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<document id="2E73DA15F5FE8F0914BA225B309C78A0" ID-CLB-Dataset="29479" ID-DOI="10.11646/zootaxa.4471.2.3" ID-GBIF-Dataset="694f85ad-7913-43ea-89b9-8eb4e20f0821" ID-ISSN="1175-5326" ID-Zenodo-Dep="1439674" ID-ZooBank="CE4C6E73-09CB-4E74-91C5-771B3696DC52" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538364447371" checkinUser="plazi" docAuthor="Smith, Andrew B. T. &amp; Evans, Arthur V." docDate="2018" docId="03AF87E48E4CFFDBFF49FEE5FA788383" docLanguage="en" docName="zootaxa.4471.2.3.pdf" docOrigin="Zootaxa 4471 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apteroathlia Smith &amp; Evans 2018, new genus" docType="key" docUuid="778D2B0C-E445-46BC-84CB-338DA4F2F7F6" docUuidSource="ZooBank" docVersion="8" lastPageNumber="282" masterDocId="FF96FF9C8E4FFFD8FFDEFFC7FFE38255" masterDocTitle="Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America" masterLastPageNumber="308" masterPageNumber="279" pageNumber="282" updateTime="1698558211898" updateUser="ExternalLinkService">
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<mods:title id="9AFEFB29A57022C64B9B315CC0F09003">Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America</mods:title>
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<mods:namePart id="D2A3D442DF5A4E8DA5C8491BABF8138E">Smith, Andrew B. T.</mods:namePart>
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<mods:namePart id="C79455C54A71A1FEEBDD8881A99A9BB0">Evans, Arthur V.</mods:namePart>
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<treatment id="03AF87E48E4CFFDBFF49FEE5FA788383" ID-DOI="http://doi.org/10.5281/zenodo.6485493" ID-GBIF-Taxon="148415675" ID-Zenodo-Dep="6485493" LSID="urn:lsid:zoobank.org:act:778D2B0C-E445-46BC-84CB-338DA4F2F7F6" httpUri="http://treatment.plazi.org/id/03AF87E48E4CFFDBFF49FEE5FA788383" lastPageNumber="282" pageId="3" pageNumber="282">
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Key to species of
<taxonomicName id="4C064D718E4CFFDBFEB3FEE5FDE68369" ID-CoL="ZCG" authority="Smith &amp; Evans, 2018" authorityName="Smith &amp; Evans" authorityYear="2018" box="[365,517,290,316]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972EAE08E4CFFDBFEB3FEE5FDE68369" bold="true" box="[365,517,290,316]" italics="true" pageId="3" pageNumber="282">Apteroathlia</emphasis>
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<taxonomicNameLabel id="A241579B8E4CFFDBFDD2FEE5FD6E8369" box="[524,653,290,316]" pageId="3" pageNumber="282" rank="genus">new genus</taxonomicNameLabel>
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1. Anterior labral margin broadly concave and not upturned medially (
<figureCitation id="133D2A778E4CFFDBFCB1FEAFFC49832B" box="[879,938,360,382]" captionStart="FIGURES 15" captionStartId="4.[151,264,1267,1289]" captionTargetBox="[318,1274,195,1255]" captionTargetId="figure@4.[318,1274,195,1255]" captionTargetPageId="4" captionText="FIGURES 15. Apteroathlia nox Smith &amp; Evans, new species, holotype. 1, dorsal habitus; 2, oblique habitus; 3, ventral habitus; 4, head and pronotum; 5, parameres." httpUri="https://zenodo.org/record/1439676/files/figure.png" pageId="3" pageNumber="282">Fig. 4</figureCitation>
), dorsal color black (pronotum sometimes medially or completely light brown/tan) (
<figureCitation id="133D2A778E4CFFDBFDCDFE42FD8D83CE" box="[531,622,389,411]" captionStart="FIGURES 15" captionStartId="4.[151,264,1267,1289]" captionTargetBox="[318,1274,195,1255]" captionTargetId="figure@4.[318,1274,195,1255]" captionTargetPageId="4" captionText="FIGURES 15. Apteroathlia nox Smith &amp; Evans, new species, holotype. 1, dorsal habitus; 2, oblique habitus; 3, ventral habitus; 4, head and pronotum; 5, parameres." httpUri="https://zenodo.org/record/1439676/files/figure.png" pageId="3" pageNumber="282">Figs. 12</figureCitation>
).............................
<taxonomicName id="4C064D718E4CFFDBFC12FE42FAFF83CE" ID-CoL="FYHY" authority="Smith and Evans" authorityName="Smith &amp; Evans" authorityYear="2018" box="[972,1308,389,411]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="nox" status="sp. nov.">
<emphasis id="B972EAE08E4CFFDBFC12FE42FB9283CE" box="[972,1137,389,411]" italics="true" pageId="3" pageNumber="282">Apteroathlia nox</emphasis>
Smith and Evans
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,
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<emphasis id="B972EAE08E4CFFDBFAF8FE41FA7883C9" bold="true" box="[1318,1435,390,412]" pageId="3" pageNumber="282">new species</emphasis>
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<keyLead id="30F795C78E4CFFDBFF49FE64FA788383" pageId="3" pageNumber="282">
- Anterior labral margin narrowly concave and upturned medially (
<figureCitation id="133D2A778E4CFFDBFC8AFE64FC6E83EC" box="[852,909,419,441]" captionStart="FIGURES 610" captionStartId="5.[151,264,1714,1736]" captionTargetBox="[335,1263,553,1700]" captionTargetId="figure@5.[335,1263,553,1700]" captionTargetPageId="5" captionText="FIGURES 610. Apteroathlia translucida Smith &amp; Evans, new species, holotype. 6, dorsal habitus; 7, oblique habitus; 8, ventral habitus; 9, head and pronotum; 10, parameres." httpUri="https://zenodo.org/record/1439678/files/figure.png" pageId="3" pageNumber="282">Fig. 9</figureCitation>
), dorsal color tan to light brown (
<figureCitation id="133D2A778E4CFFDBFB0EFE64FACF83EC" box="[1232,1324,419,441]" captionStart="FIGURES 610" captionStartId="5.[151,264,1714,1736]" captionTargetBox="[335,1263,553,1700]" captionTargetId="figure@5.[335,1263,553,1700]" captionTargetPageId="5" captionText="FIGURES 610. Apteroathlia translucida Smith &amp; Evans, new species, holotype. 6, dorsal habitus; 7, oblique habitus; 8, ventral habitus; 9, head and pronotum; 10, parameres." httpUri="https://zenodo.org/record/1439678/files/figure.png" pageId="3" pageNumber="282">Figs. 67</figureCitation>
)................................................................
<taxonomicName id="4C064D718E4CFFDBFC5BFE07FAFF8383" ID-CoL="FYHZ" authority="Smith and Evans" authorityName="Smith &amp; Evans" authorityYear="2018" box="[901,1308,448,470]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="translucida" status="sp. nov.">
<emphasis id="B972EAE08E4CFFDBFC5BFE07FB928383" box="[901,1137,448,470]" italics="true" pageId="3" pageNumber="282">Apteroathlia translucida</emphasis>
Smith and Evans
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,
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<emphasis id="B972EAE08E4CFFDBFAF8FE07FA788383" bold="true" box="[1318,1435,448,470]" pageId="3" pageNumber="282">new species</emphasis>
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