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<document id="BE9F12355F4817A8EABAE0B71272B373" ID-CLB-Dataset="3646" ID-DOI="10.11646/zootaxa.5067.1.5" ID-GBIF-Dataset="74f3ce16-7e8f-420a-b76e-d306ea2a42ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="5656552" ID-ZooBank="4763DE40-3A0D-4F23-9381-3C4754A59AE4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636445280528" checkinUser="plazi" docAuthor="Nozaki, Takao &amp; Park, Sun-Jin" docDate="2021" docId="03AF878FFFC4FFAA0C933951FE2FFAFF" docLanguage="en" docName="zootaxa.5067.1.5.pdf" docOrigin="Zootaxa 5067 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila yamamotoi Nozaki &amp; Park 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="100" masterDocId="FF96FFF7FFC6FFA90C043D33FFE8FF86" masterDocTitle="Two new species of the genus Rhyacophila Pictet (Trichoptera, Rhyacophilidae) from Korea and Japan" masterLastPageNumber="105" masterPageNumber="97" pageNumber="99" updateTime="1701999324714" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="321DE305C4E9603521196DA9830626DF">Two new species of the genus Rhyacophila Pictet (Trichoptera, Rhyacophilidae) from Korea and Japan</mods:title>
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<mods:namePart id="5842D3C459F740D4A7AD6E7BF085A693">Nozaki, Takao</mods:namePart>
<mods:affiliation id="DAB9FD0030217CC94C5DFC3064A3C9D9">3 - 16 - 15, Midorigaoka, Ninomiya-machi, Naka-gun, Kanagawa, 259 - 0132 Japan.</mods:affiliation>
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<mods:namePart id="749A79A9BE24CC5C0542F7D239BAF7A4">Park, Sun-Jin</mods:namePart>
<mods:affiliation id="ED4943053C92DD0CF05A86C9FC6083E4">Department of Life Science, Kyonggi University, Suwon 16227, Korea.</mods:affiliation>
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<treatment id="03AF878FFFC4FFAA0C933951FE2FFAFF" ID-DOI="http://doi.org/10.5281/zenodo.5690986" ID-GBIF-Taxon="190304775" ID-Zenodo-Dep="5690986" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF" httpUri="http://treatment.plazi.org/id/03AF878FFFC4FFAA0C933951FE2FFAFF" lastPageId="3" lastPageNumber="100" pageId="2" pageNumber="99">
<subSubSection id="C31C6512FFC4FFAB0C933951FD6AFBFB" box="[151,642,1122,1149]" pageId="2" pageNumber="99" type="nomenclature">
<paragraph id="8BB93699FFC4FFAB0C933951FD6AFBFB" blockId="2.[151,642,1122,1185]" box="[151,642,1122,1149]" pageId="2" pageNumber="99">
<heading id="D0F181F5FFC4FFAB0C933951FD6AFBFB" bold="true" box="[151,642,1122,1149]" fontSize="11" level="1" pageId="2" pageNumber="99" reason="1">
<emphasis id="B972EA8BFFC4FFAB0C933951FD6AFBFB" bold="true" box="[151,642,1122,1149]" pageId="2" pageNumber="99">
<taxonomicName id="4C064D1AFFC4FFAB0C933951FDF5FBFA" authority="Nozaki" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[151,541,1122,1149]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="yamamotoi" status="sp. nov.">
<emphasis id="B972EA8BFFC4FFAB0C933951FE56FBFB" bold="true" box="[151,446,1122,1149]" italics="true" pageId="2" pageNumber="99">Rhyacophila yamamotoi</emphasis>
Nozaki
</taxonomicName>
<taxonomicNameLabel id="A24157F0FFC4FFAB0E203950FD6AFBFB" box="[548,642,1123,1149]" pageId="2" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC4FFAB0C9339B4FD62FAA6" pageId="2" pageNumber="99" type="description">
<paragraph id="8BB93699FFC4FFAB0C9339B4FECFFB27" blockId="2.[151,642,1122,1185]" box="[151,295,1158,1185]" pageId="2" pageNumber="99">
(
<figureCitation id="133D2A1CFFC4FFAB0C9B39B4FF00FB27" box="[159,232,1158,1185]" captionStart="FIGURE 47" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 47. Rhyacophila spp., female. 4A4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="2" pageNumber="99">Figs 5</figureCitation>
,
<figureCitation id="133D2A1CFFC4FFAB0CF039B4FEEBFB27" box="[244,259,1159,1185]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">8</figureCitation>
,
<figureCitation id="133D2A1CFFC4FFAB0D1439B4FEF7FB27" box="[272,287,1159,1185]" captionStart="FIGURE 9" captionStartId="7.[151,250,1110,1134]" captionTargetBox="[155,1428,281,1079]" captionTargetId="figure-64@7.[151,1436,273,1087]" captionTargetPageId="7" captionText="FIGURE 9. Distributions of 4 Rhyacophila species in East Asia. Distribution areas of R. confissa and R. vicina are cited from Ivanov (2011), Park &amp; Kong (2020), and Yang et al. (2016)." figureDoi="http://doi.org/10.5281/zenodo.5656565" httpUri="https://zenodo.org/record/5656565/files/figure.png" pageId="2" pageNumber="99">9</figureCitation>
)
</paragraph>
<paragraph id="8BB93699FFC4FFAB0C9339F8FDAAFB65" blockId="2.[151,667,1227,1313]" box="[151,578,1227,1251]" pageId="2" pageNumber="99">
<taxonomicName id="4C064D1AFFC4FFAB0C9339F8FEBDFB65" box="[151,341,1227,1251]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="undefined-b">
<emphasis id="B972EA8BFFC4FFAB0C9339F8FEFFFB65" box="[151,279,1227,1251]" italics="true" pageId="2" pageNumber="99">Rhyacophila</emphasis>
sp. B
</taxonomicName>
:
<bibRefCitation id="EF974B68FFC4FFAB0D5B39F8FE37FB65" author="Morita, H." box="[351,479,1227,1251]" pageId="2" pageNumber="99" pagination="3 - 7" refId="ref5488" refString="Morita, H. (2009) Suzuka-sammyaku hokubu no tobikera-so [Trichoptera fauna of northern part of Suzuka Mountains]. Fujiwaradake, 31, 3 - 7. [in Japanese]" type="journal article" year="2009">Morita 2009</bibRefCitation>
, 67, list.
</paragraph>
<paragraph id="8BB93699FFC4FFAB0C9339D9FD73FA84" blockId="2.[151,667,1227,1313]" box="[151,667,1258,1282]" pageId="2" pageNumber="99">
<taxonomicName id="4C064D1AFFC4FFAB0C9339D9FEB9FA84" box="[151,337,1258,1282]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="undefined-4">
<emphasis id="B972EA8BFFC4FFAB0C9339D9FEFFFA84" box="[151,279,1258,1282]" italics="true" pageId="2" pageNumber="99">Rhyacophila</emphasis>
sp. 4
</taxonomicName>
:
<bibRefCitation id="EF974B68FFC4FFAB0D5F39D9FDA4FA84" author="Kawase, N. &amp; Morita, H." box="[347,588,1258,1282]" pageId="2" pageNumber="99" pagination="31 - 50" refId="ref5314" refString="Kawase, N. &amp; Morita, H. (2010) The caddisfly (Insecta, Trichoptera) fauna of the Suzuka Mounatins. Biology of Inland Waters, 25, 31 - 50. [in Japanese with English abstract]" type="journal article" year="2010">Kawase &amp; Morita 2010</bibRefCitation>
, 36, list
</paragraph>
<paragraph id="8BB93699FFC4FFAB0C93383BFD62FAA6" blockId="2.[151,667,1227,1313]" box="[151,650,1288,1313]" pageId="2" pageNumber="99">
<taxonomicName id="4C064D1AFFC4FFAB0C93383BFEB8FAA7" box="[151,336,1288,1313]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="B972EA8BFFC4FFAB0C93383BFEFFFAA6" box="[151,279,1288,1312]" italics="true" pageId="2" pageNumber="99">Rhyacophila</emphasis>
sp. 3
</taxonomicName>
:
<bibRefCitation id="EF974B68FFC4FFAB0D5E383AFDA8FAA7" author="Yamamoto, E. &amp; Ito, T." box="[346,576,1289,1313]" pageId="2" pageNumber="99" pagination="6 - 21" refId="ref5951" refString="Yamamoto, E. &amp; Ito, T. (2014) Caddisflies (Trichoptera) collected from Izugataniyama, Kumakogen-cho, Shikoku, Japan in 2012 - 2013. Shikokukogera, 14, 6 - 21. [in Japanese]." type="journal article" year="2014">Yamamoto &amp; Ito 2014</bibRefCitation>
, 7, list.
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC4FFAA0C93387DFDFBFDE7" lastPageId="3" lastPageNumber="100" pageId="2" pageNumber="99" type="diagnosis">
<paragraph id="8BB93699FFC4FFAB0C93387DFB64F90F" blockId="2.[151,1437,1358,2033]" pageId="2" pageNumber="99">
<emphasis id="B972EA8BFFC4FFAB0C93387DFEFAFAEE" bold="true" box="[151,274,1358,1384]" pageId="2" pageNumber="99">Diagnosis.</emphasis>
Male and female of this species are most similar to those of
<taxonomicName id="4C064D1AFFC4FFAB0FB3387CFBCFFAEF" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[951,1063,1359,1385]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC4FFAB0FB3387CFBCFFAEF" box="[951,1063,1359,1385]" italics="true" pageId="2" pageNumber="99">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC4FFAB0829387CFB6FFAEF" bold="true" box="[1069,1159,1359,1385]" pageId="2" pageNumber="99">
<taxonomicNameLabel id="A24157F0FFC4FFAB0829387CFB6FFAEF" box="[1069,1159,1359,1385]" pageId="2" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but can be distinguished by characters of the genitalic morphology. In males, the apex of each preanal appendage of this species is weakly bilobed apically in lateral aspect (
<figureCitation id="133D2A1CFFC4FFAB0E1038A4FD82FA37" box="[532,618,1431,1457]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8B</figureCitation>
) but is rounded in
<taxonomicName id="4C064D1AFFC4FFAB0F4438A4FC59FA37" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[832,945,1431,1457]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="kangae">
<emphasis id="B972EA8BFFC4FFAB0F4438A4FC59FA37" box="[832,945,1431,1457]" italics="true" pageId="2" pageNumber="99">R. kangae</emphasis>
</taxonomicName>
(
<figureCitation id="133D2A1CFFC4FFAB0FC538A4FBF2FA37" box="[961,1050,1431,1457]" captionStart="FIGURES 13" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 13. Rhyacophila spp., male. 1A1F, Rhyacophila kangae sp. nov. 1A1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="2" pageNumber="99">Fig. 1A</figureCitation>
). Each paramere is expanded subapically in this species (
<figureCitation id="133D2A1CFFC4FFAB0DA23888FE13FA53" box="[422,507,1467,1493]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8E</figureCitation>
) but is club-like in
<taxonomicName id="4C064D1AFFC4FFAB0EDE3888FCA2FA53" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[730,842,1467,1493]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="kangae">
<emphasis id="B972EA8BFFC4FFAB0EDE3888FCA2FA53" box="[730,842,1467,1493]" italics="true" pageId="2" pageNumber="99">R. kangae</emphasis>
</taxonomicName>
(
<figureCitation id="133D2A1CFFC4FFAB0F5E3888FC58FA53" box="[858,944,1467,1493]" captionStart="FIGURES 13" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 13. Rhyacophila spp., male. 1A1F, Rhyacophila kangae sp. nov. 1A1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="2" pageNumber="99">Fig. 1E</figureCitation>
). In the female, the posterior process of the vaginal apparatus of this species is broader and thicker than that of
<taxonomicName id="4C064D1AFFC4FFAB0F9138ECFBEEFA7F" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[917,1030,1503,1529]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="kangae">
<emphasis id="B972EA8BFFC4FFAB0F9138ECFBEEFA7F" box="[917,1030,1503,1529]" italics="true" pageId="2" pageNumber="99">R. kangae</emphasis>
</taxonomicName>
(
<figureCitation id="133D2A1CFFC4FFAB081238ECFB1FFA7F" box="[1046,1271,1502,1529]" captionStart="FIGURE 47" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 47. Rhyacophila spp., female. 4A4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="2" pageNumber="99">Figs 4E, 4F, 5E, 5F</figureCitation>
). In
<collectingCountry id="F3117609FFC4FFAB092F38ECFA86FA7F" box="[1323,1390,1503,1529]" name="Japan" pageId="2" pageNumber="99">Japan</collectingCountry>
, the male genitalia of this species are somewhat similar to that of
<taxonomicName id="4C064D1AFFC4FFAB0F3D3B30FBB8F99B" authorityName="Tsuda" authorityYear="1940" box="[825,1104,1539,1565]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="kawamurae">
<emphasis id="B972EA8BFFC4FFAB0F3D3B30FBB8F99B" box="[825,1104,1539,1565]" italics="true" pageId="2" pageNumber="99">Rhyacophila kawamurae</emphasis>
</taxonomicName>
in lateral aspect, especially in the shape of the inferior appendages, but can be easily distinguished by the shape of the complex of a pair of preanal appendages and the apicodorsal lobe of segment IX in dorsal aspect: These are bilobed from the basal 1/31/
<quantity id="4CFE9B7CFFC4FFAB096B3B78FA74F9E3" box="[1391,1436,1611,1637]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="99" unit="in" value="2.0">2 in</quantity>
this species but bearing only a minute apical notch in
<taxonomicName id="4C064D1AFFC4FFAB0EF53B5CFC7BF90F" authorityName="Tsuda" authorityYear="1940" box="[753,915,1647,1673]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="kawamurae">
<emphasis id="B972EA8BFFC4FFAB0EF53B5CFC7BF90F" box="[753,915,1647,1673]" italics="true" pageId="2" pageNumber="99">R. kawamurae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF974B68FFC4FFAB0FA53B5CFBC0F90F" author="Tsuda, M." box="[929,1064,1647,1673]" pageId="2" pageNumber="99" pagination="119 - 135" refId="ref5895" refString="Tsuda, M. (1940) Zur kenntnis der Japanischen Rhyacophilinen (Rhyacophilidae, Trichoptera). Annotationes Zoologicae Japonenses, 19, 119 - 135." type="journal article" year="1940">Tsuda 1940</bibRefCitation>
, fig. 17).
</paragraph>
<paragraph id="8BB93699FFC4FFAB0CC33BA1FC92F973" blockId="2.[151,1437,1358,2033]" pageId="2" pageNumber="99">
<emphasis id="B972EA8BFFC4FFAB0CC33BA1FEFBF92A" bold="true" box="[199,275,1682,1708]" pageId="2" pageNumber="99">Adult.</emphasis>
Specimens in alcohol mostly dark brown to black, but legs light brown. Forewings each 8.0
<quantity id="4CFE9B7CFFC4FFAB093E3BA1FA75F92B" box="[1338,1437,1682,1709]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="99" unit="mm" value="11.5">11.5 mm</quantity>
in male (n = 10), 10.211.0 mm in female (n = 7). Ratio of width of eye to distance between eyes (a/b in
<figureCitation id="133D2A1CFFC4FFAB09453B84FA7AF957" box="[1345,1426,1719,1745]" captionStart="FIGURES 13" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 13. Rhyacophila spp., male. 1A1F, Rhyacophila kangae sp. nov. 1A1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="2" pageNumber="99">Fig. 1F</figureCitation>
)
<quantity id="4CFE9B7CFFC4FFAB0C933BE8FECCF973" box="[151,292,1755,1781]" metricMagnitude="-3" metricUnit="m" metricValue="9.271" metricValueMax="10.16" metricValueMin="8.382" pageId="2" pageNumber="99" unit="in" value="0.365" valueMax="0.4" valueMin="0.33">0.330.40 in</quantity>
male (n = 10) (
<figureCitation id="133D2A1CFFC4FFAB0DD63BE8FDC4F973" box="[466,556,1755,1781]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8A</figureCitation>
),
<quantity id="4CFE9B7CFFC4FFAB0E443BE8FD25F973" box="[576,717,1755,1781]" metricMagnitude="-3" metricUnit="m" metricValue="6.858" metricValueMax="7.62" metricValueMin="6.096" pageId="2" pageNumber="99" unit="in" value="0.27" valueMax="0.3" valueMin="0.24">0.240.30 in</quantity>
female (n = 8).
</paragraph>
<paragraph id="8BB93699FFC4FFAA0CC33BCCFC30FEC7" blockId="2.[151,1437,1358,2033]" lastBlockId="3.[151,1437,151,1401]" lastPageId="3" lastPageNumber="100" pageId="2" pageNumber="99">
<emphasis id="B972EA8BFFC4FFAB0CC33BCCFE92F89E" bold="true" box="[199,378,1790,1817]" pageId="2" pageNumber="99">Male genitalia.</emphasis>
Segment IX longitudinally short in lateral aspect (
<figureCitation id="133D2A1CFFC4FFAB0FBA3BCCFBFEF89F" box="[958,1046,1791,1817]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8B</figureCitation>
), ventral half longer than dorsum; membranous along midline in dorsal aspect (
<figureCitation id="133D2A1CFFC4FFAB0E883A10FD0CF8BB" box="[652,740,1827,1853]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8C</figureCitation>
). Complex of pair of preanal appendages and apicodorsal lobe of segment IX bilobed from basal 1/31/2, widely separated, large, U-shaped in dorsal aspect (
<figureCitation id="133D2A1CFFC4FFAB08AE3A74FAE9F8E7" box="[1194,1281,1863,1889]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8C</figureCitation>
); in lateral aspect each lobe rectangular, apex slightly concave (
<figureCitation id="133D2A1CFFC4FFAB0EC33A58FCF6F803" box="[711,798,1899,1925]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8B</figureCitation>
); with 2 teeth apicomesally, dorsal one small, ventral one tiny (inset of
<figureCitation id="133D2A1CFFC4FFAB0D343ABCFE60F82F" box="[304,392,1935,1961]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8B</figureCitation>
). Anal sclerites fused basally, bilobed, each round apex with minute black denticles dorsally (
<figureCitation id="133D2A1CFFC4FFAB0CA43A80FF11F84B" box="[160,249,1971,1997]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8C</figureCitation>
). Apical band long, curved posterad, ventral margin connected to sagittal appendage of tergal band (
<figureCitation id="133D2A1CFFC4FFAB096B3A80FF52F877" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="2" pageNumber="99">Fig. 8B</figureCitation>
); sagittal appendage round trapezoidal in dorsal and ventral aspects. Basal segment of each inferior appendage thick in lateral aspect (
<figureCitation id="133D2A1CFFC5FFAA0D9D3DA4FDC0FF37" box="[409,552,151,177]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="3" pageNumber="100">Figs 8B, 8D</figureCitation>
); mesal face with ridged tendon longitudinally, extending to phallotheca (
<figureCitation id="133D2A1CFFC5FFAA096E3DA4FF05FF53" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="3" pageNumber="100">Figs 8B, 8D</figureCitation>
); posteromesal angle acute, directed mesad in dorsal aspect (
<figureCitation id="133D2A1CFFC5FFAA0FA53D88FC13FF53" box="[929,1019,187,213]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="3" pageNumber="100">Fig. 8C</figureCitation>
). Distal segment of each inferior appendage bilobed (
<figureCitation id="133D2A1CFFC5FFAA0D643DECFE1EFF7F" box="[352,502,223,249]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="3" pageNumber="100">Figs. 8B, 8D</figureCitation>
); both lobes club-like in lateral aspect, with minute spines apicomesally (
<figureCitation id="133D2A1CFFC5FFAA09303DECFA67FF7F" box="[1332,1423,223,249]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="3" pageNumber="100">Fig. 8D</figureCitation>
). In phallic apparatus, aedeagus bottle-shaped in dorsal and ventral aspects (
<figureCitation id="133D2A1CFFC5FFAA0FED3C30FBAAFE9B" box="[1001,1090,259,285]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="3" pageNumber="100">Fig. 8E</figureCitation>
); with pair of parameres, each broadened subapically, with tiny spines apically in dorsal aspect (
<figureCitation id="133D2A1CFFC5FFAA0F703C14FC23FEC7" box="[884,971,295,321]" captionStart="FIGURE 8" captionStartId="6.[151,250,1653,1677]" captionTargetBox="[154,1430,635,1623]" captionTargetId="figure-205@6.[151,1436,633,1629]" captionTargetPageId="6" captionText="FIGURE 8. Rhyacophila yamamotoi sp. nov., male. 8A, head, dorsal. 8B8E, genitalia: 8B, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 8C, dorsal; 8D, left inferior appendage, mesal; 8E, phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5656563" httpUri="https://zenodo.org/record/5656563/files/figure.png" pageId="3" pageNumber="100">Fig. 8E</figureCitation>
).
</paragraph>
<paragraph id="8BB93699FFC5FFAA0CC33C78FC52FDBB" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="100">
<emphasis id="B972EA8BFFC5FFAA0CC33C78FE7AFEE2" bold="true" box="[199,402,330,357]" pageId="3" pageNumber="100">Female genitalia.</emphasis>
Segment VIII annular, but semi-membranous along dorsal midline, with pair of dorsal weakly sclerotized extensions (
<figureCitation id="133D2A1CFFC5FFAA0DBD3C5CFDF8FE0F" box="[441,528,366,393]" captionStart="FIGURE 47" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 47. Rhyacophila spp., female. 4A4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="3" pageNumber="100">Fig. 5C</figureCitation>
); pair of apodemal rods reaching posterior end of segment VI. Segment IX membranous, with pair of dorsolateral apodemal rods extending into segment VI; fused with segment X ventrally, with pair of ventrolateral sclerotized band extending to segment X. Segment X slender, with pair of strongly sclerotized bands dorsolaterally; fused with segment XI. In vaginal apparatus, processus spermathecae claw-like in lateral aspect (
<figureCitation id="133D2A1CFFC5FFAA0CD13CCCFEC3FD9F" box="[213,299,510,537]" captionStart="FIGURE 47" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 47. Rhyacophila spp., female. 4A4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="3" pageNumber="100">Fig. 5E</figureCitation>
), semicircular in ventral aspect (
<figureCitation id="133D2A1CFFC5FFAA0E923CCCFD03FD9F" box="[662,747,510,537]" captionStart="FIGURE 47" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 47. Rhyacophila spp., female. 4A4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="3" pageNumber="100">Fig. 5F</figureCitation>
); posterior process tongue-like in ventral aspect (
<figureCitation id="133D2A1CFFC5FFAA09143CCCFA8CFD9F" box="[1296,1380,510,537]" captionStart="FIGURE 47" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 47. Rhyacophila spp., female. 4A4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="3" pageNumber="100">Fig. 5F</figureCitation>
), not more than 3 times longer than anterior width, elliptical in lateral aspect.
</paragraph>
<paragraph id="8BB93699FFC5FFAA0CC33F74FDFBFDE7" blockId="3.[151,1437,151,1401]" box="[199,531,583,609]" pageId="3" pageNumber="100">
<emphasis id="B972EA8BFFC5FFAA0CC33F74FE7EFDE7" bold="true" box="[199,406,583,609]" pageId="3" pageNumber="100">Immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC5FFAA0CC33F58FCB9FB27" pageId="3" pageNumber="100" type="materials_examined">
<paragraph id="8BB93699FFC5FFAA0CC33F58FB9FFD2F" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="100">
<materialsCitation id="3B6E3CC4FFC5FFAA0CC33F58FB9FFD2F" ID-GBIF-Occurrence="3400170318" collectingDate="2020-05-11" collectingDateMax="2020-05-20" collectingDateMin="2020-05-11" collectionCode="CBM-ZI" collectorName="E. Yamamoto" country="Japan" county="Shikoku" elevation="1225" latitude="33.56" location="Male" longLatPrecision="72" longitude="132.918" municipality="Uchiko-cho" pageId="3" pageNumber="100" specimenCode="CBM-ZI 0180217" specimenCount="1" stateProvince="Ehime" typeStatus="holotype">
<emphasis id="B972EA8BFFC5FFAA0CC33F58FED3FD03" bold="true" box="[199,315,619,645]" pageId="3" pageNumber="100">
<typeStatus id="54BD883BFFC5FFAA0CC33F58FEDEFD03" box="[199,310,619,645]" pageId="3" pageNumber="100" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8ED96042FFC5FFAA0D413F58FE97FD03" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0D413F58FE97FD03" box="[325,383,619,645]" country="Japan" county="Shikoku" latitude="33.56" longLatPrecision="72" longitude="132.918" municipality="Uchiko-cho" name="Male" pageId="3" pageNumber="100" stateProvince="Ehime">Male</location>
(in alcohol).
<location id="8ED96042FFC5FFAA0E1B3F58FD3AFD03" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0E1B3F58FD3AFD03" box="[543,722,619,645]" country="Japan" county="Shikoku" latitude="33.56" longLatPrecision="72" longitude="132.918" municipality="Uchiko-cho" name="Namakusa-dani" pageId="3" pageNumber="100" stateProvince="Ehime">Namakusa-dani</location>
,
<location id="8ED96042FFC5FFAA0EE43F58FC84FD03" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0EE43F58FC84FD03" box="[736,876,619,645]" country="Japan" county="Shikoku" latitude="33.56" longLatPrecision="72" longitude="132.918" municipality="Uchiko-cho" name="Odamiyama" pageId="3" pageNumber="100" stateProvince="Ehime">Odamiyama</location>
,
<collectingMunicipality id="6BDDACE3FFC5FFAA0F7E3F58FBE9FD03" box="[890,1025,619,645]" pageId="3" pageNumber="100">Uchiko-cho</collectingMunicipality>
,
<collectingRegion id="49C2F87BFFC5FFAA080B3F58FBB3FD03" box="[1039,1115,619,645]" country="Japan" name="Ehime" pageId="3" pageNumber="100">Ehime</collectingRegion>
,
<collectingCounty id="62D84E15FFC5FFAA086D3F58FB21FD03" box="[1129,1225,619,645]" pageId="3" pageNumber="100">Shikoku</collectingCounty>
,
<collectingCountry id="F3117609FFC5FFAA08DC3F58FAF3FD03" box="[1240,1307,619,645]" name="Japan" pageId="3" pageNumber="100">Japan</collectingCountry>
,
<geoCoordinate id="EE32505EFFC5FFAA092D3F59FA70FD03" box="[1321,1432,618,645]" degrees="33.560" direction="north" orientation="latitude" pageId="3" pageNumber="100" precision="55" value="33.56">33.560°N</geoCoordinate>
,
<geoCoordinate id="EE32505EFFC5FFAA0C933FBCFEF9FD2F" box="[151,273,655,681]" degrees="132.918" direction="east" orientation="longitude" pageId="3" pageNumber="100" precision="55" value="132.918">132.918°E</geoCoordinate>
, alt.
<quantity id="4CFE9B7CFFC5FFAA0D423FBDFE75FD2F" box="[326,413,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="3" pageNumber="100" unit="m" value="1225.0">
<elevation id="002BD1AAFFC5FFAA0D423FBDFE75FD2F" box="[326,413,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="3" pageNumber="100" unit="m" value="1225.0">1225 m</elevation>
</quantity>
,
<date id="FFB81059FFC5FFAA0DAD3FBCFDABFD2F" box="[425,579,655,681]" pageId="3" pageNumber="100" value="2020-05-11" valueMax="2020-05-20" valueMin="2020-05-11">
<collectingDate id="EFFCE9B1FFC5FFAA0DAD3FBCFDABFD2F" box="[425,579,655,681]" pageId="3" pageNumber="100" value="2020-05-11" valueMax="2020-05-20" valueMin="2020-05-11">1120.v.2020</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA0E4A3FBCFD0EFD2F" box="[590,742,655,681]" pageId="3" pageNumber="100">E. Yamamoto</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA0EF13FBCFC6BFD2F" box="[757,899,655,681]" pageId="3" pageNumber="100">Malaise trap</collectingMethod>
) (
<specimenCode id="DBA09EE2FFC5FFAA0F9D3FBCFB83FD2F" box="[921,1131,655,681]" collectionCode="CBM-ZI" pageId="3" pageNumber="100">CBM-ZI 0180217</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB93699FFC5FFAA0CC33F80FA71FCDB" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="100">
<materialsCitation id="3B6E3CC4FFC5FFAA0CC33F80FB0DFD4B" ID-GBIF-Occurrence="3400170310" box="[199,1253,691,717]" collectingDate="2020-05-11" collectingDateMax="2020-05-20" collectingDateMin="2020-05-11" collectionCode="CBM-ZI" collectorName="E. Yamamoto" country="Japan" county="Shikoku" elevation="1225" latitude="33.56" location="Male" longLatPrecision="72" longitude="132.918" municipality="Uchiko-cho" pageId="3" pageNumber="100" specimenCode="CBM-ZI 0180218-0180229" specimenCount="12" specimenCount-female="2" specimenCount-male="10" stateProvince="Ehime" typeStatus="paratype">
<emphasis id="B972EA8BFFC5FFAA0CC33F80FEAEFD4B" bold="true" box="[199,326,691,717]" pageId="3" pageNumber="100">
<typeStatus id="54BD883BFFC5FFAA0CC33F80FEA9FD4B" box="[199,321,691,717]" pageId="3" pageNumber="100" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D00FD10FFC5FFAA0D563F80FE55FD4B" box="[338,445,691,717]" pageId="3" pageNumber="100" type="male">10 males</specimenCount>
,
<specimenCount id="9D00FD10FFC5FFAA0DCA3F80FDA9FD4B" box="[462,577,691,717]" pageId="3" pageNumber="100" type="female">2 females</specimenCount>
, same data as the holotype (
<specimenCode id="DBA09EE2FFC5FFAA0F933F80FB35FD4B" box="[919,1245,691,717]" collectionCode="CBM-ZI" pageId="3" pageNumber="100">CBM-ZI 01802180180229</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B6E3CC4FFC5FFAA08F13F80FB1FFD77" ID-GBIF-Occurrence="3400170330" collectingDate="2020-06-11" collectingDateMax="2020-06-20" collectingDateMin="2020-06-11" collectionCode="CBM-ZI" collectorName="E. Yamamoto" country="Japan" county="Shikoku" elevation="1225" latitude="33.56" location="Male" longLatPrecision="72" longitude="132.918" municipality="Uchiko-cho" pageId="3" pageNumber="100" specimenCode="CBM-ZI 0180230-0180235" specimenCount="6" specimenCount-male="6" stateProvince="Ehime" typeStatus="paratype">
<specimenCount id="9D00FD10FFC5FFAA08F13F80FABBFD4B" box="[1269,1363,691,717]" pageId="3" pageNumber="100" type="male">6 males</specimenCount>
, same data as the holotype excepting the collection period
<date id="FFB81059FFC5FFAA0EFC3FE4FC70FD77" box="[760,920,727,753]" pageId="3" pageNumber="100" value="2020-06-11" valueMax="2020-06-20" valueMin="2020-06-11">
<collectingDate id="EFFCE9B1FFC5FFAA0EFC3FE4FC70FD77" box="[760,920,727,753]" pageId="3" pageNumber="100" value="2020-06-11" valueMax="2020-06-20" valueMin="2020-06-11">1120.vi.2020</collectingDate>
</date>
(
<specimenCode id="DBA09EE2FFC5FFAA0FAE3FE4FB18FD77" box="[938,1264,726,753]" collectionCode="CBM-ZI" pageId="3" pageNumber="100">CBM-ZI 01802300180235</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B6E3CC4FFC5FFAA09013FE4FDFEFCBF" ID-GBIF-Occurrence="3400170314" collectingDate="2018-05-01" collectingDateMax="2018-05-10" collectingDateMin="2018-05-01" collectionCode="NIBR" collectorName="E. Yamamoto" country="Japan" county="Kumakogen-cho" location="Izugatani-yama" municipality="Nishidani" pageId="3" pageNumber="100" specimenCode="NIBR0000935602-0000935603" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ehime" typeStatus="paratype">
<specimenCount id="9D00FD10FFC5FFAA09013FE4FABEFD77" box="[1285,1366,727,753]" pageId="3" pageNumber="100" type="male">1 male</specimenCount>
,
<specimenCount id="9D00FD10FFC5FFAA09613FE4FF38FC93" pageId="3" pageNumber="100" type="female">1 female</specimenCount>
,
<location id="8ED96042FFC5FFAA0CE73FC8FE7CFC93" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0CE73FC8FE7CFC93" box="[227,404,763,789]" country="Japan" county="Kumakogen-cho" municipality="Nishidani" name="Izugatani-yama" pageId="3" pageNumber="100" stateProvince="Ehime">Izugatani-yama</location>
,
<collectingMunicipality id="6BDDACE3FFC5FFAA0DA23FC8FDFEFC93" box="[422,534,763,789]" pageId="3" pageNumber="100">Nishidani</collectingMunicipality>
,
<collectingCounty id="62D84E15FFC5FFAA0E2C3FC8FD0EFC93" box="[552,742,763,789]" pageId="3" pageNumber="100">Kumakogen-cho</collectingCounty>
,
<collectingRegion id="49C2F87BFFC5FFAA0EFC3FC8FCACFC93" box="[760,836,763,789]" country="Japan" name="Ehime" pageId="3" pageNumber="100">Ehime</collectingRegion>
,
<collectingCountry id="F3117609FFC5FFAA0F523FC8FC71FC93" box="[854,921,763,789]" name="Japan" pageId="3" pageNumber="100">Japan</collectingCountry>
,
<date id="FFB81059FFC5FFAA0FAF3FC8FBDFFC93" box="[939,1079,763,789]" pageId="3" pageNumber="100" value="2018-05-01" valueMax="2018-05-10" valueMin="2018-05-01">
<collectingDate id="EFFCE9B1FFC5FFAA0FAF3FC8FBDFFC93" box="[939,1079,763,789]" pageId="3" pageNumber="100" value="2018-05-01" valueMax="2018-05-10" valueMin="2018-05-01">110.v.2018</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA084E3FC8FB01FC93" box="[1098,1257,763,789]" pageId="3" pageNumber="100">E. Yamamoto</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA08FB3FC8FA7CFC93" box="[1279,1428,763,789]" pageId="3" pageNumber="100">Malaise trap</collectingMethod>
) (
<specimenCode id="DBA09EE2FFC5FFAA0C9B3E2DFDE7FCBF" box="[159,527,798,825]" collectionCode="NIBR" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="3" pageNumber="100">NIBR00009356020000935603</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B6E3CC4FFC5FFAA0E203E2DFEF4FCDB" ID-GBIF-Occurrence="3400170317" collectingDate="2018-05-11" collectingDateMax="2018-05-20" collectingDateMin="2018-05-11" collectionCode="KPM-NK" collectorName="E. Yamamoto" country="Japan" county="Kumakogen-cho" location="Izugatani-yama" municipality="Nishidani" pageId="3" pageNumber="100" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Ehime" typeStatus="paratype">
<specimenCount id="9D00FD10FFC5FFAA0E203E2DFD97FCBF" box="[548,639,798,825]" pageId="3" pageNumber="100" type="male">5 males</specimenCount>
,
<specimenCount id="9D00FD10FFC5FFAA0E893E2CFD16FCBF" box="[653,766,799,825]" pageId="3" pageNumber="100" type="female">2 females</specimenCount>
, same locality,
<date id="FFB81059FFC5FFAA0FB53E2CFBA3FCBF" box="[945,1099,799,825]" pageId="3" pageNumber="100" value="2018-05-11" valueMax="2018-05-20" valueMin="2018-05-11">
<collectingDate id="EFFCE9B1FFC5FFAA0FB53E2CFBA3FCBF" box="[945,1099,799,825]" pageId="3" pageNumber="100" value="2018-05-11" valueMax="2018-05-20" valueMin="2018-05-11">1120.v.2018</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA085D3E2CFB1BFCBF" box="[1113,1267,799,825]" pageId="3" pageNumber="100">E. Yamamoto</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA09003E2CFA7CFCBF" box="[1284,1428,799,825]" pageId="3" pageNumber="100">Malaise trap</collectingMethod>
) (
<collectionCode id="ED17AE5CFFC5FFAA0C9B3E70FEFCFCDB" box="[159,276,835,861]" pageId="3" pageNumber="100">KPM-NK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6E3CC4FFC5FFAA0D233E70FA7DFCDB" ID-GBIF-Occurrence="3400170323" box="[295,1429,834,861]" collectingDate="2018-05-21" collectingDateMax="2018-05-31" collectingDateMin="2018-05-21" collectionCode="NIBR" collectorName="E. Yamamoto" country="Japan" county="Kumakogen-cho" location="Izugatani-yama" municipality="Nishidani" pageId="3" pageNumber="100" specimenCode="NIBR0000935604-0000935607" specimenCount="4" specimenCount-male="4" stateProvince="Ehime" typeStatus="paratype">
<specimenCount id="9D00FD10FFC5FFAA0D233E70FE68FCDB" box="[295,384,835,861]" pageId="3" pageNumber="100" type="male">4 males</specimenCount>
, same locality,
<date id="FFB81059FFC5FFAA0E283E70FD2FFCDB" box="[556,711,835,861]" pageId="3" pageNumber="100" value="2018-05-21" valueMax="2018-05-31" valueMin="2018-05-21">
<collectingDate id="EFFCE9B1FFC5FFAA0E283E70FD2FFCDB" box="[556,711,835,861]" pageId="3" pageNumber="100" value="2018-05-21" valueMax="2018-05-31" valueMin="2018-05-21">2131.v.2018</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA0ED63E70FC82FCDB" box="[722,874,835,861]" pageId="3" pageNumber="100">E. Yamamoto</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA0F7D3E70FBEFFCDB" box="[889,1031,835,861]" pageId="3" pageNumber="100">Malaise trap</collectingMethod>
) (
<specimenCode id="DBA09EE2FFC5FFAA08193E71FA66FCDB" box="[1053,1422,834,861]" collectionCode="NIBR" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="3" pageNumber="100">NIBR00009356040000935607</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB93699FFC5FFAA0CC33E55FCB9FB27" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="100">
<materialsCitation id="3B6E3CC4FFC5FFAA0CC33E55FC01FC23" ID-GBIF-Occurrence="3400170334" collectingDate="2006-05-05" collectionCode="T, TN" collectorName="Nozaki" country="Japan" county="Nishigochi-gawa" elevation="550" location="Aoi-ku" municipality="Yokosawa" pageId="3" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka">
<emphasis id="B972EA8BFFC5FFAA0CC33E55FD06FC07" bold="true" box="[199,750,870,897]" pageId="3" pageNumber="100">
Other specimens examined.
<collectingCountry id="F3117609FFC5FFAA0E1F3E55FD91FC06" box="[539,633,870,896]" name="Japan" pageId="3" pageNumber="100">JAPAN</collectingCountry>
: Honshu:
</emphasis>
<collectingRegion id="49C2F87BFFC5FFAA0EF23E54FC8BFC07" box="[758,867,871,897]" country="Japan" name="Shizuoka" pageId="3" pageNumber="100">Shizuoka</collectingRegion>
:
<specimenCount id="9D00FD10FFC5FFAA0F6B3E54FC56FC07" box="[879,958,871,897]" pageId="3" pageNumber="100" type="male">1 male</specimenCount>
, tributary of
<collectingCounty id="62D84E15FFC5FFAA08503E54FAFEFC07" box="[1108,1302,871,897]" pageId="3" pageNumber="100">Nishigochi-gawa</collectingCounty>
,
<collectingMunicipality id="6BDDACE3FFC5FFAA09263E54FA70FC07" box="[1314,1432,871,897]" pageId="3" pageNumber="100">Yokosawa</collectingMunicipality>
,
<location id="8ED96042FFC5FFAA0C933EB8FF01FC23" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0C933EB8FF01FC23" box="[151,233,907,933]" country="Japan" county="Nishigochi-gawa" municipality="Yokosawa" name="Aoi-ku" pageId="3" pageNumber="100" stateProvince="Shizuoka">Aoi-ku</location>
,
<location id="8ED96042FFC5FFAA0CF23EB8FE63FC23" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0CF23EB8FE63FC23" box="[246,395,907,933]" country="Japan" county="Nishigochi-gawa" municipality="Yokosawa" name="Shizuoka-shi" pageId="3" pageNumber="100" stateProvince="Shizuoka">Shizuoka-shi</location>
, alt.
<quantity id="4CFE9B7CFFC5FFAA0DC63EB9FDE4FC23" box="[450,524,906,933]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="100" unit="m" value="550.0">
<elevation id="002BD1AAFFC5FFAA0DC63EB9FDE4FC23" box="[450,524,906,933]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="100" unit="m" value="550.0">550 m</elevation>
</quantity>
,
<date id="FFB81059FFC5FFAA0E1D3EB9FD94FC23" box="[537,636,906,933]" pageId="3" pageNumber="100" value="2006-05-05">
<collectingDate id="EFFCE9B1FFC5FFAA0E1D3EB9FD94FC23" box="[537,636,906,933]" pageId="3" pageNumber="100" value="2006-05-05">5.v.2006</collectingDate>
</date>
,
<collectionCode id="ED17AE5CFFC5FFAA0E8C3EB8FD72FC23" box="[648,666,907,933]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="100">T</collectionCode>
.
<collectorName id="26F3534FFFC5FFAA0EA23EB8FD1FFC23" box="[678,759,907,933]" pageId="3" pageNumber="100">Nozaki</collectorName>
(net sweeping) (
<collectionCode id="ED17AE5CFFC5FFAA0FB33EB8FC0AFC23" box="[951,994,907,933]" pageId="3" pageNumber="100">TN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B6E3CC4FFC5FFAA0FF13EB8FCB1FC4F" ID-GBIF-Occurrence="3400170320" collectingDate="2006-05-14" collectorName="N. Kawase" country="Japan" location="Kameyamashi" municipality="Sakashita" pageId="3" pageNumber="100" specimenCount="4" specimenCount-male="4" stateProvince="Mie">
<collectingRegion id="49C2F87BFFC5FFAA0FF13EB8FBCEFC23" box="[1013,1062,907,933]" country="Japan" name="Mie" pageId="3" pageNumber="100">Mie</collectingRegion>
:
<specimenCount id="9D00FD10FFC5FFAA08363EB8FB64FC23" box="[1074,1164,907,933]" pageId="3" pageNumber="100" type="male">4 males</specimenCount>
,
<collectingMunicipality id="6BDDACE3FFC5FFAA089D3EB8FAEFFC23" box="[1177,1287,907,933]" pageId="3" pageNumber="100">Sakashita</collectingMunicipality>
,
<location id="8ED96042FFC5FFAA09173EB8FF52FC4F" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA09173EB8FF52FC4F" country="Japan" municipality="Sakashita" name="Kameyamashi" pageId="3" pageNumber="100" stateProvince="Mie">Kameyamashi</location>
,
<date id="FFB81059FFC5FFAA0CCD3E9CFED2FC4F" box="[201,314,943,969]" pageId="3" pageNumber="100" value="2006-05-14">
<collectingDate id="EFFCE9B1FFC5FFAA0CCD3E9CFED2FC4F" box="[201,314,943,969]" pageId="3" pageNumber="100" value="2006-05-14">14.v.2006</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA0D4C3E9CFE2EFC4F" box="[328,454,943,969]" pageId="3" pageNumber="100">N. Kawase</collectorName>
(net sweeping) (MITR-20090042)
</materialsCitation>
;
<materialsCitation id="3B6E3CC4FFC5FFAA0F633E9CFD8CFC6B" ID-GBIF-Occurrence="3400170329" collectingDate="2009-05-19" collectingDateMax="2009-06-08" collectingDateMin="2009-05-19" collectionCode="NK" collectorName="H. Morita" country="Japan" elevation="600" location="Taga-cho" municipality="Ojigahata" pageId="3" pageNumber="100" specimenCount="10" specimenCount-male="10" stateProvince="Honshu">
<specimenCount id="9D00FD10FFC5FFAA0F633E9CFC39FC4F" box="[871,977,943,969]" pageId="3" pageNumber="100" type="male">10 males</specimenCount>
,
<collectingMunicipality id="6BDDACE3FFC5FFAA0FDB3E9CFBA7FC4F" box="[991,1103,943,969]" pageId="3" pageNumber="100">Ojigahata</collectingMunicipality>
,
<location id="8ED96042FFC5FFAA08583E9CFB2FFC4F" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA08583E9CFB2FFC4F" box="[1116,1223,943,969]" country="Japan" municipality="Ojigahata" name="Taga-cho" pageId="3" pageNumber="100" stateProvince="Honshu">Taga-cho</location>
, alt.
<quantity id="4CFE9B7CFFC5FFAA09063E9CFAA6FC4F" box="[1282,1358,943,969]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="100" unit="m" value="600.0">
<elevation id="002BD1AAFFC5FFAA09063E9CFAA6FC4F" box="[1282,1358,943,969]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="100" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="FFB81059FFC5FFAA09593E9CFEEBFC6B" pageId="3" pageNumber="100" value="2009-05-19" valueMax="2009-06-08" valueMin="2009-05-19">
<collectingDate id="EFFCE9B1FFC5FFAA09593E9CFEEBFC6B" pageId="3" pageNumber="100" value="2009-05-19" valueMax="2009-06-08" valueMin="2009-05-19">19.v 8.vi.2009</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA0D0A3EE0FE94FC6B" box="[270,380,979,1005]" pageId="3" pageNumber="100">H. Morita</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA0D8E3EE0FDF0FC6B" box="[394,536,979,1005]" pageId="3" pageNumber="100">Malaise trap</collectingMethod>
) (
<collectionCode id="ED17AE5CFFC5FFAA0E2A3EE0FDB4FC6B" box="[558,604,979,1005]" pageId="3" pageNumber="100">NK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B6E3CC4FFC5FFAA0E6B3EE0FDFAFB97" ID-GBIF-Occurrence="3400170327" collectingDate="2009-05-12" collectingDateMax="2009-05-31" collectingDateMin="2009-05-12" collectorName="N. Kawase" country="Japan" county="Yuzurio" location="Higashi-omi" municipality="Eigenji" pageId="3" pageNumber="100" specimenCount="2" specimenCount-male="2" stateProvince="Shiga">
<collectingRegion id="49C2F87BFFC5FFAA0E6B3EE0FD5BFC6B" box="[623,691,979,1005]" country="Japan" name="Shiga" pageId="3" pageNumber="100">Shiga</collectingRegion>
:
<specimenCount id="9D00FD10FFC5FFAA0EB93EE0FCFDFC6B" box="[701,789,979,1005]" pageId="3" pageNumber="100" type="male">2 males</specimenCount>
,
<collectingCounty id="62D84E15FFC5FFAA0F1B3EE0FC93FC6B" box="[799,891,979,1005]" pageId="3" pageNumber="100">Yuzurio</collectingCounty>
,
<collectingMunicipality id="6BDDACE3FFC5FFAA0F813EE0FC31FC6B" box="[901,985,979,1005]" pageId="3" pageNumber="100">Eigenji</collectingMunicipality>
,
<location id="8ED96042FFC5FFAA0FE73EE0FB9AFC6B" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0FE73EE0FB9AFC6B" box="[995,1138,979,1005]" country="Japan" county="Yuzurio" municipality="Eigenji" name="Higashi-omi" pageId="3" pageNumber="100" stateProvince="Shiga">Higashi-omi</location>
,
<date id="FFB81059FFC5FFAA08783EE0FAFFFC6B" box="[1148,1303,979,1005]" pageId="3" pageNumber="100" value="2009-05-12" valueMax="2009-05-31" valueMin="2009-05-12">
<collectingDate id="EFFCE9B1FFC5FFAA08783EE0FAFFFC6B" box="[1148,1303,979,1005]" pageId="3" pageNumber="100" value="2009-05-12" valueMax="2009-05-31" valueMin="2009-05-12">1231.v.2009</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA09263EE0FA75FC6B" box="[1314,1437,979,1005]" pageId="3" pageNumber="100">N. Kawase</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA0C9B3EC4FEC5FB97" box="[159,301,1015,1041]" pageId="3" pageNumber="100">Malaise trap</collectingMethod>
) (MITR-20090419)
</materialsCitation>
.
<materialsCitation id="3B6E3CC4FFC5FFAA0E1F3EC5FB6AFB97" ID-GBIF-Occurrence="3400170311" box="[539,1154,1014,1041]" collectingDate="2020-05-11" collectingDateMax="2020-05-20" collectingDateMin="2020-05-11" collectionCode="TN" collectorName="E. Yamamoto" country="Japan" county="Shikoku" elevation="1225" latitude="33.56" location="Male" longLatPrecision="72" longitude="132.918" municipality="Uchiko-cho" pageId="3" pageNumber="100" specimenCount="2" specimenCount-male="2" stateProvince="Ehime">
<emphasis id="B972EA8BFFC5FFAA0E1F3EC5FD60FB96" bold="true" box="[539,648,1014,1040]" pageId="3" pageNumber="100">Shikoku:</emphasis>
<collectingRegion id="49C2F87BFFC5FFAA0E8B3EC4FD35FB97" box="[655,733,1015,1041]" country="Japan" name="Ehime" pageId="3" pageNumber="100">Ehime</collectingRegion>
:
<specimenCount id="9D00FD10FFC5FFAA0EE33EC4FCA8FB97" box="[743,832,1015,1041]" pageId="3" pageNumber="100" type="male">2 males</specimenCount>
, same data as holotype (
<collectionCode id="ED17AE5CFFC5FFAA08543EC4FB92FB97" box="[1104,1146,1015,1041]" pageId="3" pageNumber="100">TN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6E3CC4FFC5FFAA088A3EC4FC3EFBB3" ID-GBIF-Occurrence="3400170315" collectingDate="2018-06-21" collectionCode="TN" collectorName="E. Yamamoto" country="Japan" county="Izugatani-yama" location="Kumakogen-cho" municipality="Nishidani" pageId="3" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Shikoku">
<specimenCount id="9D00FD10FFC5FFAA088A3EC4FB34FB97" box="[1166,1244,1015,1041]" pageId="3" pageNumber="100" type="male">1 male</specimenCount>
,
<collectingCounty id="62D84E15FFC5FFAA08E33EC4FA70FB97" box="[1255,1432,1015,1041]" pageId="3" pageNumber="100">Izugatani-yama</collectingCounty>
,
<collectingMunicipality id="6BDDACE3FFC5FFAA0C933928FEEFFBB3" box="[151,263,1051,1077]" pageId="3" pageNumber="100">Nishidani</collectingMunicipality>
,
<location id="8ED96042FFC5FFAA0D123928FE3CFBB3" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0D123928FE3CFBB3" box="[278,468,1051,1077]" country="Japan" county="Izugatani-yama" municipality="Nishidani" name="Kumakogen-cho" pageId="3" pageNumber="100" stateProvince="Shikoku">Kumakogen-cho</location>
,
<date id="FFB81059FFC5FFAA0DE73928FDB5FBB3" box="[483,605,1051,1077]" pageId="3" pageNumber="100" value="2018-06-21">
<collectingDate id="EFFCE9B1FFC5FFAA0DE73928FDB5FBB3" box="[483,605,1051,1077]" pageId="3" pageNumber="100" value="2018-06-21">21.vi.2018</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA0E693928FCE1FBB3" box="[621,777,1051,1077]" pageId="3" pageNumber="100">E. Yamamoto</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA0F1F3928FC60FBB3" box="[795,904,1051,1077]" pageId="3" pageNumber="100">light trap</collectingMethod>
) (
<collectionCode id="ED17AE5CFFC5FFAA0FA73928FC25FBB3" box="[931,973,1051,1077]" pageId="3" pageNumber="100">TN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6E3CC4FFC5FFAA0FE23928FD4AFBDF" ID-GBIF-Occurrence="3400170325" collectingDate="2018-06-01" collectingDateMax="2018-06-30" collectingDateMin="2018-06-01" collectionCode="TN" collectorName="E. Yamamoto" country="Japan" county="Izugatani-yama" location="Kumakogen-cho" municipality="Nishidani" pageId="3" pageNumber="100" specimenCount="15" specimenCount-female="1" specimenCount-male="14" stateProvince="Shikoku">
<specimenCount id="9D00FD10FFC5FFAA0FE23928FBB9FBB3" box="[998,1105,1051,1077]" pageId="3" pageNumber="100" type="male">14 males</specimenCount>
,
<specimenCount id="9D00FD10FFC5FFAA08643928FB20FBB3" box="[1120,1224,1051,1077]" pageId="3" pageNumber="100" type="female">1 female</specimenCount>
, same locality,
<date id="FFB81059FFC5FFAA09843928FEFAFBDF" pageId="3" pageNumber="100" value="2018-06-01" valueMax="2018-06-30" valueMin="2018-06-01">
<collectingDate id="EFFCE9B1FFC5FFAA09843928FEFAFBDF" pageId="3" pageNumber="100" value="2018-06-01" valueMax="2018-06-30" valueMin="2018-06-01">1 30.vi.2018</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA0D1A390CFE50FBDF" box="[286,440,1087,1113]" pageId="3" pageNumber="100">E. Yamamoto</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA0DC3390CFDBFFBDF" box="[455,599,1087,1113]" pageId="3" pageNumber="100">Malaise trap</collectingMethod>
) (
<collectionCode id="ED17AE5CFFC5FFAA0E74390CFD72FBDF" box="[624,666,1087,1113]" pageId="3" pageNumber="100">TN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6E3CC4FFC5FFAA0EAB390CFE96FBFB" ID-GBIF-Occurrence="3400170335" collectingDate="2018-07-01" collectingDateMax="2018-08-10" collectingDateMin="2018-07-01" collectionCode="SJP" collectorName="E. Yamamoto" country="Japan" county="Izugatani-yama" location="Kumakogen-cho" municipality="Nishidani" pageId="3" pageNumber="100" specimenCount="8" specimenCount-female="1" specimenCount-male="7" stateProvince="Shikoku">
<specimenCount id="9D00FD10FFC5FFAA0EAB390CFCE2FBDF" box="[687,778,1087,1113]" pageId="3" pageNumber="100" type="male">7 males</specimenCount>
,
<specimenCount id="9D00FD10FFC5FFAA0F13390CFC94FBDF" box="[791,892,1087,1113]" pageId="3" pageNumber="100" type="female">1 female</specimenCount>
, same locality,
<date id="FFB81059FFC5FFAA0828390CFB1EFBDF" box="[1068,1270,1087,1113]" pageId="3" pageNumber="100" value="2018-07-01" valueMax="2018-08-10" valueMin="2018-07-01">
<collectingDate id="EFFCE9B1FFC5FFAA0828390CFB1EFBDF" box="[1068,1270,1087,1113]" pageId="3" pageNumber="100" value="2018-07-01" valueMax="2018-08-10" valueMin="2018-07-01">1.vii10.viii.2018</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA0907390CFA75FBDF" box="[1283,1437,1087,1113]" pageId="3" pageNumber="100">E. Yamamoto</collectorName>
(
<collectingMethod id="52474E8EFFC5FFAA0C9B3950FEC7FBFB" box="[159,303,1123,1149]" pageId="3" pageNumber="100">Malaise trap</collectingMethod>
) (
<collectionCode id="ED17AE5CFFC5FFAA0D433950FE9EFBFB" box="[327,374,1123,1149]" pageId="3" pageNumber="100">SJP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B6E3CC4FFC5FFAA0D8F3950FAD3FBFB" ID-GBIF-Occurrence="3400170301" box="[395,1339,1123,1149]" collectionCode="TN" collectorName="M. Takai" country="Japan" location="Nishikuma-rindo" pageId="3" pageNumber="100" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kochi">
<collectingRegion id="49C2F87BFFC5FFAA0D8F3950FE3CFBFB" box="[395,468,1123,1149]" country="Japan" name="Kochi" pageId="3" pageNumber="100">Kochi</collectingRegion>
:
<specimenCount id="9D00FD10FFC5FFAA0DDB3950FDC7FBFB" box="[479,559,1123,1149]" pageId="3" pageNumber="100" type="male">1 male</specimenCount>
,
<specimenCount id="9D00FD10FFC5FFAA0E383950FD49FBFB" box="[572,673,1123,1149]" pageId="3" pageNumber="100" type="female">1 female</specimenCount>
,
<location id="8ED96042FFC5FFAA0EAA3950FC98FBFB" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0EAA3950FC98FBFB" box="[686,880,1123,1149]" country="Japan" name="Nishikuma-rindo" pageId="3" pageNumber="100" stateProvince="Kochi">Nishikuma-rindo</location>
,
<collectorName id="26F3534FFFC5FFAA0F783950FBE6FBFB" box="[892,1038,1123,1149]" pageId="3" pageNumber="100">Monobe-cho</collectorName>
,
<collectorName id="26F3534FFFC5FFAA081F3950FB46FBFB" box="[1051,1198,1123,1149]" pageId="3" pageNumber="100">Kami-shi, M</collectorName>
.
<collectorName id="26F3534FFFC5FFAA08BE3950FB10FBFB" box="[1210,1272,1123,1149]" pageId="3" pageNumber="100">Takai</collectorName>
(
<collectionCode id="ED17AE5CFFC5FFAA090D3950FADBFBFB" box="[1289,1331,1123,1149]" pageId="3" pageNumber="100">TN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6E3CC4FFC5FFAA094C3950FCA4FB27" ID-GBIF-Occurrence="3400170326" collectingDate="2004-04-24" collectionCode="TN" collectorName="M. Takai" country="Japan" county="Befu-kyo" location="Kami-shi" municipality="Monobe-cho" pageId="3" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Shikoku">
<specimenCount id="9D00FD10FFC5FFAA094C3950FA70FBFB" box="[1352,1432,1123,1149]" pageId="3" pageNumber="100" type="male">1 male</specimenCount>
,
<collectingCounty id="62D84E15FFC5FFAA0C9339B4FEECFB27" box="[151,260,1159,1185]" pageId="3" pageNumber="100">Befu-kyo</collectingCounty>
,
<collectingMunicipality id="6BDDACE3FFC5FFAA0D0B39B4FE49FB27" box="[271,417,1159,1185]" pageId="3" pageNumber="100">Monobe-cho</collectingMunicipality>
,
<location id="8ED96042FFC5FFAA0DA839B4FDFFFB27" LSID="urn:lsid:plazi:treatment:03AF878FFFC4FFAA0C933951FE2FFAFF:8ED96042FFC5FFAA0DA839B4FDFFFB27" box="[428,535,1159,1185]" country="Japan" county="Befu-kyo" municipality="Monobe-cho" name="Kami-shi" pageId="3" pageNumber="100" stateProvince="Shikoku">Kami-shi</location>
,
<date id="FFB81059FFC5FFAA0E2639B4FD73FB27" box="[546,667,1159,1185]" pageId="3" pageNumber="100" value="2004-04-24">
<collectingDate id="EFFCE9B1FFC5FFAA0E2639B4FD73FB27" box="[546,667,1159,1185]" pageId="3" pageNumber="100" value="2004-04-24">24.iv.2004</collectingDate>
</date>
,
<collectorName id="26F3534FFFC5FFAA0EA239B4FCE3FB27" box="[678,779,1159,1185]" pageId="3" pageNumber="100">M. Takai</collectorName>
(
<collectionCode id="ED17AE5CFFC5FFAA0F1E39B4FCADFB27" box="[794,837,1159,1185]" pageId="3" pageNumber="100">TN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC5FFAA0CC33998FE68FB6F" pageId="3" pageNumber="100" type="etymology">
<paragraph id="8BB93699FFC5FFAA0CC33998FE68FB6F" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="100">
<emphasis id="B972EA8BFFC5FFAA0CC33998FEA6FB43" bold="true" box="[199,334,1195,1221]" pageId="3" pageNumber="100">Etymology.</emphasis>
This species is dedicated to Mr. Eiji Yamamoto, who gifted us valuable specimens, including specimens of this species.
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC5FFAA0CC339C1FD63FA88" box="[199,651,1266,1294]" pageId="3" pageNumber="100" type="distribution">
<paragraph id="8BB93699FFC5FFAA0CC339C1FD63FA88" blockId="3.[151,1437,151,1401]" box="[199,651,1266,1294]" pageId="3" pageNumber="100">
<emphasis id="B972EA8BFFC5FFAA0CC339C1FE89FA8A" bold="true" box="[199,353,1266,1292]" pageId="3" pageNumber="100">Distribution.</emphasis>
<collectingCountry id="F3117609FFC5FFAA0D6339C0FE4FFA8B" box="[359,423,1267,1293]" name="Japan" pageId="3" pageNumber="100">Japan</collectingCountry>
(Honshu, Shikoku).
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC5FFAA0CC33824FE2FFAFF" pageId="3" pageNumber="100" type="discussion">
<paragraph id="8BB93699FFC5FFAA0CC33824FE2FFAFF" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="100">
<emphasis id="B972EA8BFFC5FFAA0CC33824FED3FAB7" bold="true" box="[199,315,1303,1329]" pageId="3" pageNumber="100">Remarks.</emphasis>
This species was found from two major islands, Honshu and Shikoku, in
<collectingCountry id="F3117609FFC5FFAA08623824FB4EFAB7" box="[1126,1190,1303,1329]" name="Japan" pageId="3" pageNumber="100">Japan</collectingCountry>
(
<figureCitation id="133D2A1CFFC5FFAA08B03824FB1FFAB7" box="[1204,1271,1303,1329]" captionStart="FIGURE 9" captionStartId="7.[151,250,1110,1134]" captionTargetBox="[155,1428,281,1079]" captionTargetId="figure-64@7.[151,1436,273,1087]" captionTargetPageId="7" captionText="FIGURE 9. Distributions of 4 Rhyacophila species in East Asia. Distribution areas of R. confissa and R. vicina are cited from Ivanov (2011), Park &amp; Kong (2020), and Yang et al. (2016)." figureDoi="http://doi.org/10.5281/zenodo.5656565" httpUri="https://zenodo.org/record/5656565/files/figure.png" pageId="3" pageNumber="100">Fig. 9</figureCitation>
). The genitalic characteristics are consistent in both populations, and there are stable differences between this species and a Korean species,
<taxonomicName id="4C064D1AFFC5FFAA0CF2386CFE8EFAFF" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[246,358,1375,1401]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC5FFAA0CF2386CFE8EFAFF" box="[246,358,1375,1401]" italics="true" pageId="3" pageNumber="100">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC5FFAA0D69386CFE2FFAFF" bold="true" box="[365,455,1375,1401]" pageId="3" pageNumber="100">
<taxonomicNameLabel id="A24157F0FFC5FFAA0D69386CFE2FFAFF" box="[365,455,1375,1401]" pageId="3" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>