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<document id="C4500901B9C234FECDFE4631604A0AB7" ID-CLB-Dataset="299055" ID-DOI="10.5281/zenodo.11067355" ID-GBIF-Dataset="67ec2de4-1816-4969-8132-f6796d3afa06" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067355" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052343267" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2021" docId="03AF0B0CFFF8FF9FFF9FFF14A2F7F953" docLanguage="en" docName="ProcCalifAcadSci.67.4.85-182.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (4)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Broscosoma gongshanense Kavanaugh and Liang 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="127" masterDocId="FF967374FFDFFFB5FFE4FFCFA106FFEE" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." masterLastPageNumber="182" masterPageNumber="85" pageNumber="124" updateTime="1719540814554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C3FB24CA21934E2E1E6C71A14CB89B75">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae).</mods:title>
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<mods:name id="CFD22657163F76F1672FFE2EF042D7A6" type="personal">
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<mods:namePart id="DD86561DDD535B99A5C8B09E88D879B8">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="E575809394D700326A9A95480441E604">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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</mods:role>
<mods:namePart id="D1E84B9455773CBDBD2879B319D54B64">Liang, Hongbin</mods:namePart>
<mods:affiliation id="4A0CBE4A0383EE898C4207530B82D345">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title id="861C7B28AD9ECC63D176EAD8C61E696B">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="77995FC70BEDC73C3663B24C93A72A31">
<mods:date id="78AAC79B6BD3DE26E318E4EDC5154C1D">2021</mods:date>
<mods:detail id="C306AF3181AE305045501BE1B3089DFD" type="pubDate">
<mods:number id="17B5AE28412D95B8D3E83E3F8986872F">2021-09-30</mods:number>
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<mods:number id="621703753A3C4393D2473B337B206C15">67</mods:number>
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<mods:number id="1DC12A4FAF19017ADB6037FE02028CA7">4</mods:number>
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<treatment id="03AF0B0CFFF8FF9FFF9FFF14A2F7F953" ID-DOI="http://doi.org/10.5281/zenodo.11067413" ID-GBIF-Taxon="233814471" ID-Zenodo-Dep="11067413" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF8FF9FFF9FFF14A2F7F953" httpUri="http://treatment.plazi.org/id/03AF0B0CFFF8FF9FFF9FFF14A2F7F953" lastPageId="42" lastPageNumber="127" pageId="39" pageNumber="124">
<subSubSection id="C31CE991FFF8FF92FF9FFF14A22BFF1D" box="[123,813,219,243]" pageId="39" pageNumber="124" type="nomenclature">
<paragraph id="8BB9BA1AFFF8FF92FF9FFF14A22BFF1D" blockId="39.[123,813,219,273]" box="[123,813,219,243]" pageId="39" pageNumber="124">
<heading id="D0F10D76FFF8FF92FF9FFF14A22BFF1D" box="[123,813,219,243]" fontSize="10" level="2" pageId="39" pageNumber="124" reason="7">
<emphasis id="B9726608FFF8FF92FF9FFF14A22BFF1D" bold="true" box="[123,813,219,243]" pageId="39" pageNumber="124">
7.
<taxonomicName id="4C06C199FFF8FF92FF72FF14A3CFFF1D" authority="Kavanaugh and Liang" authorityName="Kavanaugh and Liang" authorityYear="2021" box="[150,713,219,243]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="124" phylum="Arthropoda" rank="species" species="gongshanense" status="sp. nov.">
<emphasis id="B9726608FFF8FF92FF72FF14A0C7FF1D" bold="true" box="[150,449,219,243]" italics="true" pageId="39" pageNumber="124">Broscosoma gongshanense</emphasis>
Kavanaugh and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A241DB73FFF8FF92FD33FF14A22BFF1D" box="[727,813,219,243]" pageId="39" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF8FF92FF9FFF34A09BFEFF" box="[123,413,251,273]" pageId="39" pageNumber="124" type="description">
<paragraph id="8BB9BA1AFFF8FF92FF9FFF34A09BFEFF" blockId="39.[123,813,219,273]" box="[123,413,251,273]" pageId="39" pageNumber="124">
<figureCitation id="133DA69FFFF8FF92FF9FFF34A1E6FEFF" box="[123,224,251,273]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="39" pageNumber="124">Figures 7e</figureCitation>
,
<figureCitation id="133DA69FFFF8FF92FF09FF33A003FEFF" box="[237,261,252,273]" captionStart="FIGURE" captionStartId="40.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1566]" captionTargetId="figure-56@40.[120,1176,216,1566]" captionTargetPageId="40" captionText="FIGURE 22. Broscosoma gongshanense sp. nov. a. Habitus (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067397" httpUri="https://zenodo.org/record/11067397/files/figure.png" pageId="39" pageNumber="124">22</figureCitation>
,
<figureCitation id="133DA69FFFF8FF92FEF5FF33A02FFEFF" box="[273,297,252,273]" captionStart="FIGURE" captionStartId="41.[171,183,967,986]" captionTargetBox="[157,1154,219,934]" captionTargetId="figure-1587@41.[121,1176,216,953]" captionTargetPageId="41" captionText="FIGURE 23. Male genitalia, Broscosoma gongshanense sp. nov. (CASENT1026429; 2.0 km S of Heipu Yakou, Cikai Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Medi- an lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067399" httpUri="https://zenodo.org/record/11067399/files/figure.png" pageId="39" pageNumber="124">23</figureCitation>
,
<figureCitation id="133DA69FFFF8FF92FED2FF33A05FFEFF" box="[310,345,252,273]" captionStart="FIGURE" captionStartId="82.[171,183,1585,1604]" captionTargetBox="[128,1169,216,1568]" captionTargetId="figure-57@82.[128,1169,216,1568]" captionTargetPageId="82" captionText="FIGURE 42. Photographs of habitats for Broscosoma species in the Gaoligong Shan region. a. Dulong/Gongshan Yakou area, 3300-3680m, 27.697º/098.454º, Gongshan County, Yunnan (habitat for Broscosoma bicoloratum sp. nov. and B. gongshanense sp. nov.); b. Southeastern slope of Heipu Yakou, 3365m, 27.770º/098.447º, Cikai Township, Gongshan County, Yunnan (habitat for Broscosoma bicoloratum sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067443" httpUri="https://zenodo.org/record/11067443/files/figure.png" pageId="39" pageNumber="124">42a</figureCitation>
,
<figureCitation id="133DA69FFFF8FF92FE81FF33A09BFEFF" box="[357,413,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="88.[171,183,995,1014]" captionStartId-1="88.[184,196,1682,1701]" captionStartId-2="89.[171,183,1461,1480]" captionTargetBox-0="[130,1164,233,969]" captionTargetBox-1="[130,1162,1063,1660]" captionTargetBox-2="[140,1154,249,1428]" captionTargetId-0="figure-57@88.[120,1176,215,989]" captionTargetId-1="figure-332@88.[120,1176,1054,1668]" captionTargetId-2="figure-68@89.[120,1176,216,1448]" captionTargetPageId-0="88" captionTargetPageId-1="88" captionTargetPageId-2="89" captionText-0="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." captionText-2="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi-0="http://doi.org/10.5281/zenodo.11067455" figureDoi-1="http://doi.org/10.5281/zenodo.11067457" figureDoi-2="http://doi.org/10.5281/zenodo.11067459" httpUri-0="https://zenodo.org/record/11067455/files/figure.png" httpUri-1="https://zenodo.org/record/11067457/files/figure.png" httpUri-2="https://zenodo.org/record/11067459/files/figure.png" pageId="39" pageNumber="124">48-50</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF8FF92FF4FFEE1A0F0FC81" pageId="39" pageNumber="124" type="materials_examined">
<paragraph id="8BB9BA1AFFF8FF92FF4FFEE1A379FD22" blockId="39.[123,1174,301,1726]" pageId="39" pageNumber="124">
<emphasis id="B9726608FFF8FF92FF4FFEE1A055FEA8" bold="true" box="[171,339,301,326]" pageId="39" pageNumber="124">Type material.</emphasis>
<materialsCitation id="3B6EB047FFF8FF92FEB8FEE1A34AFE29" ID-GBIF-Occurrence="4899905410" collectingDate="2006-08-12" collectionCode="IOZ" collectorName="D. H. Kavanaugh &amp; J. A. Miller" country="China" county="Gongshan County" elevation="3350" latitude="27.75442" location="Pula valley" longLatPrecision="1" longitude="98.45639" municipality="Cikai Township" pageId="39" pageNumber="124" specimenCode="CASENT 1026430" specimenCount="1" stateProvince="Yunnan Province" typeStatus="holotype">
<typeStatus id="54BD04B8FFF8FF92FEB8FEE1A0B9FEA8" box="[348,447,302,326]" pageId="39" pageNumber="124" type="holotype">Holotype</typeStatus>
, a
<specimenCount id="9D007193FFF8FF92FE00FEE1A31EFEA8" box="[484,536,302,326]" pageId="39" pageNumber="124" type="male">male</specimenCount>
, in
<collectionCode id="ED1722DFFFF8FF92FDA1FEE1A377FEA8" box="[581,625,302,326]" pageId="39" pageNumber="124">IOZ</collectionCode>
, labeled: “
<specimenCode id="DBA01261FFF8FF92FD0FFEE1A2BEFEA8" box="[747,952,302,326]" pageId="39" pageNumber="124">CASENT 1026430</specimenCode>
”/ “
<collectingCountry id="F311FA8AFFF8FF92FC05FEE1A533FEA8" box="[993,1077,302,326]" name="China" pageId="39" pageNumber="124">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF8FF92FBA0FEE1A1DCFE88" country="China" name="Yunnan" pageId="39" pageNumber="124">Yunnan Province</collectingRegion>
,
<collectingCounty id="62D8C296FFF8FF92FF0CFE81A0AFFE88" box="[232,425,334,358]" pageId="39" pageNumber="124">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF8FF92FE5CFE81A364FE88" box="[440,610,334,358]" pageId="39" pageNumber="124">Cikai Township</collectingMunicipality>
, 2.0 airkm S Heipu Yakou in cirque at head of
<location id="8ED9ECC1FFF8FF92FB82FE81A1BDFE69" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF8FF9FFF9FFF14A2F7F953:8ED9ECC1FFF8FF92FB82FE81A1BDFE69" country="China" county="Gongshan County" latitude="27.75442" longLatPrecision="1" longitude="98.45639" municipality="Cikai Township" name="Pula valley" pageId="39" pageNumber="124" stateProvince="Yunnan Province">Pula valley</location>
,
<quantity id="4CFE17FFFFF8FF92FF36FEA0A01AFE68" box="[210,284,367,391]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="39" pageNumber="124" unit="m" value="3350.0">
<elevation id="002B5D29FFF8FF92FF36FEA0A01AFE68" box="[210,284,367,391]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="39" pageNumber="124" unit="m" value="3350.0">3350m</elevation>
</quantity>
” / “
<geoCoordinate id="EE32DCDDFFF8FF92FEB9FEA0A0DAFE69" box="[349,476,367,391]" degrees="27. 75442" direction="north" orientation="latitude" pageId="39" pageNumber="124" precision="1" value="27.75442">N27.75442º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF8FF92FE00FEA0A368FE69" box="[484,622,367,391]" degrees="098. 45639" direction="east" orientation="longitude" pageId="39" pageNumber="124" precision="1" value="98.45639">E098.45639º</geoCoordinate>
,
<date id="FFB89CDAFFF8FF92FD62FEA0A245FE69" box="[646,835,367,391]" pageId="39" pageNumber="124" value="2006-08-12">
<collectingDate id="EFFC6532FFF8FF92FD62FEA0A245FE69" box="[646,835,367,391]" pageId="39" pageNumber="124" value="2006-08-12">12 August 2006</collectingDate>
</date>
, Stop #DHK2006-070,
<collectorName id="26F3DFCCFFF8FF92FB85FEA0A1F2FE49" pageId="39" pageNumber="124">D.H. Kavanaugh</collectorName>
,
<collectorName id="26F3DFCCFFF8FF92FEECFE5FA088FE49" box="[264,398,399,424]" pageId="39" pageNumber="124">J. A. Miller</collectorName>
collectors”/ “HOLOTYPE
<taxonomicName id="4C06C199FFF8FF92FD23FE5FA1BFFE26" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="124" phylum="Arthropoda" rank="species" species="gongshanense" status="sp. nov.">
<emphasis id="B9726608FFF8FF92FD23FE5FA2E8FE49" box="[711,1006,399,423]" italics="true" pageId="39" pageNumber="124">Broscosoma gongshanense</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFF8FF92FF24FE7FA012FE29" box="[192,276,432,455]" pageId="39" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [red label]
</materialsCitation>
.
<materialsCitation id="3B6EB047FFF8FF92FDBEFE7FA2E3FE07" ID-GBIF-Occurrence="4899905461" collectingDate="2000-07-16" collectingDateMax="2000-07-17" collectingDateMin="2000-07-16" collectionCode="CAS, IOZ" collectorName="C. E. Griswold &amp; H. - B. Liang &amp; D. Ubick &amp; D. - Z. Dong" country="China" county="Gaoligong Shan" elevation="3490" latitude="27.69655" location="Nujiang State Nature Reserve" longLatPrecision="1" longitude="98.45389" municipality="Nujiang Prefecture" pageId="39" pageNumber="124" specimenCode="CASENT 1026439" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54BD04B8FFF8FF92FDBEFE7FA3C4FE26" box="[602,706,432,456]" pageId="39" pageNumber="124" type="paratype">Paratypes</typeStatus>
(a total of five):
<specimenCount id="9D007193FFF8FF92FC98FE7FA2D8FE26" box="[892,990,432,456]" count="1" pageId="39" pageNumber="124" type="male">one male</specimenCount>
and
<specimenCount id="9D007193FFF8FF92FBF7FE7FA593FE26" box="[1043,1173,432,456]" count="2" pageId="39" pageNumber="124" type="female">two females</specimenCount>
(
<collectionCode id="ED1722DFFFF8FF92FF60FE1EA1BEFE07" box="[132,184,465,489]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="124" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1722DFFFF8FF92FF2EFE1EA1F0FE07" box="[202,246,465,489]" pageId="39" pageNumber="124">IOZ</collectionCode>
) labeled: same as holotype except first label “
<specimenCode id="DBA01261FFF8FF92FCEDFE1EA2DFFE06" box="[777,985,464,489]" pageId="39" pageNumber="124">CASENT 1026439</specimenCode>
</materialsCitation>
<materialsCitation id="3B6EB047FFF8FF92FC14FE1FA005FDE7" ID-GBIF-Occurrence="4899905499" collectingDate="2000-07-16" collectingDateMax="2000-07-17" collectingDateMin="2000-07-16" collectionCode="CAS, IOZ" collectorName="C. E. Griswold &amp; H. - B. Liang &amp; D. Ubick &amp; D. - Z. Dong" country="China" county="Gaoligong Shan" elevation="3490" latitude="27.69655" location="Nujiang State Nature Reserve" longLatPrecision="1" longitude="98.45389" municipality="Nujiang Prefecture" pageId="39" pageNumber="124" specimenCode="CASENT 1026439" specimenCount="1" specimenCount-female="3" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
and “
<specimenCode id="DBA01261FFF8FF92FBCAFE1EA1DEFDE7" pageId="39" pageNumber="124">CASENT 1026439</specimenCode>
” to
</materialsCitation>
<materialsCitation id="3B6EB047FFF8FF92FEE8FE3EA38CFDE7" ID-GBIF-Occurrence="4899905441" box="[268,650,497,521]" collectingDate="2000-07-16" collectingDateMax="2000-07-17" collectingDateMin="2000-07-16" collectionCode="CAS, IOZ" collectorName="C. E. Griswold &amp; H. - B. Liang &amp; D. Ubick &amp; D. - Z. Dong" country="China" county="Gaoligong Shan" elevation="3490" latitude="27.69655" location="Nujiang State Nature Reserve" longLatPrecision="1" longitude="98.45389" municipality="Nujiang Prefecture" pageId="39" pageNumber="124" specimenCode="CASENT 1026439" specimenCount="1" specimenCount-female="3" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCode id="DBA01261FFF8FF92FEFCFE3EA0E0FDE7" box="[280,486,497,521]" pageId="39" pageNumber="124">CASENT 1026439</specimenCode>
”, respectively;
</materialsCitation>
<materialsCitation id="3B6EB047FFF8FF92FD70FE3DA0A4FDC4" ID-GBIF-Occurrence="4899905364" collectingDate="2000-07-16" collectingDateMax="2000-07-17" collectingDateMin="2000-07-16" collectionCode="CAS, IOZ" collectorName="C. E. Griswold &amp; H. - B. Liang &amp; D. Ubick &amp; D. - Z. Dong" country="China" county="Gaoligong Shan" elevation="3490" latitude="27.69655" location="Nujiang State Nature Reserve" longLatPrecision="1" longitude="98.45389" municipality="Nujiang Prefecture" pageId="39" pageNumber="124" specimenCode="CASENT 1008139" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF8FF92FD70FE3DA3FFFDE7" box="[660,761,497,521]" count="1" pageId="39" pageNumber="124" type="male">one male</specimenCount>
and
<specimenCount id="9D007193FFF8FF92FCD6FE3DA2AAFDE7" box="[818,940,497,521]" count="1" pageId="39" pageNumber="124" type="female">one female</specimenCount>
(
<collectionCode id="ED1722DFFFF8FF92FC5AFE3EA2F4FDE7" box="[958,1010,497,521]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="124" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1722DFFFF8FF92FBE6FE3EA528FDE7" box="[1026,1070,497,521]" pageId="39" pageNumber="124">IOZ</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF8FF92FF63FDDDA05BFDC4" box="[135,349,530,554]" pageId="39" pageNumber="124">CASENT 1008139</specimenCode>
” and
</materialsCitation>
<materialsCitation id="3B6EB047FFF8FF92FE56FDDDA56FFD65" ID-GBIF-Occurrence="4899905549" collectingDate="2000-07-16" collectingDateMax="2000-07-17" collectingDateMin="2000-07-16" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; C. E. Griswold &amp; H. - B. Liang &amp; D. Ubick &amp; D. - Z. Dong" country="China" county="Gaoligong Shan" elevation="3490" latitude="27.69655" location="Nujiang State Nature Reserve" longLatPrecision="1" longitude="98.45389" municipality="Nujiang Prefecture" pageId="39" pageNumber="124" specimenCode="CASENT 1006244" specimenCount="1" specimenCount-female="3" specimenCount-male="2" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCode id="DBA01261FFF8FF92FE5AFDDDA393FDC7" box="[446,661,529,554]" pageId="39" pageNumber="124">CASENT 1006244</specimenCode>
”, respectively/ “
<collectingCountry id="F311FA8AFFF8FF92FC84FDDDA2B3FDC4" box="[864,949,530,554]" name="China" pageId="39" pageNumber="124">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF8FF92FC28FDDDA589FDC4" box="[972,1167,530,554]" country="China" name="Yunnan" pageId="39" pageNumber="124">Yunnan Province</collectingRegion>
,
<collectingCounty id="62D8C296FFF8FF92FF9FFDFDA020FDA4" box="[123,294,562,586]" pageId="39" pageNumber="124">Gaoligong Shan</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF8FF92FED7FDFDA0FAFDA4" box="[307,508,562,586]" pageId="39" pageNumber="124">Nujiang Prefecture</collectingMunicipality>
,
<location id="8ED9ECC1FFF8FF92FDECFDFDA245FDA4" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF8FF9FFF9FFF14A2F7F953:8ED9ECC1FFF8FF92FDECFDFDA245FDA4" box="[520,835,562,586]" country="China" county="Gaoligong Shan" latitude="27.69655" longLatPrecision="1" longitude="98.45389" municipality="Nujiang Prefecture" name="Nujiang State Nature Reserve" pageId="39" pageNumber="124" stateProvince="Yunnan Province">Nujiang State Nature Reserve</location>
,
<location id="8ED9ECC1FFF8FF92FCB4FDFDA2A6FDA4" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF8FF9FFF9FFF14A2F7F953:8ED9ECC1FFF8FF92FCB4FDFDA2A6FDA4" box="[848,928,562,586]" country="China" county="Gaoligong Shan" latitude="27.69655" longLatPrecision="1" longitude="98.45389" municipality="Nujiang Prefecture" name="Dulong" pageId="39" pageNumber="124" stateProvince="Yunnan Province">Dulong</location>
/
<location id="8ED9ECC1FFF8FF92FC43FDFDA588FDA4" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF8FF9FFF9FFF14A2F7F953:8ED9ECC1FFF8FF92FC43FDFDA588FDA4" box="[935,1166,562,586]" country="China" county="Gaoligong Shan" latitude="27.69655" longLatPrecision="1" longitude="98.45389" municipality="Nujiang Prefecture" name="Gongshan Yakou area" pageId="39" pageNumber="124" stateProvince="Yunnan Province">Gongshan Yakou area</location>
, 21 airkm
<locationDeviation id="9A0BA338FFF8FF92FF06FD9CA08AFD85" box="[226,396,595,619]" location="Gongshan" pageId="39" pageNumber="124">W of Gongshan</locationDeviation>
” / “
<geoCoordinate id="EE32DCDDFFF8FF92FE5EFD9CA33FFD85" box="[442,569,595,619]" degrees="27. 69655" direction="north" orientation="latitude" pageId="39" pageNumber="124" precision="1" value="27.69655">N27.69655º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF8FF92FDA5FD9CA3CCFD85" box="[577,714,595,619]" degrees="098. 45389" direction="east" orientation="longitude" pageId="39" pageNumber="124" precision="1" value="98.45389">E098.45389º</geoCoordinate>
,
<quantity id="4CFE17FFFFF8FF92FD3DFD9CA267FD84" box="[729,865,595,619]" metricMagnitude="3" metricUnit="m" metricValue="3.4899999999999998" metricValueMax="3.6799999999999997" metricValueMin="3.3" pageId="39" pageNumber="124" unit="m" value="3490.0" valueMax="3680.0" valueMin="3300.0">
<elevation id="002B5D29FFF8FF92FD3DFD9CA267FD84" box="[729,865,595,619]" metricMagnitude="3" metricUnit="m" metricValue="3.4899999999999998" metricValueMax="3.6799999999999997" metricValueMin="3.3" pageId="39" pageNumber="124" unit="m" value="3490.0" valueMax="3680.0" valueMin="3300.0">3300-3680m</elevation>
</quantity>
,
<date id="FFB89CDAFFF8FF92FC8BFD9CA518FD85" box="[879,1054,595,619]" pageId="39" pageNumber="124" value="2000-07-16" valueMax="2000-07-17" valueMin="2000-07-16">
<collectingDate id="EFFC6532FFF8FF92FC8BFD9CA518FD85" box="[879,1054,595,619]" pageId="39" pageNumber="124" value="2000-07-16" valueMax="2000-07-17" valueMin="2000-07-16">16-17 July 2000</collectingDate>
</date>
, Stop #00- 24C,
<collectorName id="26F3DFCCFFF8FF92FF5EFDBBA074FD65" box="[186,370,627,651]" pageId="39" pageNumber="124">D.H. Kavanaugh</collectorName>
,
<collectorName id="26F3DFCCFFF8FF92FE61FDBCA322FD65" box="[389,548,627,651]" pageId="39" pageNumber="124">C.E. Griswold</collectorName>
,
<collectorName id="26F3DFCCFFF8FF92FDD2FDBBA3BAFD65" box="[566,700,627,651]" pageId="39" pageNumber="124">H.-B. Liang</collectorName>
,
<collectorName id="26F3DFCCFFF8FF92FD2AFDBBA233FD65" box="[718,821,627,651]" pageId="39" pageNumber="124">D. Ubick</collectorName>
&amp;
<collectorName id="26F3DFCCFFF8FF92FC86FDBBA2E5FD62" box="[866,995,628,652]" pageId="39" pageNumber="124">D.-Z. Dong</collectorName>
collectors”.
</materialsCitation>
All paratypes also bear the following label: “PARATYPE
<taxonomicName id="4C06C199FFF8FF92FD34FD5AA1BFFD22" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="124" phylum="Arthropoda" rank="species" species="gongshanense" status="sp. nov.">
<emphasis id="B9726608FFF8FF92FD34FD5AA2F2FD42" box="[720,1012,660,684]" italics="true" pageId="39" pageNumber="124">Broscosoma gongshanense</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A241DB73FFF8FF92FF23FD7AA01CFD22" box="[199,282,693,716]" pageId="39" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [yellow label].
</paragraph>
<paragraph id="8BB9BA1AFFF8FF92FF4FFD1AA1A8FCE3" blockId="39.[123,1174,301,1726]" pageId="39" pageNumber="124">
<emphasis id="B9726608FFF8FF92FF4FFD1AA047FD03" bold="true" box="[171,321,725,749]" pageId="39" pageNumber="124">Type locality</emphasis>
. China, Yunnan Province, Gongshan County, Cikai Township, Heipu Yakou area.
</paragraph>
<paragraph id="8BB9BA1AFFF8FF92FF4FFCD9A0F0FC81" blockId="39.[123,1174,301,1726]" pageId="39" pageNumber="124">
<emphasis id="B9726608FFF8FF92FF4FFCD9A0E5FCC0" bold="true" box="[171,483,790,814]" pageId="39" pageNumber="124">Derivation of species name.</emphasis>
The species epithet,
<taxonomicName id="4C06C199FFF8FF92FD21FCD9A25AFCC0" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[709,860,790,814]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="124" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFF8FF92FD21FCD9A25AFCC0" box="[709,860,790,814]" italics="true" pageId="39" pageNumber="124">gongshanense</emphasis>
</taxonomicName>
, is an adjective derived from the name of the county, Gongshan, in which all members of the
<typeStatus id="54BD04B8FFF8FF92FCC3FCF8A253FCA1" box="[807,853,823,847]" pageId="39" pageNumber="124">type</typeStatus>
series were collected, and the Latin suffix,
<emphasis id="B9726608FFF8FF92FEE2FC97A042FC81" box="[262,324,856,879]" italics="true" pageId="39" pageNumber="124">-ensis</emphasis>
, denoting place.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF8FF92FF4FFCB8A06DFB5A" pageId="39" pageNumber="124" type="diagnosis">
<paragraph id="8BB9BA1AFFF8FF92FF4FFCB8A06DFB5A" blockId="39.[123,1174,301,1726]" pageId="39" pageNumber="124">
<emphasis id="B9726608FFF8FF92FF4FFCB8A019FC7E" bold="true" box="[171,287,887,912]" pageId="39" pageNumber="124">Diagnosis.</emphasis>
Adults of this species (
<figureCitation id="133DA69FFFF8FF92FDC5FCB7A37CFC61" box="[545,634,888,912]" captionStart="FIGURE" captionStartId="40.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1566]" captionTargetId="figure-56@40.[120,1176,216,1566]" captionTargetPageId="40" captionText="FIGURE 22. Broscosoma gongshanense sp. nov. a. Habitus (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067397" httpUri="https://zenodo.org/record/11067397/files/figure.png" pageId="39" pageNumber="124">Fig. 22a</figureCitation>
) can be distinguished from those of other species in the region by the following combination of character states: size moderate, BL = 9.5 to 9.
<quantity id="4CFE17FFFFF8FF92FBB4FC57A58BFC5E" box="[1104,1165,920,944]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="124" unit="mm" value="9.0">9 mm</quantity>
; head and pronotum black to piceous, without a trace of metallic reflection; head with tempora straight or only slightly convex behind eyes; genal ridge evident, extended from base of head to ventral margin of eye (as in
<figureCitation id="133DA69FFFF8FF92FE54FC35A30FFBFF" box="[432,521,1018,1042]" captionStart="FIGURE" captionStartId="13.[171,183,1441,1460]" captionTargetBox="[404,932,216,1422]" captionTargetId="figure-66@13.[404,932,216,1422]" captionTargetPageId="13" captionText="FIGURE 10. Head, Broscosoma species, lateral aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote endpoints of genal ridge." figureDoi="http://doi.org/10.5281/zenodo.11067371" httpUri="https://zenodo.org/record/11067371/files/figure.png" pageId="39" pageNumber="124">Fig. 10a</figureCitation>
); antennomeres 3 and 4 glabrous except for apical whorl of setae; pronotum anterior to sub-basal constriction slightly longitudinally ovoid in dorsal view (
<figureCitation id="133DA69FFFF8FF92FB8FFBD5A192FBBC" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="39" pageNumber="124">Fig. 7e</figureCitation>
), lateral margination absent or developed only posterior to sub-basal constriction, basolateral setae absent; elytral silhouette ovoid, with humeri indistinct, elytral microsculpture effaced; metatrochanter asetose; male protarsi with tarsomeres 1 to 3 with ventral pads of adhesive setae; male genitalia as in
<figureCitation id="133DA69FFFF8FF92FEF3FB52A062FB5B" box="[279,356,1181,1205]" captionStart="FIGURE" captionStartId="41.[171,183,967,986]" captionTargetBox="[157,1154,219,934]" captionTargetId="figure-1587@41.[121,1176,216,953]" captionTargetPageId="41" captionText="FIGURE 23. Male genitalia, Broscosoma gongshanense sp. nov. (CASENT1026429; 2.0 km S of Heipu Yakou, Cikai Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Medi- an lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067399" httpUri="https://zenodo.org/record/11067399/files/figure.png" pageId="39" pageNumber="124">Fig. 23</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF8FF9FFF4FFB72A0E3FDDB" lastPageId="42" lastPageNumber="127" pageId="39" pageNumber="124" type="description">
<paragraph id="8BB9BA1AFFF8FF92FF4FFB72A3BFFA99" blockId="39.[123,1174,301,1726]" pageId="39" pageNumber="124">
<emphasis id="B9726608FFF8FF92FF4FFB72A030FB3B" bold="true" box="[171,310,1213,1237]" pageId="39" pageNumber="124">Description.</emphasis>
<figureCitation id="133DA69FFFF8FF92FEDBFB72A09DFB3B" box="[319,411,1213,1237]" captionStart="FIGURE" captionStartId="40.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1566]" captionTargetId="figure-56@40.[120,1176,216,1566]" captionTargetPageId="40" captionText="FIGURE 22. Broscosoma gongshanense sp. nov. a. Habitus (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067397" httpUri="https://zenodo.org/record/11067397/files/figure.png" pageId="39" pageNumber="124">Fig. 22a</figureCitation>
. Size medium, BL male = 9.59.
<quantity id="4CFE17FFFFF8FF92FCF6FB72A254FB3B" box="[786,850,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="124" unit="mm" value="8.0">8 mm</quantity>
, female = 9.59.
<quantity id="4CFE17FFFFF8FF92FBF6FB72A555FB3B" box="[1042,1107,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="124" unit="mm" value="9.0">9 mm</quantity>
, ratio EL/PL male = 2.1, female = 2.22.3. Body color black or dark piceous, last visible abdominal sternite dark brown or rufous brown, antennae with antennomeres 3 and 4 black or piceous, antennomeres 1 and 2 and 5 to 11 rufous brown. Femora and tibiae piceous or dark brown, tarsi rufous brown. Head and pronotum without metallic reflection, elytra with distinct blue or greenish-blue metallic reflection, venter without metallic reflection.
</paragraph>
<paragraph id="8BB9BA1AFFF8FF9CFF4FFA4FA03EFB88" blockId="39.[123,1174,301,1726]" lastBlockId="41.[123,1174,1070,1702]" lastPageId="41" lastPageNumber="126" pageId="39" pageNumber="124">
Head.
<figureCitation id="133DA69FFFF8FF92FF16FA4FA039FA76" box="[242,319,1408,1432]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="39" pageNumber="124">Fig. 7e</figureCitation>
. Eyes medium-sized, their diameter equal to or slightly greater than length of tempora. Frontal furrows deeply impressed, narrow, short, linear, distinctly divergent posteriorly, and extended posteriorly to middle of eyes, or medium length, arcuate, slightly convergent anteriorly and posteriorly, and extended posteriorly beyond middle of eyes, faintly rugulose, impunctate. Vertex with post-temporal transverse sulcus deep, sharply defined, moderately or sparsely and finely punctate. Tempora obliquely convex. Genal ridges distinctly present from posterior region of head to ventral margins of eyes. Clypeus with one pair of setae. Supraorbital setae present, one pair. Eustipes of maxilla with two setae, the dorsobasal seta distinctly more than half as long as ventrobasal seta.
<collectingRegion id="49C274F8FFF8FF92FEE9F94AA061F972" box="[269,359,1669,1692]" country="Myanmar" name="Chin" pageId="39" pageNumber="124">Mentum</collectingRegion>
with tooth present, simple, paramedial region shallowly foveate, one pair of mental setae present. Glosal sclerite (ligula) with one pair of setae. Submentum with one or two pairs of setae. Gula without transverse grooves. Antennomeres 3 and 4 without pubescence, ratio A3/A5 = 1.11.2.
</paragraph>
<caption id="DF79EA92FFF7FF9DFF4FF9FCA0FDF997" ID-DOI="http://doi.org/10.5281/zenodo.11067397" ID-Zenodo-Dep="11067397" httpUri="https://zenodo.org/record/11067397/files/figure.png" pageId="40" pageNumber="125" startId="40.[171,183,1587,1606]" targetBox="[120,1176,216,1566]" targetPageId="40" targetType="figure">
<paragraph id="8BB9BA1AFFF7FF9DFF4FF9FCA0FDF997" blockId="40.[123,1173,1587,1657]" pageId="40" pageNumber="125">
<smallCapsWord id="8D5F2CC6FFF7FF9DFF4FF9FCA1EEF9AA" baselines="1601,1600" box="[171,232,1587,1606]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="40" pageNumber="125">FIGURE</smallCapsWord>
22.
<taxonomicName id="4C06C199FFF7FF9DFEE9F9FCA0F2F9A8" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[269,500,1587,1606]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="125" phylum="Arthropoda" rank="species" species="gongshanense" status="sp. nov.">
<emphasis id="B9726608FFF7FF9DFEE9F9FCA0F2F9A8" box="[269,500,1587,1606]" italics="true" pageId="40" pageNumber="125">Broscosoma gongshanense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFF7FF9DFE1EF9FCA33DF9A8" box="[506,571,1587,1606]" pageId="40" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
a. Habitus (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<caption id="DF79EA92FFF6FF9CFF4FFC08A227FBE3" ID-DOI="http://doi.org/10.5281/zenodo.11067399" ID-Zenodo-Dep="11067399" httpUri="https://zenodo.org/record/11067399/files/figure.png" pageId="41" pageNumber="126" startId="41.[171,183,967,986]" targetBox="[157,1154,219,934]" targetPageId="41" targetType="figure">
<paragraph id="8BB9BA1AFFF6FF9CFF4FFC08A227FBE3" blockId="41.[123,1173,966,1037]" pageId="41" pageNumber="126">
<smallCapsWord id="8D5F2CC6FFF6FF9CFF4FFC08A1EEFC36" baselines="981,980" box="[171,232,967,986]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="41" pageNumber="126">FIGURE</smallCapsWord>
23. Male genitalia,
<taxonomicName id="4C06C199FFF6FF9CFE72FC08A378FC37" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[406,638,966,985]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="126" phylum="Arthropoda" rank="species" species="gongshanense" status="sp. nov.">
<emphasis id="B9726608FFF6FF9CFE72FC08A378FC37" box="[406,638,966,985]" italics="true" pageId="41" pageNumber="126">Broscosoma gongshanense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFF6FF9CFD60FC08A3C0FC34" box="[644,710,967,986]" pageId="41" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
(CASENT1026429; 2.0 km S of Heipu Yakou, Cikai Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB9BA1AFFF6FF9CFF4FFBA0A1C0FA88" blockId="41.[123,1174,1070,1702]" pageId="41" pageNumber="126">
Pronotum.
<figureCitation id="133DA69FFFF6FF9CFEC7FBA1A069FB68" box="[291,367,1134,1158]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="41" pageNumber="126">Fig. 7e</figureCitation>
. Globose anterior to sub-basal constriction with base distinctly pedunculate, widest at middle of discal region, ratio PL/PW = 1.21.3, disk markedly convex; apical margination absent, lateral margination present, narrow, delineated only on base; basal margination absent; anterior transverse impression absent or shallow and indistinct; median longitudinal impression distinct, but fine and shallowly impressed; posterior transverse impression deep, sharply delineated; anterior region smooth; pronotal base coarsely punctate and rugulose; one pair of midlateral pronotal setae present at mid-length of discal region; basolateral pronotal setae absent.
</paragraph>
<paragraph id="8BB9BA1AFFF6FF9FFF4FFAA1A289FEDD" blockId="41.[123,1174,1070,1702]" lastBlockId="42.[123,1174,219,1725]" lastPageId="42" lastPageNumber="127" pageId="41" pageNumber="126">Elytra. Elytral silhouette ovoid, widest at middle, ratio EL/EW male = 1.41.5, female = 1.5 1.6, greatest elevation above lateral margin (in lateral view) posterior to middle; humeri narrow, sloped, indistinct; lateral margins markedly arcuate or markedly arcuate and faintly angulate sub-basally; parascutellar striole present, short, distally merged in a distinct jog with free base of stria 1. Eight striae present; stria 1 moderately deeply and sharply impressed, striae 2 to 3, 4, or 5 moderately impressed in basal two-thirds of striae and successively more shallowly impressed in apical third, striae 2 to 8 successively more shallowly impressed laterally and apically, striae obliterated apical and laterally in some specimens, striae 8 merged with lateral groove throughout its length or nearly so; stria 1 coarsely and moderately densely punctate, striae 2 and 3 or 4 moderately to coarsely punctate in basal half and successively more shallowly and sparsely punctate in apical half, striae 4 or 5 to 8 successively more finely punctate or impunctate laterally and apically. Elytral microsculpture effaced. Parascutellar seta present at base of stria 2, discal setae absent, umbilicate series comprised of one post-humeral and two preapical setae.</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FFEF3A3D8FEBD" blockId="42.[123,1174,219,1725]" box="[171,734,316,340]" pageId="42" pageNumber="127">Hindwings. Vestigial, incapable of supporting flight.</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FFE93A213FE9D" blockId="42.[123,1174,219,1725]" box="[171,789,348,372]" pageId="42" pageNumber="127">Thoracic venter. Metepisternum ratio ML/MW = 1.51.9.</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FFEB2A2E8FE3A" blockId="42.[123,1174,219,1725]" pageId="42" pageNumber="127">Legs. Hind trochanter asetose. Tarsomeres without dorsal pubescence; tarsomere 5 with two or more pairs of setae ventrally; male protarsi with pads of adhesive setae on tarsomeres 1 to 3 ventrally, male mesotarsi with pads of adhesive setae on tarsomeres 1 and 2 ventrally.</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FFE12A035FDFB" blockId="42.[123,1174,219,1725]" pageId="42" pageNumber="127">Abdomen. Sternite VI of male with two or three pairs of apicoparamedial setae, female with one pair of setae.</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FFDD2A0E3FDDB" blockId="42.[123,1174,219,1725]" box="[171,485,541,565]" pageId="42" pageNumber="127">
Male genitalia. As in
<figureCitation id="133DA69FFFF5FF9FFE76FDD2A0D9FDDB" box="[402,479,541,565]" captionStart="FIGURE" captionStartId="41.[171,183,967,986]" captionTargetBox="[157,1154,219,934]" captionTargetId="figure-1587@41.[121,1176,216,953]" captionTargetPageId="41" captionText="FIGURE 23. Male genitalia, Broscosoma gongshanense sp. nov. (CASENT1026429; 2.0 km S of Heipu Yakou, Cikai Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Medi- an lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067399" httpUri="https://zenodo.org/record/11067399/files/figure.png" pageId="42" pageNumber="127">Fig. 23</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF5FF9FFF4FFDF2A2F7F953" pageId="42" pageNumber="127" type="discussion">
<paragraph id="8BB9BA1AFFF5FF9FFF4FFDF2A0CDFBD6" blockId="42.[123,1174,219,1725]" pageId="42" pageNumber="127">
<emphasis id="B9726608FFF5FF9FFF4FFDF2A02BFDB8" bold="true" box="[171,301,573,598]" pageId="42" pageNumber="127">Comments.</emphasis>
As noted in the Comments section for
<taxonomicName id="4C06C199FFF5FF9FFD37FDF1A272FDBB" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[723,884,573,597]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFF5FF9FFD37FDF1A272FDBB" box="[723,884,573,597]" italics="true" pageId="42" pageNumber="127">B. bicoloratum</emphasis>
</taxonomicName>
, this is one of four species in the study area members of which have the elytra with distinct metallic reflection but the forebody (head and pronotum) without metallic reflection or with that reflection confined to the pronotal area anterior to the anterior transverse impression. Its members can be distinguished from those of
<taxonomicName id="4C06C199FFF5FF9FFF7CFD70A03CFD38" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[152,314,702,726]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFF5FF9FFF7CFD70A03CFD38" box="[152,314,702,726]" italics="true" pageId="42" pageNumber="127">B. viridicollare</emphasis>
</taxonomicName>
in having smaller body size, the eyes less convex and tempora straight or faintly convex (distinctly convex, subangulate
<taxonomicName id="4C06C199FFF5FF9FFDDBFD10A3E7FD19" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[575,737,735,759]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFF5FF9FFDDBFD10A3E7FD19" box="[575,737,735,759]" italics="true" pageId="42" pageNumber="127">B. viridicollare</emphasis>
</taxonomicName>
), the pronotum without a trace of metallic reflection (band of metallic green reflection present anterior to anterior transverse impression
<taxonomicName id="4C06C199FFF5FF9FFB9AFCCFA1F9FCD9" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFF5FF9FFB9AFCCFA1F9FCD9" italics="true" pageId="42" pageNumber="127">B. viridicollare</emphasis>
</taxonomicName>
), the elytral microsculpture effaced (finely but distinctly impressed in
<taxonomicName id="4C06C199FFF5FF9FFBF5FCEFA1A3FCB9" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFF5FF9FFBF5FCEFA1A3FCB9" italics="true" pageId="42" pageNumber="127">B. viridicollare</emphasis>
</taxonomicName>
), and the median lobe of the male genitalia shorter and more evenly arcuate. They differ from members of
<taxonomicName id="4C06C199FFF5FF9FFEE4FCAFA0A6FC99" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[256,416,863,887]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFF5FF9FFEE4FCAFA0A6FC99" box="[256,416,863,887]" italics="true" pageId="42" pageNumber="127">B. bicoloratum</emphasis>
</taxonomicName>
in having the eyes less convex, the tempora shorter and nearly straight (tempora longer and distinctly more convex in
<taxonomicName id="4C06C199FFF5FF9FFD88FC4EA20DFC76" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[620,779,896,920]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFF5FF9FFD88FC4EA20DFC76" box="[620,779,896,920]" italics="true" pageId="42" pageNumber="127">B. bicoloratum</emphasis>
</taxonomicName>
), and the discal portion of the pronotum very slightly more elongate (more globose
<taxonomicName id="4C06C199FFF5FF9FFD64FC6EA227FC56" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[640,801,928,952]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFF5FF9FFD64FC6EA227FC56" box="[640,801,928,952]" italics="true" pageId="42" pageNumber="127">B. bicoloratum</emphasis>
</taxonomicName>
). They differ from members of
<taxonomicName id="4C06C199FFF5FF9FFB9AFC6EA011FC36" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFF5FF9FFB9AFC6EA011FC36" italics="true" pageId="42" pageNumber="127">B. gaoligongense</emphasis>
</taxonomicName>
in having the tempora slightly less convex, the parascutellar seta inserted near the base of stria 2 (stria
<quantity id="4CFE17FFFFF5FF9FFEB8FC2EA081FC16" box="[348,391,992,1017]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="42" pageNumber="127" unit="in" value="3.0">3 in</quantity>
<emphasis id="B9726608FFF5FF9FFE6BFC2EA383FC16" box="[399,645,992,1016]" italics="true" pageId="42" pageNumber="127">
most
<taxonomicName id="4C06C199FFF5FF9FFE2EFC2EA383FC16" box="[458,645,992,1016]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="gaoligongense">B. gaoligongense</taxonomicName>
</emphasis>
), the metatrochanter asetose (unisetose
<taxonomicName id="4C06C199FFF5FF9FFBDCFC2EA1DBFBF7" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFF5FF9FFBDCFC2EA1DBFBF7" italics="true" pageId="42" pageNumber="127">B. gongshanense</emphasis>
</taxonomicName>
), and the median lobe of the male genitalia relatively shorter and with sclerites of the internal sac of different shapes.
</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FFB8EA3EBFAF4" blockId="42.[123,1174,219,1725]" pageId="42" pageNumber="127">
As noted above in the Comments sections for
<taxonomicName id="4C06C199FFF5FF9FFD4CFB8DA24FFBB7" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[680,841,1089,1113]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFF5FF9FFD4CFB8DA24FFBB7" box="[680,841,1089,1113]" italics="true" pageId="42" pageNumber="127">B. bicoloratum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C06C199FFF5FF9FFC66FB8DA538FBB7" box="[898,1086,1089,1113]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFF5FF9FFC66FB8DA538FBB7" box="[898,1086,1089,1113]" italics="true" pageId="42" pageNumber="127">B. gaoligongense</emphasis>
</taxonomicName>
, among
<taxonomicName id="4C06C199FFF5FF9FFF9FFBADA057FB97" box="[123,337,1121,1145]" pageId="42" pageNumber="127">
<emphasis id="B9726608FFF5FF9FFF9FFBADA1F8FB97" box="[123,254,1122,1145]" italics="true" pageId="42" pageNumber="127">Broscosoma</emphasis>
species
</taxonomicName>
known from outside the study area, only
<taxonomicName id="4C06C199FFF5FF9FFCE9FBADA293FB97" authorityName="Habu" authorityYear="1973" box="[781,917,1121,1145]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B9726608FFF5FF9FFCE9FBADA293FB97" box="[781,917,1121,1145]" italics="true" pageId="42" pageNumber="127">B. monticola</emphasis>
</taxonomicName>
has members of similar size and with the forebody lacking metallic reflection and the elytral with markedly metallic reflection as is seen in
<taxonomicName id="4C06C199FFF5FF9FFEDDFB6CA0F6FB54" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[313,496,1186,1210]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFF5FF9FFEDDFB6CA0F6FB54" box="[313,496,1186,1210]" italics="true" pageId="42" pageNumber="127">B. gongshanense</emphasis>
</taxonomicName>
specimens. However, they have the parascutellar seta inserted near the base of stria 3 (stria
<quantity id="4CFE17FFFFF5FF9FFE54FB0DA0DCFB34" box="[432,474,1218,1242]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="42" pageNumber="127" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C06C199FFF5FF9FFE04FB0CA393FB34" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[480,661,1218,1242]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFF5FF9FFE04FB0CA393FB34" box="[480,661,1218,1242]" italics="true" pageId="42" pageNumber="127">B. gongshanense</emphasis>
</taxonomicName>
), the femora slightly paler, and the median lobe of the male genitalia with a large protuberance on the ventral margin (see
<bibRefCitation id="EF97C7EBFFF5FF9FFC61FB2DA2FCFB14" author="Habu, A." box="[901,1018,1250,1274]" pageId="42" pageNumber="127" pagination="81 - 132" refId="ref53285" refString="Habu, A. 1973. On a collection of Carabidae from Nepal made by the Hokkaido University Scientifi c Exedition to Nepal Himalaya, 1968 (I). The Bulletin of the National Institute of Agricultural Sciences (C) 27: 81 - 132." type="journal article" year="1973">Habu 1973</bibRefCitation>
, fig. 6), which is absent from the median lobe of
<taxonomicName id="4C06C199FFF5FF9FFE0FFACCA3A6FAF4" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[491,672,1282,1306]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFF5FF9FFE0FFACCA3A6FAF4" box="[491,672,1282,1306]" italics="true" pageId="42" pageNumber="127">B. gongshanense</emphasis>
</taxonomicName>
males.
</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FFAEDA355FA75" blockId="42.[123,1174,219,1725]" pageId="42" pageNumber="127">
<emphasis id="B9726608FFF5FF9FFF4FFAEDA09DFAD5" bold="true" box="[171,411,1314,1339]" pageId="42" pageNumber="127">Habitat distribution.</emphasis>
Members of this species have been found under stones along trails through bamboo and
<taxonomicName id="4C06C199FFF5FF9FFEB8FA8CA0FCFAB5" box="[348,506,1347,1371]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="42" pageNumber="127" phylum="Tracheophyta" rank="genus">
<emphasis id="B9726608FFF5FF9FFEB8FA8CA0FCFAB5" box="[348,506,1347,1371]" italics="true" pageId="42" pageNumber="127">Rhododendron</emphasis>
</taxonomicName>
thickets and in moist alpine meadow areas (
<figureCitation id="133DA69FFFF5FF9FFC29FA8CA522FAB4" box="[973,1060,1347,1371]" captionStart="FIGURE" captionStartId="82.[171,183,1585,1604]" captionTargetBox="[128,1169,216,1568]" captionTargetId="figure-57@82.[128,1169,216,1568]" captionTargetPageId="82" captionText="FIGURE 42. Photographs of habitats for Broscosoma species in the Gaoligong Shan region. a. Dulong/Gongshan Yakou area, 3300-3680m, 27.697º/098.454º, Gongshan County, Yunnan (habitat for Broscosoma bicoloratum sp. nov. and B. gongshanense sp. nov.); b. Southeastern slope of Heipu Yakou, 3365m, 27.770º/098.447º, Cikai Township, Gongshan County, Yunnan (habitat for Broscosoma bicoloratum sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067443" httpUri="https://zenodo.org/record/11067443/files/figure.png" pageId="42" pageNumber="127">Fig. 42a</figureCitation>
) at or near the crest of the range. Our records document occurrence of this species at moderately high elevations in the
<quantity id="4CFE17FFFFF5FF9FFF1EFA4CA0A2FA75" box="[250,420,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="3.5149999999999997" metricValueMax="3.6799999999999997" metricValueMin="3.35" pageId="42" pageNumber="127" unit="m" value="3515.0" valueMax="3680.0" valueMin="3350.0">3350 to 3680 m</quantity>
range (
<figureCitation id="133DA69FFFF5FF9FFE13FA4CA345FA75" box="[503,579,1411,1435]" captionStart="FIGURE" captionStartId="88.[184,196,1682,1701]" captionTargetBox="[130,1162,1063,1660]" captionTargetId="figure-332@88.[120,1176,1054,1668]" captionTargetPageId="88" captionText="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." figureDoi="http://doi.org/10.5281/zenodo.11067457" httpUri="https://zenodo.org/record/11067457/files/figure.png" pageId="42" pageNumber="127">Fig. 49</figureCitation>
).
</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FFA6CA590F9F2" blockId="42.[123,1174,219,1725]" pageId="42" pageNumber="127">
<emphasis id="B9726608FFF5FF9FFF4FFA6CA208FA52" bold="true" box="[171,782,1443,1468]" pageId="42" pageNumber="127">
Geographical distribution within the Gaoligong
<materialsCitation id="3B6EB047FFF5FF9FFD29FA6CA588F9F2" ID-GBIF-Occurrence="4899905373" county="Gongshan County" location="Core Area" municipality="Core Area" pageId="42" pageNumber="127" specimenCount="12" specimenCount-female="3" specimenCount-male="3" stateProvince="Shan">
<collectingRegion id="49C274F8FFF5FF9FFD29FA6CA201FA55" box="[717,775,1443,1467]" country="Myanmar" name="Shan" pageId="42" pageNumber="127">Shan</collectingRegion>
.
<figureCitation id="133DA69FFFF5FF9FFCF0FA6BA26BFA52" box="[788,877,1444,1468]" captionStart="FIGURE" captionStartId="40.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1566]" captionTargetId="figure-56@40.[120,1176,216,1566]" captionTargetPageId="40" captionText="FIGURE 22. Broscosoma gongshanense sp. nov. a. Habitus (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067397" httpUri="https://zenodo.org/record/11067397/files/figure.png" pageId="42" pageNumber="127">Fig. 22b</figureCitation>
. We examined a total of
<specimenCount id="9D007193FFF5FF9FFB92FA6BA1ECFA32" count="6" pageId="42" pageNumber="127" type="generic">six specimens</specimenCount>
(
<specimenCount id="9D007193FFF5FF9FFF1EFA0BA070FA32" box="[250,374,1476,1500]" count="3" pageId="42" pageNumber="127" type="male">three males</specimenCount>
and
<specimenCount id="9D007193FFF5FF9FFE48FA0BA33AFA32" box="[428,572,1476,1500]" count="3" pageId="42" pageNumber="127" type="female">three females</specimenCount>
), all from at or near the crest of the range in
<collectingCounty id="62D8C296FFF5FF9FFBCDFA0BA1CEFA12" pageId="42" pageNumber="127">Gongshan County</collectingCounty>
. We recorded this species only from the crest and adjacent slopes to the east in
<collectingMunicipality id="6BDD2060FFF5FF9FFBCCFA2BA590FA12" box="[1064,1174,1508,1532]" pageId="42" pageNumber="127">Core Area</collectingMunicipality>
2 (
<figureCitation id="133DA69FFFF5FF9FFF7CF9CBA1E2F9F2" box="[152,228,1540,1564]" captionStart="FIGURE" captionStartId="88.[171,183,995,1014]" captionTargetBox="[130,1164,233,969]" captionTargetId="figure-57@88.[120,1176,215,989]" captionTargetPageId="88" captionText="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." figureDoi="http://doi.org/10.5281/zenodo.11067455" httpUri="https://zenodo.org/record/11067455/files/figure.png" pageId="42" pageNumber="127">Fig. 48</figureCitation>
), but it probably also occurs on the western side at equivalent elevations in
<location id="8ED9ECC1FFF5FF9FFBEAF9CBA57DF9F2" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF8FF9FFF9FFF14A2F7F953:8ED9ECC1FFF5FF9FFBEAF9CBA57DF9F2" box="[1038,1147,1540,1564]" county="Gongshan County" municipality="Core Area" name="Core Area" pageId="42" pageNumber="127" stateProvince="Shan">Core Area</location>
1
</materialsCitation>
</emphasis>
.
</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FF9EBA25FF9B2" blockId="42.[123,1174,219,1725]" pageId="42" pageNumber="127">
<emphasis id="B9726608FFF5FF9FFF4FF9EBA336F9D3" bold="true" box="[171,560,1572,1597]" pageId="42" pageNumber="127">Overall geographical distribution.</emphasis>
This species currently is known only from the northcentral part of the Gaoligong Shan, in western
<collectingRegion id="49C274F8FFF5FF9FFDA8F98AA202F9B3" box="[588,772,1605,1629]" country="China" name="Yunnan" pageId="42" pageNumber="127">Yunnan Province</collectingRegion>
,
<collectingCountry id="F311FA8AFFF5FF9FFCF6F98AA254F9B3" box="[786,850,1605,1629]" name="China" pageId="42" pageNumber="127">China</collectingCountry>
.
</paragraph>
<paragraph id="8BB9BA1AFFF5FF9FFF4FF9AAA2F7F953" blockId="42.[123,1174,219,1725]" pageId="42" pageNumber="127">
<emphasis id="B9726608FFF5FF9FFF4FF9AAA24DF993" bold="true" box="[171,843,1637,1661]" pageId="42" pageNumber="127">
Geographical relationships with other
<taxonomicName id="4C06C199FFF5FF9FFD83F9AAA24DF993" box="[615,843,1637,1661]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9726608FFF5FF9FFD83F9AAA3E8F992" bold="true" box="[615,750,1637,1660]" italics="true" pageId="42" pageNumber="127">Broscosoma</emphasis>
species.
</taxonomicName>
</emphasis>
Members of this species have been found together only with those of
<taxonomicName id="4C06C199FFF5FF9FFDD6F949A246F973" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[562,832,1669,1693]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFF5FF9FFDD6F949A246F973" box="[562,832,1669,1693]" italics="true" pageId="42" pageNumber="127">Broscosoma bicoloratum</emphasis>
</taxonomicName>
at one or more sites (
<figureCitation id="133DA69FFFF5FF9FFBDCF94AA580F973" box="[1080,1158,1669,1693]" captionStart="FIGURE" captionStartId="89.[171,183,1461,1480]" captionTargetBox="[140,1154,249,1428]" captionTargetId="figure-68@89.[120,1176,216,1448]" captionTargetPageId="89" captionText="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi="http://doi.org/10.5281/zenodo.11067459" httpUri="https://zenodo.org/record/11067459/files/figure.png" pageId="42" pageNumber="127">Fig. 50</figureCitation>
).
<taxonomicName id="4C06C199FFF5FF9FFF9FF969A043F953" authorityName="Putzeys. We" authorityYear="1877" box="[123,325,1701,1725]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="127" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFF5FF9FFF9FF969A043F953" box="[123,325,1701,1725]" italics="true" pageId="42" pageNumber="127">Broscosoma ribbei</emphasis>
</taxonomicName>
occurs in the same general area but mainly at lower elevations.
</paragraph>
</subSubSection>
</treatment>
</document>