172 lines
21 KiB
XML
172 lines
21 KiB
XML
<document id="D2A34948ED62121CA32925AC719C6A9E" ID-CLB-Dataset="29479" ID-DOI="10.11646/zootaxa.4471.2.3" ID-GBIF-Dataset="694f85ad-7913-43ea-89b9-8eb4e20f0821" ID-ISSN="1175-5326" ID-Zenodo-Dep="1439674" ID-ZooBank="CE4C6E73-09CB-4E74-91C5-771B3696DC52" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538364447371" checkinUser="plazi" docAuthor="Smith, Andrew B. T. & Evans, Arthur V." docDate="2018" docId="03AF87E48E40FFC8FF49FF50FBA783A2" docLanguage="en" docName="zootaxa.4471.2.3.pdf" docOrigin="Zootaxa 4471 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athlia rustica Erichson 1835" docType="treatment" docVersion="7" lastPageNumber="295" masterDocId="FF96FF9C8E4FFFD8FFDEFFC7FFE38255" masterDocTitle="Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America" masterLastPageNumber="308" masterPageNumber="279" pageNumber="294" updateTime="1698558211898" updateUser="ExternalLinkService">
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<mods:title id="D371ADC73DA11D0C493A0A8ABD8098BB">Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America</mods:title>
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<mods:namePart id="E026E347C12B4593C9B04A9D1DD1398A">Smith, Andrew B. T.</mods:namePart>
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<mods:namePart id="BEC46E0D2FBC83AAC6057AF6697F96FC">Evans, Arthur V.</mods:namePart>
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<mods:title id="B63961E5ED46AA9CBB292339150F94C7">Zootaxa</mods:title>
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<mods:date id="BF782F72F55D323D9AC2276214D6D627">2018</mods:date>
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<treatment id="03AF87E48E40FFC8FF49FF50FBA783A2" ID-DOI="http://doi.org/10.5281/zenodo.5958512" ID-GBIF-Taxon="148415692" ID-Zenodo-Dep="5958512" LSID="urn:lsid:plazi:treatment:03AF87E48E40FFC8FF49FF50FBA783A2" httpUri="http://treatment.plazi.org/id/03AF87E48E40FFC8FF49FF50FBA783A2" lastPageId="16" lastPageNumber="295" pageId="15" pageNumber="294">
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<subSubSection id="C31C65798E40FFD7FF49FF50FEAE8286" pageId="15" pageNumber="294" type="nomenclature">
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<paragraph id="8BB936F28E40FFD7FF49FF50FE1E82E7" blockId="15.[151,509,151,211]" box="[151,509,151,178]" pageId="15" pageNumber="294">
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<heading id="D0F1819E8E40FFD7FF49FF50FE1E82E7" bold="true" box="[151,509,151,178]" fontSize="11" level="1" pageId="15" pageNumber="294" reason="1">
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<taxonomicName id="4C064D718E40FFD7FF49FF50FE1E82E7" ID-CoL="5W8GD" authority="Erichson, 1835" authorityName="Erichson" authorityYear="1835" box="[151,509,151,178]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="species" species="rustica">
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<emphasis id="B972EAE08E40FFD7FF49FF50FE1E82E7" bold="true" box="[151,509,151,178]" pageId="15" pageNumber="294">
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<emphasis id="B972EAE08E40FFD7FF49FF50FEDF82E4" bold="true" box="[151,316,151,177]" italics="true" pageId="15" pageNumber="294">Athlia rustica</emphasis>
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<bibRefCitation id="EF974B038E40FFD7FE9AFF5FFE1E82E7" box="[324,509,152,178]" pageId="15" pageNumber="294" refId="ref10694" refString="Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270." type="journal article">Erichson, 1835</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BB936F28E40FFD7FF49FF7DFEAE8286" blockId="15.[151,509,151,211]" box="[151,333,186,211]" pageId="15" pageNumber="294">
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<figureCitation id="133D2A778E40FFD7FF49FF7DFEFD8286" box="[151,286,186,211]" captionStart="FIGURES 48–53" captionStartId="15.[151,264,1674,1696]" captionTargetBox="[283,1303,371,1652]" captionTargetId="figure@15.[283,1303,371,1652]" captionTargetPageId="15" captionText="FIGURES 48–53. Athlia rustica Erichson, 1835 lectotype. 48, dorsal habitus; 49, oblique habitus; 50, ventral habitus; 51, labels; 52, head and pronotum; 53, parameres." httpUri="https://zenodo.org/record/1439698/files/figure.png" pageId="15" pageNumber="294">Figs. 48–53</figureCitation>
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,
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<figureCitation id="133D2A778E40FFD7FEF7FF7CFEAA8286" box="[297,329,187,211]" captionStart="FIGURE 72" captionStartId="27.[151,250,1849,1871]" captionTargetBox="[153,1433,360,1800]" captionTargetId="figure@27.[153,1433,360,1800]" captionTargetPageId="27" captionText="FIGURE 72. Distribution map of Athlia rustica Erichson, 1835 (black circles)." httpUri="https://zenodo.org/record/1439718/files/figure.png" pageId="15" pageNumber="294">72</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31C65798E40FFD7FF49FEC5FCCA834E" box="[151,809,258,283]" pageId="15" pageNumber="294" type="description">
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<paragraph id="8BB936F28E40FFD7FF49FEC5FCCA834E" blockId="15.[151,809,258,319]" box="[151,809,258,283]" pageId="15" pageNumber="294">
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<emphasis id="B972EAE08E40FFD7FF49FEC5FE7B834E" bold="true" box="[151,408,258,283]" pageId="15" pageNumber="294">Original combination</emphasis>
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.
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<taxonomicName id="4C064D718E40FFD7FE78FEC4FCCA834E" authority="Erichson, 1835: 267" authorityName="Erichson" authorityPageNumber="267" authorityYear="1835" box="[422,809,258,283]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="species" species="rustica">
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<emphasis id="B972EAE08E40FFD7FE78FEC4FDA3834F" box="[422,576,259,283]" italics="true" pageId="15" pageNumber="294">Athlia rustica</emphasis>
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<bibRefCitation id="EF974B038E40FFD7FD98FEC5FD17834E" box="[582,756,258,283]" pageId="15" pageNumber="294" refId="ref10694" refString="Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270." type="journal article">Erichson, 1835</bibRefCitation>
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: 267
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31C65798E40FFC8FF19FEE1FF3D8348" lastPageId="16" lastPageNumber="295" pageId="15" pageNumber="294" type="materials_examined">
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<paragraph id="8BB936F28E40FFD7FF19FEE1FD66836A" blockId="15.[151,809,258,319]" box="[199,645,294,319]" pageId="15" pageNumber="294">
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<materialsCitation id="3B6E3CAF8E40FFD7FF19FEE1FD66836A" ID-GBIF-Occurrence="1913888852" box="[199,645,294,319]" country="Chile" location="Chile" pageId="15" pageNumber="294" specimenCount="1" typeStatus="holotype">
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<emphasis id="B972EAE08E40FFD7FF19FEE1FEBE836A" bold="true" box="[199,349,294,319]" pageId="15" pageNumber="294">
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<typeStatus id="54BD88508E40FFD7FF19FEE1FEE1836A" box="[199,258,294,319]" pageId="15" pageNumber="294">Type</typeStatus>
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locality
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</emphasis>
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. “
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<collectingCountry id="F31176628E40FFD7FEA8FEE1FE54836A" box="[374,439,294,319]" name="Chile" pageId="15" pageNumber="294">Chile</collectingCountry>
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” (
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<bibRefCitation id="EF974B038E40FFD7FE0EFEE1FD9B836A" box="[464,632,294,319]" pageId="15" pageNumber="294" refId="ref10694" refString="Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270." type="journal article">Erichson 1835</bibRefCitation>
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).
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</materialsCitation>
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</paragraph>
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<caption id="DF79667A8E40FFD7FF49F94DFD8884EB" httpUri="https://zenodo.org/record/1439698/files/figure.png" pageId="15" pageNumber="294" startId="15.[151,264,1674,1696]" targetBox="[283,1303,371,1652]" targetPageId="15">
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<paragraph id="8BB936F28E40FFD7FF49F94DFD8884EB" blockId="15.[151,1435,1674,1726]" pageId="15" pageNumber="294">
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<emphasis id="B972EAE08E40FFD7FF49F94DFEB484F5" bold="true" box="[151,343,1674,1696]" pageId="15" pageNumber="294">FIGURES 48–53.</emphasis>
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<taxonomicName id="4C064D718E40FFD7FEBEF94DFD7084F5" authority="Erichson, 1835" authorityName="Erichson" authorityYear="1835" box="[352,659,1674,1696]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="species" species="rustica">
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<emphasis id="B972EAE08E40FFD7FEBEF94DFE0E84F5" box="[352,493,1674,1696]" italics="true" pageId="15" pageNumber="294">Athlia rustica</emphasis>
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Erichson, 1835
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</taxonomicName>
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lectotype. 48, dorsal habitus; 49, oblique habitus; 50, ventral habitus; 51, labels; 52, head and pronotum; 53, parameres.
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</paragraph>
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</caption>
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<paragraph id="8BB936F28E40FFC8FF19F92CFF3D8348" blockId="15.[151,1436,1771,2012]" lastBlockId="16.[151,1436,151,503]" lastPageId="16" lastPageNumber="295" pageId="15" pageNumber="294">
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<emphasis id="B972EAE08E40FFD7FF19F92CFEA88551" bold="true" box="[199,331,1771,1796]" pageId="15" pageNumber="294">Type series</emphasis>
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. Lectotype (
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<figureCitation id="133D2A778E40FFD7FE09F92CFDBD8551" box="[471,606,1771,1796]" captionStart="FIGURES 48–53" captionStartId="15.[151,264,1674,1696]" captionTargetBox="[283,1303,371,1652]" captionTargetId="figure@15.[283,1303,371,1652]" captionTargetPageId="15" captionText="FIGURES 48–53. Athlia rustica Erichson, 1835 lectotype. 48, dorsal habitus; 49, oblique habitus; 50, ventral habitus; 51, labels; 52, head and pronotum; 53, parameres." httpUri="https://zenodo.org/record/1439698/files/figure.png" pageId="15" pageNumber="294">Figs. 48–53</figureCitation>
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) at ZMHB labeled a) “25008” (typeset), b) “Type” (typeset, orange label), c) “
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<taxonomicName id="4C064D718E40FFD7FF1EF8C8FEEE857D" authorityName="Erichson" authorityYear="1835" box="[192,269,1807,1832]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="species" species="rustica">rustica</taxonomicName>
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/ Er. / Chile Meyen.” (handwritten, green label), d) “SYNTYPUS /
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<taxonomicName id="4C064D718E40FFD7FBF6F8C8FB8D857D" authorityName="Smith & Evans" authorityYear="2018" box="[1064,1134,1807,1832]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="306" phylum="Arthropoda" rank="genus">Athlia</taxonomicName>
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/
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<taxonomicName id="4C064D718E40FFD7FB57F8C8FA69857D" authority="Erichson, 1835" authorityName="Erichson" authorityYear="1835" box="[1161,1418,1807,1832]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="species" species="rustica">
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rustica
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<bibRefCitation id="EF974B038E40FFD7FB00F8C8FA69857D" box="[1246,1418,1807,1832]" pageId="15" pageNumber="294" refId="ref10694" refString="Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270." type="journal article">Erichson, 1835</bibRefCitation>
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</taxonomicName>
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/ labelled by MNHUB 2009” (typeset, orange label), e) “
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<taxonomicName id="4C064D718E40FFD7FCD8F8F3FC8D8519" authorityName="Smith & Evans" authorityYear="2018" box="[774,878,1844,1868]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="306" phylum="Arthropoda" rank="genus">ATHLIA</taxonomicName>
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/ RUSTICA /
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<bibRefCitation id="EF974B038E40FFD7FBCDF8F3FB058519" box="[1043,1254,1843,1868]" pageId="15" pageNumber="294" refId="ref10694" refString="Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270." type="journal article">ERICHSON, 1835</bibRefCitation>
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/ LECTOTYPE / A.B.T. SMITH” (handwritten and typeset, red label).
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<emphasis id="B972EAE08E40FFD7FCDCF890FBD98525" bold="true" box="[770,1082,1879,1904]" pageId="15" pageNumber="294">Lectotype here designated</emphasis>
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. Four paralectotypes at ZMHB labeled a) “Type” (typeset, orange label), b) “Hist.-Coll. (
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<taxonomicName id="4C064D718E40FFD7FCFFF8BCFC7C85C1" authorityName="" authorityYear="1758" box="[801,927,1915,1940]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
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) / Nr. 25008 /
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<taxonomicName id="4C064D718E40FFD7FB99F8BCFB6E85C1" authorityName="Smith & Evans" authorityYear="2018" box="[1095,1165,1915,1940]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="306" phylum="Arthropoda" rank="genus">Athlia</taxonomicName>
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/
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<taxonomicName id="4C064D718E40FFD7FB7AF8BCFAA585C1" authority="Erichs." authorityName="Erichson" authorityYear="1835" box="[1188,1350,1915,1940]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="species" species="rustica">rustica Erichs.</taxonomicName>
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/ Chili, Meyen / Zool. Mus. Berlin” (typeset), c) “SYNTYPUS /
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<taxonomicName id="4C064D718E40FFD7FCF0F858FC9785ED" authorityName="Smith & Evans" authorityYear="2018" box="[814,884,1951,1976]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="306" phylum="Arthropoda" rank="genus">Athlia</taxonomicName>
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/
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<taxonomicName id="4C064D718E40FFD7FC4EF858FB7385ED" authority="Erichson, 1835" authorityName="Erichson" authorityYear="1835" box="[912,1168,1951,1976]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="species" species="rustica">
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rustica
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<bibRefCitation id="EF974B038E40FFD7FC3AF858FB7385ED" box="[996,1168,1951,1976]" pageId="15" pageNumber="294" refId="ref10694" refString="Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270." type="journal article">Erichson, 1835</bibRefCitation>
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</taxonomicName>
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/ labelled by MNHUB 2009” (typeset, orange label), d) “
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<taxonomicName id="4C064D718E40FFD7FDF0F803FD758589" authorityName="Smith & Evans" authorityYear="2018" box="[558,662,1988,2012]" class="Insecta" family="Scarabaeidae" genus="Athlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="306" phylum="Arthropoda" rank="genus">ATHLIA</taxonomicName>
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/ RUSTICA /
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<bibRefCitation id="EF974B038E40FFD7FC8EF803FBC48589" box="[848,1063,1987,2012]" pageId="15" pageNumber="294" refId="ref10694" refString="Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270." type="journal article">ERICHSON, 1835</bibRefCitation>
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/ PARALECTOTYPE / A.B.T. SMITH” (handwritten and typeset, yellow label). A lectotype is designated to preserve the stability of the nomenclature by selecting a single specimen as the name-bearing type for this species. Since
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<bibRefCitation id="EF974B038E5FFFC8FB64FF7BFA938280" box="[1210,1392,188,213]" pageId="16" pageNumber="295" refId="ref10694" refString="Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270." type="journal article">Erichson (1835)</bibRefCitation>
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did not explicitly state how many specimens he examined, it is impossible to know the exact composition of the type series.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31C65798E5FFFC8FF19FEE0FBA783A2" pageId="16" pageNumber="295" type="distribution">
|
||
<paragraph id="8BB936F28E5FFFC8FF19FEE0FBA783A2" blockId="16.[151,1436,151,503]" pageId="16" pageNumber="295">
|
||
<emphasis id="B972EAE08E5FFFC8FF19FEE0FEBA8315" bold="true" box="[199,345,295,320]" pageId="16" pageNumber="295">Distribution</emphasis>
|
||
.
|
||
<figureCitation id="133D2A778E5FFFC8FEB5FEE0FE218315" box="[363,450,295,320]" captionStart="FIGURE 72" captionStartId="27.[151,250,1849,1871]" captionTargetBox="[153,1433,360,1800]" captionTargetId="figure@27.[153,1433,360,1800]" captionTargetPageId="27" captionText="FIGURE 72. Distribution map of Athlia rustica Erichson, 1835 (black circles)." httpUri="https://zenodo.org/record/1439718/files/figure.png" pageId="16" pageNumber="295">Fig. 72</figureCitation>
|
||
.
|
||
<collectingCountry id="F31176628E5FFFC8FE0FFEE0FDF18315" box="[465,530,295,320]" name="Chile" pageId="16" pageNumber="295">Chile</collectingCountry>
|
||
: IV
|
||
<collectingRegion id="49C2F8108E5FFFC8FD97FEE0FCA08315" box="[585,835,295,320]" country="Chile" name="Coquimbo" pageId="16" pageNumber="295">Región de Coquimbo</collectingRegion>
|
||
, V
|
||
<collectingRegion id="49C2F8108E5FFFC8FCACFEE0FB8E8315" box="[882,1133,295,320]" country="Chile" name="Valparaiso" pageId="16" pageNumber="295">Región de Valparaíso</collectingRegion>
|
||
,
|
||
<collectingRegion id="49C2F8108E5FFFC8FBA2FEE0FF1F8330" country="Chile" name="Region Metropolitana" pageId="16" pageNumber="295">Región Metropolitana de Santiago</collectingRegion>
|
||
, VI
|
||
<collectingRegion id="49C2F8108E5FFFC8FEF3FE8BFC9B8330" box="[301,888,332,357]" country="Chile" name="Libertador General Bernardo O'Higgins" pageId="16" pageNumber="295">Región del Libertador General Bernardo O’Higgins</collectingRegion>
|
||
, VII
|
||
<collectingRegion id="49C2F8108E5FFFC8FC6AFE8BFB618330" box="[948,1154,332,357]" country="Chile" name="Maule" pageId="16" pageNumber="295">Región del Maule</collectingRegion>
|
||
, VIII
|
||
<collectingRegion id="49C2F8108E5FFFC8FB18FE8BFA7B8330" box="[1222,1432,332,357]" country="Chile" name="Biobio" pageId="16" pageNumber="295">Región del Biobío</collectingRegion>
|
||
, IX
|
||
<collectingRegion id="49C2F8108E5FFFC8FF60FEA8FE2D83DD" box="[190,462,367,392]" country="Chile" name="Araucania" pageId="16" pageNumber="295">Región de la Araucanía</collectingRegion>
|
||
(based on specimens from AMNH, AVEC, BMNH, CASC, CMNC, CNCI, FMNH, FSCA, INHS, LEMQ, MNHN, MNNC, UMCE, UMSP, USNM). One 19th century specimen is labeled “Valdivia” (BMNH) but we consider this record erroneous. The record from IX
|
||
<collectingRegion id="49C2F8108E5FFFC8FC77FE7DFB578386" box="[937,1204,442,467]" country="Chile" name="Araucania" pageId="16" pageNumber="295">Región de la Araucanía</collectingRegion>
|
||
is based on a single collecting event from “
|
||
<quantity id="4CFE9B178E5FFFC8FE46FE18FE0283A2" box="[408,481,478,503]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="295" unit="km" value="30.0">30 km</quantity>
|
||
NE Villarrica”, which might need further verification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |