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<mods:title id="97E6C27765C18AA6052FA33788646E2F">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<emphasis id="B960EAE26B49FFEDAF89FB71FED43E7F" box="[151,266,795,819]" italics="true" pageId="40" pageNumber="41">Omaladera</emphasis>
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<emphasis id="B960EAE26B49FFEDAC7BFB71FBFE3E7F" box="[869,1056,795,819]" italics="true" pageId="40" pageNumber="41">Amaladera diffinis</emphasis>
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<bibRefCitation id="EF854B016B49FFEDABACFB71FA9A3E7F" author="Ahrens, D." box="[1202,1348,795,819]" pageId="40" pageNumber="41" pagination="31 - 35" refId="ref329765" refString="Ahrens, D. (2004 b) Revisional notes on Sericini: The taxonomic status of the &quot; Neotropical &quot; genus Rhynchosymmela Frey, 1974. Beitrage zur Entomologie, 53, 31 - 35." type="journal article" year="2004">Ahrens 2004b</bibRefCitation>
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<bibRefCitation id="EF854B016B49FFEDAE59FB50FE7F3E1E" author="Ahrens, D." box="[327,417,826,850]" pageId="40" pageNumber="41" pagination="1 - 16" refId="ref330070" refString="Ahrens, D. (2006 d) Phylogenetic analysis of Anomalophylla Reitter, 1887 (Coleoptera, Scarabaeidae: Sericini). Insect Systematics and Evolution, 37, 1 - 16. https: // doi. org / 10.1163 / 187631206788831533" type="journal article" year="2006">2006d: 1</bibRefCitation>
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;
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<bibRefCitation id="EF854B016B49FFEDAEB0FB50FDCB3E1E" author="Ahrens, D." box="[430,533,826,850]" pageId="40" pageNumber="41" pagination="450 - 476" refId="ref330203" refString="Ahrens, D. (2007 c): Beetle evolution in the Asian highlands: insight from a phylogeny of the scarabaeid subgenus Serica (Coleoptera, Scarabaeidae). Systematic Entomology, 32, 450 - 476. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00373. x" type="journal article" year="2007">2007c: 21</bibRefCitation>
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<taxonomicName id="4C144D736B49FFEDAF89FB32FE303E3D" authority="Brenske, 1900: 79" authorityName="Brenske" authorityPageNumber="79" authorityYear="1900" box="[151,494,856,881]" class="Insecta" family="Melolonthidae" genus="Cephaloserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B49FFEDAF89FB32FEF53E3C" box="[151,299,856,880]" italics="true" pageId="40" pageNumber="41">Cephaloserica</emphasis>
<bibRefCitation id="EF854B016B49FFEDAE2DFB32FE303E3D" author="Brenske, E." box="[307,494,856,881]" pageId="40" pageNumber="41" pagination="39 - 96" refId="ref331495" refString="Brenske, E. (1900) Die Serica - Arten der Erde. IV. Berliner Entomologische Zeitschrift, 45, 39 - 96. https: // doi. org / 10.1002 / mmnd. 47919000105" type="journal article" year="1900">Brenske, 1900: 79</bibRefCitation>
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(
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species by subsequent designation:
<taxonomicName id="4C144D736B49FFEDACBEFB33FAE73E3C" authority="Brenske, 1896: 153" authorityName="Brenske" authorityPageNumber="153" authorityYear="1896" box="[928,1337,856,880]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="carinirostris">
<emphasis id="B960EAE26B49FFEDACBEFB33FBB83E3C" box="[928,1126,857,880]" italics="true" pageId="40" pageNumber="41">Serica carinirostris</emphasis>
Brenske, 1896: 153
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;
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Yu
<emphasis id="B960EAE26B49FFEDAA76FB33FA433E3C" box="[1384,1437,856,880]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
1998
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);
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, 62,
<emphasis id="B960EAE26B49FFEDAEF6FB1DFDF03EC3" bold="true" box="[488,558,887,911]" pageId="40" pageNumber="41">syn. n.</emphasis>
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<taxonomicName id="4C144D736B49FFEDAF89FBFCFE0B3EE2" authority="Brenske, 1902: 2" authorityName="Brenske" authorityPageNumber="2" authorityYear="1902" box="[151,469,918,942]" class="Insecta" family="Melolonthidae" genus="Coronoserica" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B49FFEDAF89FBFCFEFF3EE1" box="[151,289,918,941]" italics="true" pageId="40" pageNumber="41">Coronoserica</emphasis>
<bibRefCitation id="EF854B016B49FFEDAE36FBFCFE0B3EE2" author="Brenske, E." box="[296,469,918,942]" pageId="40" pageNumber="41" pagination="1 - 70" refId="ref331533" refString="Brenske, E. (1902) Die Serica - Arten der Erde. VII. Berliner Entomologische Zeitschrift, 47, 1 - 70." type="journal article" year="1902">Brenske, 1902: 2</bibRefCitation>
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(
<typeStatus id="54AF88526B49FFEDAEFCFBFCFDD13EE2" box="[482,527,918,942]" pageId="40" pageNumber="41">type</typeStatus>
species by monotypy:
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<emphasis id="B960EAE26B49FFEDADE6FBFCFC903EE2" box="[760,846,918,942]" italics="true" pageId="40" pageNumber="41">C. beata</emphasis>
<bibRefCitation id="EF854B016B49FFEDAC4AFBFCFC343EE2" author="Brenske, E." box="[852,1002,918,942]" pageId="40" pageNumber="41" pagination="1 - 70" refId="ref331533" refString="Brenske, E. (1902) Die Serica - Arten der Erde. VII. Berliner Entomologische Zeitschrift, 47, 1 - 70." type="journal article" year="1902">Brenske, 1902</bibRefCitation>
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),
<emphasis id="B960EAE26B49FFEDACE2FBFCFB9C3EE2" bold="true" box="[1020,1090,918,942]" pageId="40" pageNumber="41">syn. n.</emphasis>
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</subSubSection>
<subSubSection id="C30E657B6B49FFEDAF89FBB6FE8839CA" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BAB36F06B49FFEDAF89FBB6FE8839CA" blockId="40.[151,1437,988,1158]" pageId="40" pageNumber="41">
<emphasis id="B960EAE26B49FFEDAF89FBB6FED53EBA" bold="true" box="[151,267,988,1014]" pageId="40" pageNumber="41">Remarks.</emphasis>
According to the currently available phylogenetic hypothesis (
<bibRefCitation id="EF854B016B49FFEDACCEFBB6FB413EBA" author="Eberle, J. &amp; Fabrizi, S. &amp; Lago P. &amp; Ahrens, D." box="[976,1183,988,1014]" pageId="40" pageNumber="41" pagination="183 - 197" refId="ref331753" refString="Eberle, J., Fabrizi, S., Lago P. &amp; Ahrens, D. (2017) A historical biogeography of megadiverse Sericini-another story &quot; out of Africa &quot;? Cladistics, 33, 183 - 197. https: // doi. org / 10.1111 / cla. 12162" type="journal article" year="2017">
Eberle
<emphasis id="B960EAE26B49FFEDAB3CFBB7FB853EBA" box="[1058,1115,988,1014]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
2017
</bibRefCitation>
),
<taxonomicName id="4C144D736B49FFEDABADFBB6FAED3EBA" authorityName="Reitter" authorityYear="1896" box="[1203,1331,988,1014]" class="Insecta" family="Melolonthidae" genus="Amaladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="349" phylum="Arthropoda" rank="subGenus" subGenus="Omaladera">
<emphasis id="B960EAE26B49FFEDABADFBB6FAED3EBA" box="[1203,1331,988,1014]" italics="true" pageId="40" pageNumber="41">Omaladera</emphasis>
</taxonomicName>
is nested within
<taxonomicName id="4C144D736B49FFEDAFF7FC6AFE523956" authorityName="Brenske" authorityYear="1900" box="[233,396,1024,1050]" class="Insecta" family="Melolonthidae" genus="Cephaloserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B49FFEDAFF7FC6AFE523956" box="[233,396,1024,1050]" italics="true" pageId="40" pageNumber="41">Cephaloserica</emphasis>
</taxonomicName>
species. The proposed key character of
<taxonomicName id="4C144D736B49FFEDAC78FC6AFBD73956" authorityName="Brenske" authorityYear="1900" box="[870,1033,1024,1050]" class="Insecta" family="Melolonthidae" genus="Cephaloserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B49FFEDAC78FC6AFBD73956" box="[870,1033,1024,1050]" italics="true" pageId="40" pageNumber="41">Cephaloserica</emphasis>
</taxonomicName>
,
<emphasis id="B960EAE26B49FFEDAB05FC6BFB9D3956" box="[1051,1091,1025,1050]" italics="true" pageId="40" pageNumber="41">e.g.</emphasis>
the reduced size of the basal apodeme of the phallobase and the elongated tubular phallobase, appear to be either plesiomorphic or had multiple origins in this clade. Thus,
<taxonomicName id="4C144D736B49FFEDAEDFFC22FD9F392E" authorityName="Reitter" authorityYear="1896" box="[449,577,1096,1122]" class="Insecta" family="Melolonthidae" genus="Amaladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="349" phylum="Arthropoda" rank="subGenus" subGenus="Omaladera">
<emphasis id="B960EAE26B49FFEDAEDFFC22FD9F392E" box="[449,577,1096,1122]" italics="true" pageId="40" pageNumber="41">Omaladera</emphasis>
</taxonomicName>
renders
<taxonomicName id="4C144D736B49FFEDAD80FC22FC9F392E" authorityName="Brenske" authorityYear="1900" box="[670,833,1096,1122]" class="Insecta" family="Melolonthidae" genus="Cephaloserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B49FFEDAD80FC22FC9F392E" box="[670,833,1096,1122]" italics="true" pageId="40" pageNumber="41">Cephaloserica</emphasis>
</taxonomicName>
polyphyletic, and we synonymize
<taxonomicName id="4C144D736B49FFEDABDFFC22FABA392E" authorityName="Brenske" authorityYear="1900" box="[1217,1380,1096,1122]" class="Insecta" family="Melolonthidae" genus="Cephaloserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B49FFEDABDFFC22FABA392E" box="[1217,1380,1096,1122]" italics="true" pageId="40" pageNumber="41">Cephaloserica</emphasis>
</taxonomicName>
with
<taxonomicName id="4C144D736B49FFEDAF89FC06FEC939CA" authorityName="Reitter" authorityYear="1896" box="[151,279,1132,1158]" class="Insecta" family="Melolonthidae" genus="Amaladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="349" phylum="Arthropoda" rank="subGenus" subGenus="Omaladera">
<emphasis id="B960EAE26B49FFEDAF89FC06FEC939CA" box="[151,279,1132,1158]" italics="true" pageId="40" pageNumber="41">Omaladera</emphasis>
</taxonomicName>
here.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B49FFECAF89FCB2FA423B8C" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="key">
<paragraph id="8BAB36F06B49FFEDAF89FCB2FBDB39B9" blockId="40.[151,1029,1239,1269]" box="[151,1029,1239,1269]" pageId="40" pageNumber="41">
<emphasis id="B960EAE26B49FFEDAF89FCB2FBDB39B9" bold="true" box="[151,1029,1239,1269]" pageId="40" pageNumber="41">
Key to species of
<taxonomicName id="4C144D736B49FFEDAE70FCBDFE3D39BE" authorityName="Mulsant &amp; Rey" authorityYear="1871" box="[366,483,1239,1266]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B49FFEDAE70FCBDFE3D39BE" bold="true" box="[366,483,1239,1266]" italics="true" pageId="40" pageNumber="41">Maladera</emphasis>
</taxonomicName>
(sbg.
<taxonomicName id="4C144D736B49FFEDAD30FCBDFD6B39BE" authorityName="Reitter" authorityYear="1896" box="[558,693,1239,1266]" class="Insecta" family="Melolonthidae" genus="Amaladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="349" phylum="Arthropoda" rank="subGenus" subGenus="Omaladera">
<emphasis id="B960EAE26B49FFEDAD30FCBDFD6B39BE" bold="true" box="[558,693,1239,1266]" italics="true" pageId="40" pageNumber="41">Omaladera</emphasis>
</taxonomicName>
) occurring in
<collectingCountry id="F30376606B49FFEDAC79FCB2FC6A39BE" box="[871,948,1240,1266]" name="China" pageId="40" pageNumber="41">China</collectingCountry>
(³³):
</emphasis>
</paragraph>
<key id="FF8561C16B49FFECAF89FD77FA423B8C" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<keyStep id="30E02E556B49FFEDAF89FD77FA423803" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FD77FA42387E" blockId="40.[151,1437,1309,2035]" box="[151,1436,1309,1331]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FD77FA42387E" box="[151,1436,1309,1331]" pageId="40" pageNumber="41">1 Basal apodeme of phallobase shortened, equal to about 1/5 of phallobase length. Phallobase tube-like................. 17</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FD50FA423803" blockId="40.[151,1437,1309,2035]" box="[151,1436,1338,1360]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FD50FA423803" box="[151,1436,1338,1360]" pageId="40" pageNumber="41">1 Basal apodeme of phallobase not shortened, equal to about half of phallobase length. Phallobase not tube-like............ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FD32FA4238C6" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FD32FA423821" blockId="40.[151,1437,1309,2035]" box="[151,1436,1368,1390]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FD32FA423821" box="[151,1436,1368,1390]" pageId="40" pageNumber="41">2 Base of labroclypeus narrowly covered with dull toment..................................................... 10</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FD1FFA4238C6" blockId="40.[151,1437,1309,2035]" box="[151,1436,1397,1419]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FD1FFA4238C6" box="[151,1436,1397,1419]" pageId="40" pageNumber="41">2 Base of labroclypeus without dull toment................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FDF8FA423889" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FDF8FA4238EB" blockId="40.[151,1437,1309,2035]" box="[151,1436,1426,1448]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FDF8FA4238EB" box="[151,1436,1426,1448]" pageId="40" pageNumber="41">3 Anterior margin of metafemur with a fine, adjacent and continuously serrate line................................... 4</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FDDAFA423889" blockId="40.[151,1437,1309,2035]" box="[151,1436,1456,1478]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FDDAFA423889" box="[151,1436,1456,1478]" pageId="40" pageNumber="41">3 Anterior margin of metafemur without fine, adjacent serrate line................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FDA7FA423B4C" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FDA7FA4238AF" blockId="40.[151,1437,1309,2035]" box="[151,1436,1485,1507]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FDA7FA4238AF" box="[151,1436,1485,1507]" pageId="40" pageNumber="41">
4 Right paramere weakly bent at middle (
<figureCitation id="132F2A756B49FFEDAD40FDA7FD6938AF" box="[606,695,1485,1507]" captionStart="FIGURE 10" captionStartId="52.[151,250,1909,1933]" captionTargetBox="[207,1379,195,1873]" captionTargetId="figure-19@52.[195,1393,181,1885]" captionTargetPageId="52" captionText="FIGURE 10. AD: Maladera lignicolor (Fairmaire, 1887) (holotype Autoserica tenebrosa Frey, 1972); EH: M. orientalis (Motschulsky, 1858) (lectotype); IL: M. enigma Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496336" httpUri="https://zenodo.org/record/4496336/files/figure.png" pageId="40" pageNumber="41">Fig. 10C</figureCitation>
)..............................
<taxonomicName id="4C144D736B49FFEDAB34FDA7FA4238AF" authority="(Fairmaire, 1887)" baseAuthorityName="Fairmaire" baseAuthorityYear="1887" box="[1066,1436,1485,1507]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="lignicolor">
<emphasis id="B960EAE26B49FFEDAB34FDA7FB3338AE" box="[1066,1261,1485,1506]" italics="true" pageId="40" pageNumber="41">Maladera lignicolor</emphasis>
(
<bibRefCitation id="EF854B016B49FFEDABE7FDA7FA4B38AF" author="Fairmaire, M. L." box="[1273,1429,1485,1507]" pageId="40" pageNumber="41" pagination="87 - 136" refId="ref332020" refString="Fairmaire, M. L. (1887) Coleopteres de l'Interieur de la Chine. Annales de la Societe Entomologique de Belgique, 31, 87 - 136." type="journal article" year="1887">Fairmaire, 1887</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FD80FA423B4C" blockId="40.[151,1437,1309,2035]" box="[151,1436,1514,1536]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FD80FA423B4C" box="[151,1436,1514,1536]" pageId="40" pageNumber="41">
4 Right paramere strongly bent at middle (e.g.,
<figureCitation id="132F2A756B49FFEDAD89FD80FD3A3B4C" box="[663,740,1514,1536]" captionStart="FIGURE 9" captionStartId="46.[151,250,1909,1933]" captionTargetBox="[229,1362,188,1885]" captionTargetId="figure-19@46.[217,1371,181,1885]" captionTargetPageId="46" captionText="FIGURE 9. AE: Maladera hajeki Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); FI: M. clypeata (Fairmaire, 1887) (Vietnam: Sa Pa); JM: M. guomenshanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, G, K: parameres, dorsal view; C: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496334" httpUri="https://zenodo.org/record/4496334/files/figure.png" pageId="40" pageNumber="41">Fig. 9D</figureCitation>
)........................................................ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FE6DFA423B77" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FE6DFA433B51" blockId="40.[151,1437,1309,2035]" box="[151,1437,1543,1566]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FE6DFA433B51" box="[151,1437,1543,1566]" pageId="40" pageNumber="41">
5 Left paramere straight, bent dorsally at apex.............................
<taxonomicName id="4C144D736B49FFEDACFAFE62FABE3B52" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[996,1376,1544,1566]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hajeki" status="sp. nov.">
<emphasis id="B960EAE26B49FFEDACFAFE62FB5D3B51" box="[996,1155,1544,1565]" italics="true" pageId="40" pageNumber="41">Maladera hajeki</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B49FFEDAA77FE62FA433B51" box="[1385,1437,1544,1565]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FE4FFA423B77" blockId="40.[151,1437,1309,2035]" box="[151,1436,1573,1595]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FE4FFA423B77" box="[151,1436,1573,1595]" pageId="40" pageNumber="41">5 Left paramere bent ventrally............................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FE28FA433BDE" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FE28FA423B14" blockId="40.[151,1437,1309,2035]" box="[151,1436,1602,1624]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FE28FA423B14" box="[151,1436,1602,1624]" pageId="40" pageNumber="41">
6 Left paramere as long as the right, evenly narrowed towards pointed apex.............
<taxonomicName id="4C144D736B49FFEDAB5BFE28FA423B14" authority="(Moser, 1915)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1093,1436,1602,1624]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="cariniceps">
<emphasis id="B960EAE26B49FFEDAB5BFE28FAD03B14" box="[1093,1294,1602,1624]" italics="true" pageId="40" pageNumber="41">Maladera cariniceps</emphasis>
(Moser, 1915)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FE0AFA433BDE" blockId="40.[151,1437,1309,2035]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FE0AFA433BDE" pageId="40" pageNumber="41">
6 Left paramere distinctly shorter than the right, with a large ventral teeth before apex..................................................................................................
<taxonomicName id="4C144D736B49FFEDACEEFE17FABE3BDF" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[1008,1376,1661,1683]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="weni" status="sp. nov.">
<emphasis id="B960EAE26B49FFEDACEEFE17FB5F3BDF" box="[1008,1153,1661,1683]" italics="true" pageId="40" pageNumber="41">Maladera weni</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B49FFEDAA77FE17FA433BDE" box="[1385,1437,1661,1682]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FEF1FA423B81" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FEF1FA423BE3" blockId="40.[151,1437,1309,2035]" box="[151,1436,1690,1712]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FEF1FA423BE3" box="[151,1436,1690,1712]" pageId="40" pageNumber="41">
7 Body size larger,
<quantity id="4CEC9B156B49FFEDAE94FEF0FE123BFC" box="[394,460,1690,1712]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="40" pageNumber="41" unit="mm" value="11.0">11 mm</quantity>
or greater........................................................................ 9
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FED2FA423B81" blockId="40.[151,1437,1309,2035]" box="[151,1436,1720,1742]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FED2FA423B81" box="[151,1436,1720,1742]" pageId="40" pageNumber="41">
7 Body size smaller, &lt;
<quantity id="4CEC9B156B49FFEDAEB9FED2FE323B81" box="[423,492,1720,1741]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="41" unit="mm" value="10.0">10 mm</quantity>
............................................................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FEBFFA433A44" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FEBFFA423BA7" blockId="40.[151,1437,1309,2035]" box="[151,1436,1749,1771]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FEBFFA423BA7" box="[151,1436,1749,1771]" pageId="40" pageNumber="41">
8 Disc of frons setose...................................................
<taxonomicName id="4C144D736B49FFEDAB11FEBFFA423BA7" authority="(Motschulsky, 1858)" authorityName="Nikolaev" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[1039,1436,1749,1771]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B960EAE26B49FFEDAB11FEBFFB0E3BA6" box="[1039,1232,1749,1770]" italics="true" pageId="40" pageNumber="41">Maladera orientalis</emphasis>
(
<bibRefCitation id="EF854B016B49FFEDABC2FEBFFA4B3BA7" author="Motschulsky, V." box="[1244,1429,1749,1771]" pageId="40" pageNumber="41" pagination="25 - 41" refId="ref334477" refString="Motschulsky, V. (1858) Entomologie speciale. Insectes du Japon. Etudes Entomologiques, 6, 25 - 41. [1857]" type="journal article" year="1858">Motschulsky, 1858</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FE98FA433A44" blockId="40.[151,1437,1309,2035]" box="[151,1437,1778,1800]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FE98FA433A44" box="[151,1437,1778,1800]" pageId="40" pageNumber="41">
8 Disc of frons glabrous......................................
<taxonomicName id="4C144D736B49FFEDAC90FE98FABE3A44" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[910,1376,1778,1800]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="guangdongana" status="sp. nov.">
<emphasis id="B960EAE26B49FFEDAC90FE98FB5D3A4B" box="[910,1155,1778,1799]" italics="true" pageId="40" pageNumber="41">Maladera guangdongana</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B49FFEDAA77FE99FA433A44" box="[1385,1437,1779,1800]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FF7AFA423A32" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FF7AFA433A0E" blockId="40.[151,1437,1309,2035]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FF7AFA433A0E" pageId="40" pageNumber="41">
9 Metatibia widest at middle. Setae on head robust and long. Right paramere long, only little shorter than phallobase...............................................................
<taxonomicName id="4C144D736B49FFEDAC6AFF47FABE3A0F" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[884,1376,1837,1859]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="guomenshanensis" status="sp. nov.">
<emphasis id="B960EAE26B49FFEDAC6AFF47FB5F3A0E" box="[884,1153,1837,1858]" italics="true" pageId="40" pageNumber="41">Maladera guomenshanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B49FFEDAA77FF47FA433A0E" box="[1385,1437,1837,1858]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FF20FA423A32" blockId="40.[151,1437,1309,2035]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FF20FA423A32" pageId="40" pageNumber="41">
9 Metatibia widest at apex. Setae on head fine and short. Right paramere shorter, equals only 1/3 of phallobasal length.............................................................................
<taxonomicName id="4C144D736B49FFEDAB29FF02FA423A32" authority="(Fairmaire, 1887)" baseAuthorityName="Fairmaire" baseAuthorityYear="1887" box="[1079,1436,1896,1918]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B960EAE26B49FFEDAB29FF02FB333A31" box="[1079,1261,1896,1917]" italics="true" pageId="40" pageNumber="41">Maladera clypeata</emphasis>
(
<bibRefCitation id="EF854B016B49FFEDABE7FF02FA4B3A32" author="Fairmaire, M. L." box="[1273,1429,1896,1918]" pageId="40" pageNumber="41" pagination="87 - 136" refId="ref332020" refString="Fairmaire, M. L. (1887) Coleopteres de l'Interieur de la Chine. Annales de la Societe Entomologique de Belgique, 31, 87 - 136." type="journal article" year="1887">Fairmaire, 1887</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFEDAF89FFEFFA423AFB" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FFEFFA423AD6" blockId="40.[151,1437,1309,2035]" box="[151,1436,1925,1947]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FFEFFA423AD6" box="[151,1436,1925,1947]" pageId="40" pageNumber="41">10 Left and right parameres subequal in length............................................................... 11</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFEDAF89FFC8FA423AFB" blockId="40.[151,1437,1309,2035]" box="[151,1436,1954,1976]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FFC8FA423AFB" box="[151,1436,1954,1976]" pageId="40" pageNumber="41">10 Right paramere distinctly longer than the left one. Left paramere strongly curved.................................. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B49FFECAF89FFAAFA433DE1" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<paragraph id="8BAB36F06B49FFEDAF89FFAAFA423A99" blockId="40.[151,1437,1309,2035]" box="[151,1436,1984,2006]" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFEDAF89FFAAFA423A99" box="[151,1436,1984,2006]" pageId="40" pageNumber="41">11 Parameres and phallobase rather narrow.................................................................. 13</keyLead>
</paragraph>
<paragraph id="8BAB36F06B49FFECAF89FFB7FA433DE1" blockId="40.[151,1437,1309,2035]" lastBlockId="41.[151,1437,151,1728]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<keyLead id="30E595C56B49FFECAF89FFB7FA433DE1" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
11 Right paramere and phallobase rather robust. Left paramere straight. Right paramere distinctly concave ventrally (
<figureCitation id="132F2A756B49FFEDAA34FFB7FA5A3ABF" box="[1322,1412,2013,2035]" captionStart="FIGURE 10" captionStartId="52.[151,250,1909,1933]" captionTargetBox="[207,1379,195,1873]" captionTargetId="figure-19@52.[195,1393,181,1885]" captionTargetPageId="52" captionText="FIGURE 10. AD: Maladera lignicolor (Fairmaire, 1887) (holotype Autoserica tenebrosa Frey, 1972); EH: M. orientalis (Motschulsky, 1858) (lectotype); IL: M. enigma Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496336" httpUri="https://zenodo.org/record/4496336/files/figure.png" pageId="40" pageNumber="41">Fig. 10K</figureCitation>
)................................................................
<taxonomicName id="4C144D736B48FFECACC8F8FDFABE3DE1" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[982,1376,151,173]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
<emphasis id="B960EAE26B48FFECACC8F8FDFB5C3DE1" box="[982,1154,151,173]" italics="true" pageId="41" pageNumber="42">Maladera enigma</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77F8F2FA433DE1" box="[1385,1437,152,173]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89F8DFFA423C6F" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89F8DFFA423DA4" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89F8DFFA423DA4" pageId="41" pageNumber="42">
12 Right paramere narrow, only half as wide as phallobase at apex. Left paramere short, not exceeding basal half of the right one....................................................................
<taxonomicName id="4C144D736B48FFECAB22F8B8FA423DA4" authority="(Moser, 1915 b)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1084,1436,210,232]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="diversipes">
<emphasis id="B960EAE26B48FFECAB22F8B8FADC3DAB" box="[1084,1282,210,231]" italics="true" pageId="41" pageNumber="42">Maladera diversipes</emphasis>
(
<bibRefCitation id="EF854B016B48FFECAA10F8B8FA4B3DA4" author="Moser, J." box="[1294,1429,210,232]" pageId="41" pageNumber="42" pagination="337 - 393" refId="ref334285" refString="Moser, J. (1915 b) Neue Serica - Arten. Deutsche Entomologische Zeitschrift, 1915, 337 - 393. https: // doi. org / 10.1002 / mmnd. 48019150402" type="journal article" year="1915">Moser, 1915b</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89F89AFA423C6F" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89F89AFA423C6F" pageId="41" pageNumber="42">
12 Right paramere wide, as wide as phallobase at apex. Left paramere long, distinctly exceeding basal half of the right one.............................................................................
<taxonomicName id="4C144D736B48FFECAB5BF967FA423C6F" authority="(Moser, 1915 b)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1093,1436,269,291]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="B960EAE26B48FFECAB5BF967FADC3C6E" box="[1093,1282,269,290]" italics="true" pageId="41" pageNumber="42">Maladera infuscata</emphasis>
(
<bibRefCitation id="EF854B016B48FFECAA10F967FA4B3C6F" author="Moser, J." box="[1294,1429,269,291]" pageId="41" pageNumber="42" pagination="337 - 393" refId="ref334285" refString="Moser, J. (1915 b) Neue Serica - Arten. Deutsche Entomologische Zeitschrift, 1915, 337 - 393. https: // doi. org / 10.1002 / mmnd. 48019150402" type="journal article" year="1915">Moser, 1915b</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89F940FA423C11" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89F940FA423C0C" blockId="41.[151,1437,151,1728]" box="[151,1436,298,320]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89F940FA423C0C" box="[151,1436,298,320]" pageId="41" pageNumber="42">13 Phallobase narrowed towards apex (dorsal view)........................................................... 15</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89F922FA423C11" blockId="41.[151,1437,151,1728]" box="[151,1436,327,349]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89F922FA423C11" box="[151,1436,327,349]" pageId="41" pageNumber="42">13 Phallobase at apex as wide as at middle (dorsal view)........................................................ 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89F90FFA433CF9" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89F90FFA433C36" blockId="41.[151,1437,151,1728]" box="[151,1437,357,379]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89F90FFA433C36" box="[151,1437,357,379]" pageId="41" pageNumber="42">
14 Right paramere fused basally with the phallobase. Left paramere straight......
<taxonomicName id="4C144D736B48FFECACC4F90FFABE3C37" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[986,1376,357,379]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="riberai" status="sp. nov.">
<emphasis id="B960EAE26B48FFECACC4F90FFB5C3C36" box="[986,1154,357,378]" italics="true" pageId="41" pageNumber="42">Maladera riberai</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77F90FFA433C36" box="[1385,1437,357,378]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89F9E8FA433CF9" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89F9E8FA433CF9" pageId="41" pageNumber="42">
14 Right paramere separate basally from the phallobase. Left paramere strongly curved................................................................................................
<taxonomicName id="4C144D736B48FFECACC6F9F5FABE3CF9" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[984,1376,415,437]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="xingkei" status="sp. nov.">
<emphasis id="B960EAE26B48FFECACC6F9F5FB5F3CF8" box="[984,1153,415,436]" italics="true" pageId="41" pageNumber="42">Maladera xingkei</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77F9CAFA433CF9" box="[1385,1437,416,437]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89F9D7FA423F41" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89F9D7FA433CBC" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89F9D7FA433CBC" pageId="41" pageNumber="42">
15 Right paramere weakly curved externally. Left paramere after base strongly bent, after basal extension abruptly filiform.................................................................
<taxonomicName id="4C144D736B48FFECACA8F9B0FABE3CBC" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[950,1376,474,496]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="anhuiensis" status="sp. nov.">
<emphasis id="B960EAE26B48FFECACA8F9B0FB5C3CA3" box="[950,1154,474,495]" italics="true" pageId="41" pageNumber="42">Maladera anhuiensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77F9B1FA433CBC" box="[1385,1437,475,496]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89F99DFA423F41" blockId="41.[151,1437,151,1728]" box="[151,1436,503,525]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89F99DFA423F41" box="[151,1436,503,525]" pageId="41" pageNumber="42">15 Right paramere straight. Left paramere weakly curved, after basal extension gently and evenly narrowed............... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FA7FFA433FCE" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FA7FFA433F04" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FA7FFA433F04" pageId="41" pageNumber="42">
16 Left paramere at apex abruptly bent ventrally. Right paramere dorsally sclerotised in apical half................................................................................
<taxonomicName id="4C144D736B48FFECAC94FA58FABE3F04" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[906,1376,562,584]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dahongshanica" status="sp. nov.">
<emphasis id="B960EAE26B48FFECAC94FA58FB5C3F0B" box="[906,1154,562,583]" italics="true" pageId="41" pageNumber="42">Maladera dahongshanica</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77FA59FA433F04" box="[1385,1437,563,584]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FA25FA433FCE" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FA25FA433FCE" pageId="41" pageNumber="42">
16 Left paramere at apex straight. Right paramere dorsally not sclerotised but membraneous in apical half...........................................................................
<taxonomicName id="4C144D736B48FFECAC9BFA07FABE3FCF" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[901,1376,621,643]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="shiniushanensis" status="sp. nov.">
<emphasis id="B960EAE26B48FFECAC9BFA07FB5C3FCE" box="[901,1154,621,642]" italics="true" pageId="41" pageNumber="42">Maladera shiniushanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77FA07FA433FCE" box="[1385,1437,621,642]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FAE0FA423FF1" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FAE0FA423FD3" blockId="41.[151,1437,151,1728]" box="[151,1436,650,672]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FAE0FA423FD3" box="[151,1436,650,672]" pageId="41" pageNumber="42">17 Aedeagus apically 90º clockwise rotated, right parameres thus appears to be inserted ventrally, the left dorsally.......... 19</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FAC2FA423FF1" blockId="41.[151,1437,151,1728]" box="[151,1436,679,701]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FAC2FA423FF1" box="[151,1436,679,701]" pageId="41" pageNumber="42">17 Aedeagus apically not rotated, paramere both insert in lateral positions.......................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FAAFFA423FB4" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FAAFFA423F97" blockId="41.[151,1437,151,1728]" box="[151,1436,709,731]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FAAFFA423F97" box="[151,1436,709,731]" pageId="41" pageNumber="42">
18 Body yellowish to reddish brown. Right paramere not widened dorsally................
<taxonomicName id="4C144D736B48FFECAB53FAAFFA423F97" authority="(Brenske, 1896)" baseAuthorityName="Brenske" baseAuthorityYear="1896" box="[1101,1436,709,731]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="B960EAE26B48FFECAB53FAAFFB223F96" box="[1101,1276,709,730]" italics="true" pageId="41" pageNumber="42">Maladera cardoni</emphasis>
(Brenske, 1896)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FA88FA423FB4" blockId="41.[151,1437,151,1728]" box="[151,1436,738,760]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FA88FA423FB4" box="[151,1436,738,760]" pageId="41" pageNumber="42">
18 Body dark brown. Right paramere strongly widened dorsally.............................
<taxonomicName id="4C144D736B48FFECAB96FA88FA423FB4" authority="(Frey, 1972)" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1160,1436,738,760]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="B960EAE26B48FFECAB96FA88FAFE3FBB" box="[1160,1312,738,759]" italics="true" pageId="41" pageNumber="42">Maladera fusca</emphasis>
(Frey, 1972)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FB6AFA423E7E" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FB6AFA423E59" blockId="41.[151,1437,151,1728]" box="[151,1436,767,789]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FB6AFA423E59" box="[151,1436,767,789]" pageId="41" pageNumber="42">19 Phallobase long and narrow............................................................................ 20</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FB77FA423E7E" blockId="41.[151,1437,151,1728]" box="[151,1436,797,819]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FB77FA423E7E" box="[151,1436,797,819]" pageId="41" pageNumber="42">
19 Phallobase shorter and very robust, wide at middle..............................
<taxonomicName id="4C144D736B48FFECAB20FB77FA423E7E" authority="Nomura, 1974" authorityName="Nomura" authorityYear="1974" box="[1086,1436,797,818]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="masumotoi">
<emphasis id="B960EAE26B48FFECAB20FB77FAD53E7E" box="[1086,1291,797,818]" italics="true" pageId="41" pageNumber="42">Maladera masumotoi</emphasis>
<bibRefCitation id="EF854B016B48FFECAA0CFB77FA423E7E" author="Nomura, S." box="[1298,1436,797,818]" pageId="41" pageNumber="42" pagination="81 - 115" refId="ref335003" refString="Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115." type="journal article" year="1974">Nomura, 1974</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FB50FA423EE4" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FB50FA423E21" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FB50FA423E21" pageId="41" pageNumber="42">
20 Right paramere inclinated in right angle to longitudinal axis of phallobase. Colour of body reddish brown. Body smaller
<quantity id="4CEC9B156B48FFECAA9AFB50FED23E21" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="41" pageNumber="42" unit="mm" value="10.0">10 mm</quantity>
................................................................................................ 21
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FB1FFA423EE4" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FB1FFA423EE4" pageId="41" pageNumber="42">
20 Right paramere straight, in line with longitudinal axis of phallobase main. Colour of body brown or dark brown. Body larger
<quantity id="4CEC9B156B48FFECAFFCFBF8FEF63EE4" box="[226,296,914,936]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="41" pageNumber="42" unit="mm" value="10.0">10 mm</quantity>
............................................................................................. 25
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FBDAFA42394C" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FBDAFA423E89" blockId="41.[151,1437,151,1728]" box="[151,1436,943,965]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FBDAFA423E89" box="[151,1436,943,965]" pageId="41" pageNumber="42">21 Right paramere bent with its main surface internally, mesal side not visible in apical view........................... 22</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FBA7FA42394C" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FBA7FA42394C" pageId="41" pageNumber="42">
21 Right paramere not bent with its main surface internally, mesal side visible in apical view....................................................................................................
<taxonomicName id="4C144D736B48FFECAB5EFB80FA42394C" authority="(Fairmaire, 1891)" baseAuthorityName="Fairmaire" baseAuthorityYear="1891" box="[1088,1436,1002,1024]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="ovatula">
<emphasis id="B960EAE26B48FFECAB5EFB80FB333EB3" box="[1088,1261,1002,1023]" italics="true" pageId="41" pageNumber="42">Maladera ovatula</emphasis>
(
<bibRefCitation id="EF854B016B48FFECABE7FB80FA4B394C" author="Fairmaire, M. L." box="[1273,1429,1002,1024]" pageId="41" pageNumber="42" pagination="187 - 219" refId="ref332080" refString="Fairmaire, M. L. (1891) Coleopteres de l'Interieur de la Chine (7). Annales de la Societe Entomologique de Belgique, 35, 187 - 219." type="journal article" year="1891">Fairmaire, 1891</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FC62FA423976" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FC62FA423951" blockId="41.[151,1437,151,1728]" box="[151,1436,1031,1053]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FC62FA423951" box="[151,1436,1031,1053]" pageId="41" pageNumber="42">22 Right paramere not convexly widened ventrally, but straight................................................... 23</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FC4FFA423976" blockId="41.[151,1437,151,1728]" box="[151,1436,1061,1083]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FC4FFA423976" box="[151,1436,1061,1083]" pageId="41" pageNumber="42">22 Right paramere convexly widened ventrally............................................................... 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FC28FA42393A" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FC28FA423914" blockId="41.[151,1437,151,1728]" box="[151,1436,1090,1112]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FC28FA423914" box="[151,1436,1090,1112]" pageId="41" pageNumber="42">
23 Left paramere angled, in basal half much wider than in apical half.................
<taxonomicName id="4C144D736B48FFECAB3DFC28FA423914" authority="(Fairmaire, 1888 b)" baseAuthorityName="Fairmaire" baseAuthorityYear="1888" box="[1059,1436,1090,1112]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B960EAE26B48FFECAB3DFC28FB3F391B" box="[1059,1249,1090,1111]" italics="true" pageId="41" pageNumber="42">Maladera verticalis</emphasis>
(
<bibRefCitation id="EF854B016B48FFECABF3FC28FA573914" author="Fairmaire, M. L." box="[1261,1417,1090,1112]" pageId="41" pageNumber="42" pagination="7 - 46" refId="ref332050" refString="Fairmaire, M. L. (1888) Coleopteres de l'Interieur de la Chine. Annales de la Societe Entomologique de Belgique, 32, 7 - 46." type="journal article" year="1888">Fairmaire, 1888</bibRefCitation>
b)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FC0AFA42393A" blockId="41.[151,1437,151,1728]" box="[151,1436,1119,1142]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FC0AFA42393A" box="[151,1436,1119,1142]" pageId="41" pageNumber="42">
23 Left evenly curved, in basal half not much wider than in apical half..................
<taxonomicName id="4C144D736B48FFECAB20FC35FA42393A" authority="(Brenske, 1898)" baseAuthorityName="Brenske" baseAuthorityYear="1898" box="[1086,1436,1119,1142]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="formosae">
<emphasis id="B960EAE26B48FFECAB20FC35FB253939" box="[1086,1275,1119,1141]" italics="true" pageId="41" pageNumber="42">Maladera formosae</emphasis>
(
<bibRefCitation id="EF854B016B48FFECAA16FC35FA4B393A" author="Brenske, E." box="[1288,1429,1119,1142]" pageId="41" pageNumber="42" pagination="205 - 403" refId="ref331459" refString="Brenske, E. (1898) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205 - 403. https: // doi. org / 10.1002 / mmnd. 47918980303" type="journal article" year="1898">Brenske, 1898</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FC17FA4239A7" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FC17FA4239FC" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FC17FA4239FC" pageId="41" pageNumber="42">
24 Left paramere strongly evenly curved, in basal half strongly widenend, with a large blunt tooth in the centre of the curvation..........................................................................
<taxonomicName id="4C144D736B48FFECAB4CFCF0FA4239FC" authority="(Brenske, 1898)" baseAuthorityName="Brenske" baseAuthorityYear="1898" box="[1106,1436,1178,1200]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sontica">
<emphasis id="B960EAE26B48FFECAB4CFCF0FB2539FC" box="[1106,1275,1178,1200]" italics="true" pageId="41" pageNumber="42">Maladera sontica</emphasis>
(
<bibRefCitation id="EF854B016B48FFECAA16FCF0FA4B39FC" author="Brenske, E." box="[1288,1429,1178,1200]" pageId="41" pageNumber="42" pagination="205 - 403" refId="ref331459" refString="Brenske, E. (1898) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205 - 403. https: // doi. org / 10.1002 / mmnd. 47918980303" type="journal article" year="1898">Brenske, 1898</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FCD2FA4239A7" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FCD2FA4239A7" pageId="41" pageNumber="42">
24 Left paramere weakly unevenly curved, in basal half only little widenend, with a blunt tooth only at the very base...............................................................................
<taxonomicName id="4C144D736B48FFECAB2CFCBFFA4239A7" authority="(Brenske, 1898)" baseAuthorityName="Brenske" baseAuthorityYear="1898" box="[1074,1436,1237,1259]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="perniciosa">
<emphasis id="B960EAE26B48FFECAB2CFCBFFB2539A6" box="[1074,1275,1237,1258]" italics="true" pageId="41" pageNumber="42">Maladera perniciosa</emphasis>
(
<bibRefCitation id="EF854B016B48FFECAA16FCBFFA4B39A7" author="Brenske, E." box="[1288,1429,1237,1259]" pageId="41" pageNumber="42" pagination="205 - 403" refId="ref331459" refString="Brenske, E. (1898) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205 - 403. https: // doi. org / 10.1002 / mmnd. 47918980303" type="journal article" year="1898">Brenske, 1898</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FC98FA423869" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FC98FA42384B" blockId="41.[151,1437,151,1728]" box="[151,1436,1266,1288]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FC98FA42384B" box="[151,1436,1266,1288]" pageId="41" pageNumber="42">25 Phallobase much elongated, ratio width/ length: 1/5 (lateral view). Body dark brown............................... 27</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FD65FA423869" blockId="41.[151,1437,151,1728]" box="[151,1436,1295,1317]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FD65FA423869" box="[151,1436,1295,1317]" pageId="41" pageNumber="42">25 Phallobase less elongated, ratio width/ length: 1/4 (lateral view). Body reddish brown.............................. 26</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FD47FA4338D6" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FD47FA43382C" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FD47FA43382C" pageId="41" pageNumber="42">
26 Right paramere wide at base, sharply pointed at apex, abruptly narrowed (lateral view) at middle. Left paramere atrongly curved over its entire length...................................
<taxonomicName id="4C144D736B48FFECAC86FD20FABE382C" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[920,1376,1354,1376]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="businskyorum" status="sp. nov.">
<emphasis id="B960EAE26B48FFECAC86FD20FB5F3813" box="[920,1153,1354,1375]" italics="true" pageId="41" pageNumber="42">Maladera businskyorum</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77FD21FA43382C" box="[1385,1437,1355,1376]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FD0DFA4338D6" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FD0DFA4338D6" pageId="41" pageNumber="42">
26 Right paramere moderately wide at base, nearly rounded at apex, weakly narrowed (lateral view) at middle. Left paramere nearly straight, only weakly curved at apex....................
<taxonomicName id="4C144D736B48FFECAC6CFDEFFABE38D7" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[882,1376,1413,1435]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="wuliangshanensis" status="sp. nov.">
<emphasis id="B960EAE26B48FFECAC6CFDEFFB5C38D6" box="[882,1154,1413,1434]" italics="true" pageId="41" pageNumber="42">Maladera wuliangshanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77FDEFFA4338D6" box="[1385,1437,1413,1434]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FDC8FA423899" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FDC8FA4238F4" blockId="41.[151,1437,151,1728]" box="[151,1436,1442,1464]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FDC8FA4238F4" box="[151,1436,1442,1464]" pageId="41" pageNumber="42">
27 Right paramere wide at base, narrowed behind base and before apex..
<taxonomicName id="4C144D736B48FFECAC90FDC8FA4238F4" authority="Ahrens &amp; Fabrizi, 2016" authorityName="Ahrens &amp; Fabrizi" authorityYear="2016" box="[910,1436,1442,1464]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="pseudohongkongica">
<emphasis id="B960EAE26B48FFECAC90FDC8FB6C38FB" box="[910,1202,1442,1463]" italics="true" pageId="41" pageNumber="42">Maladera pseudohongkongica</emphasis>
<bibRefCitation id="EF854B016B48FFECABA6FDC8FA4238F4" author="Ahrens, D. &amp; Fabrizi, S." box="[1208,1436,1442,1464]" pageId="41" pageNumber="42" pagination="1 - 355" refId="ref330518" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi, 2016</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FDAAFA423899" blockId="41.[151,1437,151,1728]" box="[151,1436,1471,1493]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FDAAFA423899" box="[151,1436,1471,1493]" pageId="41" pageNumber="42">27 Right paramere at middle as narrow as at base, abruptly narrowing only shortly before apex......................... 28</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FDB7FA423B43" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FDB7FA4238BE" blockId="41.[151,1437,151,1728]" box="[151,1436,1501,1523]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FDB7FA4238BE" box="[151,1436,1501,1523]" pageId="41" pageNumber="42">28 Dorsal margin of right paramere concave (lateral view)...................................................... 29</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FD90FA423B43" blockId="41.[151,1437,151,1728]" box="[151,1436,1530,1552]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FD90FA423B43" box="[151,1436,1530,1552]" pageId="41" pageNumber="42">28 Dorsal margin of right paramere weakly convex (lateral view)................................................. 30</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FE72FA433BC9" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FE72FA423B07" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FE72FA423B07" pageId="41" pageNumber="42">
29 Right paramere shorter and reflexed, emargination on dorsal margin before apex deeper...................................................................................................
<taxonomicName id="4C144D736B48FFECAB05FE5FFA423B07" authority="(Brenske, 1898)" baseAuthorityName="Brenske" baseAuthorityYear="1898" box="[1051,1436,1589,1611]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hongkongica">
<emphasis id="B960EAE26B48FFECAB05FE5FFB223B06" box="[1051,1276,1589,1610]" italics="true" pageId="41" pageNumber="42">Maladera hongkongica</emphasis>
(
<bibRefCitation id="EF854B016B48FFECAA16FE5FFA4B3B07" author="Brenske, E." box="[1288,1429,1589,1611]" pageId="41" pageNumber="42" pagination="205 - 403" refId="ref331459" refString="Brenske, E. (1898) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205 - 403. https: // doi. org / 10.1002 / mmnd. 47918980303" type="journal article" year="1898">Brenske, 1898</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FE38FA433BC9" blockId="41.[151,1437,151,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FE38FA433BC9" pageId="41" pageNumber="42">
29 Right paramere longer and straight, emargination on dorsal margin before apex shallow.............................................................................................
<taxonomicName id="4C144D736B48FFECACD3FE05FABE3BC9" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[973,1376,1647,1669]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="juntongi" status="sp. nov.">
<emphasis id="B960EAE26B48FFECACD3FE05FB5C3BC9" box="[973,1154,1647,1669]" italics="true" pageId="41" pageNumber="42">Maladera juntongi</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77FE1AFA433BC9" box="[1385,1437,1648,1669]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B48FFECAF89FEE7FA423B8C" pageId="41" pageNumber="42">
<paragraph id="8BAB36F06B48FFECAF89FEE7FA433BEE" blockId="41.[151,1437,151,1728]" box="[151,1437,1677,1699]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FEE7FA433BEE" box="[151,1437,1677,1699]" pageId="41" pageNumber="42">
30 Right paramere tubular in apical half, pointed apex very short...............
<taxonomicName id="4C144D736B48FFECACF8FEE7FABE3BEF" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[998,1376,1677,1699]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="yibini" status="sp. nov.">
<emphasis id="B960EAE26B48FFECACF8FEE7FB5C3BEE" box="[998,1154,1677,1698]" italics="true" pageId="41" pageNumber="42">Maladera yibini</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B48FFECAA77FEE7FA433BEE" box="[1385,1437,1677,1698]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B48FFECAF89FEC0FA423B8C" blockId="41.[151,1437,151,1728]" box="[151,1436,1706,1728]" pageId="41" pageNumber="42">
<keyLead id="30E595C56B48FFECAF89FEC0FA423B8C" box="[151,1436,1706,1728]" pageId="41" pageNumber="42">
30 Right paramere parabolic in apical half, pointed apex long..................
<taxonomicName id="4C144D736B48FFECACEEFEC0FA423B8C" authority="Ahrens &amp; Fabrizi, 2016" authorityName="Ahrens &amp; Fabrizi" authorityYear="2016" box="[1008,1436,1706,1728]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="johannesi">
<emphasis id="B960EAE26B48FFECACEEFEC0FB6C3BF3" box="[1008,1202,1706,1727]" italics="true" pageId="41" pageNumber="42">Maladera johannesi</emphasis>
<bibRefCitation id="EF854B016B48FFECABA6FEC0FA423B8C" author="Ahrens, D. &amp; Fabrizi, S." box="[1208,1436,1706,1728]" pageId="41" pageNumber="42" pagination="1 - 355" refId="ref330518" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi, 2016</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>