256 lines
30 KiB
XML
256 lines
30 KiB
XML
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<mods:title id="7ACCC1A8985270F14221FD67EE6698BD">A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="3F222CB66A829054614F3CEBFD6BB3C3">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="2EC79AA4EFE005D2F702799BD546CABF">Ahrens, Dirk</mods:namePart>
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<mods:title id="184D98E74B902F2D1E231D2DE564E49B">Zootaxa</mods:title>
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<taxonomicName id="4C144D736BD9FF7DAF89FB0CFD333ECC" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[151,749,870,897]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="shiruguanensis" status="sp. nov.">
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<emphasis id="B960EAE26BD9FF7DAF89FB0CFE103ECD" bold="true" box="[151,462,870,897]" italics="true" pageId="184" pageNumber="185">Maladera shiruguanensis</emphasis>
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Ahrens, Fabrizi & Liu
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</taxonomicName>
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,
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</emphasis>
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</paragraph>
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<figureCitation id="132F2A756BD9FF7DAF89FBE1FED33EE9" box="[151,269,907,933]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="184" pageNumber="185">Figures 41</figureCitation>
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I–L, 109
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</paragraph>
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<emphasis id="B960EAE26BD9FF7DAF89FBB9FE633EA0" bold="true" box="[151,445,978,1005]" pageId="184" pageNumber="185">Type material examined.</emphasis>
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:
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, W
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/ E Chongquing,
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,
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pass ~
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NW Taiyanghe
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traps,
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leg., ~
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/ 737
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Asia spec.” (
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<collectionCode id="ED05AE356BD9FF7DABB0FB9DFADC395D" box="[1198,1282,1015,1041]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="184" pageNumber="185" type="Museum">ZFMK</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B7C3CAD6BD9FF7DAA0AFB9DFB573979" ID-GBIF-Occurrence="3034555785" collectionCode="USNM" country="China" county="Aug" elevation="763" location="Shin Kai Si Mt." municipality="Mt. Omei" pageId="184" pageNumber="185" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="54AF88526BD9FF7DAA0AFB9DFA5B395D" box="[1300,1413,1015,1041]" pageId="184" pageNumber="185" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D12FD796BD9FF7DAA90FB9DFF723979" pageId="184" pageNumber="185" type="male">1 ♂</specimenCount>
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“Szechuen
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<collectingCountry id="F30376606BD9FF7DAE2EFC71FEAD3979" box="[304,371,1051,1077]" name="China" pageId="184" pageNumber="185">China</collectingCountry>
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DC Graham/
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<location id="8ECB602B6BD9FF7DAD11FC71FD633979" LSID="urn:lsid:plazi:treatment:03BD87E66BD9FF7CAF89FB0CFD103E59:8ECB602B6BD9FF7DAD11FC71FD633979" box="[527,701,1051,1077]" country="China" county="Aug" municipality="Mt. Omei" name="Shin Kai Si Mt." pageId="184" pageNumber="185">
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Shin Kai Si
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<collectingMunicipality id="6BCFAC8A6BD9FF7DAD8AFC71FD633979" box="[660,701,1051,1077]" pageId="184" pageNumber="185">Mt.</collectingMunicipality>
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</location>
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Omei
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<collectingCounty id="62CA4E7C6BD9FF7DAC95FC71FC653979" box="[907,955,1051,1077]" pageId="184" pageNumber="185">Aug</collectingCounty>
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16, ‘33” (
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<collectionCode id="ED05AE356BD9FF7DAB34FC71FB5F3979" box="[1066,1153,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="184" pageNumber="185" type="Museum">USNM</collectionCode>
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)
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</materialsCitation>
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,
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||
<materialsCitation id="3B7C3CAD6BD9FF7DAB8DFC71FCFC3915" ID-GBIF-Occurrence="3034556267" collectionCode="ZFMK" collectorName="G. M. Franck Collector" country="China" location="Mt. Omei Kiating" pageId="184" pageNumber="185" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount id="9D12FD796BD9FF7DAB8DFC71FB623979" box="[1171,1212,1051,1077]" pageId="184" pageNumber="185" type="male">1 ♂</specimenCount>
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“
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<location id="8ECB602B6BD9FF7DABD3FC71FA463979" LSID="urn:lsid:plazi:treatment:03BD87E66BD9FF7CAF89FB0CFD103E59:8ECB602B6BD9FF7DABD3FC71FA463979" box="[1229,1432,1051,1077]" country="China" name="Mt. Omei Kiating" pageId="184" pageNumber="185">Mt. Omei Kiating</location>
|
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, Szechuan
|
||
<collectingCountry id="F30376606BD9FF7DAE17FC55FE923915" box="[265,332,1087,1113]" name="China" pageId="184" pageNumber="185">China</collectingCountry>
|
||
VII.32/
|
||
<collectorName id="26E153266BD9FF7DAEB5FC55FD733915" box="[427,685,1087,1113]" pageId="184" pageNumber="185">G.M. Franck Collector</collectorName>
|
||
” (
|
||
<collectionCode id="ED05AE356BD9FF7DADD8FC55FCC43915" box="[710,794,1087,1113]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="184" pageNumber="185" type="Museum">ZFMK</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6BD9FF7CAFD9FC08FCB53CB9" lastPageId="185" lastPageNumber="186" pageId="184" pageNumber="185" type="description">
|
||
<paragraph id="8BAB36F06BD9FF7DAFD9FC08FB5939ED" blockId="184.[151,1437,978,2013]" pageId="184" pageNumber="185">
|
||
<emphasis id="B960EAE26BD9FF7DAFD9FC08FE873930" bold="true" box="[199,345,1122,1148]" pageId="184" pageNumber="185">Description.</emphasis>
|
||
Length:
|
||
<quantity id="4CEC9B156BD9FF7DAEA2FC09FDFE3931" box="[444,544,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="184" pageNumber="185" unit="mm" value="11.3">11.3 mm</quantity>
|
||
, length of elytra:
|
||
<quantity id="4CEC9B156BD9FF7DADFDFC09FCE43931" box="[739,826,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="184" pageNumber="185" unit="mm" value="8.8">8.8 mm</quantity>
|
||
, width:
|
||
<quantity id="4CEC9B156BD9FF7DAC8CFC09FC353931" box="[914,1003,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="184" pageNumber="185" unit="mm" value="7.2">7.2 mm</quantity>
|
||
. Body wide, oval, dark brown, antenna yellowish brown, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BAB36F06BD9FF7DAFD9FCC1FC2738A9" blockId="184.[151,1437,978,2013]" pageId="184" pageNumber="185">
|
||
Labroclypeus wide, lateral margins convex and moderately convergent, lateral margins produce with ocular canthus an indistinct angle, anterior angles strongly convex, anterior margin shallowly emarginate medially, margins weakly reflexed; surface moderately shiny, at base widely dull, surface nearly flat, finely and densely punctate, distance between punctures smaller their diameter, with numerous long erect setae; smooth area in front of eyes three times as wide as long; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and short (1/3 of ocular diameter), finely and densely punctate, with a short terminal seta. Eyes moderately large, ratio diameter/ interocular width: 0.6. Frons with fine and moderately dense punctures, with a few single setae beside eyes.Antenna with ten antennomeres, club with three antennomeres and straight, little shorter than remaining antennomeres combined.
|
||
<collectingRegion id="49D0F8126BD9FF7DAEA8FDA1FDCB38A9" box="[438,533,1483,1509]" country="Myanmar" name="Chin" pageId="184" pageNumber="185">Mentum</collectingRegion>
|
||
convexly elevated and flattened anteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BAB36F06BD9FF7DAFD9FD85FABC3B39" blockId="184.[151,1437,978,2013]" pageId="184" pageNumber="185">Pronotum widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced; anterior margin convex, with complete marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only, setae of lateral and anterior margin sparse; hypomeron carinate. Scutellum wide, triangular, punctation as in pronotum, less densely punctate on midline.</paragraph>
|
||
<paragraph id="8BAB36F06BD9FF7DAFD9FE15FED33A49" blockId="184.[151,1437,978,2013]" pageId="184" pageNumber="185">Elytra convex, widest at posterior third, striae finely impressed, finely punctate, intervals flat, finely and densely punctate, with minute setae in punctures, odd intervals with a few erect short setae; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes.</paragraph>
|
||
<paragraph id="8BAB36F06BD9FF7DAFD9FF65FED13AF5" blockId="184.[151,1437,978,2013]" pageId="184" pageNumber="185">Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae wide, as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.5. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium strongly convex, coarsely and densely punctate, on apical margin with a few long setae.</paragraph>
|
||
<paragraph id="8BAB36F06BD9FF7CAFD9FFA9FAAA3C9D" blockId="184.[151,1437,978,2013]" lastBlockId="185.[151,1437,151,789]" lastPageId="185" lastPageNumber="186" pageId="184" pageNumber="185">Legs moderately long and wide; femora superficially and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae absent; posterior ventral margin almost straight, not widened in apical half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia moderately long and wide, flattened, ratio width/length: 1/3.8, sharply carinate dorsally, with two groups of spines, basal one at basal third, apical one at two thirds of metatibial length, basally with a few robust and short setae; lateral face weakly longitudinally convex, with a very superficial fine punctures; medial face impunctate, apex shallowly and concavely emarginate interiorly near tarsal articulation. Tarsomeres sparsely finely punctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate. Anterior claws symmetrical.</paragraph>
|
||
<paragraph id="8BAB36F06BD8FF7CAFD9F9B1FCB53CB9" blockId="185.[151,1437,151,789]" box="[199,875,475,501]" pageId="185" pageNumber="186">
|
||
Aedeagus:
|
||
<figureCitation id="132F2A756BD8FF7CAE5AF9B1FE4A3CB9" box="[324,404,475,501]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="185" pageNumber="186">Fig. 41</figureCitation>
|
||
I–K. Habitus:
|
||
<figureCitation id="132F2A756BD8FF7CAD2DF9B1FD493CB9" box="[563,663,475,501]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="185" pageNumber="186">
|
||
Fig.
|
||
<specimenCount id="9D12FD796BD8FF7CAD79F9B1FD493CB9" box="[615,663,475,501]" pageId="185" pageNumber="186" type="generic">41L</specimenCount>
|
||
</figureCitation>
|
||
. Female unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6BD8FF7CAFD9F994FA423FC9" pageId="185" pageNumber="186" type="diagnosis">
|
||
<paragraph id="8BAB36F06BD8FF7CAFD9F994FA423FC9" blockId="185.[151,1437,151,789]" pageId="185" pageNumber="186">
|
||
<emphasis id="B960EAE26BD8FF7CAFD9F994FE9C3F54" bold="true" box="[199,322,510,536]" pageId="185" pageNumber="186">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C144D736BD8FF7CAE52F995FC5C3F55" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[332,898,511,537]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="shiruguanensis" status="sp. nov.">
|
||
<emphasis id="B960EAE26BD8FF7CAE52F995FDAE3F55" box="[332,624,511,537]" italics="true" pageId="185" pageNumber="186">Maladera shiruguanensis</emphasis>
|
||
Ahrens, Fabrizi & Liu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25357996BD8FF7CAC8FF995FC0F3F55" box="[913,977,511,537]" pageId="185" pageNumber="186" rank="species">sp. n.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C144D736BD8FF7CAB6FF995FEE63F71" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="guanxianensis" status="sp. nov.">
|
||
<emphasis id="B960EAE26BD8FF7CAB6FF995FAE53F54" box="[1137,1339,511,536]" italics="true" pageId="185" pageNumber="186">M. guanxianensis</emphasis>
|
||
Ahrens, Fabrizi & Liu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25357996BD8FF7CAE5BFA49FE5C3F71" box="[325,386,547,573]" pageId="185" pageNumber="186" rank="species">sp. n.</taxonomicNameLabel>
|
||
in having the dorsal process of the phallobase abruptly narrowed, as well as by the distinctive shape of the parameres: the left paramere is reflexed around its own axis, narrowed apically in lateral view, and slightly widened before apex (dorsal view), rather than being narrowed at apex (as in
|
||
<taxonomicName id="4C144D736BD8FF7CABDCFA01FA523FC8" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[1218,1420,619,644]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="guanxianensis">
|
||
<emphasis id="B960EAE26BD8FF7CABDCFA01FA523FC8" box="[1218,1420,619,644]" italics="true" pageId="185" pageNumber="186">M. guanxianensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6BD8FF7CAFD9FAE5FF1B3F82" pageId="185" pageNumber="186" type="etymology">
|
||
<paragraph id="8BAB36F06BD8FF7CAFD9FAE5FF1B3F82" blockId="185.[151,1437,151,789]" pageId="185" pageNumber="186">
|
||
<emphasis id="B960EAE26BD8FF7CAFD9FAE5FE903FE5" bold="true" box="[199,334,655,681]" pageId="185" pageNumber="186">Etymology.</emphasis>
|
||
The new species is named after the
|
||
<typeStatus id="54AF88526BD8FF7CADEBFAE5FCFB3FE5" box="[757,805,655,681]" pageId="185" pageNumber="186">type</typeStatus>
|
||
locality, Shiruguan (adjective in the nominative singular).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6BD8FF7CAFD9FABCFBA03FBD" box="[199,1150,726,753]" pageId="185" pageNumber="186" type="description">
|
||
<paragraph id="8BAB36F06BD8FF7CAFD9FABCFBA03FBD" blockId="185.[151,1437,151,789]" box="[199,1150,726,753]" pageId="185" pageNumber="186">
|
||
<emphasis id="B960EAE26BD8FF7CAFD9FABCFEE13FBC" bold="true" box="[199,319,726,752]" pageId="185" pageNumber="186">Variation.</emphasis>
|
||
Length:
|
||
<quantity id="4CEC9B156BD8FF7CAEBAFABDFD963FBD" box="[420,584,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.185" metricValueMax="1.24" metricValueMin="1.13" pageId="185" pageNumber="186" unit="mm" value="11.85" valueMax="12.4" valueMin="11.3">11.3–12.4 mm</quantity>
|
||
, length of elytra:
|
||
<quantity id="4CEC9B156BD8FF7CAC13FABCFC483FBD" box="[781,918,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="9.15" metricValueMax="9.5" metricValueMin="8.8" pageId="185" pageNumber="186" unit="mm" value="9.15" valueMax="9.5" valueMin="8.8">8.8–9.5 mm</quantity>
|
||
, width: 7.0–
|
||
<quantity id="4CEC9B156BD8FF7CAB3CFABDFBA43FBD" box="[1058,1146,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="185" pageNumber="186" unit="mm" value="7.2">7.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6BD8FF7CAFD9FA90FD103E59" box="[199,718,762,789]" pageId="185" pageNumber="186" type="distribution">
|
||
<paragraph id="8BAB36F06BD8FF7CAFD9FA90FD103E59" blockId="185.[151,1437,151,789]" box="[199,718,762,789]" pageId="185" pageNumber="186">
|
||
<emphasis id="B960EAE26BD8FF7CAFD9FA90FEBF3E58" bold="true" box="[199,353,762,788]" pageId="185" pageNumber="186">Distribution.</emphasis>
|
||
See map (
|
||
<figureCitation id="132F2A756BD8FF7CAEC8FA91FDE83E59" box="[470,566,763,789]" captionStart="FIGURE 109" captionStartId="356.[151,250,1044,1068]" captionTargetBox="[203,1383,189,1012]" captionTargetId="figure-19@356.[196,1391,181,1020]" captionTargetPageId="356" captionText="FIGURE 109. Distribution of the Maladera granuligera group: Maladera maguanensis sp. n., M. mupingensis sp. n., M. shiruguanensis sp. n., M. siniaevi Ahrens, 2004." figureDoi="http://doi.org/10.5281/zenodo.4496614" httpUri="https://zenodo.org/record/4496614/files/figure.png" pageId="185" pageNumber="186">Fig. 109</figureCitation>
|
||
) and
|
||
<tableCitation id="C696034B6BD8FF7CAD6AFA91FD163E59" box="[628,712,763,789]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant & Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="185" pageNumber="186">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |