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<document id="4E883588C68D0C74ED9F0DF8B27876D0" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BFDFF58AF89F94CFB713DB6" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera rufotestacea" docType="treatment" docVersion="8" lastPageNumber="158" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="157" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="A695D168BE45A4CDFA14818038F9CA3B">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="DB83CFF962E526BF87F8E1E6D2668CB3">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="8B340A34BABDDEC33EDEBB5DF23A44BD">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="FE00B5B180A0F451592B2615ED724F45">Bai, Ming</mods:namePart>
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<mods:namePart id="277FFCB3E78EB66D62DF5671B6E4597C">Yang, Xing-Ke</mods:namePart>
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<mods:namePart id="6E27DD780AA71CBC762E94D94089C622">Ahrens, Dirk</mods:namePart>
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<treatment id="03BD87E66BFDFF58AF89F94CFB713DB6" ID-DOI="http://doi.org/10.5281/zenodo.4496620" ID-GBIF-Taxon="177323528" ID-Zenodo-Dep="4496620" LSID="urn:lsid:plazi:treatment:03BD87E66BFDFF58AF89F94CFB713DB6" httpUri="http://treatment.plazi.org/id/03BD87E66BFDFF58AF89F94CFB713DB6" lastPageId="157" lastPageNumber="158" pageId="156" pageNumber="157">
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<taxonomicName id="4C144D736BFDFF59AF89F94CFD873C0C" authority="(Moser, 1915)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[151,601,294,321]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="156" pageNumber="157" phylum="Arthropoda" rank="species" species="rufotestacea">
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<emphasis id="B960EAE26BFDFF59AF89F94CFE763C0C" bold="true" box="[151,424,294,321]" italics="true" pageId="156" pageNumber="157">Maladera rufotestacea</emphasis>
(Moser, 1915)
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<figureCitation id="132F2A756BFDFF59AF89F921FED03C29" box="[151,270,331,357]" captionStart="FIGURE 34" captionStartId="154.[151,250,1943,1967]" captionTargetBox="[243,1352,190,1910]" captionTargetId="figure-19@154.[230,1358,181,1920]" captionTargetPageId="154" captionText="FIGURE 34. A–D: Maladera modestula (Brenske, 1902) (Myanmar: Putao); E–H: M. vethi (Moser, 1916) (China: Guangxi: Banbaxiang); I–L: M. rufotestacea (Moser, 1915) (Vietnam: Sa Pa). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496396" httpUri="https://zenodo.org/record/4496396/files/figure.png" pageId="156" pageNumber="157">Figures 34</figureCitation>
I–L, 104
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<taxonomicName id="4C144D736BFDFF59AF89F9E5FD9A3CEB" authority="Moser, 1915: 342" authorityName="Moser" authorityPageNumber="342" authorityYear="1915" box="[151,580,399,423]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="156" pageNumber="157" phylum="Arthropoda" rank="species" species="rufotestacea">
<emphasis id="B960EAE26BFDFF59AF89F9E5FE563CEB" box="[151,392,399,423]" italics="true" pageId="156" pageNumber="157">Autoserica rufotestacea</emphasis>
Moser, 1915: 342
</taxonomicName>
.
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<paragraph id="8BAB36F06BFDFF59AF89F9C4FB693C8A" blockId="156.[151,1207,399,454]" box="[151,1207,430,454]" pageId="156" pageNumber="157">
<treatmentCitationGroup id="AB0411DE6BFDFF59AF89F9C4FB693C8A" box="[151,1207,430,454]" pageId="156" pageNumber="157">
<treatmentCitation id="0AB510E16BFDFF59AF89F9C4FD8E3C8A" author="Ahrens, D." box="[151,592,430,454]" page="238" pageId="156" pageNumber="157" year="2006">
<taxonomicName id="4C144D736BFDFF59AF89F9C4FD8E3C8A" authority=": Ahrens 2006 b: 238" authorityName="Ahrens" authorityPageNumber="238" authorityYear="2006" box="[151,592,430,454]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="156" pageNumber="157" phylum="Arthropoda" rank="species" species="rufotestacea">
<emphasis id="B960EAE26BFDFF59AF89F9C4FEA03C8A" box="[151,382,430,454]" italics="true" pageId="156" pageNumber="157">Maladera rufotestacea</emphasis>
:
<bibRefCitation id="EF854B016BFDFF59AE95F9C4FD8E3C8A" author="Ahrens, D." box="[395,592,430,454]" pageId="156" pageNumber="157" refId="ref329946" refString="Ahrens, D. (2006 b) Sericinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248." type="book" year="2006">Ahrens 2006b: 238</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0AB510E16BFDFF59AD44F9C4FD1D3C8A" author="Ahrens, D." box="[602,707,430,454]" page="24" pageId="156" pageNumber="157" year="2007">
<bibRefCitation id="EF854B016BFDFF59AD44F9C4FD1D3C8A" author="Ahrens, D." box="[602,707,430,454]" pageId="156" pageNumber="157" pagination="1 - 51" refId="ref330155" refString="Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1" type="journal article" year="2007">2007b: 24</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16BFDFF59ADD0F9C4FC573C8A" author="Krajcik, M." box="[718,905,430,454]" page="155" pageId="156" pageNumber="157" year="2012">
<bibRefCitation id="EF854B016BFDFF59ADD0F9C4FC573C8A" author="Krajcik, M." box="[718,905,430,454]" pageId="156" pageNumber="157" pagination="1 - 278" refId="ref332946" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement, 5, 1 - 278." type="journal article" year="2012">Krajcik 2012: 155</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16BFDFF59AC8CF9C4FB6D3C8A" author="Ahrens, D. &amp; Fabrizi, S." box="[914,1203,430,454]" page="210" pageId="156" pageNumber="157" year="2016">
<bibRefCitation id="EF854B016BFDFF59AC8CF9C4FB6D3C8A" author="Ahrens, D. &amp; Fabrizi, S." box="[914,1203,430,454]" pageId="156" pageNumber="157" pagination="1 - 355" refId="ref330518" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi 2016: 210</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C30E657B6BFDFF59AF89F99EFC583F7F" pageId="156" pageNumber="157" type="materials_examined">
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<emphasis id="B960EAE26BFDFF59AF89F99EFE613F42" bold="true" box="[151,447,500,526]" pageId="156" pageNumber="157">
<typeStatus id="54AF88526BFDFF59AF89F99EFF0C3F42" box="[151,210,500,526]" pageId="156" pageNumber="157">Type</typeStatus>
material examined.
</emphasis>
See
<bibRefCitation id="EF854B016BFDFF59AEEAF99EFCD33F43" author="Ahrens, D. &amp; Fabrizi, S." box="[500,781,500,527]" pageId="156" pageNumber="157" pagination="1 - 355" refId="ref330518" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi (2016)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAB36F06BFDFF59AFD9FA72FC583F7F" blockId="156.[151,1437,500,2038]" box="[199,902,536,563]" pageId="156" pageNumber="157">
<emphasis id="B960EAE26BFDFF59AFD9FA72FDEA3F7E" bold="true" box="[199,564,536,562]" pageId="156" pageNumber="157">Additional material examined.</emphasis>
See
<bibRefCitation id="EF854B016BFDFF59AD77FA72FC5C3F7F" author="Ahrens, D. &amp; Fabrizi, S." box="[617,898,536,563]" pageId="156" pageNumber="157" pagination="1 - 355" refId="ref330518" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BFDFF58AFD9FA56FD443DFD" lastPageId="157" lastPageNumber="158" pageId="156" pageNumber="157" type="description">
<paragraph id="8BAB36F06BFDFF59AFD9FA56FE533FD2" blockId="156.[151,1437,500,2038]" pageId="156" pageNumber="157">
<emphasis id="B960EAE26BFDFF59AFD9FA56FEAB3F1A" bold="true" box="[199,373,572,598]" pageId="156" pageNumber="157">Redescription.</emphasis>
Length:
<quantity id="4CEC9B156BFDFF59AEFCFA56FDE03F1B" box="[482,574,572,599]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="156" pageNumber="157" unit="mm" value="8.6">8.6 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156BFDFF59AC0DFA56FCB13F1B" box="[787,879,572,599]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="156" pageNumber="157" unit="mm" value="6.1">6.1 mm</quantity>
, width:
<quantity id="4CEC9B156BFDFF59ACCFFA56FBF03F1B" box="[977,1070,572,599]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="156" pageNumber="157" unit="mm" value="4.7">4.7 mm</quantity>
. Body oblong, uniform reddish brown, dorsal surface dull, except setae of lateral margins of pronotum and elytra nearly glabrous, labroclypeus, tarsi, and tibiae shiny.
</paragraph>
<paragraph id="8BAB36F06BFDFF59AFD9FAC2FAF23EAE" blockId="156.[151,1437,500,2038]" pageId="156" pageNumber="157">
Labroclypeus wide and subtrapezoidal, widest at base; anterior angles strongly convex, anterior margin nearly straight medially; lateral margins slightly convex and convergent anteriorly, producing with ocular canthus a blunt angle, not incised at transition to labrum; margins strongly reflexed; surface shiny and flat, coarsely and very densely punctate, partly punctures fusing with each other, with a number of very large punctures each bearing a single erect seta; frontoclypeal suture finely incised, weakly curved; smooth area in front of eyes twice as wide as long; ocular canthus moderately long and wide, finely and densely punctate, with a single terminal seta. Frons dull, finely and sparsely punctate, near interior margin of eyes and behind lateral frontoclypeal suture with a few single long setae. Eyes large, ratio diameter/ interocular width: 0.77. Antenna with ten antennomeres, club with three antennomeres, 1.7 times as long as remaining antennomeres combined.
<collectingRegion id="49D0F8126BFDFF59AC0CFBA3FCAF3EAF" box="[786,881,969,995]" country="Myanmar" name="Chin" pageId="156" pageNumber="157">Mentum</collectingRegion>
convexly elevated, flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06BFDFF59AFD9FB87FBDE39DB" blockId="156.[151,1437,500,2038]" pageId="156" pageNumber="157">Pronotum widest at middle, lateral margins evenly convex and moderately convergent anteriorly and posteriorly, anterior angles moderately sharp and distinctly produced, posterior angles blunt; anterior margin with fine marginal line and convexly produced medially, basal margin without marginal line; surface finely and densely punctate, with microscopic setae in punctures, otherwise glabrous. Scutellum triangular, finely and moderately densely punctate, punctures less dense at middle, with microscopic setae in punctures.</paragraph>
<paragraph id="8BAB36F06BFDFF59AFD9FCCAFAD0386A" blockId="156.[151,1437,500,2038]" pageId="156" pageNumber="157">Elytra elongate, widest in posterior third; striae finely impressed and sparsely punctate, intervals convex, finely and sparsely punctate, punctures concentrated along striae and with a few single, short setae on penultimate lateral interval, otherwise glabrous; epipleural margin robust, ending at moderately rounded external apical angle of elytra; epipleura with long and dense setae; apex of elytra membraneous, with a wide rim of microtrichomes.</paragraph>
<paragraph id="8BAB36F06BFDFF59AFD9FD5AFEB93B6F" blockId="156.[151,1437,500,2038]" pageId="156" pageNumber="157">Ventral surface dull, with moderately dense and coarse punctures, sparsely setose, with a dense erect setae on meso-and metasternum, otherwise only with microscopic setae in punctures. Mesosternum between mesocoxae slightly wider than mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 2.0. Metacoxa with a few robust setae laterally. Abdominal sternites dull, with sparse and very fine punctation and a row of coarse punctures each bearing a robust seta; penultimate sternite with a wide shiny and chitinous rim. Pygidium dull, strongly convex, coarsely and densely punctate, without impunctate midline, apically with a few longer setae, otherwise only with microscopic setae.</paragraph>
<paragraph id="8BAB36F06BFDFF59AFD9FE47FBF83ABA" blockId="156.[151,1437,500,2038]" pageId="156" pageNumber="157">Legs moderately long and moderately wide; femora nearly impunctate, with two longitudinal rows of setae. Metafemur dull, very finely and superficially punctate, anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, moderately widened in apical half, posterior margin not serrated ventrally nor dorsally. Metatibia moderately wide and long, widest at middle, weakly narrowed basally, ratio width/ length: 1/ 3.46, sharply carinate dorsally, with two groups of spines, basal one shortly before middle, apical one at three quarters of metatibial length; lateral face longitudinally convex, with moderately dense and fine punctures basally and dorsally, otherwise nearly impunctate, punctures with microscopic setae only; ventral margin with four equidistant strong spines; medial face smooth and glabrous, apex very shallowly emarginate interiorly near tarsal articulation. Tarsomeres impunctate dorsally, with dense, fine setae ventrally, circular in cross-section; metatarsomeres impunctate dorsally, with a strongly serrated ventral carina, subventrally with a second, smooth longitudinal carina; metatarsomeres glabrous ventrally, first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin weakly convex. All claws symmetrical, feebly curved and long, with normally developed basal tooth.</paragraph>
<paragraph id="8BAB36F06BFCFF58AFD9F8FDFD443DFD" blockId="157.[151,1436,151,250]" box="[199,666,151,177]" pageId="157" pageNumber="158">
Aedeagus.
<figureCitation id="132F2A756BFCFF58AE5AF8FDFE4A3DFD" box="[324,404,151,177]" captionStart="FIGURE 34" captionStartId="154.[151,250,1943,1967]" captionTargetBox="[243,1352,190,1910]" captionTargetId="figure-19@154.[230,1358,181,1920]" captionTargetPageId="154" captionText="FIGURE 34. A–D: Maladera modestula (Brenske, 1902) (Myanmar: Putao); E–H: M. vethi (Moser, 1916) (China: Guangxi: Banbaxiang); I–L: M. rufotestacea (Moser, 1915) (Vietnam: Sa Pa). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496396" httpUri="https://zenodo.org/record/4496396/files/figure.png" pageId="157" pageNumber="158">Fig. 34</figureCitation>
I–K. Habitus:
<figureCitation id="132F2A756BFCFF58AD2CF8FDFD493DFD" box="[562,663,151,177]" captionStart="FIGURE 34" captionStartId="154.[151,250,1943,1967]" captionTargetBox="[243,1352,190,1910]" captionTargetId="figure-19@154.[230,1358,181,1920]" captionTargetPageId="154" captionText="FIGURE 34. A–D: Maladera modestula (Brenske, 1902) (Myanmar: Putao); E–H: M. vethi (Moser, 1916) (China: Guangxi: Banbaxiang); I–L: M. rufotestacea (Moser, 1915) (Vietnam: Sa Pa). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496396" httpUri="https://zenodo.org/record/4496396/files/figure.png" pageId="157" pageNumber="158">
Fig.
<specimenCount id="9D12FD796BFCFF58AD78F8FDFD493DFD" box="[614,663,151,177]" pageId="157" pageNumber="158" type="generic">34L</specimenCount>
</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BFCFF58AFD9F8D0FB713DB6" pageId="157" pageNumber="158" type="distribution">
<paragraph id="8BAB36F06BFCFF58AFD9F8D0FB713DB6" blockId="157.[151,1436,151,250]" pageId="157" pageNumber="158">
<emphasis id="B960EAE26BFCFF58AFD9F8D0FEBF3D98" bold="true" box="[199,353,186,212]" pageId="157" pageNumber="158">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BFCFF58AEC9F8D1FDE93D99" box="[471,567,187,213]" captionStart="FIGURE 104" captionStartId="353.[151,250,1974,1998]" captionTargetBox="[220,1366,1143,1944]" captionTargetId="figure-63@353.[214,1374,1139,1950]" captionTargetPageId="353" captionText="FIGURE 104. Distribution of the Maladera modestula group: Maladera jinghongensis sp. n., M. modestula (Brenske, 1902), M. rufotestacea (Moser, 1915), M. taiwana Nomura, 1974, M. vethi (Moser, 1916)." figureDoi="http://doi.org/10.5281/zenodo.4496600" httpUri="https://zenodo.org/record/4496600/files/figure.png" pageId="157" pageNumber="158">Fig. 104</figureCitation>
) and
<tableCitation id="C696034B6BFCFF58AD6BF8D1FD173D99" box="[629,713,187,213]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="157" pageNumber="158">Table 1</tableCitation>
. The species was originally described from
<collectingRegion id="49D0F8126BFCFF58ABA6F8D1FAD03D99" box="[1208,1294,187,213]" country="China" name="Yunnan" pageId="157" pageNumber="158">Yunnan</collectingRegion>
(
<collectingCountry id="F30376606BFCFF58AA03F8D1FABB3D9A" box="[1309,1381,187,214]" name="China" pageId="157" pageNumber="158">China</collectingCountry>
), but was recorded also from NE
<collectingCountry id="F30376606BFCFF58AECFF8B5FDD33DB5" box="[465,525,223,249]" name="India" pageId="157" pageNumber="158">India</collectingCountry>
,
<collectingCountry id="F30376606BFCFF58AD06F8B5FDA43DB5" box="[536,634,223,249]" name="Vietnam" pageId="157" pageNumber="158">Vietnam</collectingCountry>
,
<collectingCountry id="F30376606BFCFF58AD9AF8B5FD363DB5" box="[644,744,223,249]" name="Thailand" pageId="157" pageNumber="158">Thailand</collectingCountry>
and
<collectingCountry id="F30376606BFCFF58AC00F8B5FC553DB5" box="[798,907,223,249]" name="Myanmar" pageId="157" pageNumber="158">Myanmar</collectingCountry>
(
<bibRefCitation id="EF854B016BFCFF58AC87F8B5FB7D3DB6" author="Ahrens, D. &amp; Fabrizi, S." box="[921,1187,223,250]" pageId="157" pageNumber="158" pagination="1 - 355" refId="ref330518" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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