259 lines
30 KiB
XML
259 lines
30 KiB
XML
<document id="853425E5CCA9E4C16DB727FA51607B6D" ID-CLB-Dataset="21388" ID-DOI="10.11646/zootaxa.2310.1.1" ID-GBIF-Dataset="1ffd1c6f-d374-4a5f-9d33-2c71ba54274a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3968627" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1596207919668" checkinUser="valdenar" docAuthor="Bernardes, Jorge L. C., Schwertner, Cristiano F. & Grazia, Jocélia" docDate="2009" docId="03B609203F1A894D53D2A65DFAD9E185" docLanguage="en" docName="Zootaxa.2310.1-23.pdf.imf" docOrigin="Zootaxa 2310" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thoreyella Bernardes & Schwertner & Grazia 2009" docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="FF8F71583F0B895C5345A567FF93E63D" masterDocTitle="Cladistic analysis of Thoreyella and related genera (Hemiptera: Pentatomidae: Pentatominae: Procleticini)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="18" updateTime="1698844036444" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="BAE0AA7BAA7B0F6165A7993F06DF9557" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4EC712A4DE2A19D27BFC59CE0BF0C81F">
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<mods:title id="75D71FB092F0FE14B8D676D742A6B63C">Cladistic analysis of Thoreyella and related genera (Hemiptera: Pentatomidae: Pentatominae: Procleticini)</mods:title>
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</mods:titleInfo>
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<mods:name id="73C022B67E4A0484F9C49F3A7371B19E" type="personal">
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<mods:roleTerm id="ED39C90FFC589F8B1AE7B6E1261C96B7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1523983D2FC207F54F3320041FCC1071">Bernardes, Jorge L. C.</mods:namePart>
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<mods:affiliation id="5499FA76FEB4BC62EF2A49A37466358A">Laboratório de Entomologia Sistemática, Departamento de Zoologia, Programa de Pós-Graduação em Biologia Animal, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves, 9500, Bloco IV, prédio 43435, 91501 - 970 Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="F42107DE5CF147C500A0C06530206177" type="email">jorgecabeleira@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="F259FD8A7AE87D933160FCD68DFA35F0" type="personal">
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<mods:role id="7C372B262BFBDE521999BEEB80A287D1">
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<mods:roleTerm id="540583E72A1B38C65CFABBDC1C4753DF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="07BA6962B347F15CC9979F2F4DA0EC25">Schwertner, Cristiano F.</mods:namePart>
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<mods:affiliation id="9E6E9D7636D339968F12DA21D79B32DB">Departamento de Ciências Biológicas, Universidade Federal de São Paulo, Campus Diadema, Rua Prof. Artur Riedel 275, Diadema, SP, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="D3FC266CE98A299ACB9F3EC48664A40A" type="email">schwertner@unifesp.br</mods:nameIdentifier>
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</mods:name>
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<mods:name id="110F56DC21091E27F7E4EE7BE3DA2360" type="personal">
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<mods:role id="0D32191447E8F58F2DCDB5B17E8CFFDC">
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<mods:roleTerm id="6FA8140C1594C67EE1BF88F301C86359">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D30A3E1C75E973A668E479C5F27F700F">Grazia, Jocélia</mods:namePart>
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<mods:affiliation id="470EF80035ED0FA1926523E7E1DFA296">Laboratório de Entomologia Sistemática, Departamento de Zoologia, Programa de Pós-Graduação em Biologia Animal, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves, 9500, Bloco IV, prédio 43435, 91501 - 970 Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="CAEBD8E91C445C96ACCA45D14BF4AB0A" type="email">jocelia@ufrgs.br</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="422BEA988D3AD37C8BE7F19F25C2D449">text</mods:typeOfResource>
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<mods:titleInfo id="04360D95AFBDE684F1EC6738DD439684">
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<mods:title id="B8C70A06A340DD6C47E248C3BC75E768">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3D1AB1C241860C221874A5FC8BF7B8BE">
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<mods:date id="DEB0E13767D5DF4ECC4BF9E837428336">2009</mods:date>
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<mods:detail id="ED1B1D38E18C1AE68107241B0462AA57" type="pubDate">
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<mods:number id="6AF9A0C0720C069D8F966133F8AB3DF0">2009-12-14</mods:number>
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</mods:detail>
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<mods:detail id="0CA0B3CC3605C971CB6235C6A5BCDA2B" type="volume">
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<mods:number id="CF9D3A90873BAA772C0AAD28E1826B88">2310</mods:number>
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</mods:detail>
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<mods:start id="29C941A56600381BAE038663B4AE4BCE">1</mods:start>
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<mods:end id="E8F8EFD31527B3EACF6C5BFE6C09099D">23</mods:end>
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<mods:classification id="0012A633D4D4DB245F1A6E12083E6FE9">journal article</mods:classification>
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<mods:identifier id="36610774BF74EE0FEE4AE84DDE8CA84F" type="DOI">10.11646/zootaxa.2310.1.1</mods:identifier>
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<mods:identifier id="EA7BDE160C5CD6A280B28552A33411E7" type="GBIF-Dataset">1ffd1c6f-d374-4a5f-9d33-2c71ba54274a</mods:identifier>
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<mods:identifier id="0C874C0DC01DCBF819F63358147CA443" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B609203F1A894D53D2A65DFAD9E185" ID-DOI="http://doi.org/10.5281/zenodo.4323846" ID-GBIF-Taxon="166018136" ID-Zenodo-Dep="4323846" LSID="urn:lsid:plazi:treatment:03B609203F1A894D53D2A65DFAD9E185" httpUri="http://treatment.plazi.org/id/03B609203F1A894D53D2A65DFAD9E185" lastPageNumber="18" pageId="17" pageNumber="18">
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<subSubSection id="C305EBBD3F1A894D53D2A65DFE72E569" box="[151,481,826,852]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8BA0B8363F1A894D53D2A65DFE72E569" blockId="17.[151,1437,826,1976]" box="[151,481,826,852]" pageId="17" pageNumber="18">
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Genus
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<taxonomicName id="4C1FC3B53F1A894D53ACA65CFEF4E569" authorityName="Bernardes & Schwertner & Grazia" authorityYear="2009" box="[233,359,827,852]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B96B64243F1A894D53ACA65CFEF4E569" box="[233,359,827,852]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
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</taxonomicName>
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(Clade B)
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</paragraph>
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</subSubSection>
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<subSubSection id="C305EBBD3F1A894D5383A606FAD9E185" pageId="17" pageNumber="18" type="description">
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<paragraph id="8BA0B8363F1A894D5383A606FE97E24F" blockId="17.[151,1437,826,1976]" pageId="17" pageNumber="18">
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The monophyletic condition of
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<taxonomicName id="4C1FC3B53F1A894D5103A606FD50E547" authorityName="Bernardes & Schwertner & Grazia" authorityYear="2009" box="[582,707,865,890]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B96B64243F1A894D5103A606FD50E547" box="[582,707,865,890]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
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</taxonomicName>
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was confirmed by the analysis, and was supported by three non-ambiguous synapomorphies: humeral angles not emarginated (
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<subScript id="179BBA733F1A894D5099A6EFFC67E59B" attach="none" box="[988,1012,904,934]" fontSize="6" pageId="17" pageNumber="18">81</subScript>
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); lateral margins of the scutellum straight (
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<subScript id="179BBA733F1A894D5246A6D5FEB9E5ED" attach="none" box="[259,298,946,976]" fontSize="6" pageId="17" pageNumber="18">141</subScript>
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); and apex of the abdominal spine curved (
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<subScript id="179BBA733F1A894D507FA6D5FCF1E5ED" attach="right" box="[826,866,946,976]" fontSize="6" pageId="17" pageNumber="18">241</subScript>
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). Also, three homoplasies help to diagnose the genus: lobe of the posterior margin of the bucculae rounded (
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<subScript id="179BBA733F1A894D502DA6BCFCECE5C4" attach="none" box="[872,895,987,1017]" fontSize="6" pageId="17" pageNumber="18">41</subScript>
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), also found in
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<taxonomicName id="4C1FC3B53F1A894D577FA6BCFB2FE5C9" authorityName="Berg" authorityYear="1891" box="[1082,1212,987,1012]" class="Insecta" genus="Lobepomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="peltifera">
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<emphasis id="B96B64243F1A894D577FA6BCFB2FE5C9" box="[1082,1212,987,1012]" italics="true" pageId="17" pageNumber="18">L. peltifera</emphasis>
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</taxonomicName>
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; lateral margins of propleura with two black macula (
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<subScript id="179BBA733F1A894D516DA162FDACE21E" attach="none" box="[552,575,1029,1059]" fontSize="6" pageId="17" pageNumber="18">92</subScript>
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), reversed to a black line (
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<subScript id="179BBA733F1A894D5030A162FC1EE21E" attach="right" box="[885,909,1029,1059]" fontSize="6" pageId="17" pageNumber="18">91</subScript>
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) in
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<taxonomicName id="4C1FC3B53F1A894D50F9A162FBCCE223" authorityName="Spinola" authorityYear="1852" box="[956,1119,1029,1054]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brasiliensis">
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<emphasis id="B96B64243F1A894D50F9A162FBCCE223" box="[956,1119,1029,1054]" italics="true" pageId="17" pageNumber="18">T. brasiliensis</emphasis>
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</taxonomicName>
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+
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<taxonomicName id="4C1FC3B53F1A894D5735A162FB60E223" authorityName="Berg" authorityYear="1878" box="[1136,1267,1029,1054]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trinotata">
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<emphasis id="B96B64243F1A894D5735A162FB60E223" box="[1136,1267,1029,1054]" italics="true" pageId="17" pageNumber="18">T. trinotata</emphasis>
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</taxonomicName>
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; and posterior margin of the female segment VII excavate in “U” (
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<subScript id="179BBA733F1A894D5046A149FCB9E271" attach="none" box="[771,810,1070,1100]" fontSize="6" pageId="17" pageNumber="18">251</subScript>
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), reversed to the plesiomorphic condition (
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<subScript id="179BBA733F1A894D566BA149FAC6E271" attach="none" box="[1326,1365,1070,1100]" fontSize="6" pageId="17" pageNumber="18">250</subScript>
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) in
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<taxonomicName id="4C1FC3B53F1A894D56C3A148FF6DE24C" authorityName="Berg" authorityYear="1878" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trinotata">
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<emphasis id="B96B64243F1A894D56C3A148FF6DE24C" italics="true" pageId="17" pageNumber="18">T. trinotata</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BA0B8363F1A894D5383A118FE19E32D" blockId="17.[151,1437,826,1976]" pageId="17" pageNumber="18">
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The first dichotomy among the species of
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<taxonomicName id="4C1FC3B53F1A894D51A1A118FCF1E2A5" authorityName="Bernardes & Schwertner & Grazia" authorityYear="2009" box="[740,866,1151,1176]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B96B64243F1A894D51A1A118FCF1E2A5" box="[740,866,1151,1176]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
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</taxonomicName>
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(
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<figureCitation id="1324A4B33F1A894D503DA118FC46E2A4" box="[888,981,1151,1177]" captionStart="FIGURE 54" captionStartId="16.[151,257,1708,1732]" captionTargetBox="[218,1351,1003,1672]" captionTargetId="figure@16.[188,1398,962,1718]" captionTargetPageId="16" captionText="FIGURE 54. Resulting cladogram of the cladistic analysis of Thoreyella and related genera. s: 71; ci: 0.73; ri: 0.80. (black circle = synapomorphy; white circle = homoplasy)." figureDoi="http://doi.org/10.5281/zenodo.3968650" httpUri="https://zenodo.org/record/3968650/files/figure.png" pageId="17" pageNumber="18">Fig. 54</figureCitation>
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) was supported by one homoplastic character: besides
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<taxonomicName id="4C1FC3B53F1A894D523CA1C1FE68E282" authority="Bernardes & Schwertner & Grazia, 2009" authorityName="Bernardes & Schwertner & Grazia" authorityYear="2009" box="[377,507,1190,1215]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B96B64243F1A894D523CA1C1FE68E282" box="[377,507,1190,1215]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A258D95F3F1A894D5146A1C2FDF1E282" box="[515,610,1189,1215]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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2, all other species of the genus have the surface of laterotergites 9 slightly convex (
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<subScript id="179BBA733F1A894D5224A1ABFE1BE2D7" attach="none" box="[353,392,1228,1258]" fontSize="6" pageId="17" pageNumber="18">331</subScript>
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) to strongly convex (
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<subScript id="179BBA733F1A894D51C8A1ABFD27E2D7" attach="none" box="[653,692,1228,1258]" fontSize="6" pageId="17" pageNumber="18">332</subScript>
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); the latter condition also supported the group
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<taxonomicName id="4C1FC3B53F1A894D57AAA1ABFAE6E2D8" authorityName="Berg" authorityYear="1891" box="[1263,1397,1228,1253]" class="Insecta" genus="Lobepomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="peltifera">
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<emphasis id="B96B64243F1A894D57AAA1ABFAE6E2D8" box="[1263,1397,1228,1253]" italics="true" pageId="17" pageNumber="18">L. peltifera</emphasis>
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</taxonomicName>
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+
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<taxonomicName id="4C1FC3B53F1A894D56C3A1ABFF6FE332" authorityName="Berg" authorityYear="1891" class="Insecta" genus="Procleticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="corniger">
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<emphasis id="B96B64243F1A894D56C3A1ABFF6FE332" italics="true" pageId="17" pageNumber="18">P. corniger</emphasis>
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</taxonomicName>
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+
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<taxonomicName id="4C1FC3B53F1A894D5249A191FE11E332" authorityName="Pennington" authorityYear="1922" box="[268,386,1270,1295]" class="Insecta" genus="Neoderoploa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bruchii">
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<emphasis id="B96B64243F1A894D5249A191FE11E332" box="[268,386,1270,1295]" italics="true" pageId="17" pageNumber="18">N. bruchii</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BA0B8363F1A894D5383A07BFEECE3EA" blockId="17.[151,1437,826,1976]" pageId="17" pageNumber="18">
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The clade
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<emphasis id="B96B64243F1A894D5204A07BFDA6E308" box="[321,565,1308,1333]" italics="true" pageId="17" pageNumber="18">
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<taxonomicName id="4C1FC3B53F1A894D5204A07BFE2FE308" authorityName="Berg" authorityYear="1883" box="[321,444,1308,1333]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cornuta">T. cornuta</taxonomicName>
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+
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<taxonomicName id="4C1FC3B53F1A894D5289A07BFDA6E308" authorityName="Jensen-Haarup" authorityYear="1931" box="[460,565,1308,1333]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="taurus">T. taurus</taxonomicName>
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</emphasis>
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share the dark sternum (
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<subScript id="179BBA733F1A894D501CA07BFCEDE307" attach="none" box="[857,894,1308,1338]" fontSize="6" pageId="17" pageNumber="18">151</subScript>
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), a synapomorphic condition. Its sister group, the clade including
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<taxonomicName id="4C1FC3B53F1A894D523EA021FE6AE362" authority="Bernardes & Schwertner & Grazia, 2009" authorityName="Bernardes & Schwertner & Grazia" authorityYear="2009" box="[379,505,1350,1375]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B96B64243F1A894D523EA021FE6AE362" box="[379,505,1350,1375]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A258D95F3F1A894D5145A021FDCAE35D" box="[512,601,1350,1376]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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1+
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<taxonomicName id="4C1FC3B53F1A894D513AA021FCB1E362" authorityName="Spinola" authorityYear="1852" box="[639,802,1350,1375]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brasiliensis">
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<emphasis id="B96B64243F1A894D513AA021FCB1E362" box="[639,802,1350,1375]" italics="true" pageId="17" pageNumber="18">T. brasiliensis</emphasis>
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</taxonomicName>
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+
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<emphasis id="B96B64243F1A894D5076A021FC2FE362" box="[819,956,1350,1375]" italics="true" pageId="17" pageNumber="18">
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<taxonomicName id="4C1FC3B53F1A894D5076A021FC26E362" authorityName="Berg" authorityYear="1878" box="[819,949,1350,1375]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trinotata">T. trinotata</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
shared three homoplasies: dorsal surface of the head convex (
|
|
<subScript id="179BBA733F1A894D52DDA00BFE3EE3B7" attach="none" box="[408,429,1388,1418]" fontSize="6" pageId="17" pageNumber="18">11</subScript>
|
|
) and lateral margin of the juga blackish (
|
|
<subScript id="179BBA733F1A894D50F0A00BFC5EE3B7" attach="none" box="[949,973,1388,1418]" fontSize="6" pageId="17" pageNumber="18">51</subScript>
|
|
), conditions also found in one of the outgroup taxa (
|
|
<taxonomicName id="4C1FC3B53F1A894D520EA0F1FE66E392" authorityName="Barber" authorityYear="1911" box="[331,501,1430,1455]" class="Insecta" family="Pentatomidae" genus="Dendrocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="arizonensis">
|
|
<emphasis id="B96B64243F1A894D520EA0F1FE66E392" box="[331,501,1430,1455]" italics="true" pageId="17" pageNumber="18">D. arizonensis</emphasis>
|
|
</taxonomicName>
|
|
); and laterotergites 8 entirely concave, reversed to the plesiomorphic condition (
|
|
<subScript id="179BBA733F1A894D53DAA0DAFF55E3E6" attach="none" box="[159,198,1469,1499]" fontSize="6" pageId="17" pageNumber="18">340</subScript>
|
|
) in
|
|
<taxonomicName id="4C1FC3B53F1A894D53B0A0DAFEEBE3EB" authorityName="Berg" authorityYear="1878" box="[245,376,1469,1494]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trinotata">
|
|
<emphasis id="B96B64243F1A894D53B0A0DAFEEBE3EB" box="[245,376,1469,1494]" italics="true" pageId="17" pageNumber="18">T. trinotata</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BA0B8363F1A894D5383A081FE0EE0C9" blockId="17.[151,1437,826,1976]" pageId="17" pageNumber="18">
|
|
The dichotomy between
|
|
<taxonomicName id="4C1FC3B53F1A894D52AFA080FDFBE03D" authority="Bernardes & Schwertner & Grazia, 2009" authorityName="Bernardes & Schwertner & Grazia" authorityYear="2009" box="[490,616,1511,1536]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" status="sp. nov.">
|
|
<emphasis id="B96B64243F1A894D52AFA080FDFBE03D" box="[490,616,1511,1536]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A258D95F3F1A894D5135A081FD58E03D" box="[624,715,1510,1536]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
|
1 and the clade
|
|
<taxonomicName id="4C1FC3B53F1A894D50C8A080FBA2E03D" authorityName="Spinola" authorityYear="1852" box="[909,1073,1511,1536]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brasiliensis">
|
|
<emphasis id="B96B64243F1A894D50C8A080FBA2E03D" box="[909,1073,1511,1536]" italics="true" pageId="17" pageNumber="18">T. brasiliensis</emphasis>
|
|
</taxonomicName>
|
|
+
|
|
<taxonomicName id="4C1FC3B53F1A894D5707A080FB56E03D" authorityName="Berg" authorityYear="1878" box="[1090,1221,1511,1536]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trinotata">
|
|
<emphasis id="B96B64243F1A894D5707A080FB56E03D" box="[1090,1221,1511,1536]" italics="true" pageId="17" pageNumber="18">T. trinotata</emphasis>
|
|
</taxonomicName>
|
|
was supported by one synapomorphic character: length of frena reaching basal ½ of the scutellum (
|
|
<subScript id="179BBA733F1A894D5708A36AFBE0E016" attach="none" box="[1101,1139,1549,1579]" fontSize="6" pageId="17" pageNumber="18">131</subScript>
|
|
). Also, four homoplasies supported the sister group relationship of
|
|
<emphasis id="B96B64243F1A894D51D4A350FC45E06D" box="[657,982,1591,1616]" italics="true" pageId="17" pageNumber="18">
|
|
<taxonomicName id="4C1FC3B53F1A894D51D4A350FCAEE06D" authorityName="Spinola" authorityYear="1852" box="[657,829,1591,1616]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brasiliensis">T. brasiliensis</taxonomicName>
|
|
+
|
|
<taxonomicName id="4C1FC3B53F1A894D500BA350FC45E06D" authorityName="Berg" authorityYear="1878" box="[846,982,1591,1616]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trinotata">T. trinotata</taxonomicName>
|
|
</emphasis>
|
|
: lateral margins of propleura lined in black (
|
|
<subScript id="179BBA733F1A894D53ADA33AFF6CE046" attach="none" box="[232,255,1629,1659]" fontSize="6" pageId="17" pageNumber="18">91</subScript>
|
|
), which is found in
|
|
<taxonomicName id="4C1FC3B53F1A894D52B5A339FDE7E04A" authorityName="Berg" authorityYear="1891" box="[496,628,1630,1655]" class="Insecta" genus="Lobepomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="peltifera">
|
|
<emphasis id="B96B64243F1A894D52B5A339FDE7E04A" box="[496,628,1630,1655]" italics="true" pageId="17" pageNumber="18">L. peltifera</emphasis>
|
|
</taxonomicName>
|
|
+
|
|
<taxonomicName id="4C1FC3B53F1A894D51C1A339FC94E04A" authorityName="Berg" authorityYear="1891" box="[644,775,1630,1655]" class="Insecta" genus="Procleticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="corniger">
|
|
<emphasis id="B96B64243F1A894D51C1A339FC94E04A" box="[644,775,1630,1655]" italics="true" pageId="17" pageNumber="18">P. corniger</emphasis>
|
|
</taxonomicName>
|
|
; evaporative area not reaching the pseudo-suture of the metaepimerum (
|
|
<subScript id="179BBA733F1A894D5225A3E0FE1BE098" attach="none" box="[352,392,1671,1701]" fontSize="6" pageId="17" pageNumber="18">200</subScript>
|
|
), a reversal of the condition found in the basal node of the ingroup; vesica strongly sinuous (
|
|
<subScript id="179BBA733F1A894D5247A3D6FEBAE0F2" attach="none" box="[258,297,1713,1743]" fontSize="6" pageId="17" pageNumber="18">322</subScript>
|
|
), and laterotergites 9 strongly convex (
|
|
<subScript id="179BBA733F1A894D5044A3D6FCBBE0F2" attach="none" box="[769,808,1713,1743]" fontSize="6" pageId="17" pageNumber="18">332</subScript>
|
|
), conditions also supporting Clade A (
|
|
<taxonomicName id="4C1FC3B53F1A894D57BFA3D6FAE7E0F7" authorityName="Pennington" authorityYear="1922" box="[1274,1396,1713,1738]" class="Insecta" genus="Neoderoploa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bruchii">
|
|
<emphasis id="B96B64243F1A894D57BFA3D6FAE7E0F7" box="[1274,1396,1713,1738]" italics="true" pageId="17" pageNumber="18">N. bruchii</emphasis>
|
|
</taxonomicName>
|
|
+
|
|
<taxonomicName id="4C1FC3B53F1A894D56C1A3D6FF69E0CE" authorityName="Berg" authorityYear="1891" class="Insecta" genus="Lobepomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="peltifera">
|
|
<emphasis id="B96B64243F1A894D56C1A3D6FF69E0CE" italics="true" pageId="17" pageNumber="18">L. peltifera</emphasis>
|
|
</taxonomicName>
|
|
+
|
|
<taxonomicName id="4C1FC3B53F1A894D524EA3BDFE1FE0CE" authorityName="Berg" authorityYear="1891" box="[267,396,1754,1779]" class="Insecta" genus="Procleticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="corniger">
|
|
<emphasis id="B96B64243F1A894D524EA3BDFE1FE0CE" box="[267,396,1754,1779]" italics="true" pageId="17" pageNumber="18">P. corniger</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BA0B8363F1A894D5383A266FAD9E185" blockId="17.[151,1437,826,1976]" pageId="17" pageNumber="18">
|
|
Just one of the diagnostic characters of
|
|
<taxonomicName id="4C1FC3B53F1A894D51D6A266FC83E127" authorityName="Bernardes & Schwertner & Grazia" authorityYear="2009" box="[659,784,1793,1818]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96B64243F1A894D51D6A266FC83E127" box="[659,784,1793,1818]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
|
|
</taxonomicName>
|
|
pointed out by
|
|
<bibRefCitation id="EF8EC5C73F1A894D508DA266FBE2E126" author="Rolston, L. H." box="[968,1137,1793,1819]" journalOrPublisher="Journal of the New York Entomological Society" pageId="17" pageNumber="18" pagination="20 - 36" part="86" refId="ref11075" refString="Rolston, L. H. (1978) A revision of the genus Odmalea Bergroth (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 86 (1), 20 - 36." title="A revision of the genus Odmalea Bergroth (Hemiptera: Pentatomidae)" type="journal article" year="1978">Rolston (1978</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EF8EC5C73F1A894D5738A266FB2AE126" author="Rolston, L. H." box="[1149,1209,1793,1819]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="17" pageNumber="18" pagination="826 - 834" part="86" refId="ref11114" refString="Rolston, L. H. (1984) A review of the genus Thoreyella Spinola (Hemiptera: Pentatomidae). Proceedings of the Entomological Society of Washington, 86 (4), 826 - 834." title="A review of the genus Thoreyella Spinola (Hemiptera: Pentatomidae)" type="journal article" year="1984">1984</bibRefCitation>
|
|
), was confirmed as a synapomorphy: apex of abdominal spine curved (
|
|
<subScript id="179BBA733F1A894D51B9A240FCB0E178" attach="none" box="[764,803,1831,1861]" fontSize="6" pageId="17" pageNumber="18">241</subScript>
|
|
). The ostiolar rugae acuminate (
|
|
<subScript id="179BBA733F1A894D57EFA240FB43E178" attach="none" box="[1194,1232,1831,1861]" fontSize="6" pageId="17" pageNumber="18">171</subScript>
|
|
), also pointed as diagnostic to
|
|
<taxonomicName id="4C1FC3B53F1A894D5271A236FDF8E156" authority="(Rolston 1978)" baseAuthorityName="Rolston" baseAuthorityYear="1978" box="[308,619,1873,1899]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96B64243F1A894D5271A236FE21E157" box="[308,434,1873,1898]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
|
|
(
|
|
<bibRefCitation id="EF8EC5C73F1A894D5285A236FDF1E156" author="Rolston, L. H." box="[448,610,1873,1899]" journalOrPublisher="Journal of the New York Entomological Society" pageId="17" pageNumber="18" pagination="20 - 36" part="86" refId="ref11075" refString="Rolston, L. H. (1978) A revision of the genus Odmalea Bergroth (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 86 (1), 20 - 36." title="A revision of the genus Odmalea Bergroth (Hemiptera: Pentatomidae)" type="journal article" year="1978">Rolston 1978</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
, are shared by all the ingroup taxa. Other characteristics mentioned as diagnostic to
|
|
<taxonomicName id="4C1FC3B53F1A894D5202A21FFE42E1AC" baseAuthorityName="Rolston" baseAuthorityYear="1978" box="[327,465,1912,1937]" class="Insecta" family="Pentatomidae" genus="Thoreyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96B64243F1A894D5202A21FFE42E1AC" box="[327,465,1912,1937]" italics="true" pageId="17" pageNumber="18">Thoreyella</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation id="EF8EC5C73F1A894D52A6A21FFD05E1AF" author="Rolston, L. H." box="[483,662,1912,1938]" journalOrPublisher="Journal of the New York Entomological Society" pageId="17" pageNumber="18" pagination="20 - 36" part="86" refId="ref11075" refString="Rolston, L. H. (1978) A revision of the genus Odmalea Bergroth (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 86 (1), 20 - 36." title="A revision of the genus Odmalea Bergroth (Hemiptera: Pentatomidae)" type="journal article" year="1978">Rolston 1978</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EF8EC5C73F1A894D51E1A21FFCA5E1AF" author="Rider, D. A." box="[676,822,1912,1938]" journalOrPublisher="Journal of the New York Entomological Society" pageId="17" pageNumber="18" pagination="193 - 221" part="102" refId="ref11026" refString="Rider, D. A. (1994) A generic conspectus of the tribe Procleticini Pennington (Heteroptera, Pentatomidae), with the description of Paraodmalea rubella, new genus and species. Journal of the New York Entomological Society, 102, 193 - 221." title="A generic conspectus of the tribe Procleticini Pennington (Heteroptera, Pentatomidae), with the description of Paraodmalea rubella, new genus and species" type="journal article" year="1994">Rider 1994</bibRefCitation>
|
|
) represented homoplastic characters or showed ambiguous reconstruction in the analysis, being dependent of the optimization procedures (see below).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |