186 lines
22 KiB
XML
186 lines
22 KiB
XML
<document id="DCA88D64A6EA63FE04BE6BA9C5360B5B" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk" docDate="2021" docId="03BD87E66B16FFBDAF89FBE0FD1E3FBD" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera yakouensis Ahrens, Fabrizi & Liu 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="121" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="120" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="9FA2C9288A2ED93CD73B1BF6757E5F39">A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="B6570CDBFC9D0402FCA9793C1194E973">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="583A42B9473EA6F0B3EBDC8CB19B314D">Liu, Wan-Gang</mods:namePart>
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<mods:nameIdentifier id="24767DE580545C87AEF1A47579284513" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:namePart id="7AB7B2E1AE577B4FD0068E206F9A6D0F">Bai, Ming</mods:namePart>
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<mods:namePart id="F6A6D88384F674612880B4B30C84295A">Yang, Xing-Ke</mods:namePart>
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<mods:namePart id="42E2D9292D11B460D34C02E3CC6A95DC">Ahrens, Dirk</mods:namePart>
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<mods:title id="EBE8FF24912707272CB89D44367D83D3">Zootaxa</mods:title>
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<mods:part id="77619B0FD2D54150BA08F5DFA999D637">
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<mods:date id="DD29731E10614902E9951D622E9D6B7A">2021</mods:date>
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<mods:number id="F6754AC3FD35AD294834967932E3A67D">2021-02-03</mods:number>
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<mods:number id="6B13DDFB998F407F243CC6B158065E74">4922</mods:number>
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<mods:classification id="44677BE15C80D10C250D5DE73954A036">journal article</mods:classification>
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<mods:identifier id="B128AF30D083641E2CA2611A34B490AB" type="CLB-Dataset">8257</mods:identifier>
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<mods:identifier id="634539E446DF7091FD8959A1E755C4F4" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
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<treatment id="03BD87E66B16FFBDAF89FBE0FD1E3FBD" ID-DOI="http://doi.org/10.5281/zenodo.4496526" ID-GBIF-Taxon="177323692" ID-Zenodo-Dep="4496526" LSID="urn:lsid:plazi:treatment:03BD87E66B16FFBDAF89FBE0FD1E3FBD" httpUri="http://treatment.plazi.org/id/03BD87E66B16FFBDAF89FBE0FD1E3FBD" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120">
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<subSubSection id="C30E657B6B16FFB2AF89FBE0FCD63EE9" box="[151,776,906,933]" pageId="119" pageNumber="120" type="nomenclature">
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<heading id="D0E3819C6B16FFB2AF89FBE0FCD63EE9" bold="true" box="[151,776,906,933]" fontSize="11" level="2" pageId="119" pageNumber="120" reason="4">
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<emphasis id="B960EAE26B16FFB2AF89FBE0FCD63EE9" bold="true" box="[151,776,906,933]" pageId="119" pageNumber="120">
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<taxonomicName id="4C144D736B16FFB2AF89FBE0FD673EE8" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[151,697,906,933]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="yakouensis" status="sp. nov.">
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<emphasis id="B960EAE26B16FFB2AF89FBE0FE443EE9" bold="true" box="[151,410,906,933]" italics="true" pageId="119" pageNumber="120">Maladera yakouensis</emphasis>
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Ahrens, Fabrizi & Liu
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25357996B16FFB2ADD8FBE1FCD63EE9" box="[710,776,907,933]" pageId="119" pageNumber="120" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30E657B6B16FFB2AF89FBC5FEBF3E85" box="[151,353,942,969]" pageId="119" pageNumber="120" type="description">
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<paragraph id="8BAB36F06B16FFB2AF89FBC5FEBF3E85" blockId="119.[151,776,906,969]" box="[151,353,942,969]" pageId="119" pageNumber="120">
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<figureCitation id="132F2A756B16FFB2AF89FBC5FED03E85" box="[151,270,942,969]" captionStart="FIGURE 25" captionStartId="117.[151,250,1897,1921]" captionTargetBox="[236,1354,251,1860]" captionTargetId="figure-52@117.[221,1365,237,1873]" captionTargetPageId="117" captionText="FIGURE 25.A–D: Maladera diaolinensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. yipinglangensis Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. yakouensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496370" httpUri="https://zenodo.org/record/4496370/files/figure.png" pageId="119" pageNumber="120">Figures 25</figureCitation>
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I–L, 98
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</paragraph>
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</subSubSection>
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<subSubSection id="C30E657B6B16FFB2AF89FB9DFB0A395D" box="[151,1236,1014,1041]" pageId="119" pageNumber="120" type="materials_examined">
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<paragraph id="8BAB36F06B16FFB2AF89FB9DFB0A395D" blockId="119.[151,1437,1014,2013]" box="[151,1236,1014,1041]" pageId="119" pageNumber="120">
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<emphasis id="B960EAE26B16FFB2AF89FB9DFE61395C" bold="true" box="[151,447,1014,1041]" pageId="119" pageNumber="120">Type material examined.</emphasis>
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<materialsCitation id="3B7C3CAD6B16FFB2AED8FB9DFB0A395D" ID-GBIF-Occurrence="3034556327" box="[454,1236,1014,1041]" collectingDate="2012-05-11" collectionCode="ISAZ" country="China" location="Yakou" pageId="119" pageNumber="120" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="54AF88526B16FFB2AED8FB9DFDED395D" box="[454,563,1015,1041]" pageId="119" pageNumber="120" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D12FD796B16FFB2AD23FB9DFD8C395D" box="[573,594,1015,1041]" pageId="119" pageNumber="120" type="male">♂</specimenCount>
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“[
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<collectingCountry id="F30376606B16FFB2AD74FB9CFD6A395D" box="[618,692,1014,1041]" name="China" pageId="119" pageNumber="120">China</collectingCountry>
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]
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<collectingRegion id="49D0F8126B16FFB2ADDFFB9DFCC4395D" box="[705,794,1015,1041]" country="China" name="Yunnan" pageId="119" pageNumber="120">Yunnan</collectingRegion>
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,
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<location id="8ECB602B6B16FFB2AC3DFB9DFCB0395D" LSID="urn:lsid:plazi:treatment:03BD87E66B16FFBDAF89FBE0FD1E3FBD:8ECB602B6B16FFB2AC3DFB9DFCB0395D" box="[803,878,1015,1041]" country="China" name="Yakou" pageId="119" pageNumber="120" stateProvince="Yunnan">Yakou</location>
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,
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<date id="FFAA10306B16FFB2AC67FB9DFC31395D" box="[889,1007,1015,1041]" pageId="119" pageNumber="120" value="2012-05-11">
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<collectingDate id="EFEEE9D86B16FFB2AC67FB9DFC31395D" box="[889,1007,1015,1041]" pageId="119" pageNumber="120" value="2012-05-11">2012-V-11</collectingDate>
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</date>
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/ LW-1289” (
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<collectionCode id="ED05AE356B16FFB2AB9AFB9DFB16395D" box="[1156,1224,1015,1041]" pageId="119" pageNumber="120">ISAZ</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30E657B6B16FFBDAFD9FC70FCB53F71" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120" type="description">
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<paragraph id="8BAB36F06B16FFB2AFD9FC70FD3A3915" blockId="119.[151,1437,1014,2013]" pageId="119" pageNumber="120">
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<emphasis id="B960EAE26B16FFB2AFD9FC70FE873978" bold="true" box="[199,345,1050,1076]" pageId="119" pageNumber="120">Description.</emphasis>
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Length:
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<quantity id="4CEC9B156B16FFB2AEA5FC71FDCC3979" box="[443,530,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="119" pageNumber="120" unit="mm" value="8.1">8.1 mm</quantity>
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, length of elytra:
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<quantity id="4CEC9B156B16FFB2ADCDFC71FCF43979" box="[723,810,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="119" pageNumber="120" unit="mm" value="6.8">6.8 mm</quantity>
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, width:
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<quantity id="4CEC9B156B16FFB2AC9FFC70FC073979" box="[897,985,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="119" pageNumber="120" unit="mm" value="5.8">5.8 mm</quantity>
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. Body oblong-oval, dark brown, antenna yellow, shiny iridescent, dorsal face nearly glabrous.
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</paragraph>
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<paragraph id="8BAB36F06B16FFB2AFD9FC09FBA33835" blockId="119.[151,1437,1014,2013]" pageId="119" pageNumber="120">
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Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately rounded, anterior margin distinctly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex medially, finely, densely punctate, with a few long setae anteriorly; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye convex, 1.5 times as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely densely punctate, with one terminal seta. Frons with dense and fine punctures, with a few fine setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.58. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined.
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<collectingRegion id="49D0F8126B16FFB2AD53FD35FD723835" box="[589,684,1375,1401]" country="Myanmar" name="Chin" pageId="119" pageNumber="120">Mentum</collectingRegion>
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elevated and slightly flattened anteriorly.
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</paragraph>
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<paragraph id="8BAB36F06B16FFB2AFD9FDE9FE233B1D" blockId="119.[151,1437,1014,2013]" pageId="119" pageNumber="120">Pronotum transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles moderately rounded; anterior margin convex, with fine complete marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous, with a shallow median longitudinal impression; lateral margins sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures, impunctate medially.</paragraph>
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<paragraph id="8BAB36F06B16FFB2AFD9FE31FB1B3BAE" blockId="119.[151,1437,1014,2013]" pageId="119" pageNumber="120">Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures, with minute setae in punctures, penultimate lateral interval with a few fine setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification).</paragraph>
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<paragraph id="8BAB36F06B16FFB2AFD9FE81FB233AD9" blockId="119.[151,1437,1014,2013]" pageId="119" pageNumber="120">Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.4. Pygidium dull, strongly convex, finely and densely punctate, with short smooth midline, with numerous long and short, fine setae in apical half.</paragraph>
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<paragraph id="8BAB36F06B16FFBDAFD9FFF5FE783F55" blockId="119.[151,1437,1014,2013]" lastBlockId="120.[151,1437,151,753]" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120">Legs moderately long and narrow, shiny; femora with two longitudinal rows of setae, coarsely and moderately densely punctate. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia long and wide, widest at apex, ratio of width/length: 1/3.6, moderately carinate dorsally, with two groups of spines, basal group at three fifths, apical group at four fifths of metatibial length, in basal half with a few short and fine single setae and a continuous serrated line beside entire dorsal margin until the basal group of spines; lateral face longitudinally convex, with sparse fine punctures in basal half, minutely setose; ventral margin finely serrate, with three equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere distinctly longer than following tarsomere and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
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<paragraph id="8BAB36F06B19FFBDAFD9FA49FCB53F71" blockId="120.[151,1437,151,753]" box="[199,875,546,573]" pageId="120" pageNumber="121">
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Aedeagus:
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<figureCitation id="132F2A756B19FFBDAE5AFA49FE4A3F71" box="[324,404,546,573]" captionStart="FIGURE 25" captionStartId="117.[151,250,1897,1921]" captionTargetBox="[236,1354,251,1860]" captionTargetId="figure-52@117.[221,1365,237,1873]" captionTargetPageId="117" captionText="FIGURE 25.A–D: Maladera diaolinensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. yipinglangensis Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. yakouensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496370" httpUri="https://zenodo.org/record/4496370/files/figure.png" pageId="120" pageNumber="121">Fig. 25</figureCitation>
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I–K. Habitus:
|
||
<figureCitation id="132F2A756B19FFBDAD2DFA49FD493F71" box="[563,663,546,573]" captionStart="FIGURE 25" captionStartId="117.[151,250,1897,1921]" captionTargetBox="[236,1354,251,1860]" captionTargetId="figure-52@117.[221,1365,237,1873]" captionTargetPageId="117" captionText="FIGURE 25.A–D: Maladera diaolinensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. yipinglangensis Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. yakouensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496370" httpUri="https://zenodo.org/record/4496370/files/figure.png" pageId="120" pageNumber="121">
|
||
Fig.
|
||
<specimenCount id="9D12FD796B19FFBDAD79FA48FD493F71" box="[615,663,546,573]" pageId="120" pageNumber="121" type="generic">25L</specimenCount>
|
||
</figureCitation>
|
||
. Female unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6B19FFBDAFD9FA2CFB4F3FC9" pageId="120" pageNumber="121" type="diagnosis">
|
||
<paragraph id="8BAB36F06B19FFBDAFD9FA2CFB4F3FC9" blockId="120.[151,1437,151,753]" pageId="120" pageNumber="121">
|
||
<emphasis id="B960EAE26B19FFBDAFD9FA2CFE9C3F2C" bold="true" box="[199,322,582,608]" pageId="120" pageNumber="121">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C144D736B19FFBDAE55FA2DFC973F2D" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[331,841,583,609]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="yakouensis" status="sp. nov.">
|
||
<emphasis id="B960EAE26B19FFBDAE55FA2DFDE33F2D" box="[331,573,583,609]" italics="true" pageId="120" pageNumber="121">Maladera yakouensis</emphasis>
|
||
Ahrens, Fabrizi & Liu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25357996B19FFBDAC48FA2DFC4A3F2D" box="[854,916,583,609]" pageId="120" pageNumber="121" rank="species">sp. n.</taxonomicNameLabel>
|
||
differs from all other large species of the
|
||
<taxonomicName id="4C144D736B19FFBDAA60FA2DFF323FC8" baseAuthorityName="Arrow" baseAuthorityYear="1946" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="egregia">
|
||
<emphasis id="B960EAE26B19FFBDAA60FA2DFF323FC8" italics="true" pageId="120" pageNumber="121">M. egregia</emphasis>
|
||
</taxonomicName>
|
||
group in that the fused left and right parameres have two pronounced, pointed tips.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6B19FFBDAFD9FAE5FE983F81" pageId="120" pageNumber="121" type="etymology">
|
||
<paragraph id="8BAB36F06B19FFBDAFD9FAE5FE983F81" blockId="120.[151,1437,151,753]" pageId="120" pageNumber="121">
|
||
<emphasis id="B960EAE26B19FFBDAFD9FAE5FE903FE5" bold="true" box="[199,334,655,681]" pageId="120" pageNumber="121">Etymology.</emphasis>
|
||
The name (adjective in the nominative singular case) of the new species is derived from its
|
||
<typeStatus id="54AF88526B19FFBDAA72FAE5FA423FE5" box="[1388,1436,655,681]" pageId="120" pageNumber="121">type</typeStatus>
|
||
locality, Yakou.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6B19FFBDAFD9FABCFD1E3FBD" box="[199,704,726,753]" pageId="120" pageNumber="121" type="distribution">
|
||
<paragraph id="8BAB36F06B19FFBDAFD9FABCFD1E3FBD" blockId="120.[151,1437,151,753]" box="[199,704,726,753]" pageId="120" pageNumber="121">
|
||
<emphasis id="B960EAE26B19FFBDAFD9FABCFEBF3FBC" bold="true" box="[199,353,726,752]" pageId="120" pageNumber="121">Distribution.</emphasis>
|
||
See map (
|
||
<figureCitation id="132F2A756B19FFBDAEC8FABDFDF63FBD" box="[470,552,727,753]" captionStart="FIGURE 98" captionStartId="350.[151,250,1969,1993]" captionTargetBox="[209,1379,1124,1940]" captionTargetId="figure-61@350.[202,1386,1119,1945]" captionTargetPageId="350" captionText="FIGURE 98. Distribution of the Maladera egregia group: Maladera pseudoegregia sp. n., M. tiachiensis sp. n., M. tiani sp. n., M. yakouensis sp. n., M. yipinglangensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496580" httpUri="https://zenodo.org/record/4496580/files/figure.png" pageId="120" pageNumber="121">Fig. 98</figureCitation>
|
||
) and
|
||
<tableCitation id="C696034B6B19FFBDAD78FABDFD643FBD" box="[614,698,727,753]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant & Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="120" pageNumber="121">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |