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<document id="93518E1C40C9EA749589424589DDBAF8" ID-DOI="10.5281/zenodo.5351598" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351598" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630206282112" checkinUser="tatiana" docAuthor="Jäch, Manfred A., Díaz, Juan A. &amp; Skale, Andre" docDate="2013" docId="03AF8781EA332933CCE7BCA41FA9DBD8" docLanguage="en" docName="61rbz053-071.pdf" docOrigin="Raffles Bulletin of Zoology 61 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Hydraena (Hydraenopsis) singaporensis Jäch &amp; Díaz &amp; Skale 2013, new species" docType="treatment" docVersion="3" lastPageNumber="68" masterDocId="FF96FFF9EA3D293CCC79BD781C2BDF4E" masterDocTitle="The Hydraenidae (Coleoptera) Of The Republic Of Singapore" masterLastPageNumber="71" masterPageNumber="53" pageNumber="67" updateTime="1699184966419" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0429939F184FA6F1C82352CC9E7A18F3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EA5E37A3F165D51D714E3B359E133A8F">The Hydraenidae (Coleoptera) Of The Republic Of Singapore</mods:title>
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<mods:name id="DFF5FC3167B01F825970E5B08B3F69B7" type="personal">
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<mods:namePart id="4746C6A562A636B0ACF97118C0FF8C6E">Jäch, Manfred A.</mods:namePart>
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<mods:namePart id="4EF9A19ED87890B9DEFFB9185BAACA8E">Díaz, Juan A.</mods:namePart>
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<mods:namePart id="CA3D03A85E9427A0E1C818F986319E9C">Skale, Andre</mods:namePart>
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<mods:title id="CEA3C4C2062F09494B8B966BF2366CA2">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="7F5E598D97F4619361C55C82E04B7A75">
<mods:date id="ADE8F61E916DFA836AE7EE8940E2D581">2013</mods:date>
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<mods:number id="F3C51F84FECC17137D2765DE8644BB2C">2013-02-28</mods:number>
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<mods:number id="CF6B2468888B482FC0D2BCDD929608E9">61</mods:number>
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<treatment id="03AF8781EA332933CCE7BCA41FA9DBD8" LSID="urn:lsid:plazi:treatment:03AF8781EA332933CCE7BCA41FA9DBD8" httpUri="http://treatment.plazi.org/id/03AF8781EA332933CCE7BCA41FA9DBD8" lastPageId="15" lastPageNumber="68" pageId="14" pageNumber="67">
<subSubSection id="C31C651CEA332932CCE7BCA41ED9DEBB" box="[158,754,476,501]" pageId="14" pageNumber="67" type="nomenclature">
<paragraph id="8BB93697EA332932CCE7BCA41ED9DEBB" blockId="14.[158,754,476,533]" box="[158,754,476,501]" pageId="14" pageNumber="67">
<heading id="D0F181FBEA332932CCE7BCA41ED9DEBB" bold="true" box="[158,754,476,501]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="67" reason="2">
<emphasis id="B972EA85EA332932CCE7BCA41ED9DEBB" bold="true" box="[158,754,476,501]" pageId="14" pageNumber="67">
<taxonomicName id="4C064D14EA332932CCE7BCA41E76DEBB" ID-CoL="3N3HS" authority="Jäch &amp; Díaz &amp; Skale, 2013" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[158,605,476,501]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="singaporensis" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B972EA85EA332932CCE7BCA41D27DEBB" bold="true" box="[158,268,476,501]" italics="true" pageId="14" pageNumber="67">Hydraena</emphasis>
(
<emphasis id="B972EA85EA332932CD64BCA41D9FDEBB" bold="true" box="[285,436,476,501]" italics="true" pageId="14" pageNumber="67">Hydraenopsis</emphasis>
)
<emphasis id="B972EA85EA332932CDBFBCA51E76DEBB" bold="true" box="[454,605,477,501]" italics="true" pageId="14" pageNumber="67">singaporensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24157FEEA332932CE15BCA51ED9DEBB" box="[620,754,477,501]" pageId="14" pageNumber="67" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA332932CD35BC851E68DD5B" box="[332,579,509,533]" pageId="14" pageNumber="67" type="description">
<paragraph id="8BB93697EA332932CD35BC851E68DD5B" blockId="14.[158,754,476,533]" box="[332,579,509,533]" pageId="14" pageNumber="67">
(
<figureCitation id="133D2A12EA332932CD2DBC851D8ADD5B" box="[340,417,509,533]" captionStart="Figs" captionStartId="3.[113,157,2061,2083]" captionTargetBox="[222,1331,314,1975]" captionTargetId="figure-8@3.[195,1369,301,1985]" captionTargetPageId="3" captionText="Figs. 14. Habitus of: 1, Hydraena hendrichi, paratype; 2, H. paulmoritz, paratype; 3, H. singaporensis, paratype; 4, H. yangae, holotype, right foreleg (lateral view) depicted separately (photograph taken before right elytron got unhinged)." figureDoi="http://doi.org/10.5281/zenodo.5351600" httpUri="https://zenodo.org/record/5351600/files/figure.png" pageId="14" pageNumber="67">Figs. 3</figureCitation>
,
<figureCitation id="133D2A12EA332932CDD6BC851D95DD5B" box="[431,446,509,533]" captionStart="Fig" captionStartId="4.[136,171,2060,2082]" captionTargetBox="[157,1469,185,2015]" captionTargetId="figure-8@4.[98,1492,175,2032]" captionTargetPageId="4" captionText="Fig. 5. Hydraena singaporensis: a, aedeagus in right lateral view; b, aedeagus, ventral view (flagellum not depicted completely in a, b); c, aedeagus, left lateral view; d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351602" httpUri="https://zenodo.org/record/5351602/files/figure.png" pageId="14" pageNumber="67">5</figureCitation>
,
<figureCitation id="133D2A12EA332932CDB2BC851DC3DD5B" box="[459,488,509,533]" captionStart="Figs" captionStartId="12.[137,181,2091,2113]" captionTargetBox="[138,1476,1428,2061]" captionTargetId="figure-8@12.[134,1479,1425,2064]" captionTargetPageId="12" captionText="Figs. 1214. Meso- and metaventrite of: 12, Hydraena hendrichi; 13, H. singaporensis; and 14, H. formula." figureDoi="http://doi.org/10.5281/zenodo.5351616" httpUri="https://zenodo.org/record/5351616/files/figure.png" pageId="14" pageNumber="67">13</figureCitation>
,
<figureCitation id="133D2A12EA332932CD8CBC851E39DD5B" box="[501,530,509,533]" captionStart="Fig" captionStartId="14.[137,172,2090,2112]" captionTargetBox="[137,1470,1208,2063]" captionTargetId="figure-8@14.[137,1478,1202,2067]" captionTargetPageId="14" captionText="Fig. 15. Geographical map of Singapore, showing distribution of Hydraenidae." figureDoi="http://doi.org/10.5281/zenodo.5351618" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="14" pageNumber="67">15</figureCitation>
,
<figureCitation id="133D2A12EA332932CE66BC851E11DD5B" box="[543,570,509,533]" captionStart="Fig" captionStartId="16.[136,172,2061,2083]" captionTargetBox="[279,1324,1376,2039]" captionTargetId="figure-537@16.[276,1328,1373,2041]" captionTargetPageId="16" captionText="Fig. 17. Nee Soon Swamp Forest, Singapore; C.M. Yang on right. Four species, H. formula, H. jacobsoni, H. paulmoritz, and H. singaporensis were collected from this site. (Photograph by: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5351622" httpUri="https://zenodo.org/record/5351622/files/figure.png" pageId="14" pageNumber="67">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA332932CCF1BF441E8ADB4C" pageId="14" pageNumber="67" type="materials_examined">
<paragraph id="8BB93697EA332932CCF1BF441EFFDD1C" blockId="14.[136,724,571,594]" box="[136,724,571,594]" pageId="14" pageNumber="67">
<materialsCitation id="3B6E3CCAEA332932CCF1BF441EE4DD1C" box="[136,719,571,594]" country="Singapore" location="Stream in Upper Peirce Forest" pageId="14" pageNumber="67" specimenCount="1" typeStatus="holotype">
<emphasis id="B972EA85EA332932CCF1BF441D27DD1F" bold="true" box="[136,268,571,593]" italics="true" pageId="14" pageNumber="67">
<typeStatus id="54BD8835EA332932CCF1BF441C9CDD1F" box="[136,183,572,593]" pageId="14" pageNumber="67">Type</typeStatus>
locality.
</emphasis>
<location id="8ED9604CEA332932CD4DBF441E74DD1F" LSID="urn:lsid:plazi:treatment:03AF8781EA332933CCE7BCA41FA9DBD8:8ED9604CEA332932CD4DBF441E74DD1F" box="[308,607,572,594]" country="Singapore" name="Stream in Upper Peirce Forest" pageId="14" pageNumber="67">Stream in Upper Peirce Forest</location>
,
<collectingCountry id="F3117607EA332932CE12BF441EE4DD1C" box="[619,719,572,594]" name="Singapore" pageId="14" pageNumber="67">Singapore</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB93697EA332932CCF1BF0E1E8ADB4C" blockId="14.[136,775,628,1026]" pageId="14" pageNumber="67">
<emphasis id="B972EA85EA332932CCF1BF0E1D14DDC5" bold="true" box="[136,319,628,651]" pageId="14" pageNumber="67">
<emphasis id="B972EA85EA332932CCF1BF0E1D35DDC4" bold="true" box="[136,286,628,651]" italics="true" pageId="14" pageNumber="67">Type material.</emphasis>
</emphasis>
<materialsCitation id="3B6E3CCAEA332932CD30BF0D1D38DD8B" collectingDate="1991-10-31" collectionCode="ZRC" collectorName="DH Murphy" country="Singapore" county="Forest" location="Upper Pierce" municipality="Peirce" pageId="14" pageNumber="67" specimenCount="1" typeStatus="holotype">
<emphasis id="B972EA85EA332932CD30BF0D1D82DDC4" bold="true" box="[329,425,629,650]" pageId="14" pageNumber="67">
<typeStatus id="54BD8835EA332932CD30BF0D1D82DDC4" box="[329,425,629,650]" pageId="14" pageNumber="67" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="ED17AE52EA332932CD8ABF0D1E02DDC5" box="[499,553,629,651]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="14" pageNumber="67">ZRC</collectionCode>
): “
<collectingCountry id="F3117607EA332932CE3FBF0D1EFEDDC4" box="[582,725,629,650]" name="Singapore" pageId="14" pageNumber="67">SINGAPORE</collectingCountry>
: 424 Y774
<location id="8ED9604CEA332932CCBDBFEB1D69DDE6" LSID="urn:lsid:plazi:treatment:03AF8781EA332933CCE7BCA41FA9DBD8:8ED9604CEA332932CCBDBFEB1D69DDE6" box="[196,322,658,680]" country="Singapore" county="Forest" municipality="Peirce" name="Upper Pierce" pageId="14" pageNumber="67">Upper Pierce</location>
[
<collectingMunicipality id="6BDDACEDEA332932CD37BFE91DA4DDE9" box="[334,399,657,679]" pageId="14" pageNumber="67">Peirce</collectingMunicipality>
]
<collectingCounty id="62D84E1BEA332932CDE2BFEB1DF2DDE6" box="[411,473,659,680]" pageId="14" pageNumber="67">Forest</collectingCounty>
, J stream
<date id="FFB81057EA332932CE31BFEA1E9FDDE9" box="[584,692,658,679]" pageId="14" pageNumber="67" value="1991-10-31">
<collectingDate id="EFFCE9BFEA332932CE31BFEA1E9FDDE9" box="[584,692,658,679]" pageId="14" pageNumber="67" value="1991-10-31">31.10.1991</collectingDate>
</date>
leg.
<collectorName id="26F35341EA332932CE9DBFEB1CFEDD8B" pageId="14" pageNumber="67">DH Murphy</collectorName>
et al”
</materialsCitation>
.
<materialsCitation id="3B6E3CCAEA332932CD65BFD71D00DDAC" collectingDate="1991-10-31" collectionCode="NMW, ZRC" collectorName="DH Murphy" country="Singapore" county="Forest" location="Upper Pierce" municipality="Peirce" pageId="14" pageNumber="67" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="B972EA85EA332932CD65BFD71DA8DD8A" bold="true" box="[284,387,687,708]" pageId="14" pageNumber="67">
<typeStatus id="54BD8835EA332932CD65BFD71DA8DD8A" box="[284,387,687,708]" pageId="14" pageNumber="67" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D00FD1EEA332932CDE9BFD71DC5DD8B" box="[400,494,687,709]" pageId="14" pageNumber="67" type="female">2 females</specimenCount>
(
<collectionCode id="ED17AE52EA332932CD83BFD71E12DD8A" box="[506,569,687,708]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="67">NMW</collectionCode>
,
<collectionCode id="ED17AE52EA332932CE3DBFD71E5DDD8B" box="[580,630,687,709]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="14" pageNumber="67">ZRC</collectionCode>
), same locality data as
<typeStatus id="54BD8835EA332932CCADBFB41D00DDAC" box="[212,299,716,738]" pageId="14" pageNumber="67" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="3B6E3CCAEA332932CD4EBFB41E48DDB0" collectingDate="1992-04-30" collectionCode="ZRC" collectorName="Coll. KL Yeo &amp; Bengkeng" country="Singapore" location="Nee Soon Swamp Forest" pageId="14" pageNumber="67" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D00FD1EEA332932CD4EBFB41DA7DDAC" box="[311,396,716,738]" pageId="14" pageNumber="67" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA332932CDE5BFB41DFBDDAC" box="[412,464,716,738]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="14" pageNumber="67">ZRC</collectionCode>
): “
<collectingCountry id="F3117607EA332932CD92BFB41E65DDAC" box="[491,590,716,738]" name="Singapore" pageId="14" pageNumber="67">Singapore</collectingCountry>
<location id="8ED9604CEA332932CE2EBFB41CEDDDB0" LSID="urn:lsid:plazi:treatment:03AF8781EA332933CCE7BCA41FA9DBD8:8ED9604CEA332932CE2EBFB41CEDDDB0" country="Singapore" name="Nee Soon Swamp Forest" pageId="14" pageNumber="67">Nee Soon Swamp Forest</location>
,
<date id="FFB81057EA332932CCABBF911D71DDB0" box="[210,346,745,766]" pageId="14" pageNumber="67" value="1992-04-30">
<collectingDate id="EFFCE9BFEA332932CCABBF911D71DDB0" box="[210,346,745,766]" pageId="14" pageNumber="67" value="1992-04-30">30. Apr. 1992</collectingDate>
</date>
<collectorName id="26F35341EA332932CD1BBF901DC3DDB0" box="[354,488,744,766]" pageId="14" pageNumber="67">Coll. KL Yeo</collectorName>
,
<collectorName id="26F35341EA332932CD8DBF901E71DDB0" box="[500,602,744,766]" pageId="14" pageNumber="67">Bengkeng</collectorName>
</materialsCitation>
;
<materialsCitation id="3B6E3CCAEA332932CE16BF911D6DDC76" collectingDate="1997-04-21" collectionCode="NMW, ZRC" country="Singapore" location="Nee Soon Swamp-forest" pageId="14" pageNumber="67" specimenCount="10" typeStatus="paratype">
<specimenCount id="9D00FD1EEA332932CE16BF911E9DDDB0" box="[623,694,745,766]" pageId="14" pageNumber="67" type="generic">10 exs.</specimenCount>
(
<collectionCode id="ED17AE52EA332932CEBDBF911F28DDB0" box="[708,771,745,766]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="67">NMW</collectionCode>
,
<collectionCode id="ED17AE52EA332932CCF1BE7D1C96DC55" box="[136,189,773,795]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="14" pageNumber="67">ZRC</collectionCode>
): “
<collectingCountry id="F3117607EA332932CCA3BE7D1D43DC54" box="[218,360,773,794]" name="Singapore" pageId="14" pageNumber="67">SINGAPORE</collectingCountry>
,
<location id="8ED9604CEA332932CD0DBE7E1E40DC55" LSID="urn:lsid:plazi:treatment:03AF8781EA332933CCE7BCA41FA9DBD8:8ED9604CEA332932CD0DBE7E1E40DC55" box="[372,619,773,795]" country="Singapore" name="Nee Soon Swamp-forest" pageId="14" pageNumber="67">Nee Soon Swamp-forest</location>
<date id="FFB81057EA332932CE0CBE7D1EDCDC54" box="[629,759,773,794]" pageId="14" pageNumber="67" value="1997-04-21">
<collectingDate id="EFFCE9BFEA332932CE0CBE7D1EDCDC54" box="[629,759,773,794]" pageId="14" pageNumber="67" value="1997-04-21">21 Apr 1997</collectingDate>
</date>
\ Balke &amp; Hendrich”
</materialsCitation>
;
<materialsCitation id="3B6E3CCAEA332932CD29BE5A1E67DC1B" collectingDate="1997-04-21" collectionCode="ZRC" country="Singapore" location="Nee Soon Swampforest." pageId="14" pageNumber="67" specimenCount="5" typeStatus="paratype">
<specimenCount id="9D00FD1EEA332932CD29BE5A1DA2DC76" box="[336,393,802,824]" pageId="14" pageNumber="67" type="generic">5 exs.</specimenCount>
(
<collectionCode id="ED17AE52EA332932CDECBE5A1DE2DC76" box="[405,457,802,824]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="14" pageNumber="67">ZRC</collectionCode>
): “
<collectingCountry id="F3117607EA332932CD9ABE5A1E62DC76" box="[483,585,802,824]" name="Singapore" pageId="14" pageNumber="67">Singapore</collectingCountry>
:
<location id="8ED9604CEA332932CE2BBE5B1CEDDC1B" LSID="urn:lsid:plazi:treatment:03AF8781EA332933CCE7BCA41FA9DBD8:8ED9604CEA332932CE2BBE5B1CEDDC1B" country="Singapore" name="Nee Soon Swampforest." pageId="14" pageNumber="67">Nee Soon Swampforest.</location>
<date id="FFB81057EA332932CCB4BE471D67DC1A" box="[205,332,831,852]" pageId="14" pageNumber="67" value="1997-04-21">
<collectingDate id="EFFCE9BFEA332932CCB4BE471D67DC1A" box="[205,332,831,852]" pageId="14" pageNumber="67" value="1997-04-21">21 Apr 1997</collectingDate>
</date>
. M Balke &amp; L Hendrich”
</materialsCitation>
;
<materialsCitation id="3B6E3CCAEA332932CE2EBE471E66DCC0" collectingDate="1997-04-28" collectionCode="ZSM" collectorName="Balke &amp; Hendrich" country="Singapore" location="SINGAPORE" pageId="14" pageNumber="67" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D00FD1EEA332932CE2EBE471E8EDC1B" box="[599,677,831,853]" pageId="14" pageNumber="67" type="male">2 males</specimenCount>
,
<specimenCount id="9D00FD1EEA332932CEC8BE471F2CDC1B" box="[689,775,831,853]" pageId="14" pageNumber="67" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA332932CCF6BE241CEDDC3C" box="[143,198,860,882]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="67" type="Museum">ZSM</collectionCode>
): “
<collectingCountry id="F3117607EA332932CC9ABE241D5BDC3F" box="[227,368,860,881]" name="Singapore" pageId="14" pageNumber="67">SINGAPORE</collectingCountry>
: 21.-
<date id="FFB81057EA332932CDE4BE241E2FDC3F" box="[413,516,860,881]" pageId="14" pageNumber="67" value="1997-04-28">
<collectingDate id="EFFCE9BFEA332932CDE4BE241E2FDC3F" box="[413,516,860,881]" pageId="14" pageNumber="67" value="1997-04-28">28.4.1997</collectingDate>
</date>
<collectingMethod id="52474E80EA332932CE74BE241D67DCC0" pageId="14" pageNumber="67">Nee Soon Swamp Forest flight intercept trap</collectingMethod>
leg.
<collectorName id="26F35341EA332932CDFDBE001D96DCC0" box="[388,445,888,910]" pageId="14" pageNumber="67">Balke</collectorName>
&amp;
<collectorName id="26F35341EA332932CD9ABE001E6FDCC0" box="[483,580,888,910]" pageId="14" pageNumber="67">Hendrich</collectorName>
</materialsCitation>
;
<materialsCitation id="3B6E3CCAEA332932CE22BE011C96DCAB" collectingDate="1997-04-28" collectionCode="ZSM" collectorName="Flight Intercept Trap Balke &amp; Hendrich" country="Singapore" location="Nee Soon Swamp Forest" pageId="14" pageNumber="67" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D00FD1EEA332932CE22BE011E9FDCC0" box="[603,692,888,910]" pageId="14" pageNumber="67" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA332932CEBDBE011ED7DCC1" box="[708,764,889,911]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="67" type="Museum">ZSM</collectionCode>
): “
<collectingCountry id="F3117607EA332932CCE8BEED1D09DCE4" box="[145,290,917,938]" name="Singapore" pageId="14" pageNumber="67">SINGAPORE</collectingCountry>
/
<location id="8ED9604CEA332932CD53BEED1E12DCE5" LSID="urn:lsid:plazi:treatment:03AF8781EA332933CCE7BCA41FA9DBD8:8ED9604CEA332932CD53BEED1E12DCE5" box="[298,569,917,939]" country="Singapore" name="Nee Soon Swamp Forest" pageId="14" pageNumber="67">Nee Soon Swamp Forest</location>
, small path n[ear]. pipeline, 21.-
<date id="FFB81057EA332932CD7EBECA1D58DC89" box="[263,371,946,967]" pageId="14" pageNumber="67" value="1997-04-28">
<collectingDate id="EFFCE9BFEA332932CD7EBECA1D58DC89" box="[263,371,946,967]" pageId="14" pageNumber="67" value="1997-04-28">28.4. 1997</collectingDate>
</date>
<collectorName id="26F35341EA332932CD05BECA1EA2DC86" box="[380,649,946,968]" pageId="14" pageNumber="67">Flight Intercept Trap Balke</collectorName>
&amp;
<collectorName id="26F35341EA332932CED7BECA1F2CDC86" box="[686,775,946,968]" pageId="14" pageNumber="67">Hendrich</collectorName>
leg.”
</materialsCitation>
;
<materialsCitation id="3B6E3CCAEA332932CCB5BEB71EB6DB4C" collectingDate="2002-01-19" collectionCode="NMW" collectorName="Zettel" country="Singapore" location="McRitchie Reservoir" pageId="14" pageNumber="67" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D00FD1EEA332932CCB5BEB71D3CDCAB" box="[204,279,975,997]" pageId="14" pageNumber="67" type="male">1 male</specimenCount>
,
<specimenCount id="9D00FD1EEA332932CD5FBEB71DA8DCAB" box="[294,387,975,997]" pageId="14" pageNumber="67" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA332932CDECBEB71DF7DCAB" box="[405,476,975,997]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="67">NMW</collectionCode>
): “
<collectingCountry id="F3117607EA332932CD83BEB71EA1DCAA" box="[506,650,975,996]" name="Singapore" pageId="14" pageNumber="67">SINGAPORE</collectingCountry>
<location id="8ED9604CEA332932CEE3BEB71CCEDB4C" LSID="urn:lsid:plazi:treatment:03AF8781EA332933CCE7BCA41FA9DBD8:8ED9604CEA332932CEE3BEB71CCEDB4C" country="Singapore" name="McRitchie Reservoir" pageId="14" pageNumber="67">McRitchie Reservoir</location>
preserve area,
<date id="FFB81057EA332932CD05BE941DF5DB4F" box="[380,478,1004,1025]" pageId="14" pageNumber="67" value="2002-01-19">
<collectingDate id="EFFCE9BFEA332932CD05BE941DF5DB4F" box="[380,478,1004,1025]" pageId="14" pageNumber="67" value="2002-01-19">19.1.2002</collectingDate>
</date>
, leg.
<collectorName id="26F35341EA332932CE6CBE941E66DB4C" box="[533,589,1004,1026]" pageId="14" pageNumber="67">Zettel</collectorName>
(SG4)”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA332933CCF1B9521904DC58" lastPageId="15" lastPageNumber="68" pageId="14" pageNumber="67" type="description">
<paragraph id="8BB93697EA332932CCF1B9521F2DDB2C" blockId="14.[136,775,1066,1122]" pageId="14" pageNumber="67">
<emphasis id="B972EA85EA332932CCF1B9521D25DB0C" bold="true" box="[136,270,1066,1090]" italics="true" pageId="14" pageNumber="67">Description.</emphasis>
— Habitus as in
<figureCitation id="133D2A12EA332932CDAEB9521E30DB0C" box="[471,539,1066,1090]" captionStart="Figs" captionStartId="3.[113,157,2061,2083]" captionTargetBox="[222,1331,314,1975]" captionTargetId="figure-8@3.[195,1369,301,1985]" captionTargetPageId="3" captionText="Figs. 14. Habitus of: 1, Hydraena hendrichi, paratype; 2, H. paulmoritz, paratype; 3, H. singaporensis, paratype; 4, H. yangae, holotype, right foreleg (lateral view) depicted separately (photograph taken before right elytron got unhinged)." figureDoi="http://doi.org/10.5281/zenodo.5351600" httpUri="https://zenodo.org/record/5351600/files/figure.png" pageId="14" pageNumber="67">Fig. 3</figureCitation>
.
<quantity id="4CFE9B72EA332932CE50B9521EEDDB0F" box="[553,710,1066,1090]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.25" metricValueMin="1.05" pageId="14" pageNumber="67" unit="mm" value="1.15" valueMax="1.25" valueMin="1.05">1.051.25 mm</quantity>
long. Dorsum unicoloured reddish brown to dark brown, or head
</paragraph>
<paragraph id="8BB93697EA332932CF3ABDC518A6DFBB" blockId="14.[835,1473,189,245]" pageId="14" pageNumber="67">and/or pronotum paler yellowish brown; maxillary palpi always unicoloured yellowish.</paragraph>
<paragraph id="8BB93697EA332932CF3ABC651FCFDEBB" blockId="14.[835,1474,285,501]" pageId="14" pageNumber="67">Labrum excised anteriorly, very sparsely punctate. Clypeus sparsely punctate and smooth medially. Fronto-clypeal suture slightly arcuate, slightly impressed. Frons sparsely (middle) to moderately densely (laterally) punctate, interstices shining; interocular grooves more or less obsolete. Eyes large, protruding, more than 30 facets visible in dorsal view.</paragraph>
<paragraph id="8BB93697EA332932CF3ABF661905DDBB" blockId="14.[835,1475,541,757]" pageId="14" pageNumber="67">Pronotum weakly cordiform, wider than long; anterior margin concave; anterior angles rounded; lateral rim slightly denticulate and very slightly produced at middle; disc rather flat, sparsely punctate, glabrous, disc with a pair of shallow basal impressions, which are sometimes connected by shallow transverse groove; anterior, posterior and lateral portions of pronotum more densely punctate.</paragraph>
<paragraph id="8BB93697EA332932CF3ABE6519B6DCFB" blockId="14.[835,1474,797,949]" pageId="14" pageNumber="67">Elytra suboval, with about nine rows of punctures between suture and shoulder; punctures small, not deeply impressed, arranged in more or less regular, not impressed lines; intervals and interstices flat and glabrous; explanate margin of elytra narrow, hardly noticeably denticulate apically.</paragraph>
<paragraph id="8BB93697EA332932CF3ABEA5185BDB7B" blockId="14.[835,1475,989,1077]" pageId="14" pageNumber="67">
Mesoventral process (
<figureCitation id="133D2A12EA332932C84ABEA518A9DCBB" box="[1075,1154,989,1013]" captionStart="Figs" captionStartId="12.[137,181,2091,2113]" captionTargetBox="[138,1476,1428,2061]" captionTargetId="figure-8@12.[134,1479,1425,2064]" captionTargetPageId="12" captionText="Figs. 1214. Meso- and metaventrite of: 12, Hydraena hendrichi; 13, H. singaporensis; and 14, H. formula." figureDoi="http://doi.org/10.5281/zenodo.5351616" httpUri="https://zenodo.org/record/5351616/files/figure.png" pageId="14" pageNumber="67">Fig. 13</figureCitation>
) about as wide as mesotibia. Metaventrite (
<figureCitation id="133D2A12EA332932CF97BE85186DDB5B" box="[1006,1094,1021,1045]" captionStart="Figs" captionStartId="12.[137,181,2091,2113]" captionTargetBox="[138,1476,1428,2061]" captionTargetId="figure-8@12.[134,1479,1425,2064]" captionTargetPageId="12" captionText="Figs. 1214. Meso- and metaventrite of: 12, Hydraena hendrichi; 13, H. singaporensis; and 14, H. formula." figureDoi="http://doi.org/10.5281/zenodo.5351616" httpUri="https://zenodo.org/record/5351616/files/figure.png" pageId="14" pageNumber="67">Fig. 13</figureCitation>
) moderately deeply impressed, metaventral plaques absent.
</paragraph>
<caption id="DF79661FEA332932CCF0B5521FA4D70E" ID-DOI="http://doi.org/10.5281/zenodo.5351618" ID-Zenodo-Dep="5351618" box="[137,911,2090,2112]" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="14" pageNumber="67" startId="14.[137,172,2090,2112]" targetBox="[137,1470,1208,2063]" targetPageId="14">
<paragraph id="8BB93697EA332932CCF0B5521FA4D70E" blockId="14.[137,911,2090,2112]" box="[137,911,2090,2112]" pageId="14" pageNumber="67">
Fig. 15. Geographical map of Singapore, showing distribution of
<taxonomicName id="4C064D14EA332932CF74B5521FA0D70E" box="[781,907,2090,2112]" class="Insecta" family="Hydraenidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="67" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BB93697EA322933CC08BDC51DFADE5B" blockId="15.[113,752,189,277]" pageId="15" pageNumber="68">
Male terminal sternite and spiculum (
<figureCitation id="133D2A12EA322933CE51BDC51EAADF9B" box="[552,641,189,213]" captionStart="Fig" captionStartId="4.[136,171,2060,2082]" captionTargetBox="[157,1469,185,2015]" captionTargetId="figure-8@4.[98,1492,175,2032]" captionTargetPageId="4" captionText="Fig. 5. Hydraena singaporensis: a, aedeagus in right lateral view; b, aedeagus, ventral view (flagellum not depicted completely in a, b); c, aedeagus, left lateral view; d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351602" httpUri="https://zenodo.org/record/5351602/files/figure.png" pageId="15" pageNumber="68">Fig. 5d</figureCitation>
): Sternite subrectangular, longer than wide, apical half asymmetrical, firmly connected with spiculum.
</paragraph>
<paragraph id="8BB93697EA322933CC08BC451DDDDD9B" blockId="15.[113,752,317,725]" pageId="15" pageNumber="68">
Aedeagus (
<figureCitation id="133D2A12EA322933CC9EBC451D65DE1B" box="[231,334,317,341]" captionStart="Fig" captionStartId="4.[136,171,2060,2082]" captionTargetBox="[157,1469,185,2015]" captionTargetId="figure-8@4.[98,1492,175,2032]" captionTargetPageId="4" captionText="Fig. 5. Hydraena singaporensis: a, aedeagus in right lateral view; b, aedeagus, ventral view (flagellum not depicted completely in a, b); c, aedeagus, left lateral view; d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351602" httpUri="https://zenodo.org/record/5351602/files/figure.png" pageId="15" pageNumber="68">Fig. 5ac</figureCitation>
): Main piece strongly angulate in lateral view (basal and apical part forming a right angle), apical part long and slender, almost straight, without setae, ventral margin of basal part with conspicuous spine; phallobase subsymmetrical. Distal lobe moderately large, inserted near angle between basal and apical part of main piece, not reaching apex of main piece, with an extraordinary long, thin, coiled flagellum. Parameres articulately connected with main piece, inserted near phallobase. Left paramere very short, suboval, with about five apical setae. Right paramere elongate and slender, club-shaped, reaching middle of main piece; apex with about four moderately long setae, lateral face with short spines.
</paragraph>
<paragraph id="8BB93697EA322933CC08BF851D43DC1B" blockId="15.[113,752,765,853]" pageId="15" pageNumber="68">
Gonocoxite (
<figureCitation id="133D2A12EA322933CC84BF851D60DC5B" box="[253,331,765,789]" captionStart="Fig" captionStartId="4.[136,171,2060,2082]" captionTargetBox="[157,1469,185,2015]" captionTargetId="figure-8@4.[98,1492,175,2032]" captionTargetPageId="4" captionText="Fig. 5. Hydraena singaporensis: a, aedeagus in right lateral view; b, aedeagus, ventral view (flagellum not depicted completely in a, b); c, aedeagus, left lateral view; d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351602" httpUri="https://zenodo.org/record/5351602/files/figure.png" pageId="15" pageNumber="68">Fig. 5e</figureCitation>
): Subtriangular, approximately as wide as long; lateral margins curved; inner plate hardly noticeably projecting sublaterally.
</paragraph>
<paragraph id="8BB93697EA322933CC08BE051CE2DCBB" blockId="15.[113,752,893,1013]" pageId="15" pageNumber="68">
Female tergite X (
<figureCitation id="133D2A12EA322933CD48BE051D57DCDB" box="[305,380,893,917]" captionStart="Fig" captionStartId="4.[136,171,2060,2082]" captionTargetBox="[157,1469,185,2015]" captionTargetId="figure-8@4.[98,1492,175,2032]" captionTargetPageId="4" captionText="Fig. 5. Hydraena singaporensis: a, aedeagus in right lateral view; b, aedeagus, ventral view (flagellum not depicted completely in a, b); c, aedeagus, left lateral view; d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351602" httpUri="https://zenodo.org/record/5351602/files/figure.png" pageId="15" pageNumber="68">Fig. 5f</figureCitation>
): Subsemicircular, disc moderately densely covered with trichoid setae; subapical setae also trichoid, vermiform setae absent; posterior margin excised.
</paragraph>
<paragraph id="8BB93697EA322933CF55BDC61F5EDE78" blockId="15.[812,1452,190,310]" pageId="15" pageNumber="68">
Spermatheca (
<figureCitation id="133D2A12EA322933CFBFBDC61819DF98" box="[966,1074,190,214]" captionStart="Fig" captionStartId="4.[136,171,2060,2082]" captionTargetBox="[157,1469,185,2015]" captionTargetId="figure-8@4.[98,1492,175,2032]" captionTargetPageId="4" captionText="Fig. 5. Hydraena singaporensis: a, aedeagus in right lateral view; b, aedeagus, ventral view (flagellum not depicted completely in a, b); c, aedeagus, left lateral view; d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351602" httpUri="https://zenodo.org/record/5351602/files/figure.png" pageId="15" pageNumber="68">
Fig.
<quantity id="4CFE9B72EA322933CF81BDC61838DF98" box="[1016,1043,190,214]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="15" pageNumber="68" unit="g" value="5.0">5g</quantity>
h
</figureCitation>
): Proximal portion crescentic, with sickle-shaped transverse crest in the middle; distal portion discoidal; spermathecal duct very long and conspicuously coiled.
</paragraph>
<paragraph id="8BB93697EA322933CF55BC261FAADEB8" blockId="15.[812,1452,350,502]" pageId="15" pageNumber="68">Secondary sexual characters: Male foretibia very slightly curved. Male mesoventrite more deeply impressed between mesoventral disc and mesoventral process. Male tergite X excised apically, asymmetrical (left lobe wider and longer).</paragraph>
<paragraph id="8BB93697EA322933CF55BF661904DC58" blockId="15.[812,1453,542,790]" pageId="15" pageNumber="68">
Variability: The variability in the colouration of the dorsum is quite remarkable. The
<specimenCount id="9D00FD1EEA322933C84CBF4618CBDD18" box="[1077,1248,574,598]" pageId="15" pageNumber="68" type="generic">three specimens</specimenCount>
from Peirce Forest are unicoloured, whereas all other specimens are more or less distinctly bicoloured. This may, however, be attributed to tenerality rather than to genetic differences. The
<specimenCount id="9D00FD1EEA322933C90FBFE61FB6DD98" pageId="15" pageNumber="68" type="generic">three specimens</specimenCount>
from Peirce Forest were collected in October, while the remaining specimens were collected from January to April, some of these being distinctly teneral.
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA322933CF55BE4519AADCF8" pageId="15" pageNumber="68" type="diagnosis">
<paragraph id="8BB93697EA322933CF55BE4519AADCF8" blockId="15.[812,1453,829,950]" pageId="15" pageNumber="68">
<emphasis id="B972EA85EA322933CF55BE451870DC18" bold="true" box="[812,1115,829,854]" italics="true" pageId="15" pageNumber="68">Differential diagnosis.</emphasis>
— Externally,
<taxonomicName id="4C064D14EA322933C948BE381F94DC39" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="singaporensis">
<emphasis id="B972EA85EA322933C948BE381F94DC39" italics="true" pageId="15" pageNumber="68">Hydraena singaporensis</emphasis>
</taxonomicName>
can be recognised by the combination of the following characters: small body size, mesoventral process not wider than mesotibia, metaventral plaques lacking.
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA322933CF55BEA51800DB78" pageId="15" pageNumber="68" type="distribution">
<paragraph id="8BB93697EA322933CF55BEA51800DB78" blockId="15.[812,1452,989,1078]" pageId="15" pageNumber="68">
<emphasis id="B972EA85EA322933CF55BEA51F93DCB8" bold="true" box="[812,952,989,1014]" italics="true" pageId="15" pageNumber="68">Distribution.</emphasis>
— So far known only from
<collectingCountry id="F3117607EA322933C894BEA61972DCB8" box="[1261,1369,990,1014]" name="Singapore" pageId="15" pageNumber="68">Singapore</collectingCountry>
(Upper Peirce Forest, Nee Soon Swamp Forest, MacRitchie Reservoir), see
<figureCitation id="133D2A12EA322933CFAEB966180CDB78" box="[983,1063,1054,1078]" captionStart="Fig" captionStartId="14.[137,172,2090,2112]" captionTargetBox="[137,1470,1208,2063]" captionTargetId="figure-8@14.[137,1478,1202,2067]" captionTargetPageId="14" captionText="Fig. 15. Geographical map of Singapore, showing distribution of Hydraenidae." figureDoi="http://doi.org/10.5281/zenodo.5351618" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="15" pageNumber="68">Fig. 15</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA322933CF55B9251FA9DBD8" pageId="15" pageNumber="68" type="etymology">
<paragraph id="8BB93697EA322933CF55B9251FA9DBD8" blockId="15.[812,1452,1117,1174]" pageId="15" pageNumber="68">
<emphasis id="B972EA85EA322933CF55B9251F8CDB38" bold="true" box="[812,935,1117,1142]" italics="true" pageId="15" pageNumber="68">Etymology.</emphasis>
— This species named in reference to the
<typeStatus id="54BD8835EA322933C907B9271987DB39" box="[1406,1452,1119,1143]" pageId="15" pageNumber="68">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>