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<document id="839C3604F7E6622E008CDDD17F5EA8E1" ID-CLB-Dataset="27975" ID-DOI="10.11646/zootaxa.4518.1.1" ID-GBIF-Dataset="ba2bd1cd-8a6f-4a94-93a2-96b99318a8b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2609500" ID-ZooBank="708981EF-21DC-4DC2-B1CD-8CFF4373DA8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553622948496" checkinUser="plazi" docAuthor="Harrison, Sophie E., Rix, Michael G., Harvey, Mark S. &amp; Austin, Andrew D." docDate="2018" docId="03C10411554DFFDDE1E8FF66FD66FE3E" docLanguage="en" docName="zootaxa.4518.1.1.pdf" docOrigin="Zootaxa 4518 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Blakistonia bassi Harrison &amp; Rix &amp; Harvey &amp; Austin 2018, sp. n." docType="treatment" docVersion="8" lastPageNumber="28" masterDocId="FFF87C695557FFC6E17FFFF1FF86FF92" masterDocTitle="Systematics of the Australian spiny trapdoor spiders of the genus Blakistonia Hogg (Araneae: Idiopidae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="27" updateTime="1698713611295" updateUser="ExternalLinkService">
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<mods:title id="364071EE40D3F2C7496B0354F3E60F0E">Systematics of the Australian spiny trapdoor spiders of the genus Blakistonia Hogg (Araneae: Idiopidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="4B69EC5733D8342221ED091BC93737D6">Harrison, Sophie E.</mods:namePart>
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<mods:namePart id="6BC6CA8D59754807C9DD96CF802EC507">Rix, Michael G.</mods:namePart>
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<mods:namePart id="0DD73746F90F50B23B5AB4E988B1A3FB">Harvey, Mark S.</mods:namePart>
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<mods:namePart id="74FDDDD3AE70BF668DC5B88E569B5172">Austin, Andrew D.</mods:namePart>
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<mods:date id="AE17DC428DBFE6A7737FE5C0C8C7E2A9">2018</mods:date>
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<mods:number id="496F6403970D895381DF885CE6FF38B6">2018-11-12</mods:number>
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<treatment id="03C10411554DFFDDE1E8FF66FD66FE3E" ID-DOI="http://doi.org/10.5281/zenodo.5967799" ID-GBIF-Taxon="154498537" ID-Zenodo-Dep="5967799" LSID="urn:lsid:plazi:treatment:03C10411554DFFDDE1E8FF66FD66FE3E" httpUri="http://treatment.plazi.org/id/03C10411554DFFDDE1E8FF66FD66FE3E" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<subSubSection id="C372E68C554DFFDCE1E8FF66FE28FF20" box="[151,430,151,178]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BD7B507554DFFDCE1E8FF66FE28FF20" blockId="26.[151,430,151,211]" box="[151,430,151,178]" pageId="26" pageNumber="27">
<heading id="D09F026B554DFFDCE1E8FF66FE28FF20" bold="true" box="[151,430,151,178]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B91C6915554DFFDCE1E8FF66FE28FF20" bold="true" box="[151,430,151,178]" pageId="26" pageNumber="27">
<taxonomicName id="4C68CE84554DFFDCE1E8FF66FEE3FF23" ID-CoL="LZSZ" authority="Harrison &amp; Rix &amp; Harvey &amp; Austin, 2018" authorityName="Harrison &amp; Rix &amp; Harvey &amp; Austin" authorityYear="2018" box="[151,357,151,177]" class="Arachnida" family="Idiopidae" genus="Blakistonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bassi" status="sp. nov.">
<emphasis id="B91C6915554DFFDCE1E8FF66FEE3FF23" bold="true" box="[151,357,151,177]" italics="true" pageId="26" pageNumber="27">Blakistonia bassi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22FD46E554DFFDCE013FF69FE28FF20" box="[364,430,152,178]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C372E68C554DFFDCE1E8FF4BFE91FF41" box="[151,279,186,211]" pageId="26" pageNumber="27" type="description">
<paragraph id="8BD7B507554DFFDCE1E8FF4BFE91FF41" blockId="26.[151,430,151,211]" box="[151,279,186,211]" pageId="26" pageNumber="27">
(
<figureCitation id="1353A982554DFFDCE1E0FF4BFF67FF41" box="[159,225,186,211]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7</figureCitation>
AI)
</paragraph>
</subSubSection>
<subSubSection id="C372E68C554DFFDCE1E8FEF3FDC0FE18" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BD7B507554DFFDCE1E8FEF3FDC0FE18" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1E8FEF3FD39FE89" bold="true" box="[151,703,258,283]" pageId="26" pageNumber="27">
Type material.
<materialsCitation id="3B00BF5A554DFFDCE034FEF3FD39FE89" ID-GBIF-Occurrence="2028519260" box="[331,703,258,283]" collectingDate="2014-12-22" collectionCode="SAM" collectorName="S. E. Harrison &amp; M. Harrison" country="Australia" county="Pound Road" latitude="-34.932114" location="Mount Lofty Ranges" longLatPrecision="1" longitude="138.74713" municipality="Ashton" pageId="26" pageNumber="27" specimenCode="NN29619" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<collectingCountry id="F37FF597554DFFDCE034FEF3FE72FE89" box="[331,500,258,283]" name="Australia" pageId="26" pageNumber="27">AUSTRALIA</collectingCountry>
:
<emphasis id="B91C6915554DFFDCE37FFEF3FD39FE89" bold="true" box="[512,703,258,283]" italics="true" pageId="26" pageNumber="27">
<collectingRegion id="49AC7BE5554DFFDCE37FFEF3FD3FFE89" box="[512,697,258,283]" country="Australia" name="South Australia" pageId="26" pageNumber="27">South Australia</collectingRegion>
:
</emphasis>
</materialsCitation>
</emphasis>
<typeStatus id="54D30BA5554DFFDCE3B9FEF3FCA8FE89" box="[710,814,258,283]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
female, off
<collectingCounty id="62B6CD8B554DFFDCE2C9FEF3FBC5FE89" box="[950,1091,258,283]" pageId="26" pageNumber="27">Pound Road</collectingCounty>
,
<collectingMunicipality id="6BB32F7D554DFFDCE531FEF3FB24FE89" box="[1102,1186,258,283]" pageId="26" pageNumber="27">Ashton</collectingMunicipality>
,
<location id="8EB7E3DC554DFFDCE5D3FEF2FA1EFE89" LSID="urn:lsid:plazi:treatment:03C10411554DFFDDE1E8FF66FD66FE3E:8EB7E3DC554DFFDCE5D3FEF2FA1EFE89" box="[1196,1432,258,283]" country="Australia" county="Pound Road" latitude="-34.932114" longLatPrecision="1" longitude="138.74713" municipality="Ashton" name="Mount Lofty Ranges" pageId="26" pageNumber="27" stateProvince="South Australia">Mount Lofty Ranges</location>
,,
<geoCoordinate id="EE5CD3C0554DFFDCE1DAFED7FEDAFEAD" box="[165,348,294,319]" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="1" value="-34.932114">34°5555.608”S</geoCoordinate>
,
<geoCoordinate id="EE5CD3C0554DFFDCE017FED7FDA9FEAD" box="[360,559,294,319]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="1" value="138.74713">138°4449.667”E</geoCoordinate>
,
<date id="FFD693C7554DFFDCE344FED6FC95FEAD" box="[571,787,294,319]" pageId="26" pageNumber="27" value="2014-12-22">
<collectingDate id="EF926A2F554DFFDCE344FED6FC95FEAD" box="[571,787,294,319]" pageId="26" pageNumber="27" value="2014-12-22">22 December 2014</collectingDate>
</date>
, hand collected from mossy roadside bank,
<collectorName id="269DD0D1554DFFDCE581FED6FA1EFEAD" box="[1278,1432,294,319]" pageId="26" pageNumber="27">S.E. Harrison</collectorName>
,
<collectorName id="269DD0D1554DFFDCE1E8FEBFFE99FEF4" box="[151,287,333,358]" pageId="26" pageNumber="27">M. Harrison</collectorName>
(SAM
<specimenCode id="DBCE1D7C554DFFDCE00CFEBCFE64FEF4" box="[371,482,333,358]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="26" pageNumber="27">NN29619</specimenCode>
<superScript id="7C1D184F554DFFDCE09EFEBBFD84FECA" attach="none" box="[481,514,330,344]" fontSize="6" pageId="26" pageNumber="27">DNA</superScript>
).
<materialsCitation id="3B00BF5A554DFFDCE365FEBFFDC4FE18" ID-GBIF-Occurrence="2028519279" collectingDate="2016-05-13" collectionCode="SAM" collectorName="S. E. Harrison &amp; N. Birks" country="Australia" county="Pound Road" latitude="-34.932114" location="Mount Lofty Ranges" longLatPrecision="1" longitude="138.74713" municipality="Ashton" pageId="26" pageNumber="27" specimenCode="NN28529, NN28530" specimenCount="2" specimenCount-female="2" stateProvince="South Australia" typeStatus="paratype">
<typeStatus id="54D30BA5554DFFDCE365FEBFFD0DFEF4" box="[538,651,334,358]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D6E7E8E554DFFDCE3EAFEBFFC82FEF4" box="[661,772,333,358]" pageId="26" pageNumber="27" type="female">2 females</specimenCount>
, same data as holotype except
<date id="FFD693C7554DFFDCE524FEBFFB75FEF4" box="[1115,1267,333,358]" pageId="26" pageNumber="27" value="2016-05-13">
<collectingDate id="EF926A2F554DFFDCE524FEBFFB75FEF4" box="[1115,1267,333,358]" pageId="26" pageNumber="27" value="2016-05-13">13 May 2016</collectingDate>
</date>
,
<collectorName id="269DD0D1554DFFDCE581FEBFFA1EFEF4" box="[1278,1432,333,358]" pageId="26" pageNumber="27">S.E. Harrison</collectorName>
,
<collectorName id="269DD0D1554DFFDCE1E8FE83FF70FE18" box="[151,246,369,394]" pageId="26" pageNumber="27">N. Birks</collectorName>
(
<collectionCode id="ED792DC2554DFFDCE07BFE83FEC4FE18" box="[260,322,370,394]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="26" pageNumber="27">SAM</collectionCode>
<specimenCode id="DBCE1D7C554DFFDCE035FE80FE3DFE18" box="[330,443,369,394]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="26" pageNumber="27">NN28529</specimenCode>
,
<specimenCode id="DBCE1D7C554DFFDCE0B9FE80FDBCFE18" box="[454,570,369,394]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="26" pageNumber="27">NN28530</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C372E68C554DFFDCE1B8FE64FAF4FDF0" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BD7B507554DFFDCE1B8FE64FCDCFE40" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8FE64FEC7FE3C" bold="true" box="[199,321,405,430]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
Females of
<taxonomicName id="4C68CE84554DFFDCE0B1FE67FDAEFE3C" authorityName="Harrison &amp; Rix &amp; Harvey &amp; Austin" authorityYear="2018" box="[462,552,405,430]" class="Arachnida" family="Idiopidae" genus="Blakistonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bassi">
<emphasis id="B91C6915554DFFDCE0B1FE67FE59FE3F" box="[462,479,406,429]" italics="true" pageId="26" pageNumber="27">B</emphasis>
.
<emphasis id="B91C6915554DFFDCE090FE64FDAEFE3C" box="[495,552,405,430]" italics="true" pageId="26" pageNumber="27">bassi</emphasis>
</taxonomicName>
can be distinguished from all other species of
<taxonomicName id="4C68CE84554DFFDCE541FE64FB38FE3C" authorityName="Hogg. Where" authorityYear="1902" box="[1086,1214,405,430]" class="Arachnida" family="Idiopidae" genus="Blakistonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91C6915554DFFDCE541FE64FB38FE3C" box="[1086,1214,405,430]" italics="true" pageId="26" pageNumber="27">Blakistonia</emphasis>
</taxonomicName>
by the fine, golden hairs that cover the carapace (
<figureCitation id="1353A982554DFFDCE09AFE48FDE7FE40" box="[485,609,441,466]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7A, D</figureCitation>
). Males are unknown.
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8FE2CFAF4FDF0" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
All life stages of
<taxonomicName id="4C68CE84554DFFDCE0EBFE2FFE76FE64" authorityName="Harrison &amp; Rix &amp; Harvey &amp; Austin" authorityYear="2018" box="[404,496,477,502]" class="Arachnida" family="Idiopidae" genus="Blakistonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bassi">
<emphasis id="B91C6915554DFFDCE0EBFE2FFE76FE64" box="[404,496,477,502]" italics="true" pageId="26" pageNumber="27">B. bassi</emphasis>
</taxonomicName>
can also be distinguished from those of other species with sequence data by the following nucleotide substitutions (
<emphasis id="B91C6915554DFFDCE358FDF3FDB3FD8B" box="[551,565,514,537]" italics="true" pageId="26" pageNumber="27">n</emphasis>
=
<specimenCount id="9D6E7E8E554DFFDCE336FDF3FD4AFD88" box="[585,716,513,538]" pageId="26" pageNumber="27" type="generic">1 specimen</specimenCount>
): G(3), C(87), C(102), T(111), G(199), C(205), T(207), T(255), C(339), A(390), C(462), C(479), C(481), G(546), G(573), G(591); and by the following unique molecular motifs: TT(3031), GAC(6668), GCT(367369), AAG(372374), GG(432433), TGC(456458), TGGA (468471).
</paragraph>
</subSubSection>
<subSubSection id="C372E68C554DFFDCE1B8FD9CFF7DF828" pageId="26" pageNumber="27" type="description">
<paragraph id="8BD7B507554DFFDCE1B8FD9CFA89FD14" blockId="26.[151,1437,258,1978]" box="[199,1295,621,646]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8FD9CFEDFFD14" bold="true" box="[199,345,621,646]" pageId="26" pageNumber="27">Description.</emphasis>
<emphasis id="B91C6915554DFFDCE020FD9CFD9EFD14" box="[351,536,621,646]" italics="true" pageId="26" pageNumber="27">
<typeStatus id="54D30BA5554DFFDCE020FD9CFE40FD14" box="[351,454,621,646]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
female
</emphasis>
(SAM NN29619): Medium-sized idiopid spider (total length 18.5).
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8FD60FEFAFC84" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8FD60FE90FD38" box="[199,278,657,682]" italics="true" pageId="26" pageNumber="27">Colour</emphasis>
(in ethanol;
<figureCitation id="1353A982554DFFDCE0D8FD60FE6CFD38" box="[423,490,657,682]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7</figureCitation>
AC): Carapace, legs and pedipalp dark red-brown, darker around fovea and lateral margins of caput, with darkened line from fovea to eye group (
<figureCitation id="1353A982554DFFDCE21CFD44FC38FD5C" box="[867,958,693,718]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7A</figureCitation>
); sternum, labium and maxillae uniformly golden-brown, chelicerae dark red-brown (
<figureCitation id="1353A982554DFFDCE3F2FD28FC8BFD60" box="[653,781,729,754]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7E, F</figureCitation>
); abdomen dark brown with indistinct mottled chevron pattern (
<figureCitation id="1353A982554DFFDCE18BFD0CFEE8FC84" box="[244,366,765,790]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7A, C</figureCitation>
).
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8FCD0FBAAFB54" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8FCD0FEE9FCA8" box="[199,367,801,826]" italics="true" pageId="26" pageNumber="27">Cephalothorax</emphasis>
: Carapace 7.8 long, 7.2 wide, 6.3 high, 1.1 times longer than wide; oval (
<figureCitation id="1353A982554DFFDCE5CFFCD0FA8CFCA8" box="[1200,1290,801,826]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7A</figureCitation>
), caput high, ocular area flat (
<figureCitation id="1353A982554DFFDCE026FCB4FE33FCCC" box="[345,437,837,862]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7C</figureCitation>
); cuticle smooth, with pits outward from fovea and both sides of caput, and also two diagonally inward-facing indentations posteriorly; fovea procurved; two indistinct parallel rows of setae from fovea to eye group, less noticeabe both sideson both sides of caput; smaller fine setae also scattered across carapace, concentrated and form fringe around lateral margins; one long seta in fovea; median clump of thickened setae on clypeus (
<figureCitation id="1353A982554DFFDCE01DFC24FE3BFC7C" box="[354,445,981,1006]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7D</figureCitation>
); carapace with fine cover of thin golden hairs (
<figureCitation id="1353A982554DFFDCE29EFC24FBE6FC7C" box="[993,1120,981,1006]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7A, D</figureCitation>
). Length of median clypeus less than 1.0; anterior margin slightly convex. Eye group 1.6 wide, 1.1 long, 0.2 of carapace width; anterior eye row strongly procurved; PLEPLE/ALEALE ratio 1.1; posterior eye row slightly recurved; AME about equal in size to ALE and separated by twice diameter of ALE/AME; ALE and PLE separated by just over ALE diameter; PME similar in size to ALE/AME and about half size of PLE, and separated from PLE by about its own diameter (
<figureCitation id="1353A982554DFFDCE410FB94FF3BFB30" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7D</figureCitation>
). Labium without cuspules (
<figureCitation id="1353A982554DFFDCE37AFB78FDDBFB30" box="[517,605,1161,1186]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7F</figureCitation>
). Sternum 5.0 long, 3.3 wide, evenly setose with setae becoming longer towards anterior margin. Maxillae with
<emphasis id="B91C6915554DFFDCE32BFB5FFDE9FB57" box="[596,623,1198,1221]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 35 cuspules on both sides (
<figureCitation id="1353A982554DFFDCE2D4FB5CFB99FB54" box="[939,1055,1197,1222]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7E, F</figureCitation>
).
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8FB23FD50FAE8" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8FB23FF7AFB7B" box="[199,252,1234,1257]" italics="true" pageId="26" pageNumber="27">Legs</emphasis>
: moderately setose and diffusely spinose; distinct upright setae on distal metatarsi I, II; femora I, II, and pedipalp laterally bowed; tarsi and metatarsi I, II and palpal tarsi scopulate (
<figureCitation id="1353A982554DFFDCE28CFB04FBB0FA9C" box="[1011,1078,1269,1294]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7</figureCitation>
GI). Paired tarsal claws with 1 row of ventral teeth: leg I p2 (1 large, 1 small) r1 (1 large, 1 small); leg II p2 (1 large, 1 small), r2 (1 large, 2 small); right leg III p2 (2 large), r1 (1 large); right leg IV p2 (1 large, 1 small), r1 (1 large); median tarsal claw without teeth. Pedipalp claw with 1 large and 1 small tooth.
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8FA77FAFBF998" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8FA77FEB4FA0F" box="[199,306,1414,1437]" italics="true" pageId="26" pageNumber="27">Spination</emphasis>
: Leg I: tibia p3, r4; metatarsus p2, r5; tarsus with patch of 5 short spines scattered over ventral surface (
<figureCitation id="1353A982554DFFDCE187FA58FEF5FA50" box="[248,371,1449,1474]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7G, H</figureCitation>
). Leg II: tibia p4, r4; metatarsus p4, r5; tarsus with patch of 3 short spines ventrally. Right leg III: patella p3; tibia p3, r3; metatarsus p12, r8; tarsus with patch of 8 spines scattered ventrally. Right leg IV: metatarsus p10, r6; tarsus with
<emphasis id="B91C6915554DFFDCE08AFA03FD89F99B" box="[501,527,1522,1545]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 20 short spines scattered ventrally. Palp: patella p1, tibia p8, r6; tarsus p1, r1.
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8F9E6FE4FF908" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8F9E6FDBAF9BC" box="[199,572,1557,1582]" italics="true" pageId="26" pageNumber="27">Leg and pedipalp measurements</emphasis>
: Length of legs IV&gt; II&gt; I&gt; III. Leg I: femur 4.3, patella 3.1, tibia 2.7, metatarsus 2.1, tarsus 1.7, total = 13.9. Leg II: femur 4.2, patella 3.1, tibia 2.6, metatarsus 1.9, total = 14.9. Leg IV (right): femur 5.5, patella 3.9, tibia 4.0, metatarsus 3.7, tarsus 2.2, total = 19.3. Pedipalp: femur 4, patella 2.2, tibia 2.1, tarsus 2.5, total = 10.8.
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8F954FA1AF92C" blockId="26.[151,1437,258,1978]" box="[199,1436,1701,1726]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8F954FEB7F92C" box="[199,305,1701,1726]" italics="true" pageId="26" pageNumber="27">Abdomen</emphasis>
: Setose, oval, one pair of very small, faint, unsclerotised dorsal sigilla; 10.7 long, 7.3 wide (
<figureCitation id="1353A982554DFFDCE44AF954FA09F92C" box="[1333,1423,1701,1726]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8F938FCF2F894" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8F938FEB7F970" box="[199,305,1737,1762]" italics="true" pageId="26" pageNumber="27">Genitalia</emphasis>
: Spermathecae paired, simple, unbranched, stout and outward facing, oval-shaped with lobe on anterior end, covered in opaque, mottled brown nodules (
<figureCitation id="1353A982554DFFDCE267F91CFCE1F894" box="[792,871,1773,1798]" captionStart="FIGURE 7" captionStartId="27.[151,250,1718,1740]" captionTargetBox="[151,1436,488,1696]" captionTargetId="figure@27.[151,1436,483,1696]" captionTargetPageId="27" captionText="FIGURE 7. Blakistonia bassi sp. n., holotype female (SAM NN29619): A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view; D, eye group, dorsal view; E, sternum, ventral view; F, maxillae and labium, ventral view; G, left leg I, prolateral view; H, left leg I, retrolateral view; J, spermathecae. Scale bars = 5 mm (A), 2 mm (G)." httpUri="https://zenodo.org/record/2609514/files/figure.png" pageId="26" pageNumber="27">
Fig.
<date id="FFD693C7554DFFDCE232F91FFCE1F894" box="[845,871,1774,1798]" pageId="26" pageNumber="27">7I</date>
</figureCitation>
).
</paragraph>
<paragraph id="8BD7B507554DFFDCE1B8F8E3FF7DF828" blockId="26.[151,1437,258,1978]" pageId="26" pageNumber="27">
<emphasis id="B91C6915554DFFDCE1B8F8E3FE02F8B8" box="[199,388,1810,1834]" italics="true" pageId="26" pageNumber="27">Variation (n=3):</emphasis>
Carapace 7.89.0 long, 6.08.0 wide, no labial cuspules. Spination: Leg I: tibia p34, r4; metatarsus p23, r35; tarsus with 36 spines ventrally. Leg II: tibia p34, r4; metatarsus p4, r45; tarsus with 34 spines ventrally. Leg III: patella p23; tibia p03; r23; metatarsus p613, r68; tarsus with 813 spines ventrally. Leg IV: metatarsus p812, r36; tarsus with 1020 spines ventrally. Pedipalp patella p12; tibia p78, r46; tarsus p1, r12.
</paragraph>
</subSubSection>
<subSubSection id="C372E68C554CFFDDE1B8FF66FA1EFF22" box="[199,1432,151,176]" pageId="27" pageNumber="28" type="etymology">
<paragraph id="8BD7B507554CFFDDE1B8FF66FA1EFF22" blockId="27.[151,1437,151,429]" box="[199,1432,151,176]" pageId="27" pageNumber="28">
<emphasis id="B91C6915554CFFDDE1B8FF66FED2FF22" bold="true" box="[199,340,151,176]" pageId="27" pageNumber="28">Etymology.</emphasis>
This species is named in honour of Daniel Bass, for his unwavering support of this research.
</paragraph>
</subSubSection>
<subSubSection id="C372E68C554CFFDDE1B8FF4DFDC1FED2" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BD7B507554CFFDDE1B8FF4DFDC1FED2" blockId="27.[151,1437,151,429]" pageId="27" pageNumber="28">
<emphasis id="B91C6915554CFFDDE1B8FF4DFEEEFF47" bold="true" box="[199,360,188,213]" pageId="27" pageNumber="28">Distribution.</emphasis>
<taxonomicName id="4C68CE84554CFFDDE00CFF4DFDC6FF47" authorityName="Harrison &amp; Rix &amp; Harvey &amp; Austin" authorityYear="2018" box="[371,576,188,213]" class="Arachnida" family="Idiopidae" genus="Blakistonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bassi">
<emphasis id="B91C6915554CFFDDE00CFF4DFDC6FF47" box="[371,576,188,213]" italics="true" pageId="27" pageNumber="28">Blakistonia bassi</emphasis>
</taxonomicName>
is known from only a single roadside cutting at Ashton in the Mount Lofty Ranges, (
<figureCitation id="1353A982554CFFDDE030FF2EFE21FF6A" box="[335,423,223,248]" captionStart="FIGURE 31" captionStartId="71.[151,250,1965,1987]" captionTargetBox="[269,1330,1140,1913]" captionTargetId="figure@71.[190,1434,1116,1996]" captionTargetPageId="71" captionText="FIGURE 31. Distributions of Blakistonia species: B. bassi sp. n. (black circle), B. hortoni sp. n. (white squares), B. bella sp. n. (white circles), and B. gemmelli sp. n. (black square)." httpUri="https://zenodo.org/record/2609562/files/figure.png" pageId="27" pageNumber="28">Fig. 31</figureCitation>
). A number of active burrows were found in 2017; however, the species has not been found elsewhere in the Mount Lofty Range despite extensive historical collection, and was not discovered anywhere else as part of this project.
</paragraph>
</subSubSection>
<subSubSection id="C372E68C554CFFDDE1B8FEBDFD66FE3E" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BD7B507554CFFDDE1B8FEBDFD66FE3E" blockId="27.[151,1437,151,429]" pageId="27" pageNumber="28">
<emphasis id="B91C6915554CFFDDE1B8FEBDFEC6FEF7" bold="true" box="[199,320,332,357]" pageId="27" pageNumber="28">Remarks.</emphasis>
The burrow (
<figureCitation id="1353A982554CFFDDE09EFEBDFDD9FEF6" box="[481,607,332,357]" captionStart="FIGURE 2" captionStartId="8.[151,250,1508,1530]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@8.[151,1436,193,1486]" captionTargetPageId="8" captionText="FIGURE 2. Images of Blakistonia burrows. A, B. aurea Hogg, 1902 burrow with scalloped edge from Humbug Scrub, South Australia; B, B. aurea burrow from Parra Wirra Recreation Park, South Australia; C, B. aurea burrow, Mount Lofty Ranges, South Australia; DE, B. bassi sp. n. burrow from Ashton, Mount Lofty Ranges, South Australia; F, B. birksi sp. n. burrow from Mark Oliphant Conservation Park, Mount Lofty Ranges, South Australia; G, B. mainae sp. n. burrow from Mount Ragged, Cape Arid National Park, Western Australia; HI, B. maryae sp. n. burrow showing twig-lining from Scrubby Peak, Gawler Ranges, South Australia; J, B. maryae sp. n. burrow showing twig-lining from Tumby Bay, Eyre Peninsula, South Australia; K, B. maryae sp. n. burrow from Gawler Ranges, South Australia; L, B. nullarborensis sp. n. burrow from Cocklebiddy, Western Australia. All images by S. Harrison except: C by N. Birks; H, by N. Birks; I, by N. Birks; all used with permission." httpUri="https://zenodo.org/record/2609504/files/figure.png" pageId="27" pageNumber="28">Fig. 2D, E</figureCitation>
) is similar to that of
<taxonomicName id="4C68CE84554CFFDDE227FEBDFC43FEF7" authorityName="Ausserer" authorityYear="1871" box="[856,965,332,357]" class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B91C6915554CFFDDE227FEBDFC43FEF7" box="[856,965,332,357]" italics="true" pageId="27" pageNumber="28">Idiosoma</emphasis>
</taxonomicName>
in its thin, cryptic, flap-like nature, and certainly different to the D-shaped, plug-like burrow typical of
<taxonomicName id="4C68CE84554CFFDDE2E5FE80FB86FE1A" authorityName="Hogg" authorityYear="1902" box="[922,1024,369,392]" class="Arachnida" family="Idiopidae" genus="Blakistonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="aurea">
<emphasis id="B91C6915554CFFDDE2E5FE80FB86FE1A" box="[922,1024,369,392]" italics="true" pageId="27" pageNumber="28">B. aurea</emphasis>
</taxonomicName>
. All burrows found were adorned with moss and cryptic in appearance (
<figureCitation id="1353A982554CFFDDE32CFE65FD55FE3E" box="[595,723,404,429]" captionStart="FIGURE 2" captionStartId="8.[151,250,1508,1530]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@8.[151,1436,193,1486]" captionTargetPageId="8" captionText="FIGURE 2. Images of Blakistonia burrows. A, B. aurea Hogg, 1902 burrow with scalloped edge from Humbug Scrub, South Australia; B, B. aurea burrow from Parra Wirra Recreation Park, South Australia; C, B. aurea burrow, Mount Lofty Ranges, South Australia; DE, B. bassi sp. n. burrow from Ashton, Mount Lofty Ranges, South Australia; F, B. birksi sp. n. burrow from Mark Oliphant Conservation Park, Mount Lofty Ranges, South Australia; G, B. mainae sp. n. burrow from Mount Ragged, Cape Arid National Park, Western Australia; HI, B. maryae sp. n. burrow showing twig-lining from Scrubby Peak, Gawler Ranges, South Australia; J, B. maryae sp. n. burrow showing twig-lining from Tumby Bay, Eyre Peninsula, South Australia; K, B. maryae sp. n. burrow from Gawler Ranges, South Australia; L, B. nullarborensis sp. n. burrow from Cocklebiddy, Western Australia. All images by S. Harrison except: C by N. Birks; H, by N. Birks; I, by N. Birks; all used with permission." httpUri="https://zenodo.org/record/2609504/files/figure.png" pageId="27" pageNumber="28">Fig. 2D, E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>