314,474,762,787]" class="Insecta" family="Veliidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="subFamily" subFamily="Microveliinae">Microveliinae</taxonomicName>
, in turn, also lack ocelli and a swimming fan, but can be separated from
<taxonomicName id="4C63ABD9A7009C1A460E61C9FA8A2821" box="[1308,1400,762,787]" class="Insecta" family="Veliidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="subFamily" subFamily="Veliinae">Veliinae</taxonomicName>
by the tarsal formula 1-2-2, which is a result of the fusion of tarsomeres IIII of the fore legs and III of middle and hind legs.
</paragraph>
<paragraph id="8BDCD05AA7009C1A43D56055FBFF2F01" blockId="4.[151,1437,546,1075]" pageId="4" pageNumber="595">
The type-series of the two species of
<taxonomicName id="4C63ABD9A7009C1A41776055FD21284C" box="[613,723,870,894]" class="Insecta" family="Veliidae" genus="Paravelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A41776055FD21284C" box="[613,723,870,894]" italics="true" pageId="4" pageNumber="595">Paravelia</emphasis>
</taxonomicName>
mentioned in the above paragraph were examined at the Museo de Entomología de la Universidad Nacional de
<collectingCountry id="F37490CAA7009C1A41B460B9FCE52891" box="[678,791,906,931]" name="Colombia" pageId="4" pageNumber="595">Colombia</collectingCountry>
, and it was possible to determine that the tarsal formula is 1-2-2, not 3-3-3. Therefore, the specimens cannot be members of the genus
<taxonomicName id="4C63ABD9A7009C1A4711609DFB8328F4" box="[1027,1137,942,966]" class="Insecta" family="Veliidae" genus="Paravelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A4711609DFB8328F4" box="[1027,1137,942,966]" italics="true" pageId="4" pageNumber="595">Paravelia</emphasis>
</taxonomicName>
, and belong in fact to the subfamily
<taxonomicName id="4C63ABD9A7009C1A420160E1FE4628D9" box="[275,436,978,1003]" class="Insecta" family="Veliidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="subFamily" subFamily="Microveliinae">Microveliinae</taxonomicName>
. Both species lack pretarsal leaf-like structures on the middle leg, and thus must be regarded as members of the genus
<emphasis id="B9170C48A7009C1A412960C5FD4E2F3C" box="[571,700,1014,1038]" italics="true" pageId="4" pageNumber="595">
<taxonomicName id="4C63ABD9A7009C1A412960C5FD4B2F3C" box="[571,697,1014,1038]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">Microvelia</taxonomicName>
.
</emphasis>
Accordingly, the present authors propose transferring the two species described by
<bibRefCitation id="EFF2ADABA7009C1A42966729FDA22F01" author="Padilla-Gil" box="[388,592,1050,1075]" pageId="4" pageNumber="595" refString="Padilla-Gil, D. N. (2013) Two new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3693 (4), 491 - 502. http: // dx. doi. org / 10.11646 / zootaxa. 3693.4.5" type="journal article" year="2013">Padilla-Gil (2013)</bibRefCitation>
from
<taxonomicName id="4C63ABD9A7009C1A41876728FCF12F01" box="[661,771,1051,1075]" class="Insecta" family="Veliidae" genus="Paravelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A41876728FCF12F01" box="[661,771,1051,1075]" italics="true" pageId="4" pageNumber="595">Paravelia</emphasis>
</taxonomicName>
to
<taxonomicName id="4C63ABD9A7009C1A40356728FC532F01" box="[807,929,1051,1075]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A40356728FC532F01" box="[807,929,1051,1075]" italics="true" pageId="4" pageNumber="595">Microvelia</emphasis>
</taxonomicName>
. In short:
</paragraph>
</subSubSection>
<subSubSection id="C37983D1A7009C1A43856750FCCC2E61" pageId="4" pageNumber="595" type="reference_group">
<paragraph id="8BDCD05AA7009C1A43856750FD3D2F48" blockId="4.[151,1436,1122,1255]" box="[151,719,1122,1147]" pageId="4" pageNumber="595">
<taxonomicName id="4C63ABD9A7009C1A43856750FDBE2F49" ID-CoL="439L6" authority="Padilla-Gil, 2013" authorityName="Padilla-Gil" authorityYear="2013" box="[151,588,1122,1147]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="acantha" status="comb. nov.">
<emphasis id="B9170C48A7009C1A43856750FE802F49" box="[151,370,1123,1147]" italics="true" pageId="4" pageNumber="595">Microvelia acantha</emphasis>
(
<bibRefCitation id="EFF2ADABA7009C1A42926751FDB72F49" author="Padilla-Gil" box="[384,581,1122,1147]" pageId="4" pageNumber="595" refString="Padilla-Gil, D. N. (2013) Two new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3693 (4), 491 - 502. http: // dx. doi. org / 10.11646 / zootaxa. 3693.4.5" type="journal article" year="2013">Padilla-Gil, 2013</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A224B133A7009C1A41416751FD3D2F48" box="[595,719,1122,1147]" pageId="4" pageNumber="595">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDCD05AA7009C1A438567B4FF132FD5" blockId="4.[151,1436,1122,1255]" pageId="4" pageNumber="595">
=
<taxonomicName id="4C63ABD9A7009C1A43BF67B5FDBB2FAD" ID-CoL="4DHM9" authority="Padilla-Gil, 2013" authorityName="Padilla-Gil" authorityYear="2013" box="[173,585,1158,1183]" class="Insecta" family="Veliidae" genus="Paravelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="B9170C48A7009C1A43BF67B5FE8F2FAC" box="[173,381,1158,1182]" italics="true" pageId="4" pageNumber="595">Paravelia acantha</emphasis>
<bibRefCitation id="EFF2ADABA7009C1A429667B5FDBB2FAD" author="Padilla-Gil" box="[388,585,1158,1183]" pageId="4" pageNumber="595" refString="Padilla-Gil, D. N. (2013) Two new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3693 (4), 491 - 502. http: // dx. doi. org / 10.11646 / zootaxa. 3693.4.5" type="journal article" year="2013">Padilla-Gil, 2013</bibRefCitation>
</taxonomicName>
. Zootaxa 3693(4): 495. Material examined:
<typeStatus id="54D86EF8A7009C1A426B6798FDE32FF1" box="[377,529,1195,1219]" pageId="4" pageNumber="595" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="F37490CAA7009C1A41326799FD652FF1" box="[544,663,1194,1219]" name="Colombia" pageId="4" pageNumber="595">Colombia</collectingCountry>
, Departamento Nariño, Funes,
<date id="FFDDF69AA7009C1A40E46798FB362FF1" box="[1014,1220,1194,1219]" pageId="4" pageNumber="595" value="2009-11-02">2 November 2009</date>
, leg. D. N. Padilla (ICN).
</paragraph>
<paragraph id="8BDCD05AA7009C1A43856625FD4C2E1C" blockId="4.[151,1436,1302,1435]" box="[151,702,1302,1327]" pageId="4" pageNumber="595">
<taxonomicName id="4C63ABD9A7009C1A43856625FDC92E1D" ID-CoL="439NS" authority="Padilla-Gil, 2013" authorityName="Padilla-Gil" authorityYear="2013" box="[151,571,1302,1327]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="fanera" status="comb. nov.">
<emphasis id="B9170C48A7009C1A43856625FE932E1D" box="[151,353,1302,1327]" italics="true" pageId="4" pageNumber="595">Microvelia fanera</emphasis>
(
<bibRefCitation id="EFF2ADABA7009C1A427D6625FDC62E1D" author="Padilla-Gil" box="[367,564,1302,1327]" pageId="4" pageNumber="595" refString="Padilla-Gil, D. N. (2013) Two new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3693 (4), 491 - 502. http: // dx. doi. org / 10.11646 / zootaxa. 3693.4.5" type="journal article" year="2013">Padilla-Gil, 2013</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A224B133A7009C1A41506625FD4C2E1C" box="[578,702,1302,1327]" pageId="4" pageNumber="595">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDCD05AA7009C1A43856608FCCC2E61" blockId="4.[151,1436,1302,1435]" box="[151,830,1338,1363]" pageId="4" pageNumber="595">
=
<taxonomicName id="4C63ABD9A7009C1A43BF6608FDCA2E61" ID-CoL="4DHN5" authority="Padilla-Gil, 2013" authorityName="Padilla-Gil" authorityYear="2013" box="[173,568,1338,1363]" class="Insecta" family="Veliidae" genus="Paravelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="fanera">
<emphasis id="B9170C48A7009C1A43BF6608FE9F2E61" box="[173,365,1339,1363]" italics="true" pageId="4" pageNumber="595">Paravelia fanera</emphasis>
<bibRefCitation id="EFF2ADABA7009C1A42616609FDCA2E61" author="Padilla-Gil" box="[371,568,1338,1363]" pageId="4" pageNumber="595" refString="Padilla-Gil, D. N. (2013) Two new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3693 (4), 491 - 502. http: // dx. doi. org / 10.11646 / zootaxa. 3693.4.5" type="journal article" year="2013">Padilla-Gil, 2013</bibRefCitation>
</taxonomicName>
. Zootaxa 3693(4): 491.
</paragraph>
</subSubSection>
<subSubSection id="C37983D1A7009C1B4385666DFE042ABA" lastPageId="5" lastPageNumber="596" pageId="4" pageNumber="595" type="materials_examined">
<paragraph id="8BDCD05AA7009C1A4385666DFDAA2EA9" blockId="4.[151,1436,1302,1435]" pageId="4" pageNumber="595">
Material examined:
<typeStatus id="54D86EF8A7009C1A4292666CFDEA2E45" box="[384,536,1375,1399]" pageId="4" pageNumber="595" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="F37490CAA7009C1A4138666CFD352E45" box="[554,711,1375,1399]" name="Colombia" pageId="4" pageNumber="595">COLOMBIA</collectingCountry>
, Departamento Nariño, Quebrada El Bohio,
<quantity id="4C9B7DBFA7009C1A47C2666CFAD92E44" box="[1232,1323,1375,1399]" metricMagnitude="3" metricUnit="m" metricValue="1.7289999999999999" pageId="4" pageNumber="595" unit="m" value="1729.0">1729 m</quantity>
,
<date id="FFDDF69AA7009C1A4629666CFF202EA9" pageId="4" pageNumber="595" value="2013-04-17">17 Abril 2013</date>
, leg. V. Paz &amp; C. Martínez (
<collectionCode id="ED72489FA7009C1A410466B0FDBE2EA9" box="[534,588,1411,1435]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="4" pageNumber="595">ICN</collectionCode>
).
</paragraph>
<paragraph id="8BDCD05AA7009C1A438566F8FD282ED1" blockId="4.[151,1437,1482,2011]" box="[151,730,1482,1507]" pageId="4" pageNumber="595">
A new synonym of
<taxonomicName id="4C63ABD9A7009C1A426666F8FD282ED1" authority="White, 1879" authorityName="White" authorityYear="1879" box="[372,730,1482,1507]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B9170C48A7009C1A426666F8FDBB2ED1" box="[372,585,1483,1507]" italics="true" pageId="4" pageNumber="595">Microvelia mimula</emphasis>
White, 1879
</taxonomicName>
</paragraph>
<paragraph id="8BDCD05AA7009C1A43D566DDFE1E2DA5" blockId="4.[151,1437,1482,2011]" pageId="4" pageNumber="595">
<taxonomicName id="4C63ABD9A7009C1A43D566DDFE532D34" box="[199,417,1518,1542]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B9170C48A7009C1A43D566DDFE532D34" box="[199,417,1518,1542]" italics="true" pageId="4" pageNumber="595">Microvelia mimula</emphasis>
</taxonomicName>
is a common species in still waters, ranging from
<collectingCountry id="F37490CAA7009C1A471E66DDFBB52D35" box="[1036,1095,1518,1543]" name="Cuba" pageId="4" pageNumber="595">Cuba</collectingCountry>
to
<collectingCountry id="F37490CAA7009C1A476766DCFB252D35" box="[1141,1239,1519,1543]" name="Uruguay" pageId="4" pageNumber="595">Uruguay</collectingCountry>
(
<bibRefCitation id="EFF2ADABA7009C1A47F966DDFA632D35" author="Moreira" box="[1259,1425,1518,1543]" pageId="4" pageNumber="595" refString="Moreira, F. F. F. (2015) Water Bugs Distributional Database. Available from: https: // sites. google. com / site / distributionaldatabase (accessed 13 February 2015)" type="journal article" year="2015">Moreira 2015</bibRefCitation>
). Females can easily be confused with their congeners, but males can be distinguished by the shape of genital segment I and the long horn-like lateral projections on the proctiger. Additional diagnostic features for males include a tubercle on the penultimate abdominal sternite and a row of spinules on hind femur, but these can often be underdeveloped or broken off.
</paragraph>
<paragraph id="8BDCD05AA7009C1A43D56591FECA2C5C" blockId="4.[151,1437,1482,2011]" pageId="4" pageNumber="595">
<bibRefCitation id="EFF2ADABA7009C1A43D56591FE852D89" author="Makhan" box="[199,375,1698,1723]" pageId="4" pageNumber="595" refString="Makhan, D. (2014) Microvelia rishwani sp. nov., Microvelia amrishi sp. nov. and Microvelia aschnakiranae sp. nov. (Hemiptera: Heteroptera: Veliidae) from Suriname. Calodema, 345, 1 - 7." type="journal article" year="2014">Makhan (2014)</bibRefCitation>
recently described three allegedly new species of
<taxonomicName id="4C63ABD9A7009C1A40AF6590FBC52D89" box="[957,1079,1699,1723]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7009C1A40AF6590FBC52D89" box="[957,1079,1699,1723]" italics="true" pageId="4" pageNumber="595">Microvelia</emphasis>
</taxonomicName>
from
<collectingCountry id="F37490CAA7009C1A476D6591FB1C2D89" box="[1151,1262,1698,1723]" name="Suriname" pageId="4" pageNumber="595">Suriname</collectingCountry>
. Even with the extremely poor descriptions and images provided, it is possible to determine that
<taxonomicName id="4C63ABD9A7009C1A474265F4FA8D2DED" authority="Makhan, 2014" authorityName="Makhan" authorityYear="2014" box="[1104,1407,1734,1759]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="amrishi">
<emphasis id="B9170C48A7009C1A474265F4FB3C2DEC" box="[1104,1230,1734,1758]" italics="true" pageId="4" pageNumber="595">M. amrishi</emphasis>
<bibRefCitation id="EFF2ADABA7009C1A47CB65F5FA8D2DED" author="Makhan" box="[1241,1407,1734,1759]" pageId="4" pageNumber="595" refString="Makhan, D. (2014) Microvelia rishwani sp. nov., Microvelia amrishi sp. nov. and Microvelia aschnakiranae sp. nov. (Hemiptera: Heteroptera: Veliidae) from Suriname. Calodema, 345, 1 - 7." type="journal article" year="2014">Makhan, 2014</bibRefCitation>
</taxonomicName>
is clearly a synonym of
<taxonomicName id="4C63ABD9A7009C1A428365D8FD9A2C31" box="[401,616,1771,1795]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B9170C48A7009C1A428365D8FD9A2C31" box="[401,616,1771,1795]" italics="true" pageId="4" pageNumber="595">Microvelia mimula</emphasis>
</taxonomicName>
, based on the figures of the habitus and genitalia of the male
<typeStatus id="54D86EF8A7009C1A462165D9FA6B2C31" box="[1331,1433,1770,1795]" pageId="4" pageNumber="595" type="holotype">holotype</typeStatus>
. The identity of the other two species,
<taxonomicName id="4C63ABD9A7009C1A4157643CFC8A2C15" authority="Makhan, 2014" authorityName="Makhan" authorityYear="2014" box="[581,888,1806,1831]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="rishwani">
<emphasis id="B9170C48A7009C1A4157643CFD3C2C14" box="[581,718,1806,1830]" italics="true" pageId="4" pageNumber="595">M. rishwani</emphasis>
<bibRefCitation id="EFF2ADABA7009C1A41C7643DFC8A2C15" author="Makhan" box="[725,888,1806,1831]" pageId="4" pageNumber="595" refString="Makhan, D. (2014) Microvelia rishwani sp. nov., Microvelia amrishi sp. nov. and Microvelia aschnakiranae sp. nov. (Hemiptera: Heteroptera: Veliidae) from Suriname. Calodema, 345, 1 - 7." type="journal article" year="2014">Makhan, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C63ABD9A7009C1A40A3643CFAD52C15" authority="Makhan, 2014" authorityName="Makhan" authorityYear="2014" box="[945,1319,1806,1831]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="aschnakiranae">
<emphasis id="B9170C48A7009C1A40A3643CFB8E2C14" box="[945,1148,1806,1830]" italics="true" pageId="4" pageNumber="595">M. aschnakiranae</emphasis>
<bibRefCitation id="EFF2ADABA7009C1A4796643DFAD52C15" author="Makhan" box="[1156,1319,1806,1831]" pageId="4" pageNumber="595" refString="Makhan, D. (2014) Microvelia rishwani sp. nov., Microvelia amrishi sp. nov. and Microvelia aschnakiranae sp. nov. (Hemiptera: Heteroptera: Veliidae) from Suriname. Calodema, 345, 1 - 7." type="journal article" year="2014">Makhan, 2014</bibRefCitation>
</taxonomicName>
cannot be defined based on the information and images provided by the author, so it is unclear if they indeed represent new species or not.
</paragraph>
<paragraph id="8BDCD05AA7009C1B43D56449FDCA2A2F" blockId="4.[151,1437,1482,2011]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="596" pageId="4" pageNumber="595">
Knowing the bad practices and misbehavior of Mr. Makhan in relation to other researchers and taxonomy itself (
<bibRefCitation id="EFF2ADABA7009C1A438C64ADFEE42C85" author="Jach" box="[158,278,1950,1975]" pageId="4" pageNumber="595" refString="Jach, M. A. (2006) Taxonomy and nomenclature threatened by D. Makhan. Koleopterologische Rundschau, 76, 360." type="journal article" year="2006">Jäch 2006</bibRefCitation>
;
<bibRefCitation id="EFF2ADABA7009C1A423664ACFE002C85" author="Anonymous" box="[292,498,1951,1975]" pageId="4" pageNumber="595" refString="Anonymous (2007) Vandalism in Taxonomy. Koleopterologische Rundschau, 77, 38 + 60 + 88 + 146." type="journal article" year="2007">Anonymous 2007</bibRefCitation>
), we inquired Dr. Nico Nieser (
<emphasis id="B9170C48A7009C1A404D64ACFC572C84" box="[863,933,1950,1974]" italics="true" pageId="4" pageNumber="595">in litt.</emphasis>
), a major specialist on the water bugs from
<collectingCountry id="F37490CAA7009C1A438564F1FEF72CE9" box="[151,261,1986,2011]" name="Suriname" pageId="4" pageNumber="595">Suriname</collectingCountry>
, about the identities of
<taxonomicName id="4C63ABD9A7009C1A411E64F0FD662CE9" box="[524,660,1987,2011]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="rishwani">
<emphasis id="B9170C48A7009C1A411E64F0FD662CE9" box="[524,660,1987,2011]" italics="true" pageId="4" pageNumber="595">M. rishwani</emphasis>
</taxonomicName>
and
<taxonomicName id="4C63ABD9A7009C1A41D864F0FC672CE9" box="[714,917,1987,2011]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="aschnakiranae">
<emphasis id="B9170C48A7009C1A41D864F0FC672CE9" box="[714,917,1987,2011]" italics="true" pageId="4" pageNumber="595">M. aschnakiranae</emphasis>
</taxonomicName>
. Dr. Nieser informed us that unfortunately his material of
<taxonomicName id="4C63ABD9A7019C1B420F63ABFE652B82" box="[285,407,152,176]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="B9170C48A7019C1B420F63ABFE652B82" box="[285,407,152,176]" italics="true" pageId="5" pageNumber="596">Microvelia</emphasis>
</taxonomicName>
from
<collectingCountry id="F37490CAA7019C1B42F263A4FDB92B82" box="[480,587,151,176]" name="Suriname" pageId="5" pageNumber="596">Suriname</collectingCountry>
got lost and that he never studied it, so it is not possible to make further decisions about Makhans species at the moment. Makhans
<typeStatus id="54D86EF8A7019C1B405A638EFC712BE7" box="[840,899,189,213]" pageId="5" pageNumber="596">types</typeStatus>
are supposedly deposited at the Department of Entomology, University of
<collectingCountry id="F37490CAA7019C1B42C063ECFDB22BCA" box="[466,576,223,248]" name="Suriname" pageId="5" pageNumber="596">Suriname</collectingCountry>
, and it will only be possible to give further insight into this issue with an accurate analysis of these specimens.
</paragraph>
<paragraph id="8BDCD05AA7019C1B4385627FFE042ABA" blockId="5.[151,908,332,429]" pageId="5" pageNumber="596">
The following new synonymy is therefore proposed:
<taxonomicName id="4C63ABD9A7019C1B43856243FE042ABA" authority="White 1879" authorityName="White" authorityYear="1879" box="[151,502,367,392]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B9170C48A7019C1B43856243FE9E2ABA" box="[151,364,368,392]" italics="true" pageId="5" pageNumber="596">Microvelia mimula</emphasis>
White 1879
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37983D1A7019C1B438562A6FC7E2A9F" box="[151,908,404,429]" pageId="5" pageNumber="596" type="reference_group">
<paragraph id="8BDCD05AA7019C1B438562A6FC7E2A9F" blockId="5.[151,908,332,429]" box="[151,908,404,429]" pageId="5" pageNumber="596">
=
<taxonomicName id="4C63ABD9A7019C1B43BF62A7FDC22A9E" ID-CoL="7VVL2" authority="Makhan, 2014" authorityName="Makhan" authorityYear="2014" box="[173,560,404,429]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="amrishi">
<emphasis id="B9170C48A7019C1B43BF62A7FE712A9E" box="[173,387,404,428]" italics="true" pageId="5" pageNumber="596">Microvelia amrishi</emphasis>
<bibRefCitation id="EFF2ADABA7019C1B429962A7FDC22A9E" author="Makhan" box="[395,560,404,429]" pageId="5" pageNumber="596" refString="Makhan, D. (2014) Microvelia rishwani sp. nov., Microvelia amrishi sp. nov. and Microvelia aschnakiranae sp. nov. (Hemiptera: Heteroptera: Veliidae) from Suriname. Calodema, 345, 1 - 7." type="journal article" year="2014">Makhan, 2014</bibRefCitation>
</taxonomicName>
. Calodema 345: 17.
<emphasis id="B9170C48A7019C1B403662A7FC7E2A9F" bold="true" box="[804,908,404,429]" pageId="5" pageNumber="596">syn. nov.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>