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<document id="CD08E4AE73AA0DCDB33879BF6BBE1900" ID-DOI="10.11646/phytotaxa.266.1.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13669606" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725396564280" checkinUser="felipe" docAuthor="Jiménez-Mejías, Pedro &amp; Roalson, Eric H." docDate="2016" docId="03B6B9172664FFBDFF12018EA1F10161" docLanguage="en" docName="phytotaxa.266.1.3.pdf" docOrigin="Phytotaxa 266 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.266.1.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Carex haematopus Jim., sp. nov." docType="treatment" docVersion="2" lastPageNumber="26" masterDocId="FF8FC16F2666FFB8FF9A046EA45A001F" masterDocTitle="Two new species of Carex (Cyperaceae) from northern South America" masterLastPageNumber="26" masterPageNumber="21" pageNumber="23" updateTime="1725418897945" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0FB888B6E3C06889423031EB73C94CD7">Two new species of Carex (Cyperaceae) from northern South America</mods:title>
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<mods:namePart id="A9C271A613908EE870AC8B66E3D785A3">Jiménez-Mejías, Pedro</mods:namePart>
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<mods:namePart id="66B6528FAC98AC2F477D1983D055A0C5">Roalson, Eric H.</mods:namePart>
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<mods:title id="5CDCC241968031BDF706E75E01253191">Phytotaxa</mods:title>
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<mods:date id="66D375E50A64739D7D0BABB1535BD6C4">2016</mods:date>
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<treatment id="03B6B9172664FFBDFF12018EA1F10161" ID-DOI="http://doi.org/10.5281/zenodo.13669602" ID-Zenodo-Dep="13669602" LSID="urn:lsid:plazi:treatment:03B6B9172664FFBDFF12018EA1F10161" httpUri="http://treatment.plazi.org/id/03B6B9172664FFBDFF12018EA1F10161" lastPageId="5" lastPageNumber="26" pageId="2" pageNumber="23">
<subSubSection id="C3055B8A2664FFBAFF12018EA74005E4" box="[136,794,1504,1531]" pageId="2" pageNumber="23" type="nomenclature">
<paragraph id="8BA008012664FFBAFF12018EA74005E4" blockId="2.[136,794,1504,1531]" box="[136,794,1504,1531]" pageId="2" pageNumber="23">
<heading id="D0E8BF6D2664FFBAFF12018EA74005E4" bold="true" box="[136,794,1504,1531]" fontSize="11" level="1" pageId="2" pageNumber="23" reason="1">
<emphasis id="B96BD4132664FFBAFF12018EA74005E4" bold="true" box="[136,794,1504,1531]" pageId="2" pageNumber="23">
<taxonomicName id="4C1F73822664FFBAFF12018EA5C105E4" ID-CoL="R9JN" authority="Jim." authorityName="Jim." box="[136,411,1504,1531]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="species" species="haematopus">
<emphasis id="B96BD4132664FFBAFF12018EA53B05E4" bold="true" box="[136,353,1504,1531]" italics="true" pageId="2" pageNumber="23">Carex haematopus</emphasis>
Jim.
</taxonomicName>
-Mejías &amp; E.H.Roalson,
<taxonomicNameLabel id="A25869682664FFBAFD5B018CA74005E4" box="[705,794,1506,1531]" pageId="2" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3055B8A2664FFBCFF120243A72F053E" lastPageId="4" lastPageNumber="25" pageId="2" pageNumber="23" type="diagnosis">
<paragraph id="8BA008012664FFBAFF120243A0CC06CD" blockId="2.[136,1452,1581,1818]" pageId="2" pageNumber="23">
<emphasis id="B96BD4132664FFBAFF120243A4A9065D" bold="true" box="[136,243,1581,1602]" pageId="2" pageNumber="23">Diagnosis:</emphasis>
—This new species is similar to the South American species
<taxonomicName id="4C1F73822664FFBAFCA90243A066065D" authority="Desvaux" authorityName="Desvaux" box="[819,1084,1581,1602]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="species" species="aematorhyncha">
<emphasis id="B96BD4132664FFBAFCA90243A7B9065D" box="[819,995,1581,1602]" italics="true" pageId="2" pageNumber="23">C. aematorhyncha</emphasis>
Desvaux
</taxonomicName>
(in Gay 1854: 224) and
<taxonomicName id="4C1F73822664FFBAFAB90243A53C0679" authority="Nees (1840: 398)" authorityName="Nees" authorityPageNumber="398" authorityYear="1840" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="species" species="tweedieana">
<emphasis id="B96BD4132664FFBAFAB90243A1F1065D" box="[1315,1451,1581,1602]" italics="true" pageId="2" pageNumber="23">C. tweedieana</emphasis>
<bibRefCitation id="EF8E75F02664FFBAFF27023FA53C0679" author="Nees, C. G. D." box="[189,358,1617,1638]" pageId="2" pageNumber="23" pagination="397 - 399" refId="ref4167" refString="Nees, C. G. D. (1840) Cyperaceae a Tweedie in Bonariae lectae. Journal of Botany 2: 397 - 399." type="journal article" year="1840">Nees (1840: 398)</bibRefCitation>
</taxonomicName>
. From
<taxonomicName id="4C1F73822664FFBAFE33023FA6000679" box="[425,602,1617,1638]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="species" species="aematorhyncha">
<emphasis id="B96BD4132664FFBAFE33023FA6000679" box="[425,602,1617,1638]" italics="true" pageId="2" pageNumber="23">C. aematorhyncha</emphasis>
</taxonomicName>
the new species is easily distinguished by its smaller utricles (up to
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vs.
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; see
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). From the more robust
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<emphasis id="B96BD4132664FFBAFDCA021BA6800695" box="[592,730,1653,1674]" italics="true" pageId="2" pageNumber="23">C. tweedieana</emphasis>
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it is distinguished by its narrower leaves (up to
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wide vs.
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wide) and the inflorescence (all the spikes are spread along the upper part of the stems vs. the uppermost male and female spikes congested at the top of the stems) (see
<bibRefCitation id="EF8E75F02664FFBAFDAB02D3A74E06CD" author="Myndel-Pedersen, T." box="[561,788,1725,1746]" pageId="2" pageNumber="23" pagination="315 - 421" refId="ref4126" refString="Myndel-Pedersen, T. (1968) Cyperaceae. In: Cabrera, A. L. (Ed.) Flora de la Provincia de Buenos Aires I. INTA, Buenos Aires, pp. 315 - 421." type="book chapter" year="1968">Myndel-Pedersen 1968</bibRefCitation>
;
<bibRefCitation id="EF8E75F02664FFBAFC8702D3A0D106CD" author="Hoff Silveira, G. &amp; Longhi-Wagner, H. M." box="[797,1163,1725,1746]" pageId="2" pageNumber="23" pagination="373 - 417" refId="ref3894" refString="Hoff Silveira, G. &amp; Longhi-Wagner, H. M. (2012) O genero Carex L. (Cyperaceae) no Rio Grande do Sul, Brasil. Revista Brasileira de Biociencias 10: 373 - 417." type="journal article" year="2012">Hoff Silveira &amp; Longhi-Wagner 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8BA008012664FFBAFF12028FA76A0705" blockId="2.[136,1452,1581,1818]" pageId="2" pageNumber="23">
<emphasis id="B96BD4132664FFBAFF12028FA4EF06E9" box="[136,181,1761,1782]" italics="true" pageId="2" pageNumber="23">Type</emphasis>
:—
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.
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:
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Mpio
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, corregimiento del
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,
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,
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.
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<collectingDate id="EFE5D7292664FFBAFBD4028FA09F06E9" box="[1102,1221,1761,1782]" pageId="2" pageNumber="23" value="1977-11-01">1 Nov 1977</collectingDate>
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.
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<collectionCode id="ED0E90C42664FFBAFB4A028FA0BE06E9" box="[1232,1252,1761,1782]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="2" pageNumber="23" type="Herbarium">O</collectionCode>
.
<collectorName id="26EA6DD72664FFBAFB75028FA12A06E9" box="[1263,1392,1761,1782]" pageId="2" pageNumber="23">de Benavides</collectorName>
1197
</emphasis>
. (
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:
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!;
<typeStatus id="54A4B6A32664FFBAFEF9036BA5F20705" box="[355,424,1797,1818]" pageId="2" pageNumber="23" type="isotype">isotype</typeStatus>
material presumably at
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).
<figureCitation id="132414842664FFBAFD4A036BA7760705" box="[720,812,1797,1818]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[136,229,2059,2080]" captionStartId-1="4.[136,229,1003,1024]" captionTargetBox-1="[151,1433,190,977]" captionTargetId-1="figure-18@4.[151,1436,190,978]" captionTargetPageId-1="4" captionText-0="FIGURE 2. Holotype of Carex haematopus Jim.-Mejías &amp; E.H.Roalson." captionText-1="FIGURE 3. Detailed images of the holotype of Carex haematopus Jim.-Mejías &amp; E.H.Roalson. A Basal sheaths showing the ladderfibrosille splitting structures. BC Female spikes. D Male spike. E Male glumes. F Female glumes. G Utricle. H Achenes. Scale bar 0.5 cm (AD), 1 mm (EH)." figureDoi-1="http://doi.org/10.5281/zenodo.13669612" httpUri-1="https://zenodo.org/record/13669612/files/figure.png" pageId="2" pageNumber="23">Figs. 23</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA008012664FFBBFF120324A0C702CA" blockId="2.[136,1452,1866,2071]" lastBlockId="3.[136,1452,160,725]" lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23">
Perennial; rhizomes unknown, apparently long. Stems 61
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tall, 1.2
<quantity id="4CE7A5E42664FFBAFC5B0324A04D077C" box="[961,1047,1866,1891]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="23" unit="mm" value="0.8">0.8 mm</quantity>
wide at the middle, smooth, sharply trigonous. Leaves flat or V folded, apparently longer than the stems,&gt;
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long, 2.5
<quantity id="4CE7A5E42664FFBAFB1A0300A0990798" box="[1152,1219,1902,1927]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="23" unit="mm" value="2.0">2 mm</quantity>
wide at the middle, hypostomatic, antrorsely scabrid at the margins, apparently light green; ligule 4
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long, acute to subacute, orangey; sheaths reddish tinged, more intense as lower the leaf is, with the apical opening translucent-orangey on the side opposite to the blade; basal sheaths purple reddish, present at the base of the flowering and sterile shoots, ~
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long, entire, without blade, splitting into a ladder-fibrosille structure. Inflorescence racemose, 20
<quantity id="4CE7A5E42664FFBAFB700390A1750808" box="[1258,1327,2046,2071]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="23" unit="cm" value="13.0">13 cm</quantity>
long, with 23 proximal female spikes, and a terminal male spike; lowermost bract longer than the inflorescence, 25
<quantity id="4CE7A5E42665FFBBFAB004CEA13600A6" box="[1322,1388,160,185]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="24" unit="cm" value="12.0">12 cm</quantity>
long, 2.4
<quantity id="4CE7A5E42665FFBBFF2104AAA54B00C2" box="[187,273,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="24" unit="mm" value="1.4">1.4 mm</quantity>
wide at the middle, flat to V folded, sheathless or with a sheath up to
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long, with a small pair of auricles ~
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on the side opposite to the blade, dark-brown with a hyaline margin; second lowermost bract 8.5
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long, 1
<quantity id="4CE7A5E42665FFBBFE880562A533013A" box="[274,361,268,293]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="24" unit="mm" value="0.5">0.5 mm</quantity>
wide in its middle part, flat to filiform, sheathless, also with similar auricles at its base. Male spike oblanceolate, 3.7
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long, with a maximum width of 3
<quantity id="4CE7A5E42665FFBBFC85055EA7220156" box="[799,888,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="24" unit="mm" value="2.5">2.5 mm</quantity>
, with about 80 flowers. Lowermost female spikes 1714 ×
<quantity id="4CE7A5E42665FFBBFF71053AA5770172" box="[235,301,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="24" unit="mm" value="5.0">5 mm</quantity>
, ovate to lanceolate, with a peduncle 3.5
<quantity id="4CE7A5E42665FFBBFC9A053AA7640172" box="[768,830,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="24" unit="mm" value="3.0">3 mm</quantity>
long, with ~2040 flowers, and a tubular cladoprophyll at its base. Male glumes 4.24 ×
<quantity id="4CE7A5E42665FFBBFD8E0516A635018E" box="[532,623,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="24" unit="mm" value="1.3">1.3 mm</quantity>
, narrowly obovoid, reddish brownish with a lighter 13-nerved middle longitudinal strip, obtuse to rounded, cuspidate, with a mucro 0.3
<quantity id="4CE7A5E42665FFBBFCE905F2A79201AA" box="[883,968,412,437]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="24" unit="mm" value="0.1">0.1 mm</quantity>
long. Female glumes 2.72 × 1.6
<quantity id="4CE7A5E42665FFBBFAD505F2A1FD01AA" box="[1359,1447,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="24" unit="mm" value="1.2">1.2 mm</quantity>
, oblong, acute, purplish-brown, with a lighter 13 nerved middle longitudinal strip, excurrent into an antrorsely scabrid 0.6
<quantity id="4CE7A5E42665FFBBFF23058AA54801E2" box="[185,274,484,509]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="24" unit="mm" value="0.2">0.2 mm</quantity>
mucro. Stigmas 3, purplish, hispidolous. Utricles 3.53.2 × 1.4
<quantity id="4CE7A5E42665FFBBFC64058AA00201E2" box="[1022,1112,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="24" unit="mm" value="1.2">1.2 mm</quantity>
, broadly elliptical, with thick walls, reddish-orange, densely pubescent, with nerves marked but hardly visible below the hairs, cuneated at the base, contracted at the apex into a wide 0.80.6 × 0.5
<quantity id="4CE7A5E42665FFBBFD320642A6A4025A" box="[680,766,556,581]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="24" unit="mm" value="0.4">0.4 mm</quantity>
beak, purplish tinged, deeply bifid, with the apical teeth 0.6
<quantity id="4CE7A5E42665FFBBFF12063EA4870276" box="[136,221,592,617]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="24" unit="mm" value="0.3">0.3 mm</quantity>
long, divergent, forming a wide sinus from acute to almost rounded; pubescence of whitish hairs about 0.2
<quantity id="4CE7A5E42665FFBBFA10063EA4EE0292" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="24" unit="mm" value="0.1">0.1 mm</quantity>
long, more densely distributed towards the apex of the utricle body, being the base and the beak glabrous. Achenes 2.21.6 × 0.9
<quantity id="4CE7A5E42665FFBBFEB406F6A5DC02AE" box="[302,390,664,689]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="24" unit="mm" value="0.8">0.8 mm</quantity>
, elliptical, trigonous, cuneated at the base, sometimes ±contracted forming a short stipe; style strongly lignified, leaving a long cylindrical,
<quantity id="4CE7A5E42665FFBBFD1906D2A68302CA" box="[643,729,700,725]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="24" unit="mm" value="0.2">0.2 mm</quantity>
wide remnant at the apex of the achene.
</paragraph>
<caption id="DF6058892665FFBBFF120C65A713083F" box="[136,841,2059,2080]" pageId="3" pageNumber="24" startId="3.[136,229,2059,2080]" targetType="figure">
<paragraph id="8BA008012665FFBBFF120C65A713083F" blockId="3.[136,841,2059,2080]" box="[136,841,2059,2080]" pageId="3" pageNumber="24">
<emphasis id="B96BD4132665FFBBFF120C65A4A4083F" bold="true" box="[136,254,2059,2080]" pageId="3" pageNumber="24">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="4C1F73822665FFBBFEE70C65A605083F" authority="Jim." authorityName="Jim." box="[381,607,2059,2080]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="3" pageNumber="24" phylum="Tracheophyta" rank="species" species="haematopus">
<emphasis id="B96BD4132665FFBBFEE70C65A66A083F" box="[381,560,2059,2080]" italics="true" pageId="3" pageNumber="24">Carex haematopus</emphasis>
Jim.
</taxonomicName>
-Mejías &amp; E.H.Roalson.
</paragraph>
</caption>
<caption id="DF6058892662FFBCFF120785A52C0457" ID-DOI="http://doi.org/10.5281/zenodo.13669612" ID-Zenodo-Dep="13669612" httpUri="https://zenodo.org/record/13669612/files/figure.png" pageId="4" pageNumber="25" startId="4.[136,229,1003,1024]" targetBox="[151,1433,190,977]" targetPageId="4" targetType="figure">
<paragraph id="8BA008012662FFBCFF120785A52C0457" blockId="4.[136,1451,1003,1096]" pageId="4" pageNumber="25">
<emphasis id="B96BD4132662FFBCFF120785A55A041F" bold="true" box="[136,256,1003,1024]" pageId="4" pageNumber="25">FIGURE 3.</emphasis>
Detailed images of the holotype of
<taxonomicName id="4C1F73822662FFBCFDF20785A714041F" authority="Jim." authorityName="Jim." box="[616,846,1003,1024]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="haematopus">
<emphasis id="B96BD4132662FFBCFDF20785A744041F" box="[616,798,1003,1024]" italics="true" pageId="4" pageNumber="25">Carex haematopus</emphasis>
Jim.
</taxonomicName>
-Mejías &amp; E.H.Roalson.
<emphasis id="B96BD4132662FFBCFBDE0785A00F041F" bold="true" box="[1092,1109,1003,1024]" pageId="4" pageNumber="25">A</emphasis>
Basal sheaths showing the ladderfibrosille splitting structures.
<emphasis id="B96BD4132662FFBCFE3E0061A588043B" bold="true" box="[420,466,1039,1060]" pageId="4" pageNumber="25">BC</emphasis>
Female spikes.
<emphasis id="B96BD4132662FFBCFDF70061A624043B" bold="true" box="[621,638,1039,1060]" pageId="4" pageNumber="25">D</emphasis>
Male spike.
<emphasis id="B96BD4132662FFBCFD660061A756043B" bold="true" box="[764,780,1039,1060]" pageId="4" pageNumber="25">E</emphasis>
Male glumes.
<emphasis id="B96BD4132662FFBCFC060061A7F1043B" bold="true" box="[924,939,1039,1060]" pageId="4" pageNumber="25">F</emphasis>
Female glumes.
<emphasis id="B96BD4132662FFBCFBD50061A038043B" bold="true" box="[1103,1122,1039,1060]" pageId="4" pageNumber="25">G</emphasis>
Utricle.
<emphasis id="B96BD4132662FFBCFB2D0061A090043B" bold="true" box="[1207,1226,1039,1060]" pageId="4" pageNumber="25">H</emphasis>
Achenes. Scale bar 0.5 cm (AD), 1 mm (EH).
</paragraph>
</caption>
<paragraph id="8BA008012662FFBCFF270016A0BC04E2" blockId="4.[136,1452,1144,1997]" pageId="4" pageNumber="25">
<emphasis id="B96BD4132662FFBCFF270016A5B5048E" bold="true" box="[189,495,1144,1169]" pageId="4" pageNumber="25">Habitat and distribution:</emphasis>
—Known only from the
<typeStatus id="54A4B6A32662FFBCFC960016A766048E" box="[780,828,1144,1169]" pageId="4" pageNumber="25">type</typeStatus>
collection, a population in La Corota island, at Cocha lake, on the eastern side of the southern Colombian Andes, at an altitude of
<quantity id="4CE7A5E42662FFBCFC6F00F2A01504AA" box="[1013,1103,1180,1205]" metricMagnitude="3" metricUnit="m" metricValue="2.66" pageId="4" pageNumber="25" unit="m" value="2660.0">2660 m</quantity>
. The herbarium label does not provide any indication about ecology. Attending to the typology of the plant and the putatively related species, it may be a plant from moist-wet soils. Future findings in neighboring parts of the Andes may be expected.
</paragraph>
<paragraph id="8BA008012662FFBCFF270166A72F053E" blockId="4.[136,1452,1144,1997]" box="[189,885,1288,1313]" pageId="4" pageNumber="25">
<emphasis id="B96BD4132662FFBCFF270166A505053E" bold="true" box="[189,351,1288,1313]" pageId="4" pageNumber="25">Phenology:—</emphasis>
Flowers and fruits were collected in November.
</paragraph>
</subSubSection>
<subSubSection id="C3055B8A2662FFBCFF270142A6FE07D2" pageId="4" pageNumber="25" type="etymology">
<paragraph id="8BA008012662FFBCFF270142A09E0576" blockId="4.[136,1452,1144,1997]" pageId="4" pageNumber="25">
<emphasis id="B96BD4132662FFBCFF270142A512055A" bold="true" box="[189,328,1324,1349]" pageId="4" pageNumber="25">Etymology:</emphasis>
—The species epithet
<taxonomicName id="4C1F73822662FFBCFDD80142A68C055A" box="[578,726,1324,1349]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="haematopus">
<emphasis id="B96BD4132662FFBCFDD50142A68C055A" box="[591,726,1324,1349]" italics="true" pageId="4" pageNumber="25">haematopus</emphasis>
</taxonomicName>
” is formed by the ancient Greek “
<emphasis id="B96BD4132662FFBCFBC30142A099055A" box="[1113,1219,1324,1349]" italics="true" pageId="4" pageNumber="25">haemato-</emphasis>
”, blood, and “-pus”, foot. It refers to the purple reddish sheaths present at the base of the flowering and sterile shoots.
</paragraph>
<paragraph id="8BA008012662FFBCFF27011AA656065E" blockId="4.[136,1452,1144,1997]" pageId="4" pageNumber="25">
<emphasis id="B96BD4132662FFBCFF27011AA53F0592" bold="true" box="[189,357,1396,1421]" pageId="4" pageNumber="25">Observations:</emphasis>
—The hairy utricles with deeply bifid beak, sheathless to shortly sheathing bracts, and achenes with strongly lignifies styles (
<bibRefCitation id="EF8E75F02662FFBCFE0701F6A61A05AE" author="Egorova, T. V." box="[413,576,1432,1457]" pageId="4" pageNumber="25" refId="ref3656" refString="Egorova, T. V. (1999) Sedges (Carex L.) of Russia and Adjacent States (Within the Limits of the Former USSR). Missouri Botanical Garden Press, St. Louis, 772 pp." type="book" year="1999">Egorova 1999</bibRefCitation>
;
<bibRefCitation id="EF8E75F02662FFBCFDD701F6A6A505AE" author="Ball, P. W. &amp; Reznicek, A. A." box="[589,767,1432,1457]" pageId="4" pageNumber="25" pagination="254 - 273" refId="ref3548" refString="Ball, P. W. &amp; Reznicek, A. A. (2002) Carex Linnaeus. In: Flora of North America Editorial Committee (Eds.) Flora of North America, North of Mexico, 23. Magnoliophyta: Commelinidae (in part): Cyperaceae Oxford University Press, New York, NY, and Oxford, UK., pp. 254 - 273." type="book chapter" year="2002">
Ball
<emphasis id="B96BD4132662FFBCFD1901F6A6E105AE" box="[643,699,1432,1457]" italics="true" pageId="4" pageNumber="25">et al.</emphasis>
2002
</bibRefCitation>
), place this species within the assemblage of sections
<emphasis id="B96BD4132662FFBCFAC501F6A52705CA" italics="true" pageId="4" pageNumber="25">
<taxonomicName id="4C1F73822662FFBCFAC501F6A1F805AE" box="[1375,1442,1432,1457]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="genus">Carex</taxonomicName>
- Paludosae -Vesicariae
</emphasis>
(see
<bibRefCitation id="EF8E75F02662FFBCFE2D01D2A6BC05CA" author="Global Carex Group" box="[439,742,1468,1493]" pageId="4" pageNumber="25" refId="ref3780" refString="Global Carex Group (2016) Megaphylogenetic specimen-level approaches to the Carex (Cyperaceae) phylogeny using ITS, ETS, and matK sequences: Implications for classification. Systematic Botany." type="book" year="2016">
Global
<taxonomicName id="4C1F73822662FFBCFD9001D2A61705CA" box="[522,589,1468,1493]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B96BD4132662FFBCFD9001D2A61705CA" box="[522,589,1468,1493]" italics="true" pageId="4" pageNumber="25">Carex</emphasis>
</taxonomicName>
Group, 2016
</bibRefCitation>
). The hairy utricles have been the defining character for section
<taxonomicName id="4C1F73822662FFBCFF12018EA49105E6" box="[136,203,1504,1529]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B96BD4132662FFBCFF12018EA49105E6" box="[136,203,1504,1529]" italics="true" pageId="4" pageNumber="25">Carex</emphasis>
</taxonomicName>
, so in a classic sense
<taxonomicName id="4C1F73822662FFBCFE24018EA63C05E6" box="[446,614,1504,1529]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="haematopus">
<emphasis id="B96BD4132662FFBCFE24018EA63C05E6" box="[446,614,1504,1529]" italics="true" pageId="4" pageNumber="25">C. haematopus</emphasis>
</taxonomicName>
may be placed in this section. This character seems to be homoplasic, thus not defining a monophyletic section
<taxonomicName id="4C1F73822662FFBCFDB9026AA63C0602" box="[547,614,1540,1565]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B96BD4132662FFBCFDB9026AA63C0602" box="[547,614,1540,1565]" italics="true" pageId="4" pageNumber="25">Carex</emphasis>
</taxonomicName>
, although in case of merging the three sections cited above within a single one, it would be the priority name.
</paragraph>
<paragraph id="8BA008012662FFBCFF270222A637079A" blockId="4.[136,1452,1144,1997]" pageId="4" pageNumber="25">
<bibRefCitation id="EF8E75F02662FFBCFF270222A5DE067A" author="Kukenthal, G." box="[189,388,1612,1637]" pageId="4" pageNumber="25" pagination="1 - 814" refId="ref4025" refString="Kukenthal, G. (1909) Cyperaceae-Caricoideae. In: Engler, H. G. A. (Ed.) Das Pflanzenreich: regni vegetabilis conspectus IV (volume 20). Whilhelm Engelmann, Leipzig, pp. 1 - 814." type="book chapter" year="1909">Kükenthal (1909)</bibRefCitation>
cited
<taxonomicName id="4C1F73822662FFBCFE530222A6CD067A" box="[457,663,1612,1637]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aematorhyncha">
<emphasis id="B96BD4132662FFBCFE530222A6CD067A" box="[457,663,1612,1637]" italics="true" pageId="4" pageNumber="25">C. aematorhyncha</emphasis>
</taxonomicName>
from
<collectingCountry id="F30848912662FFBCFD460222A764067A" box="[732,830,1612,1637]" name="Ecuador" pageId="4" pageNumber="25">Ecuador</collectingCountry>
: “auf Triften bei Otavalo in der Prov.
<collectingRegion id="49DBC6E32662FFBCFB7F0222A10A067A" box="[1253,1360,1612,1637]" country="Ecuador" name="Imbabura" pageId="4" pageNumber="25">Imbabura</collectingRegion>
(Sodiro 199/64)”. It is the single reported
<taxonomicName id="4C1F73822662FFBCFDBA021EA6AE0696" box="[544,756,1648,1673]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aematorhyncha">
<emphasis id="B96BD4132662FFBCFDBA021EA6AE0696" box="[544,756,1648,1673]" italics="true" pageId="4" pageNumber="25">C. aematorhyncha</emphasis>
</taxonomicName>
population outside austral South America. In the Southern Cone, the northern limit of
<taxonomicName id="4C1F73822662FFBCFE5102FAA6C606B2" box="[459,668,1684,1709]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aematorhyncha">
<emphasis id="B96BD4132662FFBCFE5102FAA6C606B2" box="[459,668,1684,1709]" italics="true" pageId="4" pageNumber="25">C. aematorhyncha</emphasis>
</taxonomicName>
seems to be in the Argentinian province of Córdoba (see Flora del Cono Sur checklist at http://www.darwin.edu.ar/Proyectos/FloraArgentina/Generos.asp). The putative Ecuadorian
<taxonomicName id="4C1F73822662FFBCFA0802D6A56C06EA" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aematorhyncha">
<emphasis id="B96BD4132662FFBCFA0802D6A56C06EA" italics="true" pageId="4" pageNumber="25">C. aematorhyncha</emphasis>
</taxonomicName>
population is about
<quantity id="4CE7A5E42662FFBCFD8002B2A63506EA" box="[538,623,1756,1781]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="25" unit="km" value="150.0">150 km</quantity>
SW in straight line from the
<typeStatus id="54A4B6A32662FFBCFC2E02B2A7BE06EA" box="[948,996,1756,1781]" pageId="4" pageNumber="25">type</typeStatus>
locality of
<taxonomicName id="4C1F73822662FFBCFBFE02B2A15106EA" box="[1124,1291,1756,1781]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="haematopus">
<emphasis id="B96BD4132662FFBCFBFE02B2A15106EA" box="[1124,1291,1756,1781]" italics="true" pageId="4" pageNumber="25">C. haematopus</emphasis>
</taxonomicName>
, in the central Ecuadorian Andes. We requested materials of this collection from the herbaria that hold Sodiros material (see
<bibRefCitation id="EF8E75F02662FFBCFAC6036EA56C0722" author="Stafleu, F. A. &amp; Cowan, R. S." pageId="4" pageNumber="25" refId="ref4258" refString="Stafleu, F. A. &amp; Cowan, R. S. (1976 - 1997) Taxonomic literature, second edition, volumes 1 - 7. Utrecht, Bohn, Scheltema, and Holkema. Regnum Vegetabile 94, 98, 105, 110, 112, 115, 116." type="book" year="1976">Stafleu &amp; Cowan 1976</bibRefCitation>
1997). Herbaria B, E, G, K,
<collectingCountry id="F30848912662FFBCFDE5034AA6FC0722" box="[639,678,1828,1853]" name="Philippines" pageId="4" pageNumber="25">PH</collectingCountry>
, and S confirmed that they do not have such a specimen collected by Sodiro. Further study is necessary to locate the specimen cited by Kükenthal and confirm whether that Ecuadorian locality actually belongs to
<taxonomicName id="4C1F73822662FFBCFE240302A63F079A" box="[446,613,1900,1925]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="haematopus">
<emphasis id="B96BD4132662FFBCFE240302A63F079A" box="[446,613,1900,1925]" italics="true" pageId="4" pageNumber="25">C. haematopus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA008012662FFBCFF2703FEA6FE07D2" blockId="4.[136,1452,1144,1997]" pageId="4" pageNumber="25">
The following artificial key helps to distinguish the South American species of the putative section
<taxonomicName id="4C1F73822662FFBCFAAA03FEA12907B6" box="[1328,1395,1936,1961]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B96BD4132662FFBCFAAA03FEA12907B6" box="[1328,1395,1936,1961]" italics="true" pageId="4" pageNumber="25">Carex</emphasis>
</taxonomicName>
, and from all the other possible co-occurring species.
</paragraph>
</subSubSection>
<subSubSection id="C3055B8A2663FFBDFF1204CEA1F10161" pageId="5" pageNumber="26" type="key">
<key id="FF8E5F302663FFBDFF1204CEA1F10161" pageId="5" pageNumber="26">
<keyStep id="30EB10A42663FFBDFF1204CEA1F000F1" pageId="5" pageNumber="26">
<paragraph id="8BA008012663FFBDFF1204CEA1F000CE" blockId="5.[136,1452,160,382]" pageId="5" pageNumber="26">
<keyLead id="30EEAB342663FFBDFF1204CEA1F000CE" pageId="5" pageNumber="26">
1. Inflorescence with more than one spike, at least the uppermost one entirely male; utricles hairy, with a deeply bifid beak; stems elongated and well developed .................................................................................................................................. section
<taxonomicName id="4C1F73822663FFBDFAD504D2A1D300CE" ID-CoL="8VRHY" box="[1359,1417,188,209]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B96BD4132663FFBDFAD504D2A1D300CE" box="[1359,1417,188,209]" italics="true" pageId="5" pageNumber="26">Carex</emphasis>
</taxonomicName>
(2)
</keyLead>
</paragraph>
<paragraph id="8BA008012663FFBDFF1204B7A1F000F1" blockId="5.[136,1452,160,382]" box="[136,1450,217,238]" pageId="5" pageNumber="26">
<keyLead id="30EEAB342663FFBDFF1204B7A1F000F1" box="[136,1450,217,238]" pageId="5" pageNumber="26">
- Without the above combination of characters .....................................................................................................other
<taxonomicName id="4C1F73822663FFBDFA8404B7A10200F1" ID-CoL="8VRHY" box="[1310,1368,217,238]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B96BD4132663FFBDFA8404B7A10200F1" box="[1310,1368,217,238]" italics="true" pageId="5" pageNumber="26">Carex</emphasis>
</taxonomicName>
sections
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EB10A42663FFBDFF120498A1F1015A" pageId="5" pageNumber="26">
<paragraph id="8BA008012663FFBDFF120498A1F00137" blockId="5.[136,1452,160,382]" pageId="5" pageNumber="26">
<keyLead id="30EEAB342663FFBDFF120498A1F00137" pageId="5" pageNumber="26">
2. Inflorescence with the uppermost male and female spikes congested at the top of the stems, the male one sessile or subssesile...... .......................................................................................................................................................................................
<taxonomicName id="4C1F73822663FFBDFABB057DA1F00137" ID-CoL="5XCG8" authorityName="Nees" authorityYear="1840" box="[1313,1450,275,296]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="species" species="tweedieana">
<emphasis id="B96BD4132663FFBDFABB057DA1F00137" box="[1313,1450,275,296]" italics="true" pageId="5" pageNumber="26">C. tweedieana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA008012663FFBDFF12055EA1F1015A" blockId="5.[136,1452,160,382]" box="[136,1451,304,325]" pageId="5" pageNumber="26">
<keyLead id="30EEAB342663FFBDFF12055EA1F1015A" box="[136,1451,304,325]" pageId="5" pageNumber="26">- Inflorescence with the spikes spread along the upper part of the stems, the male one(s) conspicuously peduncled...................... (3)</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EB10A42663FFBDFF120522A1F10161" pageId="5" pageNumber="26">
<paragraph id="8BA008012663FFBDFF120522A1F1017E" blockId="5.[136,1452,160,382]" box="[136,1451,332,353]" pageId="5" pageNumber="26">
<keyLead id="30EEAB342663FFBDFF120522A1F1017E" box="[136,1451,332,353]" pageId="5" pageNumber="26">
3. Utricles
<quantity id="4CE7A5E42663FFBDFEBF0522A52E017E" box="[293,372,332,353]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="5" pageNumber="26" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
long .............................................................................................................................................
<taxonomicName id="4C1F73822663FFBDFB600522A1F1017E" ID-CoL="R7WY" box="[1274,1451,332,353]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="species" species="aematorhyncha">
<emphasis id="B96BD4132663FFBDFB600522A1F1017E" box="[1274,1451,332,353]" italics="true" pageId="5" pageNumber="26">C. aematorhyncha</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA008012663FFBDFF120507A1F10161" blockId="5.[136,1452,160,382]" box="[136,1451,361,382]" pageId="5" pageNumber="26">
<keyLead id="30EEAB342663FFBDFF120507A1F10161" box="[136,1451,361,382]" pageId="5" pageNumber="26">
- Utricles up to
<quantity id="4CE7A5E42663FFBDFEC10507A5FE0161" box="[347,420,361,382]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="26" unit="mm" value="3.5">3.5 mm</quantity>
long...........................................................................................................................................
<taxonomicName id="4C1F73822663FFBDFA810507A1F10161" ID-CoL="R9JN" box="[1307,1451,361,382]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="species" species="haematopus">
<emphasis id="B96BD4132663FFBDFA810507A1F10161" box="[1307,1451,361,382]" italics="true" pageId="5" pageNumber="26">C. haematopus</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>