treatments-xml/data/03/AF/87/03AF87C0FFEFA92D41C6FC704D48E6E2.xml

<document id="79AE437F43B919C91DCA23CEDEF82518" ID-DOI="10.5281/zenodo.214715" ID-GBIF-Dataset="4eb4d325-9c9f-423d-b231-99d88173627a" ID-ISSN="1175-5326" ID-Zenodo-Dep="214715" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460503928248" checkinUser="plazi" docAuthor="Wolski, Andrzej &amp; Gorczyca, Jacek" docDate="2012" docId="03AF87C0FFEFA92D41C6FC704D48E6E2" docLanguage="en" docName="zt03412p041.pdf" docOrigin="Zootaxa 3412" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dashymeniella spatulatiformis Wolski &amp; Gorczyca, sp. nov." docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="FF96FFB8FFCEA90F4151F9324C70E27D" masterDocTitle="Plant bugs of the tribe Bothriomirini (Hemiptera: Heteroptera: Miridae: Cylapinae) from the Oriental Region: descriptions of eight new species and keys to Oriental genera and species of Bothriomiris Kirkaldy, Dashymenia Poppius, and Dashymeniella Poppius" masterLastPageNumber="41" masterPageNumber="1" pageNumber="34" updateTime="1698312487025" updateUser="plazi">
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<mods:title id="69BC2396458AFC83B86E26E10E106EB2">Plant bugs of the tribe Bothriomirini (Hemiptera: Heteroptera: Miridae: Cylapinae) from the Oriental Region: descriptions of eight new species and keys to Oriental genera and species of Bothriomiris Kirkaldy, Dashymenia Poppius, and Dashymeniella Poppius</mods:title>
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<mods:namePart id="F6CF7CE499633F80EAFE14DF397E144B">Wolski, Andrzej</mods:namePart>
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<mods:namePart id="744FB8497C9E2484E42B4C2C52572292">Gorczyca, Jacek</mods:namePart>
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<mods:date id="AA4A3195DB150207FDF40A38CDE835D7">2012</mods:date>
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<treatment id="03AF87C0FFEFA92D41C6FC704D48E6E2" ID-DOI="http://doi.org/10.5281/zenodo.6166325" ID-GBIF-Taxon="119522379" ID-Zenodo-Dep="6166325" LSID="urn:lsid:plazi:treatment:03AF87C0FFEFA92D41C6FC704D48E6E2" httpUri="http://treatment.plazi.org/id/03AF87C0FFEFA92D41C6FC704D48E6E2" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<subSubSection id="C31C655DFFEFA92E41C6FC704E37E700" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BB936D6FFEFA92E41C6FC704F19E721" blockId="33.[151,873,1346,1405]" box="[151,873,1346,1372]" pageId="33" pageNumber="34">
<heading id="D0F181BAFFEFA92E41C6FC704F19E721" bold="true" box="[151,873,1346,1372]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="B972EAC4FFEFA92E41C6FC704F19E721" bold="true" box="[151,873,1346,1372]" pageId="33" pageNumber="34">
<taxonomicName id="4C064D55FFEFA92E41C6FC704F74E721" ID-CoL="346G5" authority="Wolski &amp; Gorczyca" authorityName="Wolski &amp; Gorczyca" box="[151,772,1346,1372]" class="Insecta" family="Miridae" genus="Dashymeniella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="spatulatiformis" status="sp. nov.">
<emphasis id="B972EAC4FFEFA92E41C6FC704E79E721" bold="true" box="[151,521,1346,1372]" italics="true" pageId="33" pageNumber="34">Dashymeniella spatulatiformis</emphasis>
Wolski &amp; Gorczyca
</taxonomicName>
<taxonomicNameLabel id="A24157BFFFEFA92E425AFC704F19E721" box="[779,873,1346,1372]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB936D6FFEFA92E41C6FC574E37E700" blockId="33.[151,873,1346,1405]" box="[151,583,1381,1405]" pageId="33" pageNumber="34">
(
<figureCitation id="133D2A53FFEFA92E41CFFC574D33E700" box="[158,323,1381,1405]" captionStart="FIGURES 1 – 18" captionStartId="5.[151,264,1825,1848]" captionTargetBox="[186,1404,198,1790]" captionTargetId="figure@5.[175,1411,193,1805]" captionTargetPageId="5" captionText="FIGURES 1 – 18. Dorsal habitus color photographs of the Oriental species of the tribe Bothriomirini: 1, 2 — Bothriomiris dissimulans, Ƥ and 3: Malacca, Perak; 3 — Bothriomiris lugubris (lectotype); 4 — Bothriomiris sulawesicus, 3: Indonesia: Sulawesi Utara, Dumoga-Bone N. P; 5 — D. cognata (holotype); 6 — D. colubrina (holotype); 7 — D. conspersa; 8 — D. kerzhneri (paratype, 3); 9 — D. kotejai (paratype); 10 – 11 — D. tenmalai: 10 — paratype, Ƥ, 11 — holotype, 3; 12 — D. webbi (holotype); 13 – 14 — D. spatulatiformis: 13 — paratype, 14 — holotype; 15 — D. tibialis (3: Malacca, Perak); 16 — D. ulu (holotype); 17 — D. viklundi (holotype); 18 — L. scutellaris, Ƥ: Sarawak: Gunong Mulu Nat. Park; Long Pala base camp, ii – vii 1978." httpUri="https://zenodo.org/record/214716/files/figure.png" pageId="33" pageNumber="34">Figures 13–14</figureCitation>
, 
<figureCitation id="133D2A53FFEFA92E401EFC574D1DE700" box="[335,365,1381,1405]" captionStart="FIGURES 49 – 54" captionStartId="17.[151,264,1068,1091]" captionTargetBox="[151,1435,447,1047]" captionTargetId="figure@17.[151,1435,447,1047]" captionTargetPageId="17" captionText="FIGURES 49 – 54. Left lateral view: 49 — Bothriomiris sulawesicus; 50 — Dashymenia kerzhneri; 51 — D. cognata; 52 — Dashymeniella spatulatiformis; 53 — Dashymeniella ulu; 54 — Leprocapsus scutellaris." httpUri="https://zenodo.org/record/214722/files/figure.png" pageId="33" pageNumber="34">52</figureCitation>
, 
<figureCitation id="133D2A53FFEFA92E4028FC574DE7E700" box="[377,407,1381,1405]" captionStart="FIGURE 87" captionStartId="33.[151,250,1243,1266]" captionTargetBox="[167,1404,225,1200]" captionTargetId="figure@33.[151,1436,193,1222]" captionTargetPageId="33" captionText="FIGURE 87. Distribution map of Dashymeniella and Leprocapsus." httpUri="https://zenodo.org/record/214731/files/figure.png" pageId="33" pageNumber="34">87</figureCitation>
, 
<figureCitation id="133D2A53FFEFA92E40F2FC574D9CE700" box="[419,492,1381,1405]" captionStart="FIGURES 88 – 95" captionStartId="36.[151,264,1922,1945]" captionTargetBox="[227,1354,193,1884]" captionTargetId="figure@36.[227,1355,193,1885]" captionTargetPageId="36" captionText="FIGURES 88 – 95. Scanning electron micrographs of Dashymeniella ulu (88 – 93) and Dashymeniella spatulatiformis (94 – 95): 88 – 89 — structure and vestiture of hemelytron; 90 — metafemoral trichobothria; 91 — single metafemoral trichobothrium; 92 – 93 — pretarsal structure; 94 — structure and vestiture of antennal segment I; 95 — structure and vestiture of antennal segment II." httpUri="https://zenodo.org/record/214732/files/figure.png" pageId="33" pageNumber="34">94–95</figureCitation>
, 
<figureCitation id="133D2A53FFEFA92E40A6FC574E4FE700" box="[503,575,1381,1405]" captionStart="FIGURES 96 – 107" captionStartId="37.[151,264,1357,1380]" captionTargetBox="[152,1435,207,1330]" captionTargetId="figure@37.[151,1436,193,1336]" captionTargetPageId="37" captionText="FIGURES 96 – 107. Male genitalia of Dashymeniella spatulatiformis (96 – 98), Dashymeniella tibialis (99 – 101), Dashymeniella viklundi (102 – 104), and Dashymeniella ulu (105 – 107). D. spatulatiformis: 96 — endosoma, dorsal view; 97 — left paramere, right lateral view; 98 — right paramere, dorsal view. D. tibialis: 99 — endosoma, dorsal view; 100 — left paramere, right lateral view; 101 — right paramere, dorsal view. D. viklundi: 102 — endosoma, dorsal view; 103 — left paramere, right lateral view; 104 — right paramere, dorsal view. D. ulu. 105 — endosoma, dorsal view; 106 — left paramere, right lateral view; 107 — right paramere, dorsal view. AES = anterior endosomal sclerite; DSS = sclerotized portion of ductus seminis inside endosoma; MES = mesial endosomal sclerite; PES = posterior endosomal sclerite. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/214733/files/figure.png" pageId="33" pageNumber="34">96–98</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C655DFFEFA92E41C6FC9E4FD8E550" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BB936D6FFEFA92E41C6FC9E4923E470" blockId="33.[151,1436,1452,2017]" pageId="33" pageNumber="34">
<emphasis id="B972EAC4FFEFA92E41C6FC9E4D61E7B8" bold="true" box="[151,273,1452,1477]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
<taxonomicName id="4C064D55FFEFA92E404FFC9F4E0CE7B8" box="[286,636,1453,1477]" class="Insecta" family="Miridae" genus="Dashymeniella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="spatulatiformis">
<emphasis id="B972EAC4FFEFA92E404FFC9F4E0CE7B8" box="[286,636,1453,1477]" italics="true" pageId="33" pageNumber="34">Dashymeniella spatulatiformis</emphasis>
</taxonomicName>
can be recognized by the following set of features: dorsal surface varying from brown to dark brown, nearly black (
<figureCitation id="133D2A53FFEFA92E43EFFCE34F36E794" box="[702,838,1489,1513]" captionStart="FIGURES 1 – 18" captionStartId="5.[151,264,1825,1848]" captionTargetBox="[186,1404,198,1790]" captionTargetId="figure@5.[175,1411,193,1805]" captionTargetPageId="5" captionText="FIGURES 1 – 18. Dorsal habitus color photographs of the Oriental species of the tribe Bothriomirini: 1, 2 — Bothriomiris dissimulans, Ƥ and 3: Malacca, Perak; 3 — Bothriomiris lugubris (lectotype); 4 — Bothriomiris sulawesicus, 3: Indonesia: Sulawesi Utara, Dumoga-Bone N. P; 5 — D. cognata (holotype); 6 — D. colubrina (holotype); 7 — D. conspersa; 8 — D. kerzhneri (paratype, 3); 9 — D. kotejai (paratype); 10 – 11 — D. tenmalai: 10 — paratype, Ƥ, 11 — holotype, 3; 12 — D. webbi (holotype); 13 – 14 — D. spatulatiformis: 13 — paratype, 14 — holotype; 15 — D. tibialis (3: Malacca, Perak); 16 — D. ulu (holotype); 17 — D. viklundi (holotype); 18 — L. scutellaris, Ƥ: Sarawak: Gunong Mulu Nat. Park; Long Pala base camp, ii – vii 1978." httpUri="https://zenodo.org/record/214716/files/figure.png" pageId="33" pageNumber="34">Figs. 13–14</figureCitation>
); head somewhat rugose; endosoma with DSS spoonlike; PES stout, longer than other endosomal sclerites, gradually becoming narrower towards apex (
<figureCitation id="133D2A53FFEFA92E45A3FCC74936E470" box="[1266,1350,1525,1549]" captionStart="FIGURES 96 – 107" captionStartId="37.[151,264,1357,1380]" captionTargetBox="[152,1435,207,1330]" captionTargetId="figure@37.[151,1436,193,1336]" captionTargetPageId="37" captionText="FIGURES 96 – 107. Male genitalia of Dashymeniella spatulatiformis (96 – 98), Dashymeniella tibialis (99 – 101), Dashymeniella viklundi (102 – 104), and Dashymeniella ulu (105 – 107). D. spatulatiformis: 96 — endosoma, dorsal view; 97 — left paramere, right lateral view; 98 — right paramere, dorsal view. D. tibialis: 99 — endosoma, dorsal view; 100 — left paramere, right lateral view; 101 — right paramere, dorsal view. D. viklundi: 102 — endosoma, dorsal view; 103 — left paramere, right lateral view; 104 — right paramere, dorsal view. D. ulu. 105 — endosoma, dorsal view; 106 — left paramere, right lateral view; 107 — right paramere, dorsal view. AES = anterior endosomal sclerite; DSS = sclerotized portion of ductus seminis inside endosoma; MES = mesial endosomal sclerite; PES = posterior endosomal sclerite. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/214733/files/figure.png" pageId="33" pageNumber="34">Fig. 96</figureCitation>
).
</paragraph>
<paragraph id="8BB936D6FFEFA92E4196FF284FD8E550" blockId="33.[151,1436,1452,2017]" pageId="33" pageNumber="34">
<taxonomicName id="4C064D55FFEFA92E4196FF284DE5E44C" box="[199,405,1561,1585]" class="Insecta" family="Miridae" genus="Dashymeniella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="spatulatiformis">
<emphasis id="B972EAC4FFEFA92E4196FF284CABE44C" box="[199,219,1562,1585]" italics="true" pageId="33" pageNumber="34">D</emphasis>
. 
<emphasis id="B972EAC4FFEFA92E41BAFF2B4DE5E44C" box="[235,405,1561,1585]" italics="true" pageId="33" pageNumber="34">spatulatiformis</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C064D55FFEFA92E433DFF284EC4E44C" box="[620,692,1561,1585]" class="Insecta" family="Miridae" genus="Dashymeniella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B972EAC4FFEFA92E433DFF284EF0E44C" box="[620,640,1562,1585]" italics="true" pageId="33" pageNumber="34">D</emphasis>
. 
<emphasis id="B972EAC4FFEFA92E43C1FF2B4EC4E44C" box="[656,692,1561,1585]" italics="true" pageId="33" pageNumber="34">ulu</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D55FFEFA92E43BCFF284F1BE44C" box="[749,875,1561,1585]" class="Insecta" family="Miridae" genus="Dashymeniella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="viklundi">
<emphasis id="B972EAC4FFEFA92E43BCFF284F71E44C" box="[749,769,1562,1585]" italics="true" pageId="33" pageNumber="34">D</emphasis>
. 
<emphasis id="B972EAC4FFEFA92E4240FF2B4F1BE44C" box="[785,875,1561,1585]" italics="true" pageId="33" pageNumber="34">viklundi</emphasis>
</taxonomicName>
in sharing the following set of characters: dorsal surface glabrous (
<figureCitation id="133D2A53FFEFA92E4034FF0F4E39E428" box="[357,585,1597,1621]" captionStart="FIGURES 1 – 18" captionStartId="5.[151,264,1825,1848]" captionTargetBox="[186,1404,198,1790]" captionTargetId="figure@5.[175,1411,193,1805]" captionTargetPageId="5" captionText="FIGURES 1 – 18. Dorsal habitus color photographs of the Oriental species of the tribe Bothriomirini: 1, 2 — Bothriomiris dissimulans, Ƥ and 3: Malacca, Perak; 3 — Bothriomiris lugubris (lectotype); 4 — Bothriomiris sulawesicus, 3: Indonesia: Sulawesi Utara, Dumoga-Bone N. P; 5 — D. cognata (holotype); 6 — D. colubrina (holotype); 7 — D. conspersa; 8 — D. kerzhneri (paratype, 3); 9 — D. kotejai (paratype); 10 – 11 — D. tenmalai: 10 — paratype, Ƥ, 11 — holotype, 3; 12 — D. webbi (holotype); 13 – 14 — D. spatulatiformis: 13 — paratype, 14 — holotype; 15 — D. tibialis (3: Malacca, Perak); 16 — D. ulu (holotype); 17 — D. viklundi (holotype); 18 — L. scutellaris, Ƥ: Sarawak: Gunong Mulu Nat. Park; Long Pala base camp, ii – vii 1978." httpUri="https://zenodo.org/record/214716/files/figure.png" pageId="33" pageNumber="34">Figs. 13–14, 16–17</figureCitation>
); pronotum with the anterior margin separated from the remainder of the pronotum by relatively deep, transverse incision; posterior lobe of the pronotum without any swelling medially (
<figureCitation id="133D2A53FFEFA92E41CEFFB74DFAE4E0" box="[159,394,1669,1693]" captionStart="FIGURES 1 – 18" captionStartId="5.[151,264,1825,1848]" captionTargetBox="[186,1404,198,1790]" captionTargetId="figure@5.[175,1411,193,1805]" captionTargetPageId="5" captionText="FIGURES 1 – 18. Dorsal habitus color photographs of the Oriental species of the tribe Bothriomirini: 1, 2 — Bothriomiris dissimulans, Ƥ and 3: Malacca, Perak; 3 — Bothriomiris lugubris (lectotype); 4 — Bothriomiris sulawesicus, 3: Indonesia: Sulawesi Utara, Dumoga-Bone N. P; 5 — D. cognata (holotype); 6 — D. colubrina (holotype); 7 — D. conspersa; 8 — D. kerzhneri (paratype, 3); 9 — D. kotejai (paratype); 10 – 11 — D. tenmalai: 10 — paratype, Ƥ, 11 — holotype, 3; 12 — D. webbi (holotype); 13 – 14 — D. spatulatiformis: 13 — paratype, 14 — holotype; 15 — D. tibialis (3: Malacca, Perak); 16 — D. ulu (holotype); 17 — D. viklundi (holotype); 18 — L. scutellaris, Ƥ: Sarawak: Gunong Mulu Nat. Park; Long Pala base camp, ii – vii 1978." httpUri="https://zenodo.org/record/214716/files/figure.png" pageId="33" pageNumber="34">Figs. 13–14, 16–17</figureCitation>
); pronotal calli flattened or weakly raised (
<figureCitation id="133D2A53FFEFA92E42CDFFB748F6E4E0" box="[924,1158,1669,1693]" captionStart="FIGURES 1 – 18" captionStartId="5.[151,264,1825,1848]" captionTargetBox="[186,1404,198,1790]" captionTargetId="figure@5.[175,1411,193,1805]" captionTargetPageId="5" captionText="FIGURES 1 – 18. Dorsal habitus color photographs of the Oriental species of the tribe Bothriomirini: 1, 2 — Bothriomiris dissimulans, Ƥ and 3: Malacca, Perak; 3 — Bothriomiris lugubris (lectotype); 4 — Bothriomiris sulawesicus, 3: Indonesia: Sulawesi Utara, Dumoga-Bone N. P; 5 — D. cognata (holotype); 6 — D. colubrina (holotype); 7 — D. conspersa; 8 — D. kerzhneri (paratype, 3); 9 — D. kotejai (paratype); 10 – 11 — D. tenmalai: 10 — paratype, Ƥ, 11 — holotype, 3; 12 — D. webbi (holotype); 13 – 14 — D. spatulatiformis: 13 — paratype, 14 — holotype; 15 — D. tibialis (3: Malacca, Perak); 16 — D. ulu (holotype); 17 — D. viklundi (holotype); 18 — L. scutellaris, Ƥ: Sarawak: Gunong Mulu Nat. Park; Long Pala base camp, ii – vii 1978." httpUri="https://zenodo.org/record/214716/files/figure.png" pageId="33" pageNumber="34">Figs. 13–14, 16–17</figureCitation>
); DSS stout and large, occupying most of the endosoma, distinctly broadened apically; AES distinctly smaller than the DSS and not embracing the latter. 
<taxonomicName id="4C064D55FFEFA92E40DCFFFC4E2DE498" box="[397,605,1741,1765]" class="Insecta" family="Miridae" genus="Dashymeniella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="spatulatiformis">
<emphasis id="B972EAC4FFEFA92E40DCFFFC4DD1E498" box="[397,417,1742,1765]" italics="true" pageId="33" pageNumber="34">D</emphasis>
. 
<emphasis id="B972EAC4FFEFA92E40E2FFFF4E2DE498" box="[435,605,1741,1765]" italics="true" pageId="33" pageNumber="34">spatulatiformis</emphasis>
</taxonomicName>
differs, however, by having the rugose frons, the pronotal calli weakly raised, and by the differing shape of the endosomal sclerites (
<figureCitation id="133D2A53FFEFA92E4208FFC3485EE574" box="[857,1070,1777,1801]" captionStart="FIGURES 96 – 107" captionStartId="37.[151,264,1357,1380]" captionTargetBox="[152,1435,207,1330]" captionTargetId="figure@37.[151,1436,193,1336]" captionTargetPageId="37" captionText="FIGURES 96 – 107. Male genitalia of Dashymeniella spatulatiformis (96 – 98), Dashymeniella tibialis (99 – 101), Dashymeniella viklundi (102 – 104), and Dashymeniella ulu (105 – 107). D. spatulatiformis: 96 — endosoma, dorsal view; 97 — left paramere, right lateral view; 98 — right paramere, dorsal view. D. tibialis: 99 — endosoma, dorsal view; 100 — left paramere, right lateral view; 101 — right paramere, dorsal view. D. viklundi: 102 — endosoma, dorsal view; 103 — left paramere, right lateral view; 104 — right paramere, dorsal view. D. ulu. 105 — endosoma, dorsal view; 106 — left paramere, right lateral view; 107 — right paramere, dorsal view. AES = anterior endosomal sclerite; DSS = sclerotized portion of ductus seminis inside endosoma; MES = mesial endosomal sclerite; PES = posterior endosomal sclerite. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/214733/files/figure.png" pageId="33" pageNumber="34">Figs. 96, 102, 105</figureCitation>
); right paramere with the apical process narrower than remainder of the paramere (
<figureCitation id="133D2A53FFEFA92E439DFE274FECE550" box="[716,924,1813,1837]" captionStart="FIGURES 96 – 107" captionStartId="37.[151,264,1357,1380]" captionTargetBox="[152,1435,207,1330]" captionTargetId="figure@37.[151,1436,193,1336]" captionTargetPageId="37" captionText="FIGURES 96 – 107. Male genitalia of Dashymeniella spatulatiformis (96 – 98), Dashymeniella tibialis (99 – 101), Dashymeniella viklundi (102 – 104), and Dashymeniella ulu (105 – 107). D. spatulatiformis: 96 — endosoma, dorsal view; 97 — left paramere, right lateral view; 98 — right paramere, dorsal view. D. tibialis: 99 — endosoma, dorsal view; 100 — left paramere, right lateral view; 101 — right paramere, dorsal view. D. viklundi: 102 — endosoma, dorsal view; 103 — left paramere, right lateral view; 104 — right paramere, dorsal view. D. ulu. 105 — endosoma, dorsal view; 106 — left paramere, right lateral view; 107 — right paramere, dorsal view. AES = anterior endosomal sclerite; DSS = sclerotized portion of ductus seminis inside endosoma; MES = mesial endosomal sclerite; PES = posterior endosomal sclerite. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/214733/files/figure.png" pageId="33" pageNumber="34">Figs. 98, 104, 107</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31C655DFFEFA92D4198FE0A4FF0E102" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
<paragraph id="8BB936D6FFEFA92E4198FE0A49ECE59C" blockId="33.[151,1436,1452,2017]" pageId="33" pageNumber="34">
<emphasis id="B972EAC4FFEFA92E4198FE0A4D24E52C" bold="true" box="[201,340,1848,1873]" pageId="33" pageNumber="34">Description</emphasis>
. 
<emphasis id="B972EAC4FFEFA92E4033FE0B4DEFE52C" bold="true" box="[354,415,1849,1873]" pageId="33" pageNumber="34">Male</emphasis>
. 
<emphasis id="B972EAC4FFEFA92E40FCFE0A4E1DE52C" bold="true" box="[429,621,1848,1873]" pageId="33" pageNumber="34">COLORATION</emphasis>
(
<figureCitation id="133D2A53FFEFA92E432FFE0B4F78E52C" box="[638,776,1849,1873]" captionStart="FIGURES 1 – 18" captionStartId="5.[151,264,1825,1848]" captionTargetBox="[186,1404,198,1790]" captionTargetId="figure@5.[175,1411,193,1805]" captionTargetPageId="5" captionText="FIGURES 1 – 18. Dorsal habitus color photographs of the Oriental species of the tribe Bothriomirini: 1, 2 — Bothriomiris dissimulans, Ƥ and 3: Malacca, Perak; 3 — Bothriomiris lugubris (lectotype); 4 — Bothriomiris sulawesicus, 3: Indonesia: Sulawesi Utara, Dumoga-Bone N. P; 5 — D. cognata (holotype); 6 — D. colubrina (holotype); 7 — D. conspersa; 8 — D. kerzhneri (paratype, 3); 9 — D. kotejai (paratype); 10 – 11 — D. tenmalai: 10 — paratype, Ƥ, 11 — holotype, 3; 12 — D. webbi (holotype); 13 – 14 — D. spatulatiformis: 13 — paratype, 14 — holotype; 15 — D. tibialis (3: Malacca, Perak); 16 — D. ulu (holotype); 17 — D. viklundi (holotype); 18 — L. scutellaris, Ƥ: Sarawak: Gunong Mulu Nat. Park; Long Pala base camp, ii – vii 1978." httpUri="https://zenodo.org/record/214716/files/figure.png" pageId="33" pageNumber="34">Figs. 13–14</figureCitation>
). Dorsum varying from brown to dark brown. 
<emphasis id="B972EAC4FFEFA92E4446FE0B4925E52F" bold="true" box="[1303,1365,1849,1874]" italics="true" pageId="33" pageNumber="34">Head</emphasis>
. Dark brown; vertex, maxillary and mandibular plates, and inner portion of gula somewhat paler than remainder of head; antennae dull yellowish; labium varying from dull yellowish with brown areas to reddish brown. 
<emphasis id="B972EAC4FFEFA92E4585FEB34959E5E7" bold="true" box="[1236,1321,1921,1946]" italics="true" pageId="33" pageNumber="34">Thorax</emphasis>
. 
<emphasis id="B972EAC4FFEFA92E4466FEB34C90E5C0" italics="true" pageId="33" pageNumber="34">Thoracic pleura</emphasis>
. Dark brown. 
<emphasis id="B972EAC4FFEFA92E40D0FE944DC6E5C0" box="[385,438,1958,1981]" italics="true" pageId="33" pageNumber="34">Legs</emphasis>
. Coxae brown; femora brown, dull yellowish apically; pro- and mesotibia brownish, with paler dull yellowish apex; metatibia brown, with pale yellow annulation apically; tarsi dull yellowish.
</paragraph>
<paragraph id="8BB936D6FFECA92D41C6F9A5482DE3B2" blockId="34.[151,1437,151,1184]" pageId="34" pageNumber="35">
<emphasis id="B972EAC4FFECA92D41C6F9A54E42E2CD" bold="true" box="[151,562,151,176]" pageId="34" pageNumber="35">STRUCTURE AND VESTITURE</emphasis>
(
<figureCitation id="133D2A53FFECA92D4313F9A54EBBE2D2" box="[578,715,151,175]" captionStart="FIGURES 1 – 18" captionStartId="5.[151,264,1825,1848]" captionTargetBox="[186,1404,198,1790]" captionTargetId="figure@5.[175,1411,193,1805]" captionTargetPageId="5" captionText="FIGURES 1 – 18. Dorsal habitus color photographs of the Oriental species of the tribe Bothriomirini: 1, 2 — Bothriomiris dissimulans, Ƥ and 3: Malacca, Perak; 3 — Bothriomiris lugubris (lectotype); 4 — Bothriomiris sulawesicus, 3: Indonesia: Sulawesi Utara, Dumoga-Bone N. P; 5 — D. cognata (holotype); 6 — D. colubrina (holotype); 7 — D. conspersa; 8 — D. kerzhneri (paratype, 3); 9 — D. kotejai (paratype); 10 – 11 — D. tenmalai: 10 — paratype, Ƥ, 11 — holotype, 3; 12 — D. webbi (holotype); 13 – 14 — D. spatulatiformis: 13 — paratype, 14 — holotype; 15 — D. tibialis (3: Malacca, Perak); 16 — D. ulu (holotype); 17 — D. viklundi (holotype); 18 — L. scutellaris, Ƥ: Sarawak: Gunong Mulu Nat. Park; Long Pala base camp, ii – vii 1978." httpUri="https://zenodo.org/record/214716/files/figure.png" pageId="34" pageNumber="35">Figs. 13–14</figureCitation>
, 
<figureCitation id="133D2A53FFECA92D4387F9A54E85E2D2" box="[726,757,151,175]" captionStart="FIGURES 49 – 54" captionStartId="17.[151,264,1068,1091]" captionTargetBox="[151,1435,447,1047]" captionTargetId="figure@17.[151,1435,447,1047]" captionTargetPageId="17" captionText="FIGURES 49 – 54. Left lateral view: 49 — Bothriomiris sulawesicus; 50 — Dashymenia kerzhneri; 51 — D. cognata; 52 — Dashymeniella spatulatiformis; 53 — Dashymeniella ulu; 54 — Leprocapsus scutellaris." httpUri="https://zenodo.org/record/214722/files/figure.png" pageId="34" pageNumber="35">52</figureCitation>
, 
<figureCitation id="133D2A53FFECA92D4250F9A54F3CE2D2" box="[769,844,151,175]" captionStart="FIGURES 88 – 95" captionStartId="36.[151,264,1922,1945]" captionTargetBox="[227,1354,193,1884]" captionTargetId="figure@36.[227,1355,193,1885]" captionTargetPageId="36" captionText="FIGURES 88 – 95. Scanning electron micrographs of Dashymeniella ulu (88 – 93) and Dashymeniella spatulatiformis (94 – 95): 88 – 89 — structure and vestiture of hemelytron; 90 — metafemoral trichobothria; 91 — single metafemoral trichobothrium; 92 – 93 — pretarsal structure; 94 — structure and vestiture of antennal segment I; 95 — structure and vestiture of antennal segment II." httpUri="https://zenodo.org/record/214732/files/figure.png" pageId="34" pageNumber="35">94–95</figureCitation>
). Suboval, dorsal surface strongly shining, glabrous. 
<emphasis id="B972EAC4FFECA92D41C6F9894CA5E2A9" bold="true" box="[151,213,187,212]" italics="true" pageId="34" pageNumber="35">Head</emphasis>
. Strongly shining, glabrous; vertex and frons with sparse, deep punctation; frons strongly rugose; antennal segment I covered with very sparse, fine setae, narrowed basally, apical half nearly cylindrical; segment II gradually thickened towards apex, with strongly shining basal portion, covered with moderately dense, long, semirecumbent setae; apex of labium reaching metacoxae. 
<emphasis id="B972EAC4FFECA92D4262F8154FF8E33D" bold="true" box="[819,904,295,320]" italics="true" pageId="34" pageNumber="35">Thorax</emphasis>
. 
<emphasis id="B972EAC4FFECA92D42C9F81B4877E33D" box="[920,1031,297,320]" italics="true" pageId="34" pageNumber="35">Pronotum</emphasis>
. Punctation moderately dense; calli weakly raised, inpunctate; anterior margin separated from remainder of pronotum by relatively deep, transverse incision. 
<emphasis id="B972EAC4FFECA92D4053F8424D00E3F5" box="[258,368,368,392]" italics="true" pageId="34" pageNumber="35">Scutellum</emphasis>
. With moderately dense punctation and longitudinal, weakly raised elevation. 
<emphasis id="B972EAC4FFECA92D4443F84249E5E3F5" box="[1298,1429,368,392]" italics="true" pageId="34" pageNumber="35">Hemelytron</emphasis>
. Covered with moderately dense punctation. 
<emphasis id="B972EAC4FFECA92D43C3F8A74EB7E3D1" box="[658,711,405,428]" italics="true" pageId="34" pageNumber="35">Legs</emphasis>
. Short; coxae, femora, and bases of tibiae covered with sparse setae; tibiae basally shining, apically matte, covered with dense, semirecumbent setae.
</paragraph>
<paragraph id="8BB936D6FFECA92D4198F8E94D1EE0F9" blockId="34.[151,1437,151,1184]" pageId="34" pageNumber="35">
<emphasis id="B972EAC4FFECA92D4198F8E94D06E389" bold="true" box="[201,374,475,500]" italics="true" pageId="34" pageNumber="35">Male genitalia</emphasis>
. 
<emphasis id="B972EAC4FFECA92D40DAF8EE4E72E389" box="[395,514,476,500]" italics="true" pageId="34" pageNumber="35">Endosoma</emphasis>
(
<figureCitation id="133D2A53FFECA92D4346F8EE4E02E389" box="[535,626,476,500]" captionStart="FIGURES 96 – 107" captionStartId="37.[151,264,1357,1380]" captionTargetBox="[152,1435,207,1330]" captionTargetId="figure@37.[151,1436,193,1336]" captionTargetPageId="37" captionText="FIGURES 96 – 107. Male genitalia of Dashymeniella spatulatiformis (96 – 98), Dashymeniella tibialis (99 – 101), Dashymeniella viklundi (102 – 104), and Dashymeniella ulu (105 – 107). D. spatulatiformis: 96 — endosoma, dorsal view; 97 — left paramere, right lateral view; 98 — right paramere, dorsal view. D. tibialis: 99 — endosoma, dorsal view; 100 — left paramere, right lateral view; 101 — right paramere, dorsal view. D. viklundi: 102 — endosoma, dorsal view; 103 — left paramere, right lateral view; 104 — right paramere, dorsal view. D. ulu. 105 — endosoma, dorsal view; 106 — left paramere, right lateral view; 107 — right paramere, dorsal view. AES = anterior endosomal sclerite; DSS = sclerotized portion of ductus seminis inside endosoma; MES = mesial endosomal sclerite; PES = posterior endosomal sclerite. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/214733/files/figure.png" pageId="34" pageNumber="35">Fig. 96</figureCitation>
). DSS with broadened, rounded, spoon shaped apical portion; AES gradually becoming thinner towards apex; MES stout, large, hook-shaped; PES largest, stout, gradually becoming thinner toward apex. 
<emphasis id="B972EAC4FFECA92D40DBFB164E5BE041" box="[394,555,548,572]" italics="true" pageId="34" pageNumber="35">Left paramere</emphasis>
(
<figureCitation id="133D2A53FFECA92D436DFB164EE2E041" box="[572,658,548,572]" captionStart="FIGURES 96 – 107" captionStartId="37.[151,264,1357,1380]" captionTargetBox="[152,1435,207,1330]" captionTargetId="figure@37.[151,1436,193,1336]" captionTargetPageId="37" captionText="FIGURES 96 – 107. Male genitalia of Dashymeniella spatulatiformis (96 – 98), Dashymeniella tibialis (99 – 101), Dashymeniella viklundi (102 – 104), and Dashymeniella ulu (105 – 107). D. spatulatiformis: 96 — endosoma, dorsal view; 97 — left paramere, right lateral view; 98 — right paramere, dorsal view. D. tibialis: 99 — endosoma, dorsal view; 100 — left paramere, right lateral view; 101 — right paramere, dorsal view. D. viklundi: 102 — endosoma, dorsal view; 103 — left paramere, right lateral view; 104 — right paramere, dorsal view. D. ulu. 105 — endosoma, dorsal view; 106 — left paramere, right lateral view; 107 — right paramere, dorsal view. AES = anterior endosomal sclerite; DSS = sclerotized portion of ductus seminis inside endosoma; MES = mesial endosomal sclerite; PES = posterior endosomal sclerite. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/214733/files/figure.png" pageId="34" pageNumber="35">Fig. 97</figureCitation>
). Rounded; apical process somewhat sharply pointed apically. 
<emphasis id="B972EAC4FFECA92D440EFB164D73E01D" italics="true" pageId="34" pageNumber="35">Right paramere</emphasis>
(
<figureCitation id="133D2A53FFECA92D4047FB754D1FE022" box="[278,367,583,607]" captionStart="FIGURES 96 – 107" captionStartId="37.[151,264,1357,1380]" captionTargetBox="[152,1435,207,1330]" captionTargetId="figure@37.[151,1436,193,1336]" captionTargetPageId="37" captionText="FIGURES 96 – 107. Male genitalia of Dashymeniella spatulatiformis (96 – 98), Dashymeniella tibialis (99 – 101), Dashymeniella viklundi (102 – 104), and Dashymeniella ulu (105 – 107). D. spatulatiformis: 96 — endosoma, dorsal view; 97 — left paramere, right lateral view; 98 — right paramere, dorsal view. D. tibialis: 99 — endosoma, dorsal view; 100 — left paramere, right lateral view; 101 — right paramere, dorsal view. D. viklundi: 102 — endosoma, dorsal view; 103 — left paramere, right lateral view; 104 — right paramere, dorsal view. D. ulu. 105 — endosoma, dorsal view; 106 — left paramere, right lateral view; 107 — right paramere, dorsal view. AES = anterior endosomal sclerite; DSS = sclerotized portion of ductus seminis inside endosoma; MES = mesial endosomal sclerite; PES = posterior endosomal sclerite. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/214733/files/figure.png" pageId="34" pageNumber="35">Fig. 98</figureCitation>
). Body nearly parallel-sided; apical process strongly narrower than remainder of paramere, sharpened apically.
</paragraph>
<paragraph id="8BB936D6FFECA92D4198FBA248CFE0DA" blockId="34.[151,1437,151,1184]" box="[201,1215,655,680]" pageId="34" pageNumber="35">
<emphasis id="B972EAC4FFECA92D4198FBA24D50E0D5" bold="true" box="[201,288,656,680]" pageId="34" pageNumber="35">Female</emphasis>
. Similar to male in coloration, structure, and vestiture but somewhat larger in size.
</paragraph>
<paragraph id="8BB936D6FFECA92D4198FB864FF0E102" blockId="34.[151,1437,151,1184]" pageId="34" pageNumber="35">
<emphasis id="B972EAC4FFECA92D4198FB864D0FE0B1" bold="true" box="[201,383,692,716]" pageId="34" pageNumber="35">Measurements.</emphasis>
Ƥ 
<emphasis id="B972EAC4FFECA92D40CAFB814DD3E0B1" bold="true" box="[411,419,691,716]" pageId="34" pageNumber="35">/</emphasis>
3 (n=3, 
<typeStatus id="54BD8874FFECA92D435BFB864E1CE0B1" box="[522,620,692,716]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
measurements in parentheses): body length 3.9/3.6–3.7 (3.7), width 2.0/ 1.75–1.77 (1.77); length of head 0.41/0.34–0.37 (0.34), width 1.1/0.95–0.97 (0.95), diameter of eye in dorsal view 0.25/0.22–0.25 (0.22); length of antennal segment I 0.44/0.45–0.46 (0.45), II 1.1/0.95–0.97 (0.97), III (3) 0.55, IV (3) 0.9. Labium. Length segment (3, immeasurable in Ƥ) I 0.46–0.5 (0.5), II 0.45–0.5 (0.5), III 0.3–0.36 (0.3), IV 0.28–0.3 (0.3). Pronotum. Length 1.16/0.97 (0.97), width of anterior margin 0.78/0.78–0.8 (0.8), lateral margin 1.0/ 0.78–0.81 (0.78), width of posterior margin 1.84/1.62–1.65 (1.65).
</paragraph>
</subSubSection>
<subSubSection id="C31C655DFFECA92D4198FABE4E90E1BA" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BB936D6FFECA92D4198FABE4E90E1BA" blockId="34.[151,1437,151,1184]" pageId="34" pageNumber="35">
<emphasis id="B972EAC4FFECA92D4198FABE4D38E1D9" bold="true" box="[201,328,908,932]" pageId="34" pageNumber="35">Etymology</emphasis>
. The specific epithet is derived from the Latin “spatula,” (spoon) and “forma,” (shape) and is used to denote the spoon shaped endosomal sclerite DSS.
</paragraph>
</subSubSection>
<subSubSection id="C31C655DFFECA92D4198FAE14F98E191" box="[201,1000,979,1004]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BB936D6FFECA92D4198FAE14F98E191" blockId="34.[151,1437,151,1184]" box="[201,1000,979,1004]" pageId="34" pageNumber="35">
<emphasis id="B972EAC4FFECA92D4198FAE14D2BE191" bold="true" box="[201,347,979,1004]" pageId="34" pageNumber="35">Distribution</emphasis>
(
<figureCitation id="133D2A53FFECA92D403BFAE64DCEE191" box="[362,446,980,1004]" captionStart="FIGURE 87" captionStartId="33.[151,250,1243,1266]" captionTargetBox="[167,1404,225,1200]" captionTargetId="figure@33.[151,1436,193,1222]" captionTargetPageId="33" captionText="FIGURE 87. Distribution map of Dashymeniella and Leprocapsus." httpUri="https://zenodo.org/record/214731/files/figure.png" pageId="34" pageNumber="35">Fig. 87</figureCitation>
). 
<collectingCountry id="F3117646FFECA92D4083FAE64E6EE191" box="[466,542,980,1004]" name="Brunei" pageId="34" pageNumber="35">Brunei</collectingCountry>
(Temburong: 
<taxonomicName id="4C064D55FFECA92D43EFFAE64E9CE191" box="[702,748,980,1004]" class="Insecta" family="Miridae" genus="Dashymeniella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="38" phylum="Arthropoda" rank="species" species="ulu">Ulu</taxonomicName>
), 
<collectingCountry id="F3117646FFECA92D43AEFAE64F16E191" box="[767,870,980,1004]" name="Malaysia" pageId="34" pageNumber="35">Malaysia</collectingCountry>
(
<collectingCountry id="F3117646FFECA92D4224FAE64FACE191" box="[885,988,980,1004]" name="Malaysia" pageId="34" pageNumber="35">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C31C655DFFECA92D4198FACA4D48E6E2" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BB936D6FFECA92D4198FACA4D48E6E2" blockId="34.[151,1437,151,1184]" pageId="34" pageNumber="35">
<emphasis id="B972EAC4FFECA92D4198FACA4D9EE66D" bold="true" box="[201,494,1015,1040]" pageId="34" pageNumber="35">
<typeStatus id="54BD8874FFECA92D4198FACA4D74E66D" box="[201,260,1016,1040]" pageId="34" pageNumber="35">Type</typeStatus>
material. 
<typeStatus id="54BD8874FFECA92D40D0FACA4D9EE66D" box="[385,494,1016,1040]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</emphasis>
3: 
<collectingCountry id="F3117646FFECA92D434DFACA4EF9E672" box="[540,649,1016,1039]" name="Brunei" pageId="34" pageNumber="35">BRUNEI</collectingCountry>
: 
<taxonomicName id="4C064D55FFECA92D43C4FAC54EB2E672" box="[661,706,1015,1039]" class="Insecta" family="Miridae" genus="Dashymeniella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="38" phylum="Arthropoda" rank="species" species="ulu">Ulu</taxonomicName>
, Temburong ridge, M. C. Day, B. M., 1983-75; 
<emphasis id="B972EAC4FFECA92D45BBFACA4910E66D" bold="true" box="[1258,1376,1016,1040]" pageId="34" pageNumber="35">
<typeStatus id="54BD8874FFECA92D45BBFACA4910E66D" box="[1258,1376,1016,1040]" pageId="34" pageNumber="35" type="paratype">paratypes</typeStatus>
</emphasis>
: 
<emphasis id="B972EAC4FFECA92D4420FAC549F1E66D" bold="true" box="[1393,1409,1015,1040]" pageId="34" pageNumber="35">1</emphasis>
3: approx. 
<quantity id="4CFE9B33FFECA92D41A4FD2E4D4EE649" box="[245,318,1052,1076]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="m" value="150.0">150 m</quantity>
alt. 
<date id="FFB81016FFECA92D4025FD2E4E62E649" box="[372,530,1052,1076]" pageId="34" pageNumber="35" value="1992-10">October 1992</date>
, J H Martin coll. B M 1992 – 172; Malaise trap 
<collectingCountry id="F3117646FFECA92D451BFD2F48C7E649" box="[1098,1207,1053,1076]" name="Brunei" pageId="34" pageNumber="35">BRUNEI</collectingCountry>
: Temburong, Steepslope forest above Kuala Belalong; 
<emphasis id="B972EAC4FFECA92D436EFD0D4E3DE625" bold="true" box="[575,589,1087,1112]" pageId="34" pageNumber="35">1</emphasis>
Ƥ: 
<collectingCountry id="F3117646FFECA92D43D3FD0D4F7FE62A" box="[642,783,1087,1111]" name="Malaysia" pageId="34" pageNumber="35">SARAWAK</collectingCountry>
: Foot of Mt. Dulit. Junction of Rivers Tinjar &amp; Lejok, 
<date id="FFB81016FFECA92D41C6FD564D62E601" box="[151,274,1124,1148]" pageId="34" pageNumber="35" value="1932-09-22">22.ix.1932</date>
; Native collected; Old secondary forest; Oxford Univ. Exp. B. M. Hobby&amp; A. W. Moore. B.M. 1933- 254 (
<emphasis id="B972EAC4FFECA92D4180FDBA4D58E6DD" bold="true" box="[209,296,1160,1184]" pageId="34" pageNumber="35">
<collectionCode id="ED17AE13FFECA92D4180FDBA4D58E6DD" box="[209,296,1160,1184]" pageId="34" pageNumber="35">BMNH</collectionCode>
</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>