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<document id="651B6BD39290561C08BDD3FFAB9D2D84" ID-CLB-Dataset="222448" ID-DOI="10.11646/zootaxa.5227.4.3" ID-GBIF-Dataset="ea087173-f3d9-491b-8157-366b194bb519" ID-ISSN="1175-5326" ID-Zenodo-Dep="7518924" ID-ZooBank="B28C27C8-7273-4294-8487-150075ABD889" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673311993872" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Khachikov, Eduard A." docDate="2023" docId="03BD8E09FFCAF64B4F90F9E113A9A6CB" docLanguage="en" docName="zootaxa.5227.4.3.pdf" docOrigin="Zootaxa 5227 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium rugatum Mulsant &amp; Rey 1880" docType="treatment" docVersion="7" lastPageNumber="453" masterDocId="FF84F671FFC2F6414F07FFF31767A632" masterDocTitle="Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae, Omaliini)" masterLastPageNumber="458" masterPageNumber="443" pageNumber="451" updateTime="1699473653611" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="1606C3709FAC143394D230B0F4220145" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="484F28BF53FCEB1CE04E75B63CE7A18B">Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae, Omaliini)</mods:title>
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<mods:namePart id="9B02628F1DA5BD6665CC3746C0DD83D7">Shavrin, Alexey V.</mods:namePart>
<mods:nameIdentifier id="4CCD911201EACCD21F639A2C59F23AF6" type="ORCID">0000-0001-6487-4530</mods:nameIdentifier>
<mods:affiliation id="E2717F4B76B78B3354C63FB2FC1B1293">Daugavpils University, Institute of Life Sciences and Technologies, Coleopterological Research Centre, Vienibas 13, Daugavpils, LV- 5401, Latvia. ashavrin @ hotmail. com; https: // orcid. org / 0000 - 0001 - 6487 - 4530</mods:affiliation>
<mods:nameIdentifier id="E5514978C3A3272BFA897564BCA638B5" type="email">ashavrin@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="C9C5D9139B7178AC0C94DEA473F9E9FA">Khachikov, Eduard A.</mods:namePart>
<mods:affiliation id="B20996C34D7333E792160F3FE63B8355">Southern Federal University, Bolshaya Sadovaya 105 / 42, Rostov-on-Don, 34006, Russia.</mods:affiliation>
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<mods:title id="8461E1267B578833F413D65681FEAC51">Zootaxa</mods:title>
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<mods:date id="4B11DC5367B02657346CC5E56CDD6BF3">2023</mods:date>
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<mods:number id="E835A96971285D4744A22E964D9F3959">2023-01-09</mods:number>
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<mods:identifier id="DD5972E909521BF6852E6062ED182A2F" type="DOI">10.11646/zootaxa.5227.4.3</mods:identifier>
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<treatment id="03BD8E09FFCAF64B4F90F9E113A9A6CB" ID-DOI="http://doi.org/10.5281/zenodo.7524767" ID-GBIF-Taxon="208788671" ID-Zenodo-Dep="7524767" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB" httpUri="http://treatment.plazi.org/id/03BD8E09FFCAF64B4F90F9E113A9A6CB" lastPageId="10" lastPageNumber="453" pageId="8" pageNumber="451">
<subSubSection id="C30E6C94FFCAF6494F90F9E115EEA01E" box="[151,649,1554,1581]" pageId="8" pageNumber="451" type="nomenclature">
<paragraph id="8BAB3F1FFFCAF6494F90F9E115EEA01E" blockId="8.[151,649,1554,1617]" box="[151,649,1554,1581]" pageId="8" pageNumber="451">
<heading id="D0E38873FFCAF6494F90F9E115EEA01E" bold="true" box="[151,649,1554,1581]" fontSize="11" level="1" pageId="8" pageNumber="451" reason="1">
<taxonomicName id="4C14449CFFCAF6494F90F9E115EEA01E" ID-CoL="499YL" authority="Mulsant &amp; Rey, 1880" authorityName="Mulsant &amp; Rey" authorityYear="1880" box="[151,649,1554,1581]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="451" phylum="Arthropoda" rank="species" species="rugatum">
<emphasis id="B960E30DFFCAF6494F90F9E115EEA01E" bold="true" box="[151,649,1554,1581]" pageId="8" pageNumber="451">
<emphasis id="B960E30DFFCAF6494F90F9E1161EA01F" bold="true" box="[151,377,1554,1581]" italics="true" pageId="8" pageNumber="451">Omalium rugatum</emphasis>
<bibRefCitation id="EF8542EEFFCAF6494E87F9E015EEA01E" author="Mulsant, E. &amp; Rey, C." box="[384,649,1554,1581]" pageId="8" pageNumber="451" pagination="1 - 430" refId="ref11269" refString="Mulsant, E. &amp; Rey, C. (1880) Histoire naturelle des Coleopteres de France. Tribu des brevipennes. Onzieme famille: Omaliens. Douxieme famille: Pholidiens. Annales de la Societe Linneenne de Lyon, Series 2, 27, 1 - 430. https: // doi. org / 10.3406 / linly. 1880.3986" type="journal article" year="1880">Mulsant &amp; Rey, 1880</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFCAF6494F90F9C41613A063" box="[151,372,1591,1617]" pageId="8" pageNumber="451" type="description">
<paragraph id="8BAB3F1FFFCAF6494F90F9C41613A063" blockId="8.[151,649,1554,1617]" box="[151,372,1591,1617]" pageId="8" pageNumber="451">
(
<figureCitation id="132F239AFFCAF6494F98F9C41789A063" box="[159,238,1591,1617]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1864,1888]" captionTargetBox="[234,1353,195,1823]" captionTargetId="figure-19@3.[220,1367,181,1840]" captionTargetPageId="3" captionText="FIGURES 1–10. Habitus of Omalium: 1—O. caesum (Turkey, Bursa: Uludağ), 2—O. cinnamomeum (Turkey, Antakya: Kişlak- Şenköy), 3—O. excavatum (Russia, Karachay-Cherkessia: Arkhyz), 4—O. falsum (Georgia, Tskhneti), 5—O. littorale (Russia, Adygea: Abago), 6—O. riparium impar (Russia, Krasnodar Territory: Khosta), 7—O. rivulare (Armenia, Alaverdi), 8—O. rugatum (Iran, Zagros Mts.), 9—O. turcicum (Russia, Adygea: Lagonaki), 10—O. wunderlei (Turkey, Gümüşhane: Erzincan- Kelkit). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518926" httpUri="https://zenodo.org/record/7518926/files/figure.png" pageId="8" pageNumber="451">Figs. 8</figureCitation>
,
<figureCitation id="132F239AFFCAF6494FFCF9C41624A063" box="[251,323,1591,1617]" captionStart="FIGURES 27–32" captionStartId="10.[151,264,1903,1927]" captionTargetBox="[165,1422,281,1870]" captionTargetId="figure-62@10.[152,1436,273,1880]" captionTargetPageId="10" captionText="FIGURES 27–32. Aedeagus of Omalium rugatum (Figs. 27–28: Iran, Zagros), O. turcicum (Figs. 29–30: Turkey, Izmir: Bozdagh), O. wunderlei (Figs. 31–32: Turkey, Kars: Kars): 27, 29, 31—parameral view, 28, 30, 32—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7518932" httpUri="https://zenodo.org/record/7518932/files/figure.png" pageId="8" pageNumber="451">27–28</figureCitation>
,
<figureCitation id="132F239AFFCAF6494E48F9C4160AA063" box="[335,365,1591,1617]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1503,1527]" captionTargetBox="[161,1431,318,1466]" captionTargetId="figure-33@11.[151,1435,304,1480]" captionTargetPageId="11" captionText="FIGURES 33–41. Female genital segment (ventral view) and spermatheca (shown by arrow in Fig. 40) of Omalium: 33—O. caesum (Russia, North Ossetia-Alania: Zincar), 34—O. cinnamomeum (Turkey, Antakya: Kişlak-Şenköy), 35—O. excavatum (Russia, Krasnodar Territory: Caucasus Nature Reserve), 36—O. falsum (Turkey, Rize: Ardeşen), 37—O. littorale (Russia, Volgograd Area: Elton Lake), 38—O. riparium impar (Georgia, Tskhneti), 39—O. rivulare (Russia, Krasnodar Territory: Derbiy), 40—O. rugatum (Russia, North Ossetia-Alania: Zincar), 41—O. turcicum (Russia, Krasnodar Territory, Caucasus Nature Reserve). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7518934" httpUri="https://zenodo.org/record/7518934/files/figure.png" pageId="8" pageNumber="451">40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFCAF6494F90F9881502A0A1" box="[151,613,1659,1684]" pageId="8" pageNumber="451" type="reference_group">
<paragraph id="8BAB3F1FFFCAF6494F90F9881502A0A1" blockId="8.[151,613,1659,1684]" box="[151,613,1659,1684]" pageId="8" pageNumber="451">
<treatmentCitationGroup id="AB041831FFCAF6494F90F9881502A0A1" box="[151,613,1659,1684]" pageId="8" pageNumber="451">
<treatmentCitation id="0AB5190EFFCAF6494F90F9881502A0A1" author="Mulsant, E. &amp; Rey, C." box="[151,613,1659,1684]" page="217" pageId="8" pageNumber="451" year="1880">
<taxonomicName id="4C14449CFFCAF6494F90F9881502A0A1" ID-CoL="499YL" authority="Mulsant &amp; Rey, 1880: 217" authorityName="Mulsant &amp; Rey" authorityPageNumber="217" authorityYear="1880" box="[151,613,1659,1684]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="451" phylum="Arthropoda" rank="species" species="rugatum">
<emphasis id="B960E30DFFCAF6494F90F9881635A0A1" box="[151,338,1659,1683]" italics="true" pageId="8" pageNumber="451">Omalium rugatum</emphasis>
<bibRefCitation id="EF8542EEFFCAF6494E5FF9881502A0A1" author="Mulsant, E. &amp; Rey, C." box="[344,613,1659,1684]" pageId="8" pageNumber="451" pagination="1 - 430" refId="ref11269" refString="Mulsant, E. &amp; Rey, C. (1880) Histoire naturelle des Coleopteres de France. Tribu des brevipennes. Onzieme famille: Omaliens. Douxieme famille: Pholidiens. Annales de la Societe Linneenne de Lyon, Series 2, 27, 1 - 430. https: // doi. org / 10.3406 / linly. 1880.3986" type="journal article" year="1880">Mulsant &amp; Rey, 1880: 217</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFCAF6484F90F93215FAA773" lastPageId="9" lastPageNumber="452" pageId="8" pageNumber="451" type="materials_examined">
<paragraph id="8BAB3F1FFFCAF6484F90F93216F0A6CB" blockId="8.[151,1437,1729,2011]" lastBlockId="9.[151,1436,150,393]" lastPageId="9" lastPageNumber="452" pageId="8" pageNumber="451">
<emphasis id="B960E30DFFCAF6494F90F93216E4A0E9" bold="true" box="[151,387,1729,1755]" pageId="8" pageNumber="451">Material examined.</emphasis>
<materialsCitation id="3B7C3542FFCAF6494E8BF93214B4A0CD" ID-GBIF-Occurrence="4012666301" collectingDate="2007-08-22" collectorName="F. A. Martynovchenko" country="Russia" location="Malaya Khatipara Mt." municipality="Teberda State Nature Biosphere Reserve" pageId="8" pageNumber="451" specimenCount="1" stateProvince="Karachayevo-Cherkesiya">
<emphasis id="B960E30DFFCAF6494E8BF9321444A0E9" bold="true" box="[396,803,1729,1755]" pageId="8" pageNumber="451">
<collectingCountry id="F3037F8FFFCAF6494E8BF9321690A0E9" box="[396,503,1729,1755]" name="Russia" pageId="8" pageNumber="451">RUSSIA</collectingCountry>
:
<collectingRegion id="49D0F1FDFFCAF6494D02F9321478A0E9" box="[517,799,1729,1755]" country="Russia" name="Karachayevo-Cherkesiya" pageId="8" pageNumber="451">KARACHAY-CHeRKeSSIA</collectingRegion>
:
</emphasis>
1 ♁:
<collectingMunicipality id="6BCFA565FFCAF6494C6EF932125CA0EE" box="[873,1339,1729,1756]" pageId="8" pageNumber="451">Teberda State Nature Biosphere Reserve</collectingMunicipality>
,
<location id="8ECB69C4FFCAF6494A4FF9321653A132" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB:8ECB69C4FFCAF6494A4FF9321653A132" country="Russia" municipality="Teberda State Nature Biosphere Reserve" name="Malaya Khatipara Mt." pageId="8" pageNumber="451" stateProvince="Karachayevo-Cherkesiya">Malaya Khatipara Mt.</location>
10–
<date id="FFAA19DFFFCAF6494E64F9161681A0CD" box="[355,486,1765,1791]" pageId="8" pageNumber="451" value="2007-08-22">
<collectingDate id="EFEEE037FFCAF6494E64F9161681A0CD" box="[355,486,1765,1791]" pageId="8" pageNumber="451" value="2007-08-22">22.08.2007</collectingDate>
</date>
.
<collectingMethod id="52554708FFCAF6494EF7F9161539A132" box="[496,606,1765,1792]" pageId="8" pageNumber="451">Soil traps</collectingMethod>
.
<collectorName id="26E15AC9FFCAF6494D6FF915143AA0CD" box="[616,861,1765,1792]" pageId="8" pageNumber="451">F.A. Martynovchenko</collectorName>
leg. (cSh)
</materialsCitation>
;
<materialsCitation id="3B7C3542FFCAF6494CD8F91614A2A111" ID-GBIF-Occurrence="4012666327" collectingDate="1984-04-13" collectionCode="ZMM" collectorName="S. K. Alexeev" country="Russia" location="Zincar. Forest" municipality="Ardon River" pageId="8" pageNumber="451" specimenCount="1" specimenCount-female="1" stateProvince="North Ossetia">
<emphasis id="B960E30DFFCAF6494CD8F9161265A0CD" bold="true" box="[991,1282,1765,1791]" pageId="8" pageNumber="451">
<collectingRegion id="49D0F1FDFFCAF6494CD8F9161399A0CD" box="[991,1278,1765,1791]" country="Russia" name="North Ossetia" pageId="8" pageNumber="451">NORTH OSSeTIA-ALANIA</collectingRegion>
:
</emphasis>
1 ♁,
<specimenCount id="9D12F496FFCAF6494A38F916120CA132" box="[1343,1387,1765,1792]" pageId="8" pageNumber="451" type="female">1 ♀</specimenCount>
:
<collectingMunicipality id="6BCFA565FFCAF6494A71F915166FA111" pageId="8" pageNumber="451">Ardon River</collectingMunicipality>
,
<location id="8ECB69C4FFCAF6494E14F8FA16C8A116" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB:8ECB69C4FFCAF6494E14F8FA16C8A116" box="[275,431,1801,1828]" country="Russia" municipality="Ardon River" name="Zincar. Forest" pageId="8" pageNumber="451" stateProvince="North Ossetia">Zincar. Forest</location>
swamp.
<date id="FFAA19DFFFCAF6494D13F8FA15F2A111" box="[532,661,1801,1827]" pageId="8" pageNumber="451" value="1984-04-13">
<collectingDate id="EFEEE037FFCAF6494D13F8FA15F2A111" box="[532,661,1801,1827]" pageId="8" pageNumber="451" value="1984-04-13">13.04.1984</collectingDate>
</date>
.
<collectorName id="26E15AC9FFCAF6494DA7F8FA1453A111" box="[672,820,1801,1827]" pageId="8" pageNumber="451">S.K. Alexeev</collectorName>
leg. (
<collectionCode id="ED05A7DAFFCAF6494C73F8FA14DAA111" box="[884,957,1801,1827]" pageId="8" pageNumber="451">ZMM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3542FFCAF6494CD7F8FA1629A175" ID-GBIF-Occurrence="4012666313" collectorName="Vladikavkaz &amp; Dzaudzhikau" country="Russia" location="Ordzhonikidze" pageId="8" pageNumber="451" specimenCount="1" specimenCount-female="1" stateProvince="NORTH OSSeTIA-ALANIA">
1 ♁,
<specimenCount id="9D12F496FFCAF6494B0FF8FA1353A116" box="[1032,1076,1801,1828]" pageId="8" pageNumber="451" type="female">1 ♀</specimenCount>
:
<location id="8ECB69C4FFCAF6494B46F8FA1380A111" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB:8ECB69C4FFCAF6494B46F8FA1380A111" box="[1089,1255,1801,1827]" country="Russia" name="Ordzhonikidze" pageId="8" pageNumber="451" stateProvince="NORTH OSSeTIA-ALANIA">Ordzhonikidze</location>
(=
<collectorName id="26E15AC9FFCAF6494A02F8FA12FFA111" box="[1285,1432,1801,1827]" pageId="8" pageNumber="451">Vladikavkaz</collectorName>
, =
<collectorName id="26E15AC9FFCAF6494FAAF8DE1621A175" box="[173,326,1837,1863]" pageId="8" pageNumber="451">Dzaudzhikau</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7C3542FFCAF6494E50F8DE143CA175" ID-GBIF-Occurrence="4012666310" box="[343,859,1837,1864]" collectingDate="1978-05-28" collectionCode="NMPC" collectorName="Z. et J. Novotny" country="Russia" pageId="8" pageNumber="451" specimenCount="1" stateProvince="NORTH OSSeTIA-ALANIA">
26–
<date id="FFAA19DFFFCAF6494E85F8DE1563A17A" box="[386,516,1837,1864]" pageId="8" pageNumber="451" value="1978-05-28">
<collectingDate id="EFEEE037FFCAF6494E85F8DE1563A17A" box="[386,516,1837,1864]" pageId="8" pageNumber="451" value="1978-05-28">28.05.1978</collectingDate>
</date>
.
<collectorName id="26E15AC9FFCAF6494D0AF8DD15A7A175" box="[525,704,1837,1864]" pageId="8" pageNumber="451">Z. et J. Novotný</collectorName>
leg. (
<collectionCode id="ED05A7DAFFCAF6494DFAF8DE1434A175" box="[765,851,1837,1863]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="451">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3542FFCAF6494C62F8DE12F0A159" ID-GBIF-Occurrence="4012666328" collectingDate="2011-06-23" collectorName="A. S. Prosvirov" country="Georgia" elevation="1585" location="Khilsta River" municipality="Bzyb Range" pageId="8" pageNumber="451" specimenCount="1" stateProvince="NORTH OSSeTIA-ALANIA">
<emphasis id="B960E30DFFCAF6494C62F8DE17A1A159" bold="true" pageId="8" pageNumber="451">
<collectingCountry id="F3037F8FFFCAF6494C62F8DE1496A175" box="[869,1009,1837,1863]" name="Georgia" pageId="8" pageNumber="451">GEORGIA</collectingCountry>
: AUTONOMOUS RePUBLIC OF ABKHAZIA:
</emphasis>
1 ♁:
<collectingMunicipality id="6BCFA565FFCAF6494E0CF8A216FFA15E" box="[267,408,1873,1900]" pageId="8" pageNumber="451">Bzyb Range</collectingMunicipality>
, rise on gorge of
<location id="8ECB69C4FFCAF6494D64F8A2159AA159" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB:8ECB69C4FFCAF6494D64F8A2159AA159" box="[611,765,1873,1899]" country="Georgia" municipality="Bzyb Range" name="Khilsta River" pageId="8" pageNumber="451" stateProvince="NORTH OSSeTIA-ALANIA">Khilsta River</location>
.
<elevation id="0039D82CFFCAF6494C0EF8A21484A159" box="[777,995,1873,1900]" metricMagnitude="3" metricUnit="m" metricValue="1.585" metricValueMax="1.6" metricValueMin="1.5699999999999998" pageId="8" pageNumber="451" unit="m" value="1585.0" valueMax="1600.0" valueMin="1570.0">
<quantity id="4CEC92FAFFCAF6494C0EF8A214C1A15E" box="[777,934,1873,1900]" metricMagnitude="3" metricUnit="m" metricValue="1.585" metricValueMax="1.6" metricValueMin="1.5699999999999998" pageId="8" pageNumber="451" unit="m" value="1585.0" valueMax="1600.0" valueMin="1570.0">1570–1600 m</quantity>
a.s.l.
</elevation>
<date id="FFAA19DFFFCAF6494CECF8A2130BA159" box="[1003,1132,1873,1899]" pageId="8" pageNumber="451" value="2011-06-23">
<collectingDate id="EFEEE037FFCAF6494CECF8A2130BA159" box="[1003,1132,1873,1899]" pageId="8" pageNumber="451" value="2011-06-23">23.06.2011</collectingDate>
</date>
.
<collectorName id="26E15AC9FFCAF6494B71F8A2127AA159" box="[1142,1309,1873,1899]" pageId="8" pageNumber="451">A.S. Prosvirov</collectorName>
leg. (cSh)
</materialsCitation>
;
<materialsCitation id="3B7C3542FFCAF6494F90F88616DAA181" ID-GBIF-Occurrence="4012666305" collectingDate="1987-10-05" collectorName="S. I. Golovatch &amp; K. Yu. Eskov" country="Georgia" elevation="1050" location="Bikhura." municipality="Tbilisi" pageId="8" pageNumber="451" specimenCount="1" specimenCount-female="1" stateProvince="Tbilisi">
<emphasis id="B960E30DFFCAF6494F90F8861794A1BD" bold="true" box="[151,243,1909,1935]" pageId="8" pageNumber="451">
<collectingRegion id="49D0F1FDFFCAF6494F90F8861788A1BD" box="[151,239,1909,1935]" country="Georgia" name="Tbilisi" pageId="8" pageNumber="451">TBILISI</collectingRegion>
:
</emphasis>
1 ♁,
<specimenCount id="9D12F496FFCAF6494E3FF8861601A1A2" box="[312,358,1909,1936]" pageId="8" pageNumber="451" type="female">1 ♀</specimenCount>
: pass between
<collectingMunicipality id="6BCFA565FFCAF6494D14F886153BA1BD" box="[531,604,1909,1935]" pageId="8" pageNumber="451">Tbilisi</collectingMunicipality>
and
<location id="8ECB69C4FFCAF6494D91F886159FA1BD" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB:8ECB69C4FFCAF6494D91F886159FA1BD" box="[662,760,1909,1935]" country="Georgia" municipality="Tbilisi" name="Bikhura." pageId="8" pageNumber="451" stateProvince="Tbilisi">Bikhura.</location>
<elevation id="0039D82CFFCAF6494C07F88614F1A1BD" box="[768,918,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="451" unit="m" value="1050.0">
<quantity id="4CEC92FAFFCAF6494C07F8861430A1A2" box="[768,855,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="451" unit="m" value="1050.0">1050 m</quantity>
a.s.l.
</elevation>
, in litter and soil.
<date id="FFAA19DFFFCAF6494B6DF886138CA1A2" box="[1130,1259,1909,1936]" pageId="8" pageNumber="451" value="1987-10-05">
<collectingDate id="EFEEE037FFCAF6494B6DF886138CA1A2" box="[1130,1259,1909,1936]" pageId="8" pageNumber="451" value="1987-10-05">10.05.1987</collectingDate>
</date>
.
<collectorName id="26E15AC9FFCAF6494BFFF88612FBA1BD" box="[1272,1436,1909,1935]" pageId="8" pageNumber="451">S.I. Golovatch</collectorName>
&amp;
<collectorName id="26E15AC9FFCAF6494FB2F8691622A181" box="[181,325,1945,1972]" pageId="8" pageNumber="451">K.Yu. Eskov</collectorName>
leg. (cSh)
</materialsCitation>
;
<materialsCitation id="3B7C3542FFCAF6494ECDF86A155AA1E5" ID-GBIF-Occurrence="4012666304" collectingDate="2015-09-05" collectorName="S. Anlas" country="Turkey" elevation="1893" latitude="38.88889" location="Emir Dadlari" longLatPrecision="19" longitude="31.195555" pageId="8" pageNumber="451" specimenCount="1" stateProvince="Afyonkarahisar">
<emphasis id="B960E30DFFCAF6494ECDF86A144AA181" bold="true" box="[458,813,1945,1971]" pageId="8" pageNumber="451">
<collectingCountry id="F3037F8FFFCAF6494ECDF86A1523A181" box="[458,580,1945,1971]" name="Turkey" pageId="8" pageNumber="451">TURKEY</collectingCountry>
:
<collectingRegion id="49D0F1FDFFCAF6494D56F86A144EA181" box="[593,809,1945,1971]" country="Turkey" name="Afyonkarahisar" pageId="8" pageNumber="451">AFYONKARAHISAR</collectingRegion>
:
</emphasis>
4 ♁♁:
<location id="8ECB69C4FFCAF6494C82F86A137BA186" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB:8ECB69C4FFCAF6494C82F86A137BA186" box="[901,1052,1945,1972]" country="Turkey" latitude="38.88889" longLatPrecision="19" longitude="31.195555" name="Emir Dadlari" pageId="8" pageNumber="451" stateProvince="Afyonkarahisar">Emir Daðlari</location>
,
<geoCoordinate id="EE2059D8FFCAF6494B2FF86813CAA184" box="[1064,1197,1947,1974]" degrees="38" direction="north" minutes="53" orientation="latitude" pageId="8" pageNumber="451" precision="15" seconds="20" value="38.88889">38°53′20′′N</geoCoordinate>
<geoCoordinate id="EE2059D8FFCAF6494BB1F868125CA184" box="[1206,1339,1947,1974]" degrees="31" direction="east" minutes="11" orientation="longitude" pageId="8" pageNumber="451" precision="15" seconds="44" value="31.195555">31°11′44′′E</geoCoordinate>
.
<elevation id="0039D82CFFCAF6494A41F86A17ABA1E5" metricMagnitude="3" metricUnit="m" metricValue="1.893" pageId="8" pageNumber="451" unit="m" value="1893.0">
<quantity id="4CEC92FAFFCAF6494A41F86A12FBA186" box="[1350,1436,1945,1972]" metricMagnitude="3" metricUnit="m" metricValue="1.893" pageId="8" pageNumber="451" unit="m" value="1893.0">1893 m</quantity>
a.s.l.
</elevation>
<date id="FFAA19DFFFCAF6494FD3F84E1632A1EA" box="[212,341,1981,2008]" pageId="8" pageNumber="451" value="2015-09-05">
<collectingDate id="EFEEE037FFCAF6494FD3F84E1632A1EA" box="[212,341,1981,2008]" pageId="8" pageNumber="451" value="2015-09-05">09.05.2015</collectingDate>
</date>
.
<collectorName id="26E15AC9FFCAF6494E66F84E16D8A1E5" box="[353,447,1981,2007]" pageId="8" pageNumber="451">S. Anlaş</collectorName>
leg. (cAn)
</materialsCitation>
;
<materialsCitation id="3B7C3542FFCAF6484D4DF84E14D8A683" ID-GBIF-Occurrence="4012666312" collectingDate="1978-06-15" collectionCode="SMNS" collectorName="J. Martens &amp; R. Pieper" country="Iran" elevation="1725" lastPageId="9" lastPageNumber="452" location="Arasbaran Biosphere Reserve" pageId="8" pageNumber="451" specimenCount="2" specimenCount-female="2" stateProvince="AFYONKARAHISAR">
<emphasis id="B960E30DFFCAF6494D4DF84E14D3A1E5" bold="true" box="[586,948,1981,2011]" pageId="8" pageNumber="451">
<collectingCountry id="F3037F8FFFCAF6494D4DF84E15F1A1E5" box="[586,662,1981,2007]" name="Iran" pageId="8" pageNumber="451">IRAN</collectingCountry>
: ÂZARBẲIJẲN-e SHARQI:
</emphasis>
1 ♁,
<specimenCount id="9D12F496FFCAF6494CF0F84E135DA1EA" box="[1015,1082,1981,2008]" pageId="8" pageNumber="451" type="female">2 ♀♀</specimenCount>
:
<location id="8ECB69C4FFCAF6494B41F84E12FEA1EA" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB:8ECB69C4FFCAF6494B41F84E12FEA1EA" box="[1094,1433,1981,2008]" country="Iran" name="Arasbaran Biosphere Reserve" pageId="8" pageNumber="451" stateProvince="AFYONKARAHISAR">Arasbaran Biosphere Reserve</location>
.
<elevation id="0039D82CFFCBF6484F90FF65160AA683" box="[151,365,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="9" pageNumber="452" unit="m" value="1725.0" valueMax="1800.0" valueMin="1650.0">
<quantity id="4CEC92FAFFCBF6484F90FF651655A683" box="[151,306,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="9" pageNumber="452" unit="m" value="1725.0" valueMax="1800.0" valueMin="1650.0">1650–1800 m</quantity>
a.s.l.
</elevation>
04–
<date id="FFAA19DFFFCBF6484E99FF651578A683" box="[414,543,150,177]" pageId="9" pageNumber="452" value="1978-06-15">
<collectingDate id="EFEEE037FFCBF6484E99FF651578A683" box="[414,543,150,177]" pageId="9" pageNumber="452" value="1978-06-15">15.06.1978</collectingDate>
</date>
.
<collectorName id="26E15AC9FFCBF6484D2FFF6415FAA683" box="[552,669,151,177]" pageId="9" pageNumber="452">J. Martens</collectorName>
&amp;
<collectorName id="26E15AC9FFCBF6484DB8FF641440A683" box="[703,807,151,177]" pageId="9" pageNumber="452">R. Pieper</collectorName>
leg. (
<collectionCode id="ED05A7DAFFCBF6484C62FF6414D0A683" box="[869,951,151,177]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="9" pageNumber="452">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3542FFCBF6484CCDFF6B16F4A6CB" ID-GBIF-Occurrence="4012666314" collectingDate="2018-06-24" collectionCode="NME" collectorName="M. Hartmann" country="Iran" elevation="1850" latitude="30.704445" location="Zagros Mts." longLatPrecision="20" longitude="51.586945" pageId="9" pageNumber="452" specimenCount="5" specimenCount-female="5" stateProvince="AFYONKARAHISAR">
<emphasis id="B960E30DFFCBF6484CCDFF6B12FBA686" bold="true" box="[970,1436,150,180]" pageId="9" pageNumber="452">OSTẲN-e KOHGÎLŰYe VA BŰYeR-AHMAD:</emphasis>
10 ♁♁,
<specimenCount id="9D12F496FFCBF6484FF6FF491655A6E7" box="[241,306,186,213]" pageId="9" pageNumber="452" type="female">5 ♀♀</specimenCount>
:
<location id="8ECB69C4FFCBF6484E39FF4816AFA6E7" LSID="urn:lsid:plazi:treatment:03BD8E09FFCAF64B4F90F9E113A9A6CB:8ECB69C4FFCBF6484E39FF4816AFA6E7" box="[318,456,187,213]" country="Iran" latitude="30.704445" longLatPrecision="20" longitude="51.586945" name="Zagros Mts." pageId="9" pageNumber="452" stateProvince="AFYONKARAHISAR">Zagros Mts.</location>
,
<locationDeviation id="9A19263DFFCBF6484ED5FF481500A6E7" box="[466,615,187,213]" location="Yasoj" pageId="9" pageNumber="452">
<quantity id="4CEC92FAFFCBF6484ED5FF48156CA6E7" box="[466,523,187,213]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="452" unit="km" value="4.0">4 km</quantity>
S Yasoj
</locationDeviation>
,
<geoCoordinate id="EE2059D8FFCBF6484D75FF4F159FA6E5" box="[626,760,188,215]" degrees="30" direction="north" minutes="42" orientation="latitude" pageId="9" pageNumber="452" precision="15" seconds="16" value="30.704445">30°42′16′′N</geoCoordinate>
<geoCoordinate id="EE2059D8FFCBF6484DF8FF4F14E2A6E5" box="[767,901,188,215]" degrees="51" direction="east" minutes="35" orientation="longitude" pageId="9" pageNumber="452" precision="15" seconds="13" value="51.586945">51°35′13′′E</geoCoordinate>
.
<elevation id="0039D82CFFCBF6484C88FF491378A6E7" box="[911,1055,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="452" unit="m" value="1850.0">
<quantity id="4CEC92FAFFCBF6484C88FF491484A6E7" box="[911,995,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="452" unit="m" value="1850.0">1850 m</quantity>
a.s.l.
</elevation>
, river gravelbank.
<date id="FFAA19DFFFCBF6484BF6FF481215A6E7" box="[1265,1394,187,213]" pageId="9" pageNumber="452" value="2018-06-24">
<collectingDate id="EFEEE037FFCBF6484BF6FF481215A6E7" box="[1265,1394,187,213]" pageId="9" pageNumber="452" value="2018-06-24">24.06.2018</collectingDate>
</date>
.
<collectorName id="26E15AC9FFCBF6484A7BFF481660A6CB" pageId="9" pageNumber="452">M. Hartmann</collectorName>
leg. (
<collectionCode id="ED05A7DAFFCBF6484E41FF2C16EDA6CB" box="[326,394,223,249]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="9" pageNumber="452">NME</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAB3F1FFFCBF6484FC0FEF115FAA773" blockId="9.[151,1436,150,393]" pageId="9" pageNumber="452">
<emphasis id="B960E30DFFCBF6484FC0FEF1159BA72E" bold="true" box="[199,764,258,285]" pageId="9" pageNumber="452">Published records. NAGORNO-KARABAKH:</emphasis>
<bibRefCitation id="EF8542EEFFCBF6484C03FEF01330A72F" author="Assing, V. &amp; Schulke, M." box="[772,1111,259,285]" pageId="9" pageNumber="452" pagination="91 - 173" refId="ref10354" refString="Assing, V. &amp; Schulke, M. (2019) The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera). Contributions to Entomology, 69 (1), 91 - 173. https: // doi. org / 10.21248 / contrib. entomol. 69.1.091 - 173" type="journal article" year="2019">Assing &amp; Schülke 2019: 100</bibRefCitation>
;
<emphasis id="B960E30DFFCBF6484B62FEF01383A72F" bold="true" box="[1125,1252,259,285]" pageId="9" pageNumber="452">
<collectingCountry id="F3037F8FFFCBF6484B62FEF013B8A72F" box="[1125,1247,259,285]" name="Turkey" pageId="9" pageNumber="452">TURKEY</collectingCountry>
:
</emphasis>
<bibRefCitation id="EF8542EEFFCBF6484BE9FEF012F0A72F" author="Korge, H." box="[1262,1431,258,285]" pageId="9" pageNumber="452" pagination="1 - 68" refId="ref10998" refString="Korge, H. (1971) Beitrage zur Kenntnis der Koleopterenfauna Kleinasiens. Annotationes Zoologicae et Botanicae, 67, 1 - 68." type="journal article" year="1971">Korge 1971: 5</bibRefCitation>
,
<bibRefCitation id="EF8542EEFFCBF6484F90FED41635A773" author="Zanetti, A." box="[151,338,294,321]" pageId="9" pageNumber="452" pagination="45 - 63" refId="ref11791" refString="Zanetti, A. (2002) Studies on Omalium Gravenhorst, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). Bollettino del Museo Civivo di Storia Naturale di Verona. Botanica Zoologia, 26, 45 - 63." type="journal article" year="2002">Zanetti 2002: 59</bibRefCitation>
, Anlaş 2019: 218, 2013: 106.
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFCBF6484FC0FEB913A5A7BB" pageId="9" pageNumber="452" type="distribution">
<paragraph id="8BAB3F1FFFCBF6484FC0FEB913A5A7BB" blockId="9.[151,1436,150,393]" pageId="9" pageNumber="452">
<emphasis id="B960E30DFFCBF6484FC0FEB91606A756" bold="true" box="[199,353,330,356]" pageId="9" pageNumber="452">Distribution.</emphasis>
<taxonomicName id="4C14449CFFCBF6484E6CFEB8155AA756" authorityName="Mulsant &amp; Rey" authorityYear="1880" box="[363,573,331,357]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="452" phylum="Arthropoda" rank="species" species="rugatum">
<emphasis id="B960E30DFFCBF6484E6CFEB8155AA756" box="[363,573,331,357]" italics="true" pageId="9" pageNumber="452">Omalium rugatum</emphasis>
</taxonomicName>
is widely distributed in the Western Palaearctic Region, and known from North Africa (
<collectingCountry id="F3037F8FFFCBF6484E33FE9C16E9A7BB" box="[308,398,367,393]" name="Algeria" pageId="9" pageNumber="452">Algeria</collectingCountry>
), Europe and Western Asia (
<collectingCountry id="F3037F8FFFCBF6484DCCFE9C1447A7BB" box="[715,800,367,393]" name="Cyprus" pageId="9" pageNumber="452">Cyprus</collectingCountry>
,
<collectingCountry id="F3037F8FFFCBF6484C2DFE9C141BA7BB" box="[810,892,367,393]" name="Turkey" pageId="9" pageNumber="452">Turkey</collectingCountry>
) (
<bibRefCitation id="EF8542EEFFCBF6484C95FE9C13D1A7BB" author="Schulke, M. &amp; Smetana, A." box="[914,1206,366,393]" pageId="9" pageNumber="452" refId="ref11577" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF6B6F97FFCBF6484F90F8AC1644A1D1" ID-DOI="http://doi.org/10.5281/zenodo.7518930" ID-Zenodo-Dep="7518930" httpUri="https://zenodo.org/record/7518930/files/figure.png" pageId="9" pageNumber="452" startId="9.[151,264,1887,1911]" targetBox="[178,1428,438,1853]" targetPageId="9">
<paragraph id="8BAB3F1FFFCBF6484F90F8AC1644A1D1" blockId="9.[151,1436,1887,2020]" pageId="9" pageNumber="452">
<emphasis id="B960E30DFFCBF6484F90F8AC1630A145" bold="true" box="[151,343,1887,1911]" pageId="9" pageNumber="452">FIGURES 19–26.</emphasis>
Aedeagus of
<taxonomicName id="4C14449CFFCBF6484EEEF8AC15F2A145" authorityName="Eppelsheim" authorityYear="1889" box="[489,661,1887,1911]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="452" phylum="Arthropoda" rank="species" species="falsum">
<emphasis id="B960E30DFFCBF6484EEEF8AC15F2A145" box="[489,661,1887,1911]" italics="true" pageId="9" pageNumber="452">Omalium falsum</emphasis>
</taxonomicName>
(Figs. 19–20: Georgia, Tskhneti),
<taxonomicName id="4C14449CFFCBF6484CF8F8931314A145" authorityName="Kraatz" authorityYear="1857" box="[1023,1139,1887,1911]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="452" phylum="Arthropoda" rank="species" species="littorale">
<emphasis id="B960E30DFFCBF6484CF8F8931376A145" box="[1023,1041,1888,1911]" italics="true" pageId="9" pageNumber="452">O</emphasis>
.
<emphasis id="B960E30DFFCBF6484B26F8AC1314A145" box="[1057,1139,1887,1911]" italics="true" pageId="9" pageNumber="452">littorale</emphasis>
</taxonomicName>
(Figs. 21–22: Russia, North Ossetia-Alania: Kabardino-Sunzhenskiy Mts.),
<taxonomicName id="4C14449CFFCBF6484D7CF877145EA1A9" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[635,825,1924,1947]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="riparium" subSpecies="impar">
<emphasis id="B960E30DFFCBF6484D7CF877145EA1A9" box="[635,825,1924,1947]" italics="true" pageId="9" pageNumber="452">O. riparium impar</emphasis>
</taxonomicName>
(Figs. 23–24: Russia, Kransnodar Territory: Sochi National Park),
<taxonomicName id="4C14449CFFCBF6484FDBF85B162AA18D" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[220,333,1959,1983]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="452" phylum="Arthropoda" rank="species" species="rivulare">
<emphasis id="B960E30DFFCBF6484FDBF85B1789A18D" box="[220,238,1960,1983]" italics="true" pageId="9" pageNumber="452">O</emphasis>
.
<emphasis id="B960E30DFFCBF6484FFBF854162AA18D" box="[252,333,1959,1983]" italics="true" pageId="9" pageNumber="452">rivulare</emphasis>
</taxonomicName>
(Figs. 25–26: Turkey, Artvin: Artvin): 19, 21, 23, 25—parameral view, 20, 22, 24, 26— lateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C30E6C94FFC8F64B4FC0FF6413A9A6CB" pageId="10" pageNumber="453" type="discussion">
<paragraph id="8BAB3F1FFFC8F64B4FC0FF641789A6E7" blockId="10.[151,1436,151,249]" pageId="10" pageNumber="453">
<emphasis id="B960E30DFFC8F64B4FC0FF64165CA683" bold="true" box="[199,315,151,177]" pageId="10" pageNumber="453">Remarks.</emphasis>
Habitus as in
<figureCitation id="132F239AFFC8F64B4ED9FF641545A683" box="[478,546,151,177]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1864,1888]" captionTargetBox="[234,1353,195,1823]" captionTargetId="figure-19@3.[220,1367,181,1840]" captionTargetPageId="3" captionText="FIGURES 1–10. Habitus of Omalium: 1—O. caesum (Turkey, Bursa: Uludağ), 2—O. cinnamomeum (Turkey, Antakya: Kişlak- Şenköy), 3—O. excavatum (Russia, Karachay-Cherkessia: Arkhyz), 4—O. falsum (Georgia, Tskhneti), 5—O. littorale (Russia, Adygea: Abago), 6—O. riparium impar (Russia, Krasnodar Territory: Khosta), 7—O. rivulare (Armenia, Alaverdi), 8—O. rugatum (Iran, Zagros Mts.), 9—O. turcicum (Russia, Adygea: Lagonaki), 10—O. wunderlei (Turkey, Gümüşhane: Erzincan- Kelkit). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518926" httpUri="https://zenodo.org/record/7518926/files/figure.png" pageId="10" pageNumber="453">Fig. 8</figureCitation>
. Aedeagus as in
<figureCitation id="132F239AFFC8F64B4DE7FF64140EA683" box="[736,873,151,177]" captionStart="FIGURES 27–32" captionStartId="10.[151,264,1903,1927]" captionTargetBox="[165,1422,281,1870]" captionTargetId="figure-62@10.[152,1436,273,1880]" captionTargetPageId="10" captionText="FIGURES 27–32. Aedeagus of Omalium rugatum (Figs. 27–28: Iran, Zagros), O. turcicum (Figs. 29–30: Turkey, Izmir: Bozdagh), O. wunderlei (Figs. 31–32: Turkey, Kars: Kars): 27, 29, 31—parameral view, 28, 30, 32—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7518932" httpUri="https://zenodo.org/record/7518932/files/figure.png" pageId="10" pageNumber="453">Figs. 27–28</figureCitation>
. Female genital segment (with spermatheca) as in
<figureCitation id="132F239AFFC8F64B4F90FF48178DA6E7" box="[151,234,187,213]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1503,1527]" captionTargetBox="[161,1431,318,1466]" captionTargetId="figure-33@11.[151,1435,304,1480]" captionTargetPageId="11" captionText="FIGURES 33–41. Female genital segment (ventral view) and spermatheca (shown by arrow in Fig. 40) of Omalium: 33—O. caesum (Russia, North Ossetia-Alania: Zincar), 34—O. cinnamomeum (Turkey, Antakya: Kişlak-Şenköy), 35—O. excavatum (Russia, Krasnodar Territory: Caucasus Nature Reserve), 36—O. falsum (Turkey, Rize: Ardeşen), 37—O. littorale (Russia, Volgograd Area: Elton Lake), 38—O. riparium impar (Georgia, Tskhneti), 39—O. rivulare (Russia, Krasnodar Territory: Derbiy), 40—O. rugatum (Russia, North Ossetia-Alania: Zincar), 41—O. turcicum (Russia, Krasnodar Territory, Caucasus Nature Reserve). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7518934" httpUri="https://zenodo.org/record/7518934/files/figure.png" pageId="10" pageNumber="453">Fig. 40</figureCitation>
.
</paragraph>
<paragraph id="8BAB3F1FFFC8F64B4FC0FF2C13A9A6CB" blockId="10.[151,1436,151,249]" box="[199,1230,223,249]" pageId="10" pageNumber="453">
It is here recorded from south-western regions of
<collectingCountry id="F3037F8FFFC8F64B4DF5FF2C1427A6CB" box="[754,832,223,249]" name="Russia" pageId="10" pageNumber="453">Russia</collectingCountry>
,
<collectingCountry id="F3037F8FFFC8F64B4C4CFF2C14C2A6CB" box="[843,933,223,249]" name="Georgia" pageId="10" pageNumber="453">Georgia</collectingCountry>
and
<collectingCountry id="F3037F8FFFC8F64B4CDCFF2C136FA6CB" box="[987,1032,223,249]" name="Iran" pageId="10" pageNumber="453">Iran</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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