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<document id="0A2750CE48481E0411438CD7C65F5815" ID-DOI="10.5281/zenodo.5351598" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351598" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630206282112" checkinUser="tatiana" docAuthor="Jäch, Manfred A., Díaz, Juan A. &amp; Skale, Andre" docDate="2013" docId="03AF8781EA32292DCF1EB9A51D0DDFBB" docLanguage="en" docName="61rbz053-071.pdf" docOrigin="Raffles Bulletin of Zoology 61 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Hydraena (Hydraenopsis) yangae Jäch &amp; Díaz &amp; Skale 2013, new species" docType="treatment" docVersion="3" lastPageNumber="70" masterDocId="FF96FFF9EA3D293CCC79BD781C2BDF4E" masterDocTitle="The Hydraenidae (Coleoptera) Of The Republic Of Singapore" masterLastPageNumber="71" masterPageNumber="53" pageNumber="68" updateTime="1699184966419" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="C92FA2CB922612E7C8F332ACFEFD6BA6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A35E9E5D8194BEE683C96B59A612F874">
<mods:title id="7C8433DE83837BEB15532EE84302803D">The Hydraenidae (Coleoptera) Of The Republic Of Singapore</mods:title>
</mods:titleInfo>
<mods:name id="171B1E6B1B41DA1810FB0C334B18011C" type="personal">
<mods:role id="0F02AB5C776FD7F975DE7ED55D324AA8">
<mods:roleTerm id="C5235966FE35F5A40DDF38EE7B7CA822">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70F979C54F37FF89CA968050E0738685">Jäch, Manfred A.</mods:namePart>
</mods:name>
<mods:name id="723E1A40C11983E863EE5EE9C78EF79A" type="personal">
<mods:role id="4A28E2D78F9F84445D97CE6C6543357F">
<mods:roleTerm id="58FB3E68C6AD1A3A619837A65547942C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99E719AE75B475998FA895F7F93DA0F1">Díaz, Juan A.</mods:namePart>
</mods:name>
<mods:name id="1C73C04EDE55CFA9FA204FD29D2F8A02" type="personal">
<mods:role id="E7D9E3EA204CADEE0AA15D4CBA924A6D">
<mods:roleTerm id="4A30A5685BD714E6F1CCA6155DA9812C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="52398CD76344197F96A29861847B4C26">Skale, Andre</mods:namePart>
</mods:name>
<mods:typeOfResource id="5825B8BBDB4C9651B367DAA249C3FA51">text</mods:typeOfResource>
<mods:relatedItem id="06BD31941967A7D8EF21A219D191EA41" type="host">
<mods:titleInfo id="4BB23AEB2646F929CC15A891386974DC">
<mods:title id="F305AF470A11D8F934CBF3E0392C5606">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="76C7F0C9EF2C84DE4A84A1F5D9199F75">
<mods:date id="B9A7BC4FB785171FD937B4882CA6E3FA">2013</mods:date>
<mods:detail id="F811396A57C8098B7E6667C03306F3F6" type="pubDate">
<mods:number id="38C8F57AAAEF4E262172F766D99C35D5">2013-02-28</mods:number>
</mods:detail>
<mods:detail id="EB4614C278089493D543D66273AE5FAD" type="volume">
<mods:number id="2F4FE845132A71A09C5844B9561C1967">61</mods:number>
</mods:detail>
<mods:detail id="767943C059F6C45CD1A4E031D1E6D915" type="issue">
<mods:number id="CDA36315CB25E28229C944547BCD1D57">1</mods:number>
</mods:detail>
<mods:extent id="218AEAEF1FE2CC3B598D0E908A5803E9" unit="page">
<mods:start id="3F0994811D12E57B6FAAA38617071CCC">53</mods:start>
<mods:end id="24A9609F49BE7975D947E2E43A7B449B">71</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="788EC296D7379AEED9BF8F504EE142F7">journal article</mods:classification>
<mods:identifier id="9AEB0F894EB3CA6EF78990421EF2E431" type="DOI">10.5281/zenodo.5351598</mods:identifier>
<mods:identifier id="DA84A324877C2991815B5BD46B334783" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="C5F19701FFE7E8FBD7EAC6E6FA6F258C" type="Zenodo-Dep">5351598</mods:identifier>
</mods:mods>
<treatment id="03AF8781EA32292DCF1EB9A51D0DDFBB" LSID="urn:lsid:plazi:treatment:03AF8781EA32292DCF1EB9A51D0DDFBB" httpUri="http://treatment.plazi.org/id/03AF8781EA32292DCF1EB9A51D0DDFBB" lastPageId="17" lastPageNumber="70" pageId="15" pageNumber="68">
<subSubSection id="C31C651CEA322933CF1EB9A51959DBB8" box="[871,1394,1245,1270]" pageId="15" pageNumber="68" type="nomenclature">
<paragraph id="8BB93697EA322933CF1EB9A51959DBB8" blockId="15.[871,1394,1245,1302]" box="[871,1394,1245,1270]" pageId="15" pageNumber="68">
<heading id="D0F181FBEA322933CF1EB9A51959DBB8" bold="true" box="[871,1394,1245,1270]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="68" reason="2">
<emphasis id="B972EA85EA322933CF1EB9A51959DBB8" bold="true" box="[871,1394,1245,1270]" pageId="15" pageNumber="68">
<taxonomicName id="4C064D14EA322933CF1EB9A518F5DBB8" ID-CoL="3N3KV" authority="Jäch &amp; Díaz &amp; Skale, 2013" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[871,1246,1245,1270]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="yangae" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B972EA85EA322933CF1EB9A51FFEDBB8" bold="true" box="[871,981,1245,1270]" italics="true" pageId="15" pageNumber="68">Hydraena</emphasis>
(
<emphasis id="B972EA85EA322933CF9FB9A51856DBB8" bold="true" box="[998,1149,1245,1270]" italics="true" pageId="15" pageNumber="68">Hydraenopsis</emphasis>
)
<emphasis id="B972EA85EA322933C8F6B9A718F5DBB8" bold="true" box="[1167,1246,1247,1270]" italics="true" pageId="15" pageNumber="68">yangae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24157FEEA322933C895B9A61959DBB8" box="[1260,1394,1246,1270]" pageId="15" pageNumber="68" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA322933C862B9861896DA58" box="[1051,1213,1278,1302]" pageId="15" pageNumber="68" type="description">
<paragraph id="8BB93697EA322933C862B9861896DA58" blockId="15.[871,1394,1245,1302]" box="[1051,1213,1278,1302]" pageId="15" pageNumber="68">
(
<figureCitation id="133D2A12EA322933C85AB986185BDA58" box="[1059,1136,1278,1302]" captionStart="Figs" captionStartId="3.[113,157,2061,2083]" captionTargetBox="[222,1331,314,1975]" captionTargetId="figure-8@3.[195,1369,301,1985]" captionTargetPageId="3" captionText="Figs. 1–4. Habitus of: 1, Hydraena hendrichi, paratype; 2, H. paulmoritz, paratype; 3, H. singaporensis, paratype; 4, H. yangae, holotype, right foreleg (lateral view) depicted separately (photograph taken before right elytron got unhinged)." figureDoi="http://doi.org/10.5281/zenodo.5351600" httpUri="https://zenodo.org/record/5351600/files/figure.png" pageId="15" pageNumber="68">Figs. 4</figureCitation>
,
<figureCitation id="133D2A12EA322933C807B98718A6DA58" box="[1150,1165,1279,1302]" captionStart="Fig" captionStartId="6.[136,171,2060,2082]" captionTargetBox="[166,1446,190,2011]" captionTargetId="figure-8@6.[43,1493,170,2020]" captionTargetPageId="6" captionText="Fig. 7. Hydraena yangae: a, aedeagus in lateral view; b, aedeagus, dorsal view (setae of left paramere omitted in a, b); c, aedeagus, ventral view (setae of right paramere omitted); d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351606" httpUri="https://zenodo.org/record/5351606/files/figure.png" pageId="15" pageNumber="68">7</figureCitation>
,
<figureCitation id="133D2A12EA322933C8E3B986189EDA58" box="[1178,1205,1278,1302]" captionStart="Fig" captionStartId="14.[137,172,2090,2112]" captionTargetBox="[137,1470,1208,2063]" captionTargetId="figure-8@14.[137,1478,1202,2067]" captionTargetPageId="14" captionText="Fig. 15. Geographical map of Singapore, showing distribution of Hydraenidae." figureDoi="http://doi.org/10.5281/zenodo.5351618" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="15" pageNumber="68">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA322933CF55B8451FA6D9AB" pageId="15" pageNumber="68" type="materials_examined">
<paragraph id="8BB93697EA322933CF55B845184DDAC2" blockId="15.[812,1453,1340,1420]" pageId="15" pageNumber="68">
<materialsCitation id="3B6E3CCAEA322933CF55B845184ADAC2" country="Singapore" latitude="1.355" location="Rifle Range Road" longLatPrecision="21" longitude="103.7975" municipality="Bukit Timah" pageId="15" pageNumber="68" specimenCount="1" typeStatus="holotype">
<emphasis id="B972EA85EA322933CF55B8451F85DA1C" bold="true" box="[812,942,1340,1362]" italics="true" pageId="15" pageNumber="68">
<typeStatus id="54BD8835EA322933CF55B8451F70DA1C" box="[812,859,1341,1362]" pageId="15" pageNumber="68">Type</typeStatus>
locality.
</emphasis>
— Sandy margin of a very shallow and slow flowing, shaded stream,
<geoCoordinate id="EE325050EA322933CFB1B8221805DA21" box="[968,1070,1370,1391]" degrees="1" direction="north" minutes="21" orientation="latitude" pageId="15" pageNumber="68" precision="15" seconds="18" value="1.355">1°21'18&quot;N</geoCoordinate>
<geoCoordinate id="EE325050EA322933C84EB822189EDA3E" box="[1079,1205,1370,1392]" degrees="103" direction="east" minutes="47" orientation="longitude" pageId="15" pageNumber="68" precision="15" seconds="51" value="103.7975">103°47'51&quot;E</geoCoordinate>
, near
<location id="8ED9604CEA322933C88DB8221983DA3E" LSID="urn:lsid:plazi:treatment:03AF8781EA32292DCF1EB9A51D0DDFBB:8ED9604CEA322933C88DB8221983DA3E" box="[1268,1448,1370,1392]" country="Singapore" latitude="1.355" longLatPrecision="21" longitude="103.7975" municipality="Bukit Timah" name="Rifle Range Road" pageId="15" pageNumber="68">Rifle Range Road</location>
, east of
<collectingMunicipality id="6BDDACEDEA322933CF0CB80E1FD9DAC2" box="[885,1010,1398,1420]" pageId="15" pageNumber="68">Bukit Timah</collectingMunicipality>
,
<collectingCountry id="F3117607EA322933CF84B80E184ADAC2" box="[1021,1121,1398,1420]" name="Singapore" pageId="15" pageNumber="68">Singapore</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB93697EA322933CF55B8C818A2D977" blockId="15.[812,1453,1455,1593]" pageId="15" pageNumber="68">
<emphasis id="B972EA85EA322933CF55B8C81FFDDA88" bold="true" box="[812,982,1455,1478]" pageId="15" pageNumber="68">
<emphasis id="B972EA85EA322933CF55B8C81F91DA8B" bold="true" box="[812,954,1455,1477]" italics="true" pageId="15" pageNumber="68">Type material.</emphasis>
—
</emphasis>
<materialsCitation id="3B6E3CCAEA322933CFA2B8C818A2D977" collectingDate="1993-05-27" collectionCode="ZRC" collectorName="CM Yang &amp; The" country="Singapore" location="Rifle Range Rd" pageId="15" pageNumber="68" specimenCount="2" typeStatus="holotype">
<emphasis id="B972EA85EA322933CFA2B8C81813DA8B" bold="true" box="[987,1080,1456,1477]" pageId="15" pageNumber="68">
<typeStatus id="54BD8835EA322933CFA2B8C81813DA8B" box="[987,1080,1456,1477]" pageId="15" pageNumber="68" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="ED17AE52EA322933C800B8C81886DA88" box="[1145,1197,1456,1478]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="68">ZRC</collectionCode>
): “
<collectingCountry id="F3117607EA322933C8BCB8C8197ADA8B" box="[1221,1361,1456,1477]" name="Singapore" pageId="15" pageNumber="68">SINGAPORE</collectingCountry>
: (NS102) 417 ZRC.6.18870
<location id="8ED9604CEA322933CF9FB8B518A8DAAD" LSID="urn:lsid:plazi:treatment:03AF8781EA32292DCF1EB9A51D0DDFBB:8ED9604CEA322933CF9FB8B518A8DAAD" box="[998,1155,1485,1507]" country="Singapore" name="Rifle Range Rd" pageId="15" pageNumber="68">Rifle Range Rd</location>
,
<date id="FFB81057EA322933C8E9B8B518D7DAAD" box="[1168,1276,1485,1507]" pageId="15" pageNumber="68" value="1993-05-27">
<collectingDate id="EFFCE9BFEA322933C8E9B8B518D7DAAD" box="[1168,1276,1485,1507]" pageId="15" pageNumber="68" value="1993-05-27">27.05.1993</collectingDate>
</date>
leg.
<collectorName id="26F35341EA322933C94BB8B519B9DAAC" box="[1330,1426,1485,1506]" pageId="15" pageNumber="68">CM Yang</collectorName>
et al.”
<collectorName id="26F35341EA322933CF20B8921F54D94E" box="[857,895,1514,1536]" pageId="15" pageNumber="68">The</collectorName>
<typeStatus id="54BD8835EA322933CFF0B8921FCBD94E" box="[905,992,1514,1536]" pageId="15" pageNumber="68" type="holotype">holotype</typeStatus>
is teneral; right foreleg detached and glued separately; apical tarsal segments of both forelegs lacking; right elytron not firmly attached to body.
</materialsCitation>
</paragraph>
<paragraph id="8BB93697EA322933CF55BB241FA6D9AB" blockId="15.[812,1453,1628,1765]" pageId="15" pageNumber="68">
<emphasis id="B972EA85EA322933CF55BB24187CD93C" bold="true" box="[812,1111,1628,1650]" italics="true" pageId="15" pageNumber="68">Additional material examined.</emphasis>
—
<materialsCitation id="3B6E3CCAEA322933C800BB251876D9E2" collectingDate="1993-02-16" collectionCode="NMW" collectorName="Small &amp; Jungle Lodge, N" country="Malaysia" location="Gombak River" pageId="15" pageNumber="68" specimenCount="1" specimenCount-female="1" stateProvince="Kuala Lumpur">
<collectingCountry id="F3117607EA322933C800BB2518D2D93C" box="[1145,1273,1629,1650]" name="Malaysia" pageId="15" pageNumber="68">MALAYSIA</collectingCountry>
:
<specimenCount id="9D00FD1EEA322933C978BB251978D93D" box="[1281,1363,1629,1651]" pageId="15" pageNumber="68" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA322933C926BB251989D93D" box="[1375,1442,1629,1651]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="68">NMW</collectionCode>
):
<collectorName id="26F35341EA322933CF55BB021F4FD9DE" box="[812,868,1658,1680]" pageId="15" pageNumber="68">Small</collectorName>
tributary of
<location id="8ED9604CEA322933CF9CBB02185CD9DE" LSID="urn:lsid:plazi:treatment:03AF8781EA32292DCF1EB9A51D0DDFBB:8ED9604CEA322933CF9CBB02185CD9DE" box="[997,1143,1658,1680]" country="Malaysia" name="Gombak River" pageId="15" pageNumber="68" stateProvince="Kuala Lumpur">Gombak River</location>
, above
<collectorName id="26F35341EA322933C8BFBB021941D9C1" box="[1222,1386,1658,1680]" pageId="15" pageNumber="68">Jungle Lodge, N</collectorName>
<collectingRegion id="49C2F875EA322933C90ABB021F50D9E2" country="Malaysia" name="Kuala Lumpur" pageId="15" pageNumber="68">Kuala Lumpur</collectingRegion>
,
<collectingRegion id="49C2F875EA322933CFFDBBEE1FF0D9E2" box="[900,987,1686,1708]" country="Malaysia" name="Selangor" pageId="15" pageNumber="68">Selangor</collectingRegion>
,
<date id="FFB81057EA322933CF9ABBEE1876D9E2" box="[995,1117,1686,1708]" pageId="15" pageNumber="68" value="1993-02-16">
<collectingDate id="EFFCE9BFEA322933CF9ABBEE1876D9E2" box="[995,1117,1686,1708]" pageId="15" pageNumber="68" value="1993-02-16">16 Feb.1993</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B6E3CCAEA322933C81CBBEF1FA2D9AB" collectingDate="1997-04-10" collectionCode="ZSM" collectorName="Bekok &amp; M. Balke &amp; L. Hendrich" country="Malaysia" elevation="100" location="Johor" pageId="15" pageNumber="68" specimenCount="1" specimenCount-female="1" stateProvince="Johor">
<specimenCount id="9D00FD1EEA322933C81CBBEF189CD9E2" box="[1125,1207,1686,1708]" pageId="15" pageNumber="68" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA322933C8BDBBEF18D0D9E3" box="[1220,1275,1687,1709]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="15" pageNumber="68" type="Museum">ZSM</collectionCode>
): Path to waterfall,
<quantity id="4CFE9B72EA322933CF55BBCB1FB2D987" box="[812,921,1715,1737]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="15" pageNumber="68" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<elevation id="002BD1A4EA322933CF55BBCB1FB2D987" box="[812,921,1715,1737]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="15" pageNumber="68" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150 m</elevation>
</quantity>
,
<collectorName id="26F35341EA322933CFDEBBCB1FC7D987" box="[935,1004,1715,1737]" pageId="15" pageNumber="68">Bekok</collectorName>
,
<collectingRegion id="49C2F875EA322933CF83BBCB181ED987" box="[1018,1077,1715,1737]" country="Malaysia" name="Johor" pageId="15" pageNumber="68">Johor</collectingRegion>
, coll.
<collectorName id="26F35341EA322933C803BBCC18F7D987" box="[1146,1244,1715,1737]" pageId="15" pageNumber="68">M. Balke</collectorName>
&amp;
<collectorName id="26F35341EA322933C97ABBCC19AED987" box="[1283,1413,1715,1737]" pageId="15" pageNumber="68">L. Hendrich</collectorName>
,
<date id="FFB81057EA322933C9EABBCB1FA2D9AB" pageId="15" pageNumber="68" value="1997-04-10">
<collectingDate id="EFFCE9BFEA322933C9EABBCB1FA2D9AB" pageId="15" pageNumber="68" value="1997-04-10">10 Apr.1997</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA32292CCF55BA761939DB7B" lastPageId="16" lastPageNumber="69" pageId="15" pageNumber="68" type="diagnosis">
<paragraph id="8BB93697EA32292CCF55BA761D15DFBB" blockId="15.[812,1453,1806,2086]" lastBlockId="16.[136,775,189,245]" lastPageId="16" lastPageNumber="69" pageId="15" pageNumber="68">
<emphasis id="B972EA85EA322933CF55BA761F8BD868" bold="true" box="[812,928,1806,1830]" italics="true" pageId="15" pageNumber="68">Diagnosis.</emphasis>
— Habitus as in
<figureCitation id="133D2A12EA322933C827BA7618B7D868" box="[1118,1180,1806,1830]" captionStart="Figs" captionStartId="3.[113,157,2061,2083]" captionTargetBox="[222,1331,314,1975]" captionTargetId="figure-8@3.[195,1369,301,1985]" captionTargetPageId="3" captionText="Figs. 1–4. Habitus of: 1, Hydraena hendrichi, paratype; 2, H. paulmoritz, paratype; 3, H. singaporensis, paratype; 4, H. yangae, holotype, right foreleg (lateral view) depicted separately (photograph taken before right elytron got unhinged)." figureDoi="http://doi.org/10.5281/zenodo.5351600" httpUri="https://zenodo.org/record/5351600/files/figure.png" pageId="15" pageNumber="68">Fig. 4</figureCitation>
(
<typeStatus id="54BD8835EA322933C8D2BA761924D868" box="[1195,1295,1806,1830]" pageId="15" pageNumber="68" type="holotype">holotype</typeStatus>
). Body length:
<typeStatus id="54BD8835EA322933CF55BA561FA2D808" box="[812,905,1838,1862]" pageId="15" pageNumber="68" type="holotype">holotype</typeStatus>
<quantity id="4CFE9B72EA322933CFEABA561FD3D808" box="[915,1016,1838,1862]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="15" pageNumber="68" unit="mm" value="1.05">1.05 mm</quantity>
, females from
<collectingCountry id="F3117607EA322933C8D9BA561929D808" box="[1184,1282,1838,1862]" name="Malaysia" pageId="15" pageNumber="68">Malaysia</collectingCountry>
<quantity id="4CFE9B72EA322933C972BA56195BD808" box="[1291,1392,1838,1862]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="68" unit="mm" value="1.2">1.20 mm</quantity>
. This species is very closely related with
<taxonomicName id="4C064D14EA322933C8BFBA281FA9D8C8" authority="Jach &amp; Diaz" authorityName="Jach &amp; Diaz" authorityYear="1999" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B972EA85EA322933C8BFBA28195AD829" box="[1222,1393,1872,1895]" italics="true" pageId="15" pageNumber="68">H. okinawensis</emphasis>
Jäch &amp; Díaz
</taxonomicName>
and its allies (
<taxonomicName id="4C064D14EA322933C840BA081912D8C8" authority="Jach &amp; Diaz" authorityName="Jach &amp; Diaz" authorityYear="1999" box="[1081,1337,1902,1927]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B972EA85EA322933C840BA0818BCD8C9" box="[1081,1175,1904,1927]" italics="true" pageId="15" pageNumber="68">H. satoi</emphasis>
Jäch &amp; Díaz
</taxonomicName>
,
<taxonomicName id="4C064D14EA322933C930BA081F92D8E8" authority="Jach &amp; Diaz" authorityName="Jach &amp; Diaz" authorityYear="1999" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="iheya">
<emphasis id="B972EA85EA322933C930BA081987D8C9" box="[1353,1452,1904,1927]" italics="true" pageId="15" pageNumber="68">H. iheya</emphasis>
Jäch &amp; Díaz
</taxonomicName>
,
<taxonomicName id="4C064D14EA322933CFBEBAE818F4D8E8" authority="Jach &amp; Diaz" authorityName="Jach &amp; Diaz" authorityYear="1999" box="[967,1247,1934,1959]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="victoriae">
<emphasis id="B972EA85EA322933CFBEBAE81861D8E9" box="[967,1098,1936,1959]" italics="true" pageId="15" pageNumber="68">H. victoriae</emphasis>
Jäch &amp; Díaz
</taxonomicName>
) from the Ryukyu Archipelago (see Jäch &amp; Díaz, 1999). Colouration yellowish to reddish brown, labrum, clypeus and pronotum paler than frons and elytra. The
<typeStatus id="54BD8835EA322933C836BA961887D748" box="[1103,1196,2030,2054]" pageId="15" pageNumber="68" type="holotype">holotype</typeStatus>
of the new species can be distinguished from the Ryukyu species mainly by the lacking metaventral plaques, by the pro- and metatibiae and by the aedeagus.
</paragraph>
<caption id="DF79661FEA322933CC08BA891EA2D70F" ID-DOI="http://doi.org/10.5281/zenodo.5351620" ID-Zenodo-Dep="5351620" httpUri="https://zenodo.org/record/5351620/files/figure.png" pageId="15" pageNumber="68" startId="15.[113,148,2033,2055]" targetBox="[114,753,1130,2009]" targetPageId="15">
<paragraph id="8BB93697EA322933CC08BA891EA2D70F" blockId="15.[113,753,2033,2113]" pageId="15" pageNumber="68">
Fig. 16. Taban Valley, Bukit Timah Nature Reserve, Singapore. Three species,
<taxonomicName id="4C064D14EA322933CD79B5771D51D76D" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[256,378,2063,2083]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="hendrichi">
<emphasis id="B972EA85EA322933CD79B5771D51D76D" box="[256,378,2063,2083]" italics="true" pageId="15" pageNumber="68">H. hendrichi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C064D14EA322933CDFFB5771E2BD76D" authorityName="Orchymont" authorityYear="1932" box="[390,512,2062,2083]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B972EA85EA322933CDFFB5771DBCD76D" box="[390,407,2063,2083]" italics="true" pageId="15" pageNumber="68">H</emphasis>
.
<emphasis id="B972EA85EA322933CDDAB5771E2BD76D" box="[419,512,2063,2083]" italics="true" pageId="15" pageNumber="68">jacobsoni</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C064D14EA322933CE4FB5771E90D76D" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[566,699,2062,2083]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="paulmoritz">
<emphasis id="B972EA85EA322933CE4FB5771E6CD76D" box="[566,583,2063,2083]" italics="true" pageId="15" pageNumber="68">H</emphasis>
.
<emphasis id="B972EA85EA322933CE2AB5771E90D76D" box="[595,699,2063,2083]" italics="true" pageId="15" pageNumber="68">paulmoritz</emphasis>
</taxonomicName>
were collected from this site. (Photograph by: L. Hendrich).
</paragraph>
</caption>
<paragraph id="8BB93697EA2D292CCCF1BC651D2CDEDB" blockId="16.[136,775,285,405]" pageId="16" pageNumber="69">
Ventral face of protibia of
<typeStatus id="54BD8835EA2D292CCDD1BC651E2EDE7B" box="[424,517,285,309]" pageId="16" pageNumber="69" type="holotype">holotype</typeStatus>
(see
<figureCitation id="133D2A12EA2D292CCE39BC651EABDE7B" box="[576,640,285,309]" captionStart="Figs" captionStartId="3.[113,157,2061,2083]" captionTargetBox="[222,1331,314,1975]" captionTargetId="figure-8@3.[195,1369,301,1985]" captionTargetPageId="3" captionText="Figs. 1–4. Habitus of: 1, Hydraena hendrichi, paratype; 2, H. paulmoritz, paratype; 3, H. singaporensis, paratype; 4, H. yangae, holotype, right foreleg (lateral view) depicted separately (photograph taken before right elytron got unhinged)." figureDoi="http://doi.org/10.5281/zenodo.5351600" httpUri="https://zenodo.org/record/5351600/files/figure.png" pageId="16" pageNumber="69">Fig. 4</figureCitation>
) widened in apical half, very slightly tapering to apex; metatibia hardly noticeably curved, posterior margin very slightly crested subapically.
</paragraph>
<paragraph id="8BB93697EA2D292CCCF1BCC51E7DDD1B" blockId="16.[136,775,445,597]" pageId="16" pageNumber="69">
Male terminal sternite and spiculum (
<figureCitation id="133D2A12EA2D292CCE61BCC51E43DE9B" box="[536,616,445,469]" captionStart="Fig" captionStartId="6.[136,171,2060,2082]" captionTargetBox="[166,1446,190,2011]" captionTargetId="figure-8@6.[43,1493,170,2020]" captionTargetPageId="6" captionText="Fig. 7. Hydraena yangae: a, aedeagus in lateral view; b, aedeagus, dorsal view (setae of left paramere omitted in a, b); c, aedeagus, ventral view (setae of right paramere omitted); d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351606" httpUri="https://zenodo.org/record/5351606/files/figure.png" pageId="16" pageNumber="69">Fig. 7d</figureCitation>
): Quite similar to those of
<taxonomicName id="4C064D14EA2D292CCD78BCA61DA3DEBB" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[257,392,478,501]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="hendrichi">
<emphasis id="B972EA85EA2D292CCD78BCA61DA3DEBB" box="[257,392,478,501]" italics="true" pageId="16" pageNumber="69">H. hendrichi</emphasis>
</taxonomicName>
; sternite more regularly pentagonal than in the latter, more or less symmetrical, apex straight, base V-shaped, not firmly connected with spiculum, which is thinner and longer than in
<taxonomicName id="4C064D14EA2D292CCDBEBF461E64DD1B" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[455,591,574,597]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="hendrichi">
<emphasis id="B972EA85EA2D292CCDBEBF461E64DD1B" box="[455,591,574,597]" italics="true" pageId="16" pageNumber="69">H. hendrichi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB93697EA2D292CCCF1BF051EF6DB5B" blockId="16.[136,775,637,1045]" pageId="16" pageNumber="69">
Aedeagus (
<figureCitation id="133D2A12EA2D292CCC87BF051D4EDDDB" box="[254,357,637,661]" captionStart="Fig" captionStartId="6.[136,171,2060,2082]" captionTargetBox="[166,1446,190,2011]" captionTargetId="figure-8@6.[43,1493,170,2020]" captionTargetPageId="6" captionText="Fig. 7. Hydraena yangae: a, aedeagus in lateral view; b, aedeagus, dorsal view (setae of left paramere omitted in a, b); c, aedeagus, ventral view (setae of right paramere omitted); d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351606" httpUri="https://zenodo.org/record/5351606/files/figure.png" pageId="16" pageNumber="69">Fig. 7a–c</figureCitation>
): Remarkably similar to
<taxonomicName id="4C064D14EA2D292CCE1FBF061F2DDDDB" authorityName="Jach &amp; Diaz" authorityYear="1999" box="[614,774,638,661]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B972EA85EA2D292CCE1FBF061F2DDDDB" box="[614,774,638,661]" italics="true" pageId="16" pageNumber="69">H. okinawensis</emphasis>
</taxonomicName>
and its allies from the Ryukyu Archipelago (see Jäch &amp; Díaz, 1999). Main piece conspicuously hooked apically (ventral/ dorsal view), with one long dorsal seta; phallobase almost symmetrical, forming a closed ring. Distal lobe elongate, large, not clearly delimited from main piece, with numerous conspicuous spinules. Parameres articulately connected with main piece, inserted in basal half of aedeagus. Left paramere elongate, slender, with two groups of setae on apex and ventral margin in apical third, some of these being furcate; right paramere oval in dorsal view, rather narrow and acute in lateral view, with two groups of setae, one on apex and one along ventral margin in basal two thirds.
</paragraph>
<paragraph id="8BB93697EA2D292CCCF1B9451E34DBBB" blockId="16.[136,776,1085,1270]" pageId="16" pageNumber="69">
The aedeagus of
<taxonomicName id="4C064D14EA2D292CCD3DB9461D9FDB1B" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[324,436,1086,1109]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="yangae">
<emphasis id="B972EA85EA2D292CCD3DB9461D9FDB1B" box="[324,436,1086,1109]" italics="true" pageId="16" pageNumber="69">H. yangae</emphasis>
</taxonomicName>
can primarily be distinguished from the aedeagi of its relatives from Ryukyu by the main piece, which is wider in apical 0.3. Furthermore, it lacks a flagellum. However, the aedeagus of the
<typeStatus id="54BD8835EA2D292CCE1CB9E51EE9DBFB" box="[613,706,1181,1205]" pageId="16" pageNumber="69" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C064D14EA2D292CCE94B9E61CFEDB9B" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="yangae">
<emphasis id="B972EA85EA2D292CCE94B9E61CFEDB9B" italics="true" pageId="16" pageNumber="69">H. yangae</emphasis>
</taxonomicName>
is quite teneral and it cannot be excluded that the flagellum got lost during preparation.
</paragraph>
<paragraph id="8BB93697EA2D292CCF3ABDC51FFCDE7B" blockId="16.[835,1474,189,309]" pageId="16" pageNumber="69">
Gonocoxite (
<figureCitation id="133D2A12EA2D292CCFAFBDC51800DF9B" box="[982,1067,189,213]" captionStart="Fig" captionStartId="6.[136,171,2060,2082]" captionTargetBox="[166,1446,190,2011]" captionTargetId="figure-8@6.[43,1493,170,2020]" captionTargetPageId="6" captionText="Fig. 7. Hydraena yangae: a, aedeagus in lateral view; b, aedeagus, dorsal view (setae of left paramere omitted in a, b); c, aedeagus, ventral view (setae of right paramere omitted); d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351606" httpUri="https://zenodo.org/record/5351606/files/figure.png" pageId="16" pageNumber="69">Fig. 7e</figureCitation>
): Closely resembling that of the
<taxonomicName id="4C064D14EA2D292CC9D1BDC71FE3DFB8" authorityName="Jach &amp; Diaz" authorityYear="1999" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B972EA85EA2D292CC9D1BDC71FE3DFB8" italics="true" pageId="16" pageNumber="69">H. okinawensis</emphasis>
</taxonomicName>
and its allies from the Ryukyu Archipelago, especially
<taxonomicName id="4C064D14EA2D292CCFC2BD871830DE58" authorityName="Jach &amp; Diaz" authorityYear="1999" box="[955,1051,255,278]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="iheya">
<emphasis id="B972EA85EA2D292CCFC2BD871830DE58" box="[955,1051,255,278]" italics="true" pageId="16" pageNumber="69">H. iheya</emphasis>
</taxonomicName>
. It is, however, more slender than in these species.
</paragraph>
<paragraph id="8BB93697EA2D292CCF3ABC2519B7DE9B" blockId="16.[835,1474,349,469]" pageId="16" pageNumber="69">
Female tergite X (
<figureCitation id="133D2A12EA2D292CCF82BC25186FDE3B" box="[1019,1092,349,373]" captionStart="Fig" captionStartId="6.[136,171,2060,2082]" captionTargetBox="[166,1446,190,2011]" captionTargetId="figure-8@6.[43,1493,170,2020]" captionTargetPageId="6" captionText="Fig. 7. Hydraena yangae: a, aedeagus in lateral view; b, aedeagus, dorsal view (setae of left paramere omitted in a, b); c, aedeagus, ventral view (setae of right paramere omitted); d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351606" httpUri="https://zenodo.org/record/5351606/files/figure.png" pageId="16" pageNumber="69">Fig. 7f</figureCitation>
): Closely resembling
<taxonomicName id="4C064D14EA2D292CC95BBC2719E9DE38" authorityName="Jach &amp; Diaz" authorityYear="1999" box="[1314,1474,351,374]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B972EA85EA2D292CC95BBC2719E9DE38" box="[1314,1474,351,374]" italics="true" pageId="16" pageNumber="69">H. okinawensis</emphasis>
</taxonomicName>
and its allies from the Ryukyu Archipelago, especially by the subacuminate apex and the conspicuous subapical sclerotisation. However, the disc lacks squamose setae.
</paragraph>
<paragraph id="8BB93697EA2D292CCF3ABC851FB3DD7B" blockId="16.[835,1474,509,565]" pageId="16" pageNumber="69">
Spermatheca (
<figureCitation id="133D2A12EA2D292CCF9BBC851878DD5B" box="[994,1107,509,533]" captionStart="Fig" captionStartId="6.[136,171,2060,2082]" captionTargetBox="[166,1446,190,2011]" captionTargetId="figure-8@6.[43,1493,170,2020]" captionTargetPageId="6" captionText="Fig. 7. Hydraena yangae: a, aedeagus in lateral view; b, aedeagus, dorsal view (setae of left paramere omitted in a, b); c, aedeagus, ventral view (setae of right paramere omitted); d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351606" httpUri="https://zenodo.org/record/5351606/files/figure.png" pageId="16" pageNumber="69">
Fig.
<quantity id="4CFE9B72EA2D292CC861BC85181FDD5B" box="[1048,1076,509,533]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="16" pageNumber="69" unit="g" value="7.0">7g</quantity>
–h
</figureCitation>
): More or less as in the Ryukyu species.
</paragraph>
<paragraph id="8BB93697EA2D292CCF3ABF251863DC1B" blockId="16.[835,1475,605,853]" pageId="16" pageNumber="69">
Discussion: The females from
<collectingCountry id="F3117607EA2D292CC8BFBF251918DD3B" box="[1222,1331,605,629]" name="Malaysia" pageId="16" pageNumber="69">Malaysia</collectingCountry>
agree very well with the
<typeStatus id="54BD8835EA2D292CCF9CBF051862DDDB" box="[997,1097,637,661]" pageId="16" pageNumber="69" type="holotype">holotype</typeStatus>
, with which they share the wide mesosternal process and the lack of metaventral plaques. They are, however, more vividly coloured, because they are not teneral. They are very probably conspecific with the
<typeStatus id="54BD8835EA2D292CCF17BF851FE4DC5B" box="[878,975,765,789]" pageId="16" pageNumber="69" type="holotype">holotype</typeStatus>
, because it is unlikely, that two very closely related species occur in the same area. Bekok lies only
<quantity id="4CFE9B72EA2D292CC9E3BE651F4EDC1B" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="69" unit="km" value="100.0">100 km</quantity>
NNW of
<collectingCountry id="F3117607EA2D292CCFACBE45186EDC1B" box="[981,1093,829,853]" name="Singapore" pageId="16" pageNumber="69">Singapore</collectingCountry>
.
</paragraph>
<paragraph id="8BB93697EA2D292CCF3ABE041939DB7B" blockId="16.[835,1475,892,1077]" pageId="16" pageNumber="69">
<emphasis id="B972EA85EA2D292CCF3ABE041815DCDB" bold="true" box="[835,1086,892,917]" italics="true" pageId="16" pageNumber="69">Differential diagnosis.</emphasis>
— Externally, this species closely resembles
<taxonomicName id="4C064D14EA2D292CCFCCBEE71891DCF8" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[949,1210,927,950]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="singaporensis">
<emphasis id="B972EA85EA2D292CCFCCBEE71891DCF8" box="[949,1210,927,950]" italics="true" pageId="16" pageNumber="69">Hydraena singaporensis</emphasis>
</taxonomicName>
, especially in body size, body shape, colouration and lack of metaventral plaques. However, these two species are clearly distinguished by the width of the mesosternal process, by the male protibia, and by the male and female genital characters.
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA2D292CCF3AB924197FDBDB" pageId="16" pageNumber="69" type="distribution">
<paragraph id="8BB93697EA2D292CCF3AB924197FDBDB" blockId="16.[835,1474,1116,1173]" pageId="16" pageNumber="69">
<emphasis id="B972EA85EA2D292CCF3AB9241FE4DB3B" bold="true" box="[835,975,1116,1141]" italics="true" pageId="16" pageNumber="69">Distribution.</emphasis>
—
<collectingCountry id="F3117607EA2D292CCF8FB9251849DB3B" box="[1014,1122,1117,1141]" name="Singapore" pageId="16" pageNumber="69">Singapore</collectingCountry>
(Rifle Range Road, east of Bukit Timah), see
<figureCitation id="133D2A12EA2D292CCFB2B9051830DBDB" box="[971,1051,1149,1173]" captionStart="Fig" captionStartId="14.[137,172,2090,2112]" captionTargetBox="[137,1470,1208,2063]" captionTargetId="figure-8@14.[137,1478,1202,2067]" captionTargetPageId="14" captionText="Fig. 15. Geographical map of Singapore, showing distribution of Hydraenidae." figureDoi="http://doi.org/10.5281/zenodo.5351618" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="16" pageNumber="69">Fig. 15</figureCitation>
,
<collectingCountry id="F3117607EA2D292CC851B90518A1DBDB" box="[1064,1162,1149,1173]" name="Malaysia" pageId="16" pageNumber="69">Malaysia</collectingCountry>
(
<collectingRegion id="49C2F875EA2D292CC8E3B90518F3DBDB" box="[1178,1240,1149,1173]" country="Malaysia" name="Johor" pageId="16" pageNumber="69">Johor</collectingRegion>
,
<collectingRegion id="49C2F875EA2D292CC89CB9051962DBDB" box="[1253,1353,1149,1173]" country="Malaysia" name="Selangor" pageId="16" pageNumber="69">Selangor</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA2D292DCF3AB9C41D0DDFBB" lastPageId="17" lastPageNumber="70" pageId="16" pageNumber="69" type="etymology">
<paragraph id="8BB93697EA2D292CCF3AB9C419E9DBBB" blockId="16.[835,1475,1212,1270]" pageId="16" pageNumber="69">
<emphasis id="B972EA85EA2D292CCF3AB9C41F95DB9B" bold="true" box="[835,958,1212,1237]" italics="true" pageId="16" pageNumber="69">Etymology.</emphasis>
— Named for Chang-Man Yang, former curator of
<taxonomicName id="4C064D14EA2D292CCF1BB9A61F85DBBB" authorityName="Linnaeus" authorityYear="1758" box="[866,942,1246,1269]" class="Insecta" kingdom="Animalia" pageId="16" pageNumber="69" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
at the ZRC. She is doing a great job, even after
</paragraph>
<caption id="DF79661FEA2D292CCCF1B5751F48D70E" ID-DOI="http://doi.org/10.5281/zenodo.5351622" ID-Zenodo-Dep="5351622" httpUri="https://zenodo.org/record/5351622/files/figure.png" pageId="16" pageNumber="69" startId="16.[136,172,2061,2083]" targetBox="[279,1324,1376,2039]" targetPageId="16">
<paragraph id="8BB93697EA2D292CCCF1B5751F48D70E" blockId="16.[136,1474,2061,2112]" pageId="16" pageNumber="69">
Fig. 17. Nee Soon Swamp Forest, Singapore; C.M. Yang on right. Four species,
<taxonomicName id="4C064D14EA2D292CCFB7B5761815D76C" authorityName="Orchymont" authorityYear="1932" box="[974,1086,2062,2082]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="formula">
<emphasis id="B972EA85EA2D292CCFB7B5761815D76C" box="[974,1086,2062,2082]" italics="true" pageId="16" pageNumber="69">H. formula</emphasis>
</taxonomicName>
,
<taxonomicName id="4C064D14EA2D292CC837B57618E4D76C" authorityName="Orchymont" authorityYear="1932" box="[1102,1231,2061,2082]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B972EA85EA2D292CC837B5761874D76C" box="[1102,1119,2062,2082]" italics="true" pageId="16" pageNumber="69">H</emphasis>
.
<emphasis id="B972EA85EA2D292CC816B57618E4D76C" box="[1135,1231,2062,2082]" italics="true" pageId="16" pageNumber="69">jacobsoni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C064D14EA2D292CC899B5761947D76C" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[1248,1388,2061,2082]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="paulmoritz">
<emphasis id="B972EA85EA2D292CC899B57618DAD76C" box="[1248,1265,2062,2082]" italics="true" pageId="16" pageNumber="69">H</emphasis>
.
<emphasis id="B972EA85EA2D292CC978B5761947D76C" box="[1281,1388,2062,2082]" italics="true" pageId="16" pageNumber="69">paulmoritz</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C064D14EA2D292CC9D3B5761D27D771" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="singaporensis">
<emphasis id="B972EA85EA2D292CC9D3B5761D27D771" italics="true" pageId="16" pageNumber="69">H. singaporensis</emphasis>
</taxonomicName>
were collected from this site. (Photograph by: L. Hendrich).
</paragraph>
</caption>
<paragraph id="8BB93697EA2C292DCC08BDC51D0DDFBB" blockId="17.[113,752,189,245]" pageId="17" pageNumber="70">
retirement. The
<typeStatus id="54BD8835EA2C292DCD67BDC51D50DF9B" box="[286,379,189,213]" pageId="17" pageNumber="70" type="holotype">holotype</typeStatus>
of this obviously rare species was collected by her.
</paragraph>
</subSubSection>
</treatment>
</document>
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