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<document id="6DADC61855E55A149F108DDFF3CB6472" ID-CLB-Dataset="3511" ID-DOI="10.11646/zootaxa.5068.2.6" ID-GBIF-Dataset="e17326ea-9ad5-4f11-b8d0-3e865248d420" ID-ISSN="1175-5326" ID-Zenodo-Dep="5704375" ID-ZooBank="6385D6D2-9442-4626-BA3C-65504B22CD24" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637052795992" checkinUser="plazi" docAuthor="Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M. &amp; Binoy, C." docDate="2021" docId="03BA4074F366FF9EFF168CC5B68DE396" docLanguage="en" docName="zootaxa.5068.2.6.pdf" docOrigin="Zootaxa 5068 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spilomena Shuckard 1838" docType="key" docVersion="7" lastPageNumber="265" masterDocId="FF83380CF367FF9CFF818861B311E371" masterDocTitle="A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India" masterLastPageNumber="276" masterPageNumber="263" pageNumber="264" updateTime="1699468628337" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A9DFA1BDEC06FD920F31483450C8F17F">A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India</mods:title>
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<mods:namePart id="AB56EF37221FB93C79F52917B63AC26B">Rajan, Tessy</mods:namePart>
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<mods:affiliation id="8D094535CF1C223CC7A6E79C39B80BC0">Western Ghat Regional Centre, Zoological Survey of India, Eranhipalam, Kozhikode, Kerala- 673006, India. kpgiris @ gmail. com; https: // orcid. org / 0000 - 0003 - 2121 - 0165</mods:affiliation>
<mods:nameIdentifier id="A76A95C5BAC1E647AFADC200841F3866" type="email">kpgiris@gmail.com</mods:nameIdentifier>
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<mods:namePart id="831867C79DDEDE107FEA5A2DF28AF1CA">Kumar, P. Girish</mods:namePart>
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<mods:affiliation id="EB6E4F81B7346C235C1827859905A6A2">Western Ghat Regional Centre, Zoological Survey of India, Eranhipalam, Kozhikode, Kerala- 673006, India. kpgiris @ gmail. com; https: // orcid. org / 0000 - 0003 - 2121 - 0165</mods:affiliation>
<mods:nameIdentifier id="53AC4164303B6FAA85452B76512B5D56" type="email">kpgiris@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FDF2F057C5A64B4EEC870A354E2C35B6">Sureshan, P. M.</mods:namePart>
<mods:nameIdentifier id="AAE55CD2CB34B66C0E9ACEED59B8E564" type="ORCID">0000-0003-2121-0165</mods:nameIdentifier>
<mods:affiliation id="893C0A9CDEE8C0A16C974BA6092F97B8">Western Ghat Regional Centre, Zoological Survey of India, Eranhipalam, Kozhikode, Kerala- 673006, India. kpgiris @ gmail. com; https: // orcid. org / 0000 - 0003 - 2121 - 0165</mods:affiliation>
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<mods:namePart id="5F43AF13034CC7306BC6015EE4615FC5">Binoy, C.</mods:namePart>
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<mods:affiliation id="05445518F48A36DC2A15F4EFF570B00A">Insect Ecology and Ethology Laboratory, Department of Zoology, University of Calicut, Kerala- 673635, India. binoy _ doz @ uoc. ac. in; https: // orcid. org / 0000 - 0003 - 4309 - 5460 &amp; Systematic Entomology Laboratory, Malabar Christian College, Kozhikode, Kerala- 673001, India, Affiliated to University of Calicut.</mods:affiliation>
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<mods:title id="B0E8274C19A0F45A351B79FA7D7CF48C">Zootaxa</mods:title>
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<mods:date id="5B2BD7C76750A1247AD5F2CA7D5D964E">2021</mods:date>
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<treatment id="03BA4074F366FF9EFF168CC5B68DE396" ID-DOI="http://doi.org/10.5281/zenodo.5705420" ID-GBIF-Taxon="190304477" ID-Zenodo-Dep="5705420" LSID="urn:lsid:plazi:treatment:03BA4074F366FF9EFF168CC5B68DE396" httpUri="http://treatment.plazi.org/id/03BA4074F366FF9EFF168CC5B68DE396" lastPageId="2" lastPageNumber="265" pageId="1" pageNumber="264">
<subSubSection id="C309A2E9F366FF9DFF168CC5B1E8E7CF" box="[151,761,1187,1214]" pageId="1" pageNumber="264" type="nomenclature">
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<heading id="D0E4460EF366FF9DFF168CC5B1E8E7CF" bold="true" box="[151,761,1187,1214]" fontSize="11" level="1" pageId="1" pageNumber="264" reason="1">
<emphasis id="B9672D70F366FF9DFF168CC5B1E8E7CF" bold="true" box="[151,761,1187,1214]" pageId="1" pageNumber="264">
Key to the Indian species of
<taxonomicName id="4C138AE1F366FF9DFE768CC2B1E8E7CF" ID-CoL="7KZM" authority="Shuckard" authorityName="Shuckard" authorityYear="1838" box="[503,761,1187,1214]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9672D70F366FF9DFE768CC2B169E7CF" bold="true" box="[503,632,1187,1214]" italics="true" pageId="1" pageNumber="264">Spilomena</emphasis>
Shuckard
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BACF162F366FF9DFF168C88B68DE78F" blockId="1.[151,1437,1257,2042]" box="[151,1436,1257,1279]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168C88B68DE78F" box="[151,1436,1257,1279]" pageId="1" pageNumber="264">1. Inner mandibular tooth broad and blunt.................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BACF162F366FF9DFF168D66B68CE66D" blockId="1.[151,1437,1257,2042]" box="[151,1437,1286,1308]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168D66B68CE66D" box="[151,1437,1286,1308]" pageId="1" pageNumber="264">
— Inner mandibular tooth pointed.............
<taxonomicName id="4C138AE1F366FF9DFD2B8D67B79BE66A" authority="Tessy, Sureshan et Girish Kumar" authorityName="Tessy, Sureshan et Girish Kumar" box="[682,1162,1286,1308]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="keralaensis">
<emphasis id="B9672D70F366FF9DFD2B8D67B79BE66A" bold="true" box="[682,1162,1286,1308]" pageId="1" pageNumber="264">
<emphasis id="B9672D70F366FF9DFD2B8D67B022E66D" bold="true" box="[682,819,1286,1308]" italics="true" pageId="1" pageNumber="264">S. keralaensis</emphasis>
Tessy, Sureshan et Girish Kumar
</emphasis>
</taxonomicName>
(
<figureCitation id="1328EDE7F366FF9DFB168D67B7C1E66A" box="[1175,1232,1286,1308]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Fig. 2</figureCitation>
of
<bibRefCitation id="EF828C93F366FF9DFB718D67B684E66D" author="Rajan, T. &amp; Sureshan, P. M. &amp; Girish Kumar, P. &amp; Mawadda, A. N. V." box="[1264,1429,1286,1308]" pageId="1" pageNumber="264" pagination="157 - 162" refId="ref6878" refString="Rajan, T., Sureshan, P. M., Girish Kumar, P. &amp; Mawadda, A. N. V. (2018) A taxonomic review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae: Pemphredoninae) from the Indian subcontinent with the description of a new species from Southern Western Ghats. Journal of Insect Biodiversity and Systematics, 4 (3), 157 - 162." type="journal article" year="2018">
Rajan
<emphasis id="B9672D70F366FF9DFAAF8D66B648E66A" box="[1326,1369,1286,1308]" italics="true" pageId="1" pageNumber="264">et al</emphasis>
. 2018
</bibRefCitation>
)
</keyLead>
</paragraph>
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<keyStep id="30E7E9C7F366FF9DFF168D45B68DE6E0" pageId="1" pageNumber="264">
<paragraph id="8BACF162F366FF9DFF168D45B68DE627" blockId="1.[151,1437,1257,2042]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168D45B68DE627" pageId="1" pageNumber="264">
2. Interantennal tubercle not distinctly pointed (
<figureCitation id="1328EDE7F366FF9DFD0E8D45B1DCE648" box="[655,717,1316,1338]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Figs 2</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFD578D45B1E1E648" box="[726,752,1316,1337]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1822,1846]" captionTargetBox="[164,1429,357,1794]" captionTargetId="figure-62@4.[159,1429,344,1800]" captionTargetPageId="4" captionText="FIGURES 9–16. Spilomena fulvopleuris Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 9. Habitus, lateral view; 10. Head, frontal view; 11. Head and mesosoma (in part), dorsal view; 12. Mesosoma, lateral view; 13. Propodeum, dorsal view; 14. Metasoma, dorsal view; 15. Fore wing; 16. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704381" httpUri="https://zenodo.org/record/5704381/files/figure.png" pageId="1" pageNumber="264">10</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFD788D45B002E648" box="[761,787,1316,1337]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1878,1902]" captionTargetBox="[174,1411,611,1834]" captionTargetId="figure-105@5.[151,1435,586,1856]" captionTargetPageId="5" captionText="FIGURES 17–20. Spilomena indostana Turner. 17. Habitus, lateral view with Type Label; 18. Head, frontal view; 19. Head, dorsal view; 20. Mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5704385" httpUri="https://zenodo.org/record/5704385/files/figure.png" pageId="1" pageNumber="264">18</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFC9D8D45B027E648" box="[796,822,1316,1337]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1631,1655]" captionTargetBox="[189,1398,208,1600]" captionTargetId="figure-17@7.[182,1404,201,1607]" captionTargetPageId="7" captionText="FIGURES 21–28. Spilomena reticularis Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 21. Habitus, lateral view; 22. Head, frontal view; 23. Head and mesosoma (in part), dorsal view; 24. Mesosoma, lateral view; 25. Propodeum, dorsal view; 26. Metasoma, dorsal view; 27. Fore wing; 28. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704387" httpUri="https://zenodo.org/record/5704387/files/figure.png" pageId="1" pageNumber="264">22</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFCC18D45B049E648" box="[832,856,1316,1337]" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">30</figureCitation>
&amp;
<figureCitation id="1328EDE7F366FF9DFCF48D45B081E64B" box="[885,912,1316,1338]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="1" pageNumber="264">38</figureCitation>
); mesosoma black or brownish black (
<figureCitation id="1328EDE7F366FF9DFB7D8D45B629E648" box="[1276,1336,1316,1338]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Figs 1</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFAC58D45B643E648" box="[1348,1362,1316,1337]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1822,1846]" captionTargetBox="[164,1429,357,1794]" captionTargetId="figure-62@4.[159,1429,344,1800]" captionTargetPageId="4" captionText="FIGURES 9–16. Spilomena fulvopleuris Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 9. Habitus, lateral view; 10. Head, frontal view; 11. Head and mesosoma (in part), dorsal view; 12. Mesosoma, lateral view; 13. Propodeum, dorsal view; 14. Metasoma, dorsal view; 15. Fore wing; 16. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704381" httpUri="https://zenodo.org/record/5704381/files/figure.png" pageId="1" pageNumber="264">9</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFADA8D45B664E648" box="[1371,1397,1316,1337]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1878,1902]" captionTargetBox="[174,1411,611,1834]" captionTargetId="figure-105@5.[151,1435,586,1856]" captionTargetPageId="5" captionText="FIGURES 17–20. Spilomena indostana Turner. 17. Habitus, lateral view with Type Label; 18. Head, frontal view; 19. Head, dorsal view; 20. Mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5704385" httpUri="https://zenodo.org/record/5704385/files/figure.png" pageId="1" pageNumber="264">17</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFAFF8D45B689E648" box="[1406,1432,1316,1337]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1631,1655]" captionTargetBox="[189,1398,208,1600]" captionTargetId="figure-17@7.[182,1404,201,1607]" captionTargetPageId="7" captionText="FIGURES 21–28. Spilomena reticularis Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 21. Habitus, lateral view; 22. Head, frontal view; 23. Head and mesosoma (in part), dorsal view; 24. Mesosoma, lateral view; 25. Propodeum, dorsal view; 26. Metasoma, dorsal view; 27. Fore wing; 28. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704387" httpUri="https://zenodo.org/record/5704387/files/figure.png" pageId="1" pageNumber="264">21</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFF638D20B3EBE627" box="[226,250,1345,1366]" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">29</figureCitation>
&amp;
<figureCitation id="1328EDE7F366FF9DFE988D20B225E626" box="[281,308,1345,1367]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="1" pageNumber="264">37</figureCitation>
); body length
<quantity id="4CEB5C87F366FF9DFE3C8D20B12EE627" box="[445,575,1345,1367]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="4.6" metricValueMin="2.5" pageId="1" pageNumber="264" unit="mm" value="3.55" valueMax="4.6" valueMin="2.5">2.5 – 4.6 mm</quantity>
....................................................................... 3
</keyLead>
</paragraph>
<paragraph id="8BACF162F366FF9DFF168D3EB68DE6E0" blockId="1.[151,1437,1257,2042]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168D3EB68DE6E0" pageId="1" pageNumber="264">
— Interantennal tubercle acutely pointed (
<figureCitation id="1328EDE7F366FF9DFDDB8D3FB1B0E605" box="[602,673,1374,1396]" captionStart="FIGURES 45–52" captionStartId="11.[151,264,1683,1707]" captionTargetBox="[159,1428,186,1651]" captionTargetId="figure-17@11.[151,1435,179,1657]" captionTargetPageId="11" captionText="FIGURES 45–52. Spilomena tuberculata Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 45. Habitus, lateral view; 46. Head, frontal view; 47. Head and mesosoma (in part), dorsal view; 48. Mesosoma, lateral view; 49. Propodeum, dorsal view; 50. Metasoma, dorsal view; 51. Fore wing; 52. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704397" httpUri="https://zenodo.org/record/5704397/files/figure.png" pageId="1" pageNumber="264">Fig. 46</figureCitation>
); mesosoma brownish yellow with few black patches (
<figureCitation id="1328EDE7F366FF9DFB2C8D3FB7E4E605" box="[1197,1269,1374,1396]" captionStart="FIGURES 45–52" captionStartId="11.[151,264,1683,1707]" captionTargetBox="[159,1428,186,1651]" captionTargetId="figure-17@11.[151,1435,179,1657]" captionTargetPageId="11" captionText="FIGURES 45–52. Spilomena tuberculata Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 45. Habitus, lateral view; 46. Head, frontal view; 47. Head and mesosoma (in part), dorsal view; 48. Mesosoma, lateral view; 49. Propodeum, dorsal view; 50. Metasoma, dorsal view; 51. Fore wing; 52. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704397" httpUri="https://zenodo.org/record/5704397/files/figure.png" pageId="1" pageNumber="264">Fig. 49</figureCitation>
); body length
<quantity id="4CEB5C87F366FF9DFAFF8D3FB21DE6E0" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="264" unit="mm" value="5.6">5.6 mm</quantity>
..................................................
<emphasis id="B9672D70F366FF9DFCE28D1AB68DE6E0" bold="true" box="[867,1436,1403,1425]" pageId="1" pageNumber="264">
<taxonomicName id="4C138AE1F366FF9DFCE28D1AB654E6E0" authority="Tessy, Sureshan et Girish Kumar" authorityName="Tessy, Sureshan et Girish Kumar" authorityYear="2021" box="[867,1349,1403,1425]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="B9672D70F366FF9DFCE28D1AB0FCE6E0" bold="true" box="[867,1005,1403,1425]" italics="true" pageId="1" pageNumber="264">S. tuberculata</emphasis>
Tessy, Sureshan et Girish Kumar
</taxonomicName>
,
<taxonomicNameLabel id="A254900BF366FF9DFAD18D1DB68DE6E0" box="[1360,1436,1404,1425]" pageId="1" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<keyStep id="30E7E9C7F366FF9DFF168DF8B68DE577" pageId="1" pageNumber="264">
<paragraph id="8BACF162F366FF9DFF168DF8B68DE6BA" blockId="1.[151,1437,1257,2042]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168DF8B68DE6BA" pageId="1" pageNumber="264">
3. Posterior surface of propodeum without median carina (
<figureCitation id="1328EDE7F366FF9DFC828DF8B050E6DE" box="[771,833,1433,1455]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Figs 5</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFCCC8DF8B076E6DF" box="[845,871,1433,1454]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1822,1846]" captionTargetBox="[164,1429,357,1794]" captionTargetId="figure-62@4.[159,1429,344,1800]" captionTargetPageId="4" captionText="FIGURES 9–16. Spilomena fulvopleuris Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 9. Habitus, lateral view; 10. Head, frontal view; 11. Head and mesosoma (in part), dorsal view; 12. Mesosoma, lateral view; 13. Propodeum, dorsal view; 14. Metasoma, dorsal view; 15. Fore wing; 16. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704381" httpUri="https://zenodo.org/record/5704381/files/figure.png" pageId="1" pageNumber="264">13</figureCitation>
,
<figureCitation id="1328EDE7F366FF9DFCF08DF8B098E6DF" box="[881,905,1433,1454]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1878,1902]" captionTargetBox="[174,1411,611,1834]" captionTargetId="figure-105@5.[151,1435,586,1856]" captionTargetPageId="5" captionText="FIGURES 17–20. Spilomena indostana Turner. 17. Habitus, lateral view with Type Label; 18. Head, frontal view; 19. Head, dorsal view; 20. Mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5704385" httpUri="https://zenodo.org/record/5704385/files/figure.png" pageId="1" pageNumber="264">20</figureCitation>
&amp;
<figureCitation id="1328EDE7F366FF9DFC288DF8B0D5E6DE" box="[937,964,1433,1455]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="1" pageNumber="264">37</figureCitation>
); recurrent vein of fore wing interstitial (
<figureCitation id="1328EDE7F366FF9DFACE8DF8B685E6DE" box="[1359,1428,1433,1455]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1822,1846]" captionTargetBox="[164,1429,357,1794]" captionTargetId="figure-62@4.[159,1429,344,1800]" captionTargetPageId="4" captionText="FIGURES 9–16. Spilomena fulvopleuris Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 9. Habitus, lateral view; 10. Head, frontal view; 11. Head and mesosoma (in part), dorsal view; 12. Mesosoma, lateral view; 13. Propodeum, dorsal view; 14. Metasoma, dorsal view; 15. Fore wing; 16. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704381" httpUri="https://zenodo.org/record/5704381/files/figure.png" pageId="1" pageNumber="264">Fig. 15</figureCitation>
) or nearly so (
<figureCitation id="1328EDE7F366FF9DFEE18DD7B28DE6BD" box="[352,412,1462,1484]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Figs 7</figureCitation>
&amp;
<figureCitation id="1328EDE7F366FF9DFE3A8DD7B2C5E6BD" box="[443,468,1462,1484]" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">31</figureCitation>
) with first transverse cubital vein.................................................... 4
</keyLead>
</paragraph>
<paragraph id="8BACF162F366FF9DFF168DB5B68DE577" blockId="1.[151,1437,1257,2042]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168DB5B68DE577" pageId="1" pageNumber="264">
— Posterior surface of propodeum with median carina (
<figureCitation id="1328EDE7F366FF9DFD608DB5B03BE698" box="[737,810,1491,1514]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1631,1655]" captionTargetBox="[189,1398,208,1600]" captionTargetId="figure-17@7.[182,1404,201,1607]" captionTargetPageId="7" captionText="FIGURES 21–28. Spilomena reticularis Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 21. Habitus, lateral view; 22. Head, frontal view; 23. Head and mesosoma (in part), dorsal view; 24. Mesosoma, lateral view; 25. Propodeum, dorsal view; 26. Metasoma, dorsal view; 27. Fore wing; 28. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704387" httpUri="https://zenodo.org/record/5704387/files/figure.png" pageId="1" pageNumber="264">Figs 25</figureCitation>
&amp;
<figureCitation id="1328EDE7F366FF9DFCC98DB5B072E69B" box="[840,867,1492,1514]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="1" pageNumber="264">41</figureCitation>
); recurrent vein of fore wing received well within first submarginal cell (
<figureCitation id="1328EDE7F366FF9DFEED8D90B2A4E577" box="[364,437,1521,1543]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1631,1655]" captionTargetBox="[189,1398,208,1600]" captionTargetId="figure-17@7.[182,1404,201,1607]" captionTargetPageId="7" captionText="FIGURES 21–28. Spilomena reticularis Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 21. Habitus, lateral view; 22. Head, frontal view; 23. Head and mesosoma (in part), dorsal view; 24. Mesosoma, lateral view; 25. Propodeum, dorsal view; 26. Metasoma, dorsal view; 27. Fore wing; 28. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704387" httpUri="https://zenodo.org/record/5704387/files/figure.png" pageId="1" pageNumber="264">Figs 27</figureCitation>
&amp;
<figureCitation id="1328EDE7F366FF9DFE528D90B2FFE576" box="[467,494,1521,1543]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="1" pageNumber="264">43</figureCitation>
)............................................................................. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E7E9C7F366FF9DFF168E6FB68DE533" pageId="1" pageNumber="264">
<paragraph id="8BACF162F366FF9DFF168E6FB68DE555" blockId="1.[151,1437,1257,2042]" box="[151,1436,1550,1572]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168E6FB68DE555" box="[151,1436,1550,1572]" pageId="1" pageNumber="264">
4. Recurrent vein of fore wing interstitial with first transverse cubital vein (
<figureCitation id="1328EDE7F366FF9DFC238E6FB0F8E555" box="[930,1001,1550,1572]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1822,1846]" captionTargetBox="[164,1429,357,1794]" captionTargetId="figure-62@4.[159,1429,344,1800]" captionTargetPageId="4" captionText="FIGURES 9–16. Spilomena fulvopleuris Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 9. Habitus, lateral view; 10. Head, frontal view; 11. Head and mesosoma (in part), dorsal view; 12. Mesosoma, lateral view; 13. Propodeum, dorsal view; 14. Metasoma, dorsal view; 15. Fore wing; 16. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704381" httpUri="https://zenodo.org/record/5704381/files/figure.png" pageId="1" pageNumber="264">Fig. 15</figureCitation>
).................................. 5
</keyLead>
</paragraph>
<paragraph id="8BACF162F366FF9DFF168E4DB68DE533" blockId="1.[151,1437,1257,2042]" box="[151,1436,1579,1602]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168E4DB68DE533" box="[151,1436,1579,1602]" pageId="1" pageNumber="264">
— Recurrent vein of fore wing nearly interstitial with first transverse cubital vein (
<figureCitation id="1328EDE7F366FF9DFC658E4DB70CE530" box="[996,1053,1580,1602]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Fig. 7</figureCitation>
&amp;
<figureCitation id="1328EDE7F366FF9DFBBD8E4AB746E530" box="[1084,1111,1579,1601]" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">35</figureCitation>
).......................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E7E9C7F366FF9DFF168E28B68DE5E8" pageId="1" pageNumber="264">
<paragraph id="8BACF162F366FF9DFF168E28B68DE52F" blockId="1.[151,1437,1257,2042]" box="[151,1436,1608,1631]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168E28B68DE52F" box="[151,1436,1608,1631]" pageId="1" pageNumber="264">
5. Body black (
<figureCitation id="1328EDE7F366FF9DFEE28E28B2BBE52E" box="[355,426,1609,1631]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1878,1902]" captionTargetBox="[174,1411,611,1834]" captionTargetId="figure-105@5.[151,1435,586,1856]" captionTargetPageId="5" captionText="FIGURES 17–20. Spilomena indostana Turner. 17. Habitus, lateral view with Type Label; 18. Head, frontal view; 19. Head, dorsal view; 20. Mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5704385" httpUri="https://zenodo.org/record/5704385/files/figure.png" pageId="1" pageNumber="264">Fig. 17</figureCitation>
); propodeum rugose (
<figureCitation id="1328EDE7F366FF9DFDFC8E28B1D2E52E" box="[637,707,1609,1631]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1878,1902]" captionTargetBox="[174,1411,611,1834]" captionTargetId="figure-105@5.[151,1435,586,1856]" captionTargetPageId="5" captionText="FIGURES 17–20. Spilomena indostana Turner. 17. Habitus, lateral view with Type Label; 18. Head, frontal view; 19. Head, dorsal view; 20. Mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5704385" httpUri="https://zenodo.org/record/5704385/files/figure.png" pageId="1" pageNumber="264">Fig. 20</figureCitation>
)..........................................
<taxonomicName id="4C138AE1F366FF9DFB528E28B68DE52F" ID-CoL="4Z453" authority="Turner" authorityName="Turner" box="[1235,1436,1608,1630]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="indostana">
<emphasis id="B9672D70F366FF9DFB528E28B68DE52F" bold="true" box="[1235,1436,1608,1630]" pageId="1" pageNumber="264">
<emphasis id="B9672D70F366FF9DFB528E28B65CE52F" bold="true" box="[1235,1357,1608,1630]" italics="true" pageId="1" pageNumber="264">S. indostana</emphasis>
Turner
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BACF162F366FF9DFF168E06B68DE5E8" blockId="1.[151,1437,1257,2042]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168E06B68DE5E8" pageId="1" pageNumber="264">
— Mesosoma black with mesoscutum laterally and mesopleuron yellowish brown (
<figureCitation id="1328EDE7F366FF9DFC6A8E07B737E50D" box="[1003,1062,1638,1660]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1822,1846]" captionTargetBox="[164,1429,357,1794]" captionTargetId="figure-62@4.[159,1429,344,1800]" captionTargetPageId="4" captionText="FIGURES 9–16. Spilomena fulvopleuris Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 9. Habitus, lateral view; 10. Head, frontal view; 11. Head and mesosoma (in part), dorsal view; 12. Mesosoma, lateral view; 13. Propodeum, dorsal view; 14. Metasoma, dorsal view; 15. Fore wing; 16. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704381" httpUri="https://zenodo.org/record/5704381/files/figure.png" pageId="1" pageNumber="264">Fig. 9</figureCitation>
); propodeum entirely transversely striate (
<figureCitation id="1328EDE7F366FF9DFE8A8EE5B242E5EB" box="[267,339,1668,1690]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1822,1846]" captionTargetBox="[164,1429,357,1794]" captionTargetId="figure-62@4.[159,1429,344,1800]" captionTargetPageId="4" captionText="FIGURES 9–16. Spilomena fulvopleuris Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 9. Habitus, lateral view; 10. Head, frontal view; 11. Head and mesosoma (in part), dorsal view; 12. Mesosoma, lateral view; 13. Propodeum, dorsal view; 14. Metasoma, dorsal view; 15. Fore wing; 16. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704381" httpUri="https://zenodo.org/record/5704381/files/figure.png" pageId="1" pageNumber="264">Fig. 12</figureCitation>
)..................................................
<emphasis id="B9672D70F366FF9DFC328EE2B68DE5E8" bold="true" box="[947,1436,1667,1689]" pageId="1" pageNumber="264">
<taxonomicName id="4C138AE1F366FF9DFC328EE2B654E5E8" authority="Tessy, Sureshan et Binoy" authorityName="Tessy, Sureshan et Binoy" authorityYear="2021" box="[947,1349,1667,1689]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="fulvopleuris" status="sp. nov.">
<emphasis id="B9672D70F366FF9DFC328EE2B752E5E8" bold="true" box="[947,1091,1667,1689]" italics="true" pageId="1" pageNumber="264">S. fulvopleuris</emphasis>
Tessy, Sureshan et Binoy
</taxonomicName>
,
<taxonomicNameLabel id="A254900BF366FF9DFAD18EE5B68DE5E8" box="[1360,1436,1668,1689]" pageId="1" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E7E9C7F366FF9DFF168EC0B68DE4D0" pageId="1" pageNumber="264">
<paragraph id="8BACF162F366FF9DFF168EC0B68DE45A" blockId="1.[151,1437,1257,2042]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168EC0B68DE45A" pageId="1" pageNumber="264">
6. Interantennal tubercle extending into long median carina reaching up to mid frons (
<figureCitation id="1328EDE7F366FF9DFB938EC0B75CE5C6" box="[1042,1101,1697,1719]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Fig. 2</figureCitation>
); anterior carina of pronotal collar deeply emarginate (
<figureCitation id="1328EDE7F366FF9DFE228EDFB2CFE5A5" box="[419,478,1726,1748]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Fig. 3</figureCitation>
); inner eye margin slightly concave (
<figureCitation id="1328EDE7F366FF9DFCCF8EDFB09AE5A5" box="[846,907,1726,1748]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Fig. 2</figureCitation>
); upper frons and vertex without setigerous punctures (
<figureCitation id="1328EDE7F366FF9DFF688EBDB235E583" box="[233,292,1756,1778]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Fig. 2</figureCitation>
); propodeal dorsum transversely striate, with two median longitudinal carinae; posterior surface of propodeum with irregular transverse ridges; lateral surface of propodeum finely aciculate (
<figureCitation id="1328EDE7F366FF9DFC148E98B0EFE47E" box="[917,1022,1785,1807]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1926,1950]" captionTargetBox="[183,1397,505,1891]" captionTargetId="figure-119@2.[183,1404,496,1900]" captionTargetPageId="2" captionText="FIGURES 1–8. Spilomena attenboroughi Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♂. 1. Habitus, lateral view; 2. Head, frontal view; 3. Head and mesosoma (in part), dorsal view; 4. Mesosoma, lateral view; 5. Propodeum, dorsal view; 6. Metasoma, dorsal view; 7. Fore wing; 8. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704379" httpUri="https://zenodo.org/record/5704379/files/figure.png" pageId="1" pageNumber="264">Figs 5 &amp; 4</figureCitation>
); body length
<quantity id="4CEB5C87F366FF9DFB088E98B7C4E47F" box="[1161,1237,1785,1807]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="264" unit="mm" value="3.5">3.5 mm</quantity>
..........................................................................
<emphasis id="B9672D70F366FF9DFC1C8F77B68DE45A" bold="true" box="[925,1436,1814,1836]" pageId="1" pageNumber="264">
<taxonomicName id="4C138AE1F366FF9DFC1C8F77B654E45D" authority="Tessy, Sureshan et Binoy" authorityName="Tessy, Sureshan et Binoy" authorityYear="2021" box="[925,1349,1814,1836]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="B9672D70F366FF9DFC1C8F77B752E45D" bold="true" box="[925,1091,1814,1836]" italics="true" pageId="1" pageNumber="264">S. attenboroughi</emphasis>
Tessy, Sureshan et Binoy
</taxonomicName>
,
<taxonomicNameLabel id="A254900BF366FF9DFAD18F77B68DE45A" box="[1360,1436,1814,1835]" pageId="1" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BACF162F366FF9DFF168F55B68DE4D0" blockId="1.[151,1437,1257,2042]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168F55B68DE4D0" pageId="1" pageNumber="264">
— Interantennal tubercle extending into short median carina, at most reaching lower frons (
<figureCitation id="1328EDE7F366FF9DFBC08F55B79BE43B" box="[1089,1162,1844,1866]" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">Fig. 30</figureCitation>
); anterior carina of pronotal collar straight (
<figureCitation id="1328EDE7F366FF9DFEF28F30B2ABE416" box="[371,442,1873,1895]" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">Fig. 31</figureCitation>
); inner eye margin straight (
<figureCitation id="1328EDE7F366FF9DFD4B8F30B000E416" box="[714,785,1873,1895]" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">Fig. 30</figureCitation>
); upper frons and vertex with setigerous punctures (
<figureCitation id="1328EDE7F366FF9DFA828F30B65AE416" box="[1283,1355,1873,1895]" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">Fig. 30</figureCitation>
); propodeal dorsum and posterior surface of propodeum rugose and rugulose, lateral surface of propodeum transversely striate (
<figureCitation id="1328EDE7F366FF9DFAF38F0FB225E4D3" captionStart="FIGURES 29–36" captionStartId="9.[151,264,1677,1701]" captionTargetBox="[159,1420,193,1648]" captionTargetId="figure-108@9.[151,1427,186,1653]" captionTargetPageId="9" captionText="FIGURES 29–36. Spilomena sahyadriensis Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♀. 29. Habitus, lateral view; 30. Head, frontal view; 31. Head and mesosoma (in part), dorsal view; 32. Mesosoma, lateral view; 33. Propodeum, dorsal view; 34. Metasoma, dorsal view; 35. Fore wing; 36. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704391" httpUri="https://zenodo.org/record/5704391/files/figure.png" pageId="1" pageNumber="264">Figs 33 &amp; 32</figureCitation>
); body length
<quantity id="4CEB5C87F366FF9DFE3C8FEDB118E4D0" box="[445,521,1932,1954]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="264" unit="mm" value="4.6">4.6 mm</quantity>
...........................
<emphasis id="B9672D70F366FF9DFCCF8FEAB68DE4D0" bold="true" box="[846,1436,1931,1953]" pageId="1" pageNumber="264">
<taxonomicName id="4C138AE1F366FF9DFCCF8FEAB654E4D0" authority="Tessy, Sureshan et Girish Kumar" authorityName="Tessy, Sureshan et Girish Kumar" authorityYear="2021" box="[846,1349,1931,1953]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="263" phylum="Arthropoda" rank="species" species="sahyadriensis" status="sp. nov.">
S.
<emphasis id="B9672D70F366FF9DFCE68FEAB0FFE4D0" bold="true" box="[871,1006,1931,1953]" italics="true" pageId="1" pageNumber="264">sahyadriensis</emphasis>
Tessy, Sureshan et Girish Kumar
</taxonomicName>
,
<taxonomicNameLabel id="A254900BF366FF9DFAD18FEDB68DE4D0" box="[1360,1436,1932,1953]" pageId="1" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E7E9C7F366FF9EFF168FC8B68DE396" lastPageId="2" lastPageNumber="265" pageId="1" pageNumber="264">
<paragraph id="8BACF162F366FF9DFF168FC8B68DE488" blockId="1.[151,1437,1257,2042]" pageId="1" pageNumber="264">
<keyLead id="30E25257F366FF9DFF168FC8B68DE488" pageId="1" pageNumber="264">
7. Mesopleuron finely reticulate, with several well-defined punctures (
<figureCitation id="1328EDE7F366FF9DFCF68FC8B0D1E4CE" box="[887,960,1961,1983]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1631,1655]" captionTargetBox="[189,1398,208,1600]" captionTargetId="figure-17@7.[182,1404,201,1607]" captionTargetPageId="7" captionText="FIGURES 21–28. Spilomena reticularis Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 21. Habitus, lateral view; 22. Head, frontal view; 23. Head and mesosoma (in part), dorsal view; 24. Mesosoma, lateral view; 25. Propodeum, dorsal view; 26. Metasoma, dorsal view; 27. Fore wing; 28. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704387" httpUri="https://zenodo.org/record/5704387/files/figure.png" pageId="1" pageNumber="264">Fig. 24</figureCitation>
); gena imbricate; frons reticulate; anterior carina of pronotal collar slightly emarginate (
<figureCitation id="1328EDE7F366FF9DFDD78FA7B18FE4AD" box="[598,670,1990,2012]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1631,1655]" captionTargetBox="[189,1398,208,1600]" captionTargetId="figure-17@7.[182,1404,201,1607]" captionTargetPageId="7" captionText="FIGURES 21–28. Spilomena reticularis Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 21. Habitus, lateral view; 22. Head, frontal view; 23. Head and mesosoma (in part), dorsal view; 24. Mesosoma, lateral view; 25. Propodeum, dorsal view; 26. Metasoma, dorsal view; 27. Fore wing; 28. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704387" httpUri="https://zenodo.org/record/5704387/files/figure.png" pageId="1" pageNumber="264">Fig. 23</figureCitation>
); stigma elongate (
<figureCitation id="1328EDE7F366FF9DFCD48FA7B08CE4AD" box="[853,925,1990,2012]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1631,1655]" captionTargetBox="[189,1398,208,1600]" captionTargetId="figure-17@7.[182,1404,201,1607]" captionTargetPageId="7" captionText="FIGURES 21–28. Spilomena reticularis Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 21. Habitus, lateral view; 22. Head, frontal view; 23. Head and mesosoma (in part), dorsal view; 24. Mesosoma, lateral view; 25. Propodeum, dorsal view; 26. Metasoma, dorsal view; 27. Fore wing; 28. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704387" httpUri="https://zenodo.org/record/5704387/files/figure.png" pageId="1" pageNumber="264">Fig. 27</figureCitation>
); propodeal enclosure with delimiting furrow; lateral surface of propodeum with transverse ridges (
<figureCitation id="1328EDE7F366FF9DFD1B8F85B00DE48B" box="[666,796,2019,2042]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1631,1655]" captionTargetBox="[189,1398,208,1600]" captionTargetId="figure-17@7.[182,1404,201,1607]" captionTargetPageId="7" captionText="FIGURES 21–28. Spilomena reticularis Tessy, Sureshan &amp; Binoy, sp. nov., Holotype ♀. 21. Habitus, lateral view; 22. Head, frontal view; 23. Head and mesosoma (in part), dorsal view; 24. Mesosoma, lateral view; 25. Propodeum, dorsal view; 26. Metasoma, dorsal view; 27. Fore wing; 28. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704387" httpUri="https://zenodo.org/record/5704387/files/figure.png" pageId="1" pageNumber="264">Figs 25 &amp; 24</figureCitation>
)..............
<emphasis id="B9672D70F366FF9DFC458F82B68DE488" bold="true" box="[964,1436,2019,2041]" pageId="1" pageNumber="264">
<taxonomicName id="4C138AE1F366FF9DFC458F82B654E488" authority="Tessy, Sureshan et Binoy" authorityName="Tessy, Sureshan et Binoy" authorityYear="2021" box="[964,1349,2019,2041]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="reticularis" status="sp. nov.">
<emphasis id="B9672D70F366FF9DFC458F82B753E488" bold="true" box="[964,1090,2019,2041]" italics="true" pageId="1" pageNumber="264">S. reticularis</emphasis>
Tessy, Sureshan et Binoy
</taxonomicName>
,
<taxonomicNameLabel id="A254900BF366FF9DFAD18F85B68DE488" box="[1360,1436,2020,2041]" pageId="1" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BACF162F365FF9EFF1688F9B68DE396" blockId="2.[151,1436,151,232]" pageId="2" pageNumber="265">
<keyLead id="30E25257F365FF9EFF1688F9B68DE396" pageId="2" pageNumber="265">
— Mesopleuron finely aciculate, with ill-defined punctures (
<figureCitation id="1328EDE7F365FF9EFC8788F6B05CE3DF" box="[774,845,151,174]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="2" pageNumber="265">Fig. 40</figureCitation>
); gena finely aciculate; frons imbricate; anterior carina of pronotal collar deeply emarginate (
<figureCitation id="1328EDE7F365FF9EFD9288D4B14AE3BA" box="[531,603,181,203]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="2" pageNumber="265">Fig. 39</figureCitation>
); stigma not elongate (
<figureCitation id="1328EDE7F365FF9EFCB688D4B06EE3BA" box="[823,895,181,203]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="2" pageNumber="265">Fig. 43</figureCitation>
); propodeal enclosure without delimiting furrow; lateral surface of propodeum finely aciculate (
<figureCitation id="1328EDE7F365FF9EFDDB88B3B1CAE399" box="[602,731,210,232]" captionStart="FIGURES 37–44" captionStartId="10.[151,264,1664,1688]" captionTargetBox="[158,1419,187,1639]" captionTargetId="figure-104@10.[151,1424,181,1645]" captionTargetPageId="10" captionText="FIGURES 37–44. Spilomena tsunekii Tessy, Sureshan &amp; Girish Kumar, sp. nov., Holotype ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Head and mesosoma (in part), dorsal view; 40. Mesosoma, lateral view; 41. Propodeum, dorsal view; 42. Metasoma, dorsal view; 43. Fore wing; 44. Hind wing." figureDoi="http://doi.org/10.5281/zenodo.5704393" httpUri="https://zenodo.org/record/5704393/files/figure.png" pageId="2" pageNumber="265">Figs 41 &amp; 40</figureCitation>
) ..............
<emphasis id="B9672D70F365FF9EFC0788B3B68DE396" bold="true" box="[902,1436,210,232]" pageId="2" pageNumber="265">
<taxonomicName id="4C138AE1F365FF9EFC0788B3B654E396" authority="Tessy, Sureshan et Girish Kumar" authorityName="Tessy, Sureshan et Girish Kumar" authorityYear="2021" box="[902,1349,210,232]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="tsunekii" status="sp. nov.">
S.
<emphasis id="B9672D70F365FF9EFC1E88B3B0FFE399" bold="true" box="[927,1006,210,232]" italics="true" pageId="2" pageNumber="265">tsunekii</emphasis>
Tessy, Sureshan et Girish Kumar
</taxonomicName>
,
<taxonomicNameLabel id="A254900BF365FF9EFAD188B3B68DE396" box="[1360,1436,210,231]" pageId="2" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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