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<document id="100DA9AA1A594F822683A4C014256C3D" ID-CLB-Dataset="299055" ID-DOI="10.5281/zenodo.11067355" ID-GBIF-Dataset="67ec2de4-1816-4969-8132-f6796d3afa06" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067355" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052343267" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2021" docId="03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD" docLanguage="en" docName="ProcCalifAcadSci.67.4.85-182.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (4)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Broscosoma holomarginatum Kavanaugh and Liang 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="132" masterDocId="FF967374FFDFFFB5FFE4FFCFA106FFEE" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." masterLastPageNumber="182" masterPageNumber="85" pageNumber="128" updateTime="1719540814554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="750A878C374F291AFE96745E79CED5CE">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae).</mods:title>
</mods:titleInfo>
<mods:name id="17B28D5ABBE5362CD133234AC5588682" type="personal">
<mods:role id="172DF76BD75563A68EB1D117BDC99565">
<mods:roleTerm id="CA41B68ABAB80262389456EEF793E3BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="664FCC409D7DBECF60FA76D13B2DF196">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="61A466DD424FF36466C5BCE136CC38B1">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="31E8CDB56E59AF75BE8017E74CBDEB2C">Liang, Hongbin</mods:namePart>
<mods:affiliation id="DF5CC7DE32D37A12308233AEBDE5C520">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
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<mods:title id="82E1215F2DB7023F8D8AF2AE984F970C">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="37DCF2AA462ED7ECB0DABC7B3EF23B34">
<mods:date id="C59910846CBDE50DA4A2865FF1D5C77B">2021</mods:date>
<mods:detail id="4FD94A474D0D20785BBF7B300F1BB1A1" type="pubDate">
<mods:number id="DD7BB02A3AE6BD08130CFA5412652CA9">2021-09-30</mods:number>
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<mods:detail id="F4B32A3D85EDB472F2220D3EBC38D5FD" type="volume">
<mods:number id="3BB1EE52FACD7F491DDEA78660D24EF3">67</mods:number>
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<mods:number id="A1D2B71F001CC1251FA5A9FCFDCBAAE8">4</mods:number>
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<mods:classification id="CC46AFFEFC351EA5BE639066F3CC2995">journal article</mods:classification>
<mods:identifier id="C4C9301E56993C2C059EA1DD3009D4E6" type="CLB-Dataset">299055</mods:identifier>
<mods:identifier id="D0CCF2B0AF9D9ACFB2C0F51DD915857E" type="DOI">10.5281/zenodo.11067355</mods:identifier>
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<treatment id="03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD" ID-DOI="http://doi.org/10.5281/zenodo.11067423" ID-GBIF-Taxon="233814476" ID-Zenodo-Dep="11067423" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD" httpUri="http://treatment.plazi.org/id/03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD" lastPageId="47" lastPageNumber="132" pageId="43" pageNumber="128">
<subSubSection id="C31CE991FFF4FF9EFF9FFF14A24FFF1D" box="[123,841,219,243]" pageId="43" pageNumber="128" type="nomenclature">
<paragraph id="8BB9BA1AFFF4FF9EFF9FFF14A24FFF1D" blockId="43.[123,841,219,273]" box="[123,841,219,243]" pageId="43" pageNumber="128">
<heading id="D0F10D76FFF4FF9EFF9FFF14A24FFF1D" box="[123,841,219,243]" fontSize="10" level="2" pageId="43" pageNumber="128" reason="7">
<emphasis id="B9726608FFF4FF9EFF9FFF14A24FFF1D" bold="true" box="[123,841,219,243]" pageId="43" pageNumber="128">
8.
<taxonomicName id="4C06C199FFF4FF9EFF72FF14A3E3FF1D" authority="Kavanaugh and Liang" authorityName="Kavanaugh and Liang" authorityYear="2021" box="[150,741,219,243]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="128" phylum="Arthropoda" rank="species" species="holomarginatum" status="sp. nov.">
<emphasis id="B9726608FFF4FF9EFF72FF14A0DBFF1D" bold="true" box="[150,477,219,243]" italics="true" pageId="43" pageNumber="128">Broscosoma holomarginatum</emphasis>
Kavanaugh and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A241DB73FFF4FF9EFD17FF14A24FFF1D" box="[755,841,219,243]" pageId="43" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF4FF9EFF9FFF34A0C4FEFF" box="[123,450,251,273]" pageId="43" pageNumber="128" type="description">
<paragraph id="8BB9BA1AFFF4FF9EFF9FFF34A0C4FEFF" blockId="43.[123,841,219,273]" box="[123,450,251,273]" pageId="43" pageNumber="128">
<figureCitation id="133DA69FFFF4FF9EFF9FFF34A1E6FEFF" box="[123,224,251,273]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="43" pageNumber="128">Figures 6a</figureCitation>
,
<figureCitation id="133DA69FFFF4FF9EFF09FF33A002FEFF" box="[237,260,252,273]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="43" pageNumber="128">9a</figureCitation>
,
<figureCitation id="133DA69FFFF4FF9EFEF4FF33A02EFEFF" box="[272,296,252,273]" captionStart="FIGURE" captionStartId="44.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1568]" captionTargetId="figure-56@44.[120,1176,216,1568]" captionTargetPageId="44" captionText="FIGURE 24. Broscosoma holomarginatum sp. nov. a. Habitus (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067401" httpUri="https://zenodo.org/record/11067401/files/figure.png" pageId="43" pageNumber="128">24</figureCitation>
,
<figureCitation id="133DA69FFFF4FF9EFED0FF34A04AFEFF" box="[308,332,251,273]" captionStart="FIGURE" captionStartId="45.[171,183,939,958]" captionTargetBox="[137,1161,228,920]" captionTargetId="figure-1604@45.[120,1176,215,935]" captionTargetPageId="45" captionText="FIGURE 25. Male genitalia, Broscosoma holomarginatum sp. nov. (CASENT1016743; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067405" httpUri="https://zenodo.org/record/11067405/files/figure.png" pageId="43" pageNumber="128">25</figureCitation>
,
<figureCitation id="133DA69FFFF4FF9EFEBDFF33A07BFEFF" box="[345,381,251,273]" captionStart="FIGURE" captionStartId="84.[171,183,1585,1604]" captionTargetBox="[147,1149,216,1572]" captionTargetId="figure-57@84.[147,1149,216,1572]" captionTargetPageId="84" captionText="FIGURE 44. Photographs of habitats for Broscosoma species in the Gaoligong Shan region. a. 0.5 km E of Fengxue Yakou, 3150 m, 25.972º/098.683 º, Luzhang Township, Lushui County, Yunnan (habitat for Broscosoma gaoligongense sp. nov.); b. Xiao Jiang at Gangfang Sancha Lukou, 1500 m, 26.122º/98.573º, Lushui County, Yunnan (habitat for Broscosoma holomarginatum sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067447" httpUri="https://zenodo.org/record/11067447/files/figure.png" pageId="43" pageNumber="128">44b</figureCitation>
,
<figureCitation id="133DA69FFFF4FF9EFE6EFF33A0C4FEFF" box="[394,450,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStartId-0="87.[171,183,970,989]" captionStartId-1="88.[171,183,995,1014]" captionStartId-2="88.[184,196,1682,1701]" captionStartId-3="89.[171,183,1461,1480]" captionTargetBox-0="[120,1180,216,954]" captionTargetBox-1="[130,1164,233,969]" captionTargetBox-2="[130,1162,1063,1660]" captionTargetBox-3="[140,1154,249,1428]" captionTargetId-0="figure-67@87.[120,1180,216,954]" captionTargetId-1="figure-57@88.[120,1176,215,989]" captionTargetId-2="figure-332@88.[120,1176,1054,1668]" captionTargetId-3="figure-68@89.[120,1176,216,1448]" captionTargetPageId-0="87" captionTargetPageId-1="88" captionTargetPageId-2="88" captionTargetPageId-3="89" captionText-0="FIGURE 47. Map showing approximate known overall geographical distributions of broscine species occurring in the Gaoligong Shan region as well as outside the study area. a. Broscus punctatus (Dejean); b. Eobroscus bhutanensis Morvan; c. Broscosoma ribbei Putzeys; d. Broscosoma holomarginatum sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 k" captionText-1="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-2="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." captionText-3="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi-0="http://doi.org/10.5281/zenodo.11067453" figureDoi-1="http://doi.org/10.5281/zenodo.11067455" figureDoi-2="http://doi.org/10.5281/zenodo.11067457" figureDoi-3="http://doi.org/10.5281/zenodo.11067459" httpUri-0="https://zenodo.org/record/11067453/files/figure.png" httpUri-1="https://zenodo.org/record/11067455/files/figure.png" httpUri-2="https://zenodo.org/record/11067457/files/figure.png" httpUri-3="https://zenodo.org/record/11067459/files/figure.png" pageId="43" pageNumber="128">47-50</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF4FF9EFF4FFEE1A3DAFAD8" pageId="43" pageNumber="128" type="materials_examined">
<paragraph id="8BB9BA1AFFF4FF9EFF4FFEE1A1E8FB7A" blockId="43.[123,1174,301,1726]" pageId="43" pageNumber="128">
<emphasis id="B9726608FFF4FF9EFF4FFEE1A054FEA8" bold="true" box="[171,338,301,326]" pageId="43" pageNumber="128">Type material.</emphasis>
<materialsCitation id="3B6EB047FFF4FF9EFEBEFEE1A27FFE49" ID-GBIF-Occurrence="4899905501" collectingDate="2004-11-05" collectionCode="IOZ" collectorName="H. - B. Liang" country="China" county="Gongshan County" elevation="1525" latitude="27.87696" location="Kongdang" longLatPrecision="1" longitude="98.33587" municipality="Dulongjiang Township" pageId="43" pageNumber="128" specimenCode="CASENT 1015162" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54BD04B8FFF4FF9EFEBEFEE1A0BBFEA8" box="[346,445,302,326]" pageId="43" pageNumber="128" type="holotype">Holotype</typeStatus>
: a male, in
<collectionCode id="ED1722DFFFF4FF9EFDA6FEE1A368FEA8" box="[578,622,302,326]" pageId="43" pageNumber="128">IOZ</collectionCode>
, labeled: “
<specimenCode id="DBA01261FFF4FF9EFD02FEE1A2B5FEA8" box="[742,947,302,326]" pageId="43" pageNumber="128">CASENT 1015162</specimenCode>
”/ “
<collectingCountry id="F311FA8AFFF4FF9EFC3FFEE1A529FEA8" box="[987,1071,302,326]" name="China" pageId="43" pageNumber="128">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF4FF9EFBD9FEE1A588FEAB" box="[1085,1166,302,325]" country="China" name="Yunnan" pageId="43" pageNumber="128">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF4FF9EFF9FFE81A039FE88" box="[123,319,334,358]" pageId="43" pageNumber="128">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF4FF9EFEB4FE81A34FFE88" box="[336,585,334,358]" pageId="43" pageNumber="128">Dulongjiang Township</collectingMunicipality>
,
<location id="8ED9ECC1FFF4FF9EFDBEFE81A3CFFE88" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD:8ED9ECC1FFF4FF9EFDBEFE81A3CFFE88" box="[602,713,334,358]" country="China" county="Gongshan County" latitude="27.87696" longLatPrecision="1" longitude="98.33587" municipality="Dulongjiang Township" name="Kongdang" pageId="43" pageNumber="128" stateProvince="Yunnan">Kongdang</location>
,
<quantity id="4CFE17FFFFF4FF9EFD3FFE81A236FE88" box="[731,816,334,358]" metricMagnitude="3" metricUnit="m" metricValue="1.525" pageId="43" pageNumber="128" unit="m" value="1525.0">
<elevation id="002B5D29FFF4FF9EFD3FFE81A236FE88" box="[731,816,334,358]" metricMagnitude="3" metricUnit="m" metricValue="1.525" pageId="43" pageNumber="128" unit="m" value="1525.0">1525 m</elevation>
</quantity>
,
<geoCoordinate id="EE32DCDDFFF4FF9EFCA6FE80A2C2FE88" box="[834,964,334,358]" degrees="27. 87696" direction="north" orientation="latitude" pageId="43" pageNumber="128" precision="1" value="27.87696">N27.87696°</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF4FF9EFC2FFE80A55EFE88" box="[971,1112,334,358]" degrees="098. 33587" direction="east" orientation="longitude" pageId="43" pageNumber="128" precision="1" value="98.33587">E098.33587°</geoCoordinate>
,”/ “
<date id="FFB89CDAFFF4FF9EFB6CFE81A021FE69" pageId="43" pageNumber="128" value="2004-11-05">
<collectingDate id="EFFC6532FFF4FF9EFB6CFE81A021FE69" pageId="43" pageNumber="128" value="2004-11-05">5 November 2004</collectingDate>
</date>
, Stop # LHB-2004-047,
<collectorName id="26F3DFCCFFF4FF9EFDD0FEA0A3B2FE69" box="[564,692,367,391]" pageId="43" pageNumber="128">H.-B. Liang</collectorName>
collector”/ “HOLOTYPE
<taxonomicName id="4C06C199FFF4FF9EFC37FEBFA0E7FE49" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="128" phylum="Arthropoda" rank="species" species="holomarginatum" status="sp. nov.">
<emphasis id="B9726608FFF4FF9EFC37FEBFA1FAFE49" italics="true" pageId="43" pageNumber="128">Broscosoma holomarginatum</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFF4FF9EFE0DFE5FA33BFE49" box="[489,573,400,423]" pageId="43" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [red label]
</materialsCitation>
.
<materialsCitation id="3B6EB047FFF4FF9EFC63FE5FA560FE26" ID-GBIF-Occurrence="4899905414" collectingDate="2004-11-05" collectionCode="CAS, IOZ" collectorName="H. - B. Liang" country="China" county="Gongshan County" elevation="1525" latitude="27.87696" location="Kongdang" longLatPrecision="1" longitude="98.33587" municipality="Dulongjiang Township" pageId="43" pageNumber="128" specimenCode="CASENT 1015163" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54BD04B8FFF4FF9EFC63FE5FA2E9FE46" box="[903,1007,400,424]" pageId="43" pageNumber="128" type="paratype">Paratypes</typeStatus>
(a total of 19):
<specimenCount id="9D007193FFF4FF9EFF9FFE7FA1F9FE26" box="[123,255,432,456]" count="2" pageId="43" pageNumber="128" type="female">two females</specimenCount>
(
<collectionCode id="ED1722DFFFF4FF9EFEF4FE7FA042FE26" box="[272,324,432,456]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="43" pageNumber="128" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1722DFFFF4FF9EFEB0FE7FA086FE26" box="[340,384,432,456]" pageId="43" pageNumber="128">IOZ</collectionCode>
), labeled: same as holotype except first label: “
<specimenCode id="DBA01261FFF4FF9EFC69FE7FA55CFE26" box="[909,1114,432,456]" pageId="43" pageNumber="128">CASENT 1015163</specimenCode>
”
</materialsCitation>
<materialsCitation id="3B6EB047FFF4FF9EFB8BFE7FA0F3FE06" ID-GBIF-Occurrence="4899905362" collectionCode="CAS, IOZ" country="China" pageId="43" pageNumber="128" specimenCode="CASENT 1015164" specimenCount="1" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
and “
<specimenCode id="DBA01261FFF4FF9EFF63FE1EA050FE06" box="[135,342,464,489]" pageId="43" pageNumber="128">CASENT 1015164</specimenCode>
”, respectively
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF4FF9EFDE3FE1EA0F7FDA4" ID-GBIF-Occurrence="4899905548" collectingDate="2004-10-25" collectionCode="IOZ" collectorName="H. - B. Liang &amp; D. - Z. Dong" country="China" county="Gongshan County" elevation="1500" latitude="27.88111" location="Dulongjiang Township" longLatPrecision="1" longitude="98.34063" pageId="43" pageNumber="128" specimenCode="CASENT 1016688" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF4FF9EFDE3FE1EA387FE06" box="[519,641,464,488]" count="1" pageId="43" pageNumber="128" type="female">one female</specimenCount>
(
<collectionCode id="ED1722DFFFF4FF9EFD71FE1EA3C7FE07" box="[661,705,465,489]" pageId="43" pageNumber="128">IOZ</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF4FF9EFCA5FE1EA516FE06" box="[833,1040,464,489]" pageId="43" pageNumber="128">CASENT 1016688</specimenCode>
”/ “
<collectingCountry id="F311FA8AFFF4FF9EFBDEFE1EA588FE07" box="[1082,1166,465,489]" name="China" pageId="43" pageNumber="128">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF4FF9EFF9FFE3DA1C9FDE7" box="[123,207,498,521]" country="China" name="Yunnan" pageId="43" pageNumber="128">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF4FF9EFF0FFE3EA0C6FDE7" box="[235,448,497,521]" pageId="43" pageNumber="128">Gongshan County</collectingCounty>
,
<location id="8ED9ECC1FFF4FF9EFE38FE3EA3EEFDE7" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD:8ED9ECC1FFF4FF9EFE38FE3EA3EEFDE7" box="[476,744,497,521]" country="China" county="Gongshan County" latitude="27.88111" longLatPrecision="1" longitude="98.34063" name="Dulongjiang Township" pageId="43" pageNumber="128" stateProvince="Yunnan">Dulongjiang Township</location>
,
<locationDeviation id="9A0BA338FFF4FF9EFCE0FE3EA520FDE7" box="[772,1062,497,521]" location="Kongdang" pageId="43" pageNumber="128">
<quantity id="4CFE17FFFFF4FF9EFCE0FE3EA24CFDE7" box="[772,842,497,521]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="128" unit="km" value="0.5">0.5km</quantity>
N of Kongdang
</locationDeviation>
,
<quantity id="4CFE17FFFFF4FF9EFBA6FE3EA588FDE7" box="[1090,1166,497,521]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="128" unit="m" value="1500.0">
<elevation id="002B5D29FFF4FF9EFBA6FE3EA588FDE7" box="[1090,1166,497,521]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="128" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<geoCoordinate id="EE32DCDDFFF4FF9EFF9FFDDDA1FEFDC4" box="[123,248,530,554]" degrees="27. 88111" direction="north" orientation="latitude" pageId="43" pageNumber="128" precision="1" value="27.88111">N27.88111º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF4FF9EFEE4FDDDA08FFDC4" box="[256,393,529,554]" degrees="098. 34063" direction="east" orientation="longitude" pageId="43" pageNumber="128" precision="1" value="98.34063">E098.34063º</geoCoordinate>
,
<date id="FFB89CDAFFF4FF9EFE7EFDDEA352FDC4" box="[410,596,529,554]" pageId="43" pageNumber="128" value="2004-10-25">
<collectingDate id="EFFC6532FFF4FF9EFE7EFDDEA352FDC4" box="[410,596,529,554]" pageId="43" pageNumber="128" value="2004-10-25">25 October 2004</collectingDate>
</date>
”/ “Stop #DHK-2004-057B, D.H. Kavanaugh,
<collectorName id="26F3DFCCFFF4FF9EFBBEFDDDA1BFFDA4" pageId="43" pageNumber="128">H.-B. Liang</collectorName>
, &amp;
<collectorName id="26F3DFCCFFF4FF9EFF0AFDFCA076FDA5" box="[238,368,563,587]" pageId="43" pageNumber="128">D.-Z. Dong</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF4FF9EFDE1FDFCA590FDA4" ID-GBIF-Occurrence="4899905370" box="[517,1174,562,586]" collectingDate="2004-10-25" collectionCode="CAS, IOZ" collectorName="Q. B. Hou &amp; H. B. Liang &amp; D. Z. Dong &amp; G. Tang" country="China" county="Gongshan County" elevation="1500" latitude="27.88111" location="Dulongjiang Township" longLatPrecision="1" longitude="98.34063" pageId="43" pageNumber="128" specimenCode="CASENT 1014588" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF4FF9EFDE1FDFCA37EFDA4" box="[517,632,562,586]" count="2" pageId="43" pageNumber="128" type="male">two males</specimenCount>
(
<collectionCode id="ED1722DFFFF4FF9EFD6AFDFDA3C4FDA4" box="[654,706,562,586]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="43" pageNumber="128" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1722DFFFF4FF9EFD31FDFDA207FDA4" box="[725,769,562,586]" pageId="43" pageNumber="128">IOZ</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF4FF9EFC61FDFDA550FDA4" box="[901,1110,562,586]" pageId="43" pageNumber="128">CASENT 1014588</specimenCode>
” and
</materialsCitation>
<materialsCitation id="3B6EB047FFF4FF9EFF9FFD9CA2F2FD42" ID-GBIF-Occurrence="4899905507" collectingDate="2004-10-25" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; Q. B. Hou &amp; H. B. Liang &amp; D. Z. Dong &amp; G. Tang" country="China" county="Gongshan County" elevation="1500" latitude="27.88111" location="Dulongjiang Township" longLatPrecision="1" longitude="98.34063" pageId="43" pageNumber="128" specimenCode="CASENT 1014589" specimenCount="1" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
“
<specimenCode id="DBA01261FFF4FF9EFF63FD9CA054FD85" box="[135,338,595,619]" pageId="43" pageNumber="128">CASENT 1014589</specimenCode>
”, respectively/ “
<collectingCountry id="F311FA8AFFF4FF9EFDEDFD9CA35BFD85" box="[521,605,595,619]" name="China" pageId="43" pageNumber="128">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF4FF9EFD8FFD9CA3BAFD84" box="[619,700,595,618]" country="China" name="Yunnan" pageId="43" pageNumber="128">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF4FF9EFD2EFD9CA28DFD85" box="[714,907,595,619]" pageId="43" pageNumber="128">Gongshan County</collectingCounty>
,
<location id="8ED9ECC1FFF4FF9EFC7DFD9CA589FD85" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD:8ED9ECC1FFF4FF9EFC7DFD9CA589FD85" box="[921,1167,595,619]" country="China" county="Gongshan County" latitude="27.88111" longLatPrecision="1" longitude="98.34063" name="Dulongjiang Township" pageId="43" pageNumber="128" stateProvince="Yunnan">Dulongjiang Township</location>
,
<locationDeviation id="9A0BA338FFF4FF9EFF9FFDBCA068FD65" box="[123,366,627,651]" location="Kongdang" pageId="43" pageNumber="128">
<quantity id="4CFE17FFFFF4FF9EFF9FFDBCA1B8FD65" box="[123,190,627,651]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="128" unit="km" value="0.5">0.5km</quantity>
N of Kongdang
</locationDeviation>
,
<quantity id="4CFE17FFFFF4FF9EFE98FDBCA0C1FD65" box="[380,455,627,651]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="128" unit="m" value="1500.0">
<elevation id="002B5D29FFF4FF9EFE98FDBCA0C1FD65" box="[380,455,627,651]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="128" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<geoCoordinate id="EE32DCDDFFF4FF9EFE31FDBBA354FD65" box="[469,594,627,651]" degrees="27. 88111" direction="north" orientation="latitude" pageId="43" pageNumber="128" precision="1" value="27.88111">N27.88111º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF4FF9EFDBEFDBBA3E5FD65" box="[602,739,627,651]" degrees="098. 34063" direction="east" orientation="longitude" pageId="43" pageNumber="128" precision="1" value="98.34063">E098.34063º</geoCoordinate>
,
<date id="FFB89CDAFFF4FF9EFD16FDBCA2A1FD65" box="[754,935,627,651]" pageId="43" pageNumber="128" value="2004-10-25">
<collectingDate id="EFFC6532FFF4FF9EFD16FDBCA2A1FD65" box="[754,935,627,651]" pageId="43" pageNumber="128" value="2004-10-25">25 October 2004</collectingDate>
</date>
”/ “Stop #DHK-2004- 057C,
<collectorName id="26F3DFCCFFF4FF9EFF27FD5BA07FFD42" box="[195,377,660,684]" pageId="43" pageNumber="128">D.H. Kavanaugh</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFE6CFD5BA0F7FD45" box="[392,497,660,684]" pageId="43" pageNumber="128">Q.B. Hou</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFDE4FD5BA37EFD42" box="[512,632,660,684]" pageId="43" pageNumber="128">H.B. Liang</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFD6CFD5BA3FAFD42" box="[648,764,660,684]" pageId="43" pageNumber="128">D.Z. Dong</collectorName>
&amp;
<collectorName id="26F3DFCCFFF4FF9EFCC6FD5BA27EFD42" box="[802,888,660,684]" pageId="43" pageNumber="128">G. Tang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF4FF9EFBE0FD5BA20AFD22" ID-GBIF-Occurrence="4899905522" collectingDate="2004-10-25" collectionCode="CAS, IOZ" collectorName="Q. B. Hou &amp; H. B. Liang" country="China" county="Gongshan County" elevation="1500" latitude="27.88111" location="Dulongjiang Township" longLatPrecision="1" longitude="98.34063" pageId="43" pageNumber="128" specimenCode="CASENT 1016748" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF4FF9EFBE0FD5BA561FD42" box="[1028,1127,660,684]" count="1" pageId="43" pageNumber="128" type="male">one male</specimenCount>
and
<specimenCount id="9D007193FFF4FF9EFF9FFD7BA005FD22" box="[123,259,692,716]" count="4" pageId="43" pageNumber="128" type="female">four females</specimenCount>
(
<collectionCode id="ED1722DFFFF4FF9EFEF2FD7BA04CFD22" box="[278,330,692,716]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="43" pageNumber="128" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1722DFFFF4FF9EFEBEFD7BA080FD22" box="[346,390,692,716]" pageId="43" pageNumber="128">IOZ</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF4FF9EFDE6FD7BA3D6FD22" box="[514,720,692,716]" pageId="43" pageNumber="128">CASENT 1016748</specimenCode>
” and
</materialsCitation>
<materialsCitation id="3B6EB047FFF4FF9EFCF1FD7BA51FFD22" ID-GBIF-Occurrence="4899905510" box="[789,1049,692,716]" collectingDate="2004-10-25" collectionCode="CAS, IOZ" collectorName="Q. B. Hou &amp; H. B. Liang" country="China" county="Gongshan County" elevation="1500" latitude="27.88111" location="Dulongjiang Township" longLatPrecision="1" longitude="98.34063" pageId="43" pageNumber="128" specimenCode="CASENT 1016744" specimenCount="1" specimenCount-female="4" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
“
<specimenCode id="DBA01261FFF4FF9EFCC5FD7BA2E9FD22" box="[801,1007,692,716]" pageId="43" pageNumber="128">CASENT 1016744</specimenCode>
” to
</materialsCitation>
<materialsCitation id="3B6EB047FFF4FF9EFBC6FD7BA37FFCC0" ID-GBIF-Occurrence="4899905537" collectingDate="2004-10-25" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; Q. B. Hou &amp; H. B. Liang" country="China" county="Gongshan County" elevation="1500" latitude="27.88111" location="Dulongjiang Township" longLatPrecision="1" longitude="98.34063" pageId="43" pageNumber="128" specimenCode="CASENT 1016747" specimenCount="1" specimenCount-female="4" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
“
<specimenCode id="DBA01261FFF4FF9EFBCAFD7BA1DEFD03" pageId="43" pageNumber="128">CASENT 1016747</specimenCode>
”, respectively/ “
<collectingCountry id="F311FA8AFFF4FF9EFE75FD1AA0E3FD03" box="[401,485,725,749]" name="China" pageId="43" pageNumber="128">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF4FF9EFE15FD1AA344FD02" box="[497,578,725,748]" country="China" name="Yunnan" pageId="43" pageNumber="128">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF4FF9EFDB4FD1AA209FD03" box="[592,783,725,749]" pageId="43" pageNumber="128">Gongshan County</collectingCounty>
,
<location id="8ED9ECC1FFF4FF9EFCF8FD1AA516FD03" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD:8ED9ECC1FFF4FF9EFCF8FD1AA516FD03" box="[796,1040,725,749]" country="China" county="Gongshan County" latitude="27.88111" longLatPrecision="1" longitude="98.34063" name="Dulongjiang Township" pageId="43" pageNumber="128" stateProvince="Yunnan">Dulongjiang Township</location>
,
<locationDeviation id="9A0BA338FFF4FF9EFBF8FD1AA1ECFCE3" location="Kongdang" pageId="43" pageNumber="128">
<quantity id="4CFE17FFFFF4FF9EFBF8FD1AA559FD03" box="[1052,1119,725,749]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="128" unit="km" value="0.5">0.5km</quantity>
N of Kongdang
</locationDeviation>
,
<quantity id="4CFE17FFFFF4FF9EFF1DFD3AA045FCE3" box="[249,323,757,781]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="128" unit="m" value="1500.0">
<elevation id="002B5D29FFF4FF9EFF1DFD3AA045FCE3" box="[249,323,757,781]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="128" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<geoCoordinate id="EE32DCDDFFF4FF9EFEB6FD39A0D6FCE0" box="[338,464,758,782]" degrees="27. 88111" direction="north" orientation="latitude" pageId="43" pageNumber="128" precision="1" value="27.88111">N27.88111º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF4FF9EFE33FD39A367FCE0" box="[471,609,757,782]" degrees="098. 34063" direction="east" orientation="longitude" pageId="43" pageNumber="128" precision="1" value="98.34063">E098.34063º</geoCoordinate>
,
<date id="FFB89CDAFFF4FF9EFD94FD3AA221FCE0" box="[624,807,757,782]" pageId="43" pageNumber="128" value="2004-10-25">
<collectingDate id="EFFC6532FFF4FF9EFD94FD3AA221FCE0" box="[624,807,757,782]" pageId="43" pageNumber="128" value="2004-10-25">25 October 2004</collectingDate>
</date>
”/ “Stop #DHK-2004-057D,
<collectorName id="26F3DFCCFFF4FF9EFB85FD39A1F2FCC0" pageId="43" pageNumber="128">D.H. Kavanaugh</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFEE1FCD9A076FCC3" box="[261,368,790,814]" pageId="43" pageNumber="128">Q.B. Hou</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFE64FCD9A0FDFCC0" box="[384,507,790,814]" pageId="43" pageNumber="128">H.B. Liang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF4FF9EFD6EFCD9A372FC7E" ID-GBIF-Occurrence="4899905415" collectingDate="2006-09-03" collectionCode="CAS" collectorName="D. H. Kavanaugh &amp; Y. Liu &amp; S. Z Ma &amp; P. Hu" country="China" county="Gongshan County" elevation="1615" latitude="27.68804" location="Maku village" longLatPrecision="1" longitude="98.30758" municipality="Dulongjiang Township" pageId="43" pageNumber="128" specimenCode="CASENT 1025538" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF4FF9EFD6EFCD9A3E9FCC0" box="[650,751,790,814]" count="1" pageId="43" pageNumber="128" type="male">one male</specimenCount>
(
<collectionCode id="ED1722DFFFF4FF9EFCE6FCD9A230FCC0" box="[770,822,790,814]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="43" pageNumber="128" type="Museum">CAS</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF4FF9EFC57FCD9A584FCC0" box="[947,1154,790,814]" pageId="43" pageNumber="128">CASENT 1025538</specimenCode>
”/ “
<collectingCountry id="F311FA8AFFF4FF9EFF63FCF8A1DDFCA1" box="[135,219,823,847]" name="China" pageId="43" pageNumber="128">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF4FF9EFF03FCF8A03EFCA0" box="[231,312,823,846]" country="China" name="Yunnan" pageId="43" pageNumber="128">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF4FF9EFEA1FCF9A302FCA1" box="[325,516,822,847]" pageId="43" pageNumber="128">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF4FF9EFDF4FCF9A202FCA0" box="[528,772,822,846]" pageId="43" pageNumber="128">Dulongjiang Township</collectingMunicipality>
,
<location id="8ED9ECC1FFF4FF9EFCF5FCF9A298FCA0" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD:8ED9ECC1FFF4FF9EFCF5FCF9A298FCA0" box="[785,926,822,846]" country="China" county="Gongshan County" latitude="27.68804" longLatPrecision="1" longitude="98.30758" municipality="Dulongjiang Township" name="Maku village" pageId="43" pageNumber="128" stateProvince="Yunnan">Maku village</location>
along old road through village,
<geoCoordinate id="EE32DCDDFFF4FF9EFF3DFC97A05FFC81" box="[217,345,855,879]" degrees="27. 68804" direction="north" orientation="latitude" pageId="43" pageNumber="128" precision="1" value="27.68804">N27.68804º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF4FF9EFE84FC97A0ECFC81" box="[352,490,855,879]" degrees="098. 30758" direction="east" orientation="longitude" pageId="43" pageNumber="128" precision="1" value="98.30758">E098.30758º</geoCoordinate>
”/ “
<quantity id="4CFE17FFFFF4FF9EFDFDFC98A365FC81" box="[537,611,855,879]" metricMagnitude="3" metricUnit="m" metricValue="1.6149999999999998" pageId="43" pageNumber="128" unit="m" value="1615.0">
<elevation id="002B5D29FFF4FF9EFDFDFC98A365FC81" box="[537,611,855,879]" metricMagnitude="3" metricUnit="m" metricValue="1.6149999999999998" pageId="43" pageNumber="128" unit="m" value="1615.0">1615m</elevation>
</quantity>
,
<date id="FFB89CDAFFF4FF9EFD9DFC98A24AFC81" box="[633,844,855,879]" pageId="43" pageNumber="128" value="2006-09-03">
<collectingDate id="EFFC6532FFF4FF9EFD9DFC98A24AFC81" box="[633,844,855,879]" pageId="43" pageNumber="128" value="2006-09-03">3 September 2006</collectingDate>
</date>
, Stop DHK-2006-120,
<collectorName id="26F3DFCCFFF4FF9EFB85FC97A1F2FC7E" pageId="43" pageNumber="128">D.H. Kavanaugh</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFEE4FCB7A044FC7E" box="[256,322,888,912]" pageId="43" pageNumber="128">Y. Liu</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFEABFCB7A099FC61" box="[335,415,888,912]" pageId="43" pageNumber="128">S.Z Ma</collectorName>
&amp;
<collectorName id="26F3DFCCFFF4FF9EFE25FCB7A0FDFC61" box="[449,507,888,911]" pageId="43" pageNumber="128">P. Hu</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF4FF9EFD66FCB7A07BFC5E" ID-GBIF-Occurrence="4899905369" collectingDate="1998-10-14" collectingDateMax="1998-10-15" collectingDateMin="1998-10-14" collectionCode="CAS, IOZ" collectorName="C. E. Griswold &amp; C. Ferraris" country="China" county="Gaoligongshan Mountains" elevation="1500" location="Gangfang Sancha Lukou" municipality="Nujiang Prefecture" pageId="43" pageNumber="128" specimenCode="CASENT 1000600" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF4FF9EFD66FCB7A3F5FC7E" box="[642,755,888,912]" count="4" pageId="43" pageNumber="128" type="male">four males</specimenCount>
and
<specimenCount id="9D007193FFF4FF9EFCC2FCB7A2B3FC7E" box="[806,949,888,912]" count="3" pageId="43" pageNumber="128" type="female">three females</specimenCount>
(
<collectionCode id="ED1722DFFFF4FF9EFC20FCB7A2FEFC7E" box="[964,1016,888,912]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="43" pageNumber="128" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1722DFFFF4FF9EFBE1FCB7A537FC7E" box="[1029,1073,888,912]" pageId="43" pageNumber="128">IOZ</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF4FF9EFF63FC57A055FC5E" box="[135,339,920,944]" pageId="43" pageNumber="128">CASENT 1000600</specimenCode>
” to
</materialsCitation>
<materialsCitation id="3B6EB047FFF4FF9EFE61FC57A39EFC5E" ID-GBIF-Occurrence="4899905408" box="[389,664,920,944]" collectingDate="1998-10-14" collectingDateMax="1998-10-15" collectingDateMin="1998-10-14" collectionCode="CAS, IOZ" collectorName="C. E. Griswold &amp; C. Ferraris" country="China" county="Gaoligongshan Mountains" elevation="1500" location="Gangfang Sancha Lukou" municipality="Nujiang Prefecture" pageId="43" pageNumber="128" specimenCode="CASENT 1000603" specimenCount="1" specimenCount-female="3" specimenCount-male="4" stateProvince="Yunnan" typeStatus="paratype">
“
<specimenCode id="DBA01261FFF4FF9EFE74FC57A35BFC5E" box="[400,605,920,944]" pageId="43" pageNumber="128">CASENT 1000603</specimenCode>
” and
</materialsCitation>
<materialsCitation id="3B6EB047FFF4FF9EFD44FC57A2A4FC5E" ID-GBIF-Occurrence="4899905399" box="[672,930,920,944]" collectingDate="1998-10-14" collectingDateMax="1998-10-15" collectingDateMin="1998-10-14" collectionCode="CAS, IOZ" collectorName="C. E. Griswold &amp; C. Ferraris" country="China" county="Gaoligongshan Mountains" elevation="1500" location="Gangfang Sancha Lukou" municipality="Nujiang Prefecture" pageId="43" pageNumber="128" specimenCode="CASENT 1000604" specimenCount="1" specimenCount-female="3" specimenCount-male="4" stateProvince="Yunnan" typeStatus="paratype">
“
<specimenCode id="DBA01261FFF4FF9EFD48FC57A27FFC5E" box="[684,889,920,944]" pageId="43" pageNumber="128">CASENT 1000604</specimenCode>
” to
</materialsCitation>
<materialsCitation id="3B6EB047FFF4FF9EFC4EFC57A228FBFC" ID-GBIF-Occurrence="4899905502" collectingDate="1998-10-14" collectingDateMax="1998-10-15" collectingDateMin="1998-10-14" collectionCode="CAS, IOZ" collectorName=". H. Kavanaugh &amp; C. E. Griswold &amp; C. Ferraris &amp; C. - L. Long" country="China" county="Gaoligongshan Mountains" elevation="1500" location="Gangfang Sancha Lukou" longitude="98.575" municipality="Nujiang Prefecture" pageId="43" pageNumber="128" specimenCode="CASENT 1000606" specimenCount="1" specimenCount-female="3" specimenCount-male="4" stateProvince="Yunnan Province" typeStatus="paratype">
“
<specimenCode id="DBA01261FFF4FF9EFC52FC57A584FC5E" box="[950,1154,920,944]" pageId="43" pageNumber="128">CASENT 1000606</specimenCode>
”, respectively/ “
<collectingCountry id="F311FA8AFFF4FF9EFEFDFC76A06BFC3F" box="[281,365,953,977]" name="China" pageId="43" pageNumber="128">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF4FF9EFE98FC76A331FC3F" box="[380,567,953,977]" country="China" name="Yunnan" pageId="43" pageNumber="128">Yunnan Province</collectingRegion>
,
<collectingCounty id="62D8C296FFF4FF9EFDA3FC76A262FC3F" box="[583,868,953,977]" pageId="43" pageNumber="128">Gaoligongshan Mountains</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF4FF9EFC91FC76A545FC3F" box="[885,1091,953,977]" pageId="43" pageNumber="128">Nujiang Prefecture</collectingMunicipality>
,
<location id="8ED9ECC1FFF4FF9EFBB7FC76A057FC1F" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD:8ED9ECC1FFF4FF9EFBB7FC76A057FC1F" country="China" county="Gaoligongshan Mountains" longitude="98.575" municipality="Nujiang Prefecture" name="Gangfang Sancha Lukou" pageId="43" pageNumber="128" stateProvince="Yunnan Province">Gangfang Sancha Lukou</location>
, 26º073´N/98º34.5´E”/ “
<quantity id="4CFE17FFFFF4FF9EFD80FC16A3A8FC1F" box="[612,686,985,1009]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="128" unit="m" value="1500.0">
<elevation id="002B5D29FFF4FF9EFD80FC16A3A8FC1F" box="[612,686,985,1009]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="128" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="FFB89CDAFFF4FF9EFD5AFC16A29DFC1F" box="[702,923,985,1009]" pageId="43" pageNumber="128" value="1998-10-14" valueMax="1998-10-15" valueMin="1998-10-14">
<collectingDate id="EFFC6532FFF4FF9EFD5AFC16A29DFC1F" box="[702,923,985,1009]" pageId="43" pageNumber="128" value="1998-10-14" valueMax="1998-10-15" valueMin="1998-10-14">14-15 October 1998</collectingDate>
</date>
, Stop #98-117C, D
<collectorName id="26F3DFCCFFF4FF9EFB91FC15A1F2FBFC" pageId="43" pageNumber="128">.H. Kavanaugh</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFEE5FC35A09CFBFC" box="[257,410,1018,1042]" pageId="43" pageNumber="128">C.E. Griswold</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFE4CFC35A31CFBFC" box="[424,538,1018,1042]" pageId="43" pageNumber="128">C. Ferraris</collectorName>
&amp;
<collectorName id="26F3DFCCFFF4FF9EFDD8FC35A3B5FBFC" box="[572,691,1018,1042]" pageId="43" pageNumber="128">C.-L. Long</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF4FF9EFCD8FC35A38DFBBC" ID-GBIF-Occurrence="4899905338" collectingDate="2020-09-17" collectionCode="IOZ" collectorName="Liang H. B &amp; Xu Y." country="China" elevation="2023" latitude="29.6565" location="Medog" longLatPrecision="1" longitude="95.49348" municipality="Medog" pageId="43" pageNumber="128" specimenCount="1" specimenCount-male="1" stateProvince="Tibet" typeStatus="paratype">
<specimenCount id="9D007193FFF4FF9EFCD8FC35A29BFBFC" box="[828,925,1018,1042]" count="1" pageId="43" pageNumber="128" type="male">one male</specimenCount>
(
<collectionCode id="ED1722DFFFF4FF9EFC48FC35A2DEFBFC" box="[940,984,1018,1042]" pageId="43" pageNumber="128">IOZ</collectionCode>
) labeled: “
<collectingCountry id="F311FA8AFFF4FF9EFBABFC35A589FBFC" box="[1103,1167,1018,1042]" name="China" pageId="43" pageNumber="128">China</collectingCountry>
,
<collectingRegion id="49C274F8FFF4FF9EFF9FFBD5A1B4FBDC" box="[123,178,1050,1074]" country="China" name="Xizang" pageId="43" pageNumber="128">Tibet</collectingRegion>
,
<collectingMunicipality id="6BDD2060FFF4FF9EFF5BFBD5A00DFBDC" box="[191,267,1050,1074]" pageId="43" pageNumber="128">Medog</collectingMunicipality>
,
<quantity id="4CFE17FFFFF4FF9EFEF3FBD4A061FBDC" box="[279,359,1050,1075]" metricMagnitude="4" metricUnit="m" metricValue="8.15" pageId="43" pageNumber="128" unit="km" value="81.5">
<locationDeviation id="9A0BA338FFF4FF9EFEF3FBD4A061FBDC" box="[279,359,1050,1075]" metricMagnitude="4" metricUnit="m" metricValue="8.15" pageId="43" pageNumber="128" unit="km" value="81.5">81.5km</locationDeviation>
</quantity>
on
<location id="8ED9ECC1FFF4FF9EFE6AFBD5A0DCFBDC" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD:8ED9ECC1FFF4FF9EFE6AFBD5A0DCFBDC" box="[398,474,1050,1074]" country="China" latitude="29.6565" longLatPrecision="1" longitude="95.49348" municipality="Medog" name="Medog" pageId="43" pageNumber="128" stateProvince="Tibet">Medog</location>
road,
<geoCoordinate id="EE32DCDDFFF4FF9EFDFFFBD4A395FBDC" box="[539,659,1050,1075]" degrees="29. 65650" direction="north" orientation="latitude" pageId="43" pageNumber="128" precision="1" value="29.6565">N29.65650</geoCoordinate>
,
<geoCoordinate id="EE32DCDDFFF4FF9EFD7BFBD4A215FBDD" box="[671,787,1050,1075]" degrees="95. 49348" direction="east" orientation="longitude" pageId="43" pageNumber="128" precision="1" value="95.49348">E95.49348</geoCoordinate>
”/ “
<quantity id="4CFE17FFFFF4FF9EFCDCFBD4A284FBDC" box="[824,898,1051,1075]" metricMagnitude="3" metricUnit="m" metricValue="2.023" pageId="43" pageNumber="128" unit="m" value="2023.0">
<elevation id="002B5D29FFF4FF9EFCDCFBD4A284FBDC" box="[824,898,1051,1075]" metricMagnitude="3" metricUnit="m" metricValue="2.023" pageId="43" pageNumber="128" unit="m" value="2023.0">2023m</elevation>
</quantity>
,
<date id="FFB89CDAFFF4FF9EFC6BFBD4A2FCFBDD" box="[911,1018,1051,1075]" pageId="43" pageNumber="128" value="2020-09-17">
<collectingDate id="EFFC6532FFF4FF9EFC6BFBD4A2FCFBDD" box="[911,1018,1051,1075]" pageId="43" pageNumber="128" value="2020-09-17">2020.9.17</collectingDate>
</date>
N,
<collectorName id="26F3DFCCFFF4FF9EFBFBFBD5A589FBDC" box="[1055,1167,1050,1074]" pageId="43" pageNumber="128">Liang H.B</collectorName>
,
<collectorName id="26F3DFCCFFF4FF9EFF9FFBF4A1C6FBBC" box="[123,192,1083,1106]" pageId="43" pageNumber="128">Xu Y.</collectorName>
coll., Inst. Zool., Chinese Acad. Sci.”.
</materialsCitation>
All
<typeStatus id="54BD04B8FFF4FF9EFD2DFBF4A229FBBD" box="[713,815,1083,1107]" pageId="43" pageNumber="128" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54BD04B8FFF4FF9EFF63FB93A00DFB9A" box="[135,267,1116,1140]" pageId="43" pageNumber="128" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C06C199FFF4FF9EFEF7FB93A232FB9D" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[275,820,1115,1140]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="128" phylum="Arthropoda" rank="species" species="holomarginatum" status="sp. nov.">
<emphasis id="B9726608FFF4FF9EFEF7FB93A349FB9D" box="[275,591,1115,1139]" italics="true" pageId="43" pageNumber="128">Broscosoma holomarginatum</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A241DB73FFF4FF9EFCA7FB93A291FB9D" box="[835,919,1116,1139]" pageId="43" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [yellow label].
</paragraph>
<paragraph id="8BB9BA1AFFF4FF9EFF4FFB52A555FB5A" blockId="43.[123,1174,301,1726]" box="[171,1107,1180,1205]" pageId="43" pageNumber="128">
<materialsCitation id="3B6EB047FFF4FF9EFF4FFB52A54AFB5B" ID-GBIF-Occurrence="4899905329" box="[171,1100,1180,1205]" country="China" county="Gongshan County" location="Kongdang" municipality="Dulongjiang Township" pageId="43" pageNumber="128" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B9726608FFF4FF9EFF4FFB52A03BFB5A" bold="true" box="[171,317,1180,1205]" pageId="43" pageNumber="128">Type locality</emphasis>
.
<collectingCountry id="F311FA8AFFF4FF9EFEAFFB52A099FB5B" box="[331,415,1181,1205]" name="China" pageId="43" pageNumber="128">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF4FF9EFE48FB52A0FBFB5A" box="[428,509,1181,1204]" country="China" name="Yunnan" pageId="43" pageNumber="128">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF4FF9EFDE8FB52A3CAFB5B" box="[524,716,1181,1205]" pageId="43" pageNumber="128">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF4FF9EFD3EFB52A2C9FB5B" box="[730,975,1181,1205]" pageId="43" pageNumber="128">Dulongjiang Township</collectingMunicipality>
,
<location id="8ED9ECC1FFF4FF9EFC39FB52A54AFB5B" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF4FF9AFF9FFF14A2FAFEBD:8ED9ECC1FFF4FF9EFC39FB52A54AFB5B" box="[989,1100,1181,1205]" country="China" county="Gongshan County" municipality="Dulongjiang Township" name="Kongdang" pageId="43" pageNumber="128" stateProvince="Yunnan">Kongdang</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9BA1AFFF4FF9EFF4FFB72A3DAFAD8" blockId="43.[123,1174,301,1726]" pageId="43" pageNumber="128">
<emphasis id="B9726608FFF4FF9EFF4FFB72A0EAFB3B" bold="true" box="[171,492,1213,1237]" pageId="43" pageNumber="128">Derivation of species name.</emphasis>
The species epithet,
<taxonomicName id="4C06C199FFF4FF9EFD3EFB72A28DFB3B" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[730,907,1213,1237]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="128" phylum="Arthropoda" rank="species" species="holomarginatum">
<emphasis id="B9726608FFF4FF9EFD3EFB72A28DFB3B" box="[730,907,1213,1237]" italics="true" pageId="43" pageNumber="128">holomarginatum</emphasis>
</taxonomicName>
, is an adjective derived from the Greek word,
<emphasis id="B9726608FFF4FF9EFE94FB11A0ACFB18" box="[368,426,1246,1270]" italics="true" pageId="43" pageNumber="128">holos</emphasis>
, meaning whole or entire, and the Latin word,
<emphasis id="B9726608FFF4FF9EFC56FB11A52DFB1B" box="[946,1067,1246,1269]" italics="true" pageId="43" pageNumber="128">marginatus</emphasis>
, meaning bordered or enclosed with a border. The name refers the complete lateral margination (or lateral bead) of the pronotum found in members of this species.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF4FF9EFF4FFAF0A3C9F9D5" pageId="43" pageNumber="128" type="diagnosis">
<paragraph id="8BB9BA1AFFF4FF9EFF4FFAF0A3C9F9D5" blockId="43.[123,1174,301,1726]" pageId="43" pageNumber="128">
<emphasis id="B9726608FFF4FF9EFF4FFAF0A019FAB9" bold="true" box="[171,287,1343,1367]" pageId="43" pageNumber="128">Diagnosis.</emphasis>
Adults of this species (
<figureCitation id="133DA69FFFF4FF9EFDC5FAF0A37CFAB9" box="[545,634,1343,1367]" captionStart="FIGURE" captionStartId="44.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1568]" captionTargetId="figure-56@44.[120,1176,216,1568]" captionTargetPageId="44" captionText="FIGURE 24. Broscosoma holomarginatum sp. nov. a. Habitus (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067401" httpUri="https://zenodo.org/record/11067401/files/figure.png" pageId="43" pageNumber="128">Fig. 24a</figureCitation>
) can be distinguished from those of other species in the region by the following combination of character states: size moderate, BL = 8.1 to 9.
<quantity id="4CFE17FFFFF4FF9EFBB4FAAFA58BFA99" box="[1104,1165,1376,1400]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="128" unit="mm" value="5.0">5 mm</quantity>
; dorsal surface with distinct blue or blue-green luster, legs pale rufous; antennomeres 3 and 4 glabrous except for apical whorl of setae; pronotum with lateral margination (lateral bead) complete between apical and basal margins (
<figureCitation id="133DA69FFFF4FF9EFDD3FA0DA385FA37" box="[567,643,1474,1498]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="43" pageNumber="128">Fig. 9a</figureCitation>
), basolateral setae absent; elytral silhouette subparallel to subovoid, with humeri distinct, elytral microsculpture distinct, comprised of isodiametric to slightly longitudinally stretched sculpticells; male protarsi with tarsomeres 1 to 3 with ventral pads of adhesive setae; male genitalia as in
<figureCitation id="133DA69FFFF4FF9EFD98F9ECA3CEF9D5" box="[636,712,1571,1595]" captionStart="FIGURE" captionStartId="45.[171,183,939,958]" captionTargetBox="[137,1161,228,920]" captionTargetId="figure-1604@45.[120,1176,215,935]" captionTargetPageId="45" captionText="FIGURE 25. Male genitalia, Broscosoma holomarginatum sp. nov. (CASENT1016743; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067405" httpUri="https://zenodo.org/record/11067405/files/figure.png" pageId="43" pageNumber="128">Fig. 25</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF4FF9BFF4FF98BA0E3FC99" lastPageId="46" lastPageNumber="131" pageId="43" pageNumber="128" type="description">
<paragraph id="8BB9BA1AFFF4FF98FF4FF98BA076FBB1" blockId="43.[123,1174,301,1726]" lastBlockId="45.[123,1174,1062,1725]" lastPageId="45" lastPageNumber="130" pageId="43" pageNumber="128">
<emphasis id="B9726608FFF4FF9EFF4FF98BA030F9B2" bold="true" box="[171,310,1604,1628]" pageId="43" pageNumber="128">Description.</emphasis>
<figureCitation id="133DA69FFFF4FF9EFED9F98BA093F9B5" box="[317,405,1604,1628]" captionStart="FIGURE" captionStartId="44.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1568]" captionTargetId="figure-56@44.[120,1176,216,1568]" captionTargetPageId="44" captionText="FIGURE 24. Broscosoma holomarginatum sp. nov. a. Habitus (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067401" httpUri="https://zenodo.org/record/11067401/files/figure.png" pageId="43" pageNumber="128">Fig. 24a</figureCitation>
. Size small to medium, BL male = 8.5–9.
<quantity id="4CFE17FFFFF4FF9EFCBEF98BA29EF9B5" box="[858,920,1604,1628]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="43" pageNumber="128" unit="mm" value="3.0">3 mm</quantity>
, female = 8.1–9.
<quantity id="4CFE17FFFFF4FF9EFBB4F98BA588F9B5" box="[1104,1166,1604,1628]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="128" unit="mm" value="5.0">5 mm</quantity>
, ratio EL/PL male = 2.1–2.3, female =2.1–2.3. Body color black or dark piceous, last visible abdominal sternite rufous brown, or rufous basally graded to pale yellow apically, antennae with all antennomeres pale rufous, femora, tibiae and tarsi pale rufous. Head with green, pronotum with blue or bluish-green, and elytra with marked blue metallic reflection, venter with metallic reflection absent or only faintly present.
</paragraph>
<caption id="DF79EA92FFF3FF99FF4FF9FCA3FBF997" ID-DOI="http://doi.org/10.5281/zenodo.11067401" ID-Zenodo-Dep="11067401" httpUri="https://zenodo.org/record/11067401/files/figure.png" pageId="44" pageNumber="129" startId="44.[171,183,1587,1606]" targetBox="[120,1176,216,1568]" targetPageId="44" targetType="figure">
<paragraph id="8BB9BA1AFFF3FF99FF4FF9FCA3FBF997" blockId="44.[123,1174,1587,1657]" pageId="44" pageNumber="129">
<smallCapsWord id="8D5F2CC6FFF3FF99FF4FF9FCA1EEF9AA" baselines="1601,1600" box="[171,232,1587,1606]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="44" pageNumber="129">FIGURE</smallCapsWord>
24.
<taxonomicName id="4C06C199FFF3FF99FEEFF9FCA300F9A8" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[267,518,1587,1606]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="129" phylum="Arthropoda" rank="species" species="holomarginatum" status="sp. nov.">
<emphasis id="B9726608FFF3FF99FEEFF9FCA300F9A8" box="[267,518,1587,1606]" italics="true" pageId="44" pageNumber="129">Broscosoma holomarginatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFF3FF99FDEFF9FCA34DF9A8" box="[523,587,1587,1606]" pageId="44" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
a. Habitus (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<caption id="DF79EA92FFF2FF98FF4FFC64A2A2FC1F" ID-DOI="http://doi.org/10.5281/zenodo.11067405" ID-Zenodo-Dep="11067405" httpUri="https://zenodo.org/record/11067405/files/figure.png" pageId="45" pageNumber="130" startId="45.[171,183,939,958]" targetBox="[137,1161,228,920]" targetPageId="45" targetType="figure">
<paragraph id="8BB9BA1AFFF2FF98FF4FFC64A2A2FC1F" blockId="45.[123,1173,938,1009]" pageId="45" pageNumber="130">
<smallCapsWord id="8D5F2CC6FFF2FF98FF4FFC64A1EEFC52" baselines="953,952" box="[171,232,939,958]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="45" pageNumber="130">FIGURE</smallCapsWord>
25. Male genitalia,
<taxonomicName id="4C06C199FFF2FF98FE42FC64A3A0FC53" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[422,678,938,957]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="130" phylum="Arthropoda" rank="species" species="holomarginatum" status="sp. nov.">
<emphasis id="B9726608FFF2FF98FE42FC64A3A0FC53" box="[422,678,938,957]" italics="true" pageId="45" pageNumber="130">Broscosoma holomarginatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFF2FF98FD55FC64A3F1FC50" box="[689,759,939,958]" pageId="45" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
(CASENT1016743; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB9BA1AFFF2FF98FF4FFBA6A1D4FA3F" blockId="45.[123,1174,1062,1725]" pageId="45" pageNumber="130">
Head.
<figureCitation id="133DA69FFFF2FF98FF11FBA6A042FB6F" box="[245,324,1129,1153]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="45" pageNumber="130">Fig. 6a</figureCitation>
. Eyes large, convex, their diameter slightly greater than length of tempora. Frontal furrows deeply impressed, narrow, arcuate, slightly convergent anteriorly and posteriorly, medium length, extended posteriorly to middle of eyes, rugulose and sparsely and coarsely punctate. Vertex with post-temporal transverse sulcus shallow, densely and coarsely punctate. Tempora straight, not convex. Genal ridges present, restricted to head region distinctly behind post-temporal constriction. Clypeus with one pair of setae. Supraorbital setae present, one pair. Eustipes of maxilla with two setae, the dorsobasal seta distinctly more than half as long as ventrobasal seta.
<collectingRegion id="49C274F8FFF2FF98FF9FFA9AA1D3FA82" box="[123,213,1365,1388]" country="Myanmar" name="Chin" pageId="45" pageNumber="130">Mentum</collectingRegion>
with tooth present, simple, paramedial region shallowly foveate, one pair of mental setae present. Glossal sclerite (ligula) with one pair of setae. Submentum with one or two pairs of setae. Gula without transverse grooves. Antennomeres 3 and 4 without pubescence, ratio A3/A5 = 1.2–1.3.
</paragraph>
<paragraph id="8BB9BA1AFFF2FF9BFF4FFA13A1C0FEFD" blockId="45.[123,1174,1062,1725]" lastBlockId="46.[123,1174,219,1726]" lastPageId="46" lastPageNumber="131" pageId="45" pageNumber="130">
Pronotum.
<figureCitation id="133DA69FFFF2FF98FEC5FA14A06CFA1D" box="[289,362,1499,1523]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="45" pageNumber="130">Fig. 6a</figureCitation>
. Globose anterior to sub-basal constriction with base distinctly pedunculate, widest at middle of discal region, PL/PW = 1.1, disk markedly convex; apical margination absent; lateral margination present, narrow, distinctly delineated throughout pronotal length (Fig. FX09a); basal margination absent; anterior transverse impression shallow, broad and vaguely delineated, coarsely punctate both at bottom and on slopes of depression; median longitudinal impression moderately impressed; posterior transverse impression deep, sharply delineated; anterior region moderately to densely and coarsely punctate; pronotal base coarsely punctate and rugulose; one pair of midlateral pronotal setae present at mid-length of discal region; basolateral pronotal setae absent.
</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FFED4A518FD7B" blockId="46.[123,1174,219,1726]" pageId="46" pageNumber="131">Elytra. Elytral silhouette subovoid, slightly narrowed in basal half, widest at or slightly posterior to middle, ratio EL/EW in males = 1.5, in female = EL/EW 1.6, greatest elevation above lateral margin (in lateral view) posterior to middle; humeri distinctly present, roundly obtuse; lateral margins moderately arcuate and nearly straight in anterior half sub-basally; parascutellar striole present, short, distally merged in a distinct jog with free base of stria 1. Eight elytral striae present; stria 1 deeply and sharply impressed, striae 2 to 8 indistinct, obliterated or represented by extremely fine and shallow linear depressions, successively less evident laterally and apically, striae 8 merged with lateral groove throughout its length or nearly so; stria 1 moderately punctate at least in basal half, striae 2 to 8 densely and moderately punctate, but with punctures increasingly shallower and sparser laterally and apically. Elytral microsculpture comprised of distinct, deeply to shallowly impressed isodiameteric sculpticells. Parascutellar seta present at base of stria 2, discal setae absent, umbilicate series comprised of one post-humeral and two preapical setae.</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FFD51A31FFD58" blockId="46.[123,1174,219,1726]" box="[171,537,670,694]" pageId="46" pageNumber="131">Hindwings. Full-sized, functional.</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FFD71A213FD38" blockId="46.[123,1174,219,1726]" box="[171,789,702,727]" pageId="46" pageNumber="131">Thoracic venter. Metepisternum ratio ML/MW = 2.2–2.3.</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FFD10A2E8FCD9" blockId="46.[123,1174,219,1726]" pageId="46" pageNumber="131">Legs. Hind trochanter asetose. Tarsomeres without dorsal pubescence; tarsomere 5 with two or more pairs of setae ventrally; male protarsi with pads of adhesive setae on tarsomeres 1 to 3 ventrally, male mesotarsi with pads of adhesive setae on tarsomeres 1 and 2 ventrally.</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FFCF0A555FCB9" blockId="46.[123,1174,219,1726]" box="[171,1107,831,855]" pageId="46" pageNumber="131">Abdomen. Sternite VI of both male and female with one pair of apicoparamedial setae.</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FFC90A0E3FC99" blockId="46.[123,1174,219,1726]" box="[171,485,863,888]" pageId="46" pageNumber="131">
Male genitalia. As in
<figureCitation id="133DA69FFFF1FF9BFE76FC90A0D9FC99" box="[402,479,863,887]" captionStart="FIGURE" captionStartId="45.[171,183,939,958]" captionTargetBox="[137,1161,228,920]" captionTargetId="figure-1604@45.[120,1176,215,935]" captionTargetPageId="45" captionText="FIGURE 25. Male genitalia, Broscosoma holomarginatum sp. nov. (CASENT1016743; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067405" httpUri="https://zenodo.org/record/11067405/files/figure.png" pageId="46" pageNumber="131">Fig. 25</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF1FF9AFF4FFC4FA2FAFEBD" lastPageId="47" lastPageNumber="132" pageId="46" pageNumber="131" type="discussion">
<paragraph id="8BB9BA1AFFF1FF9BFF4FFC4FA394FB77" blockId="46.[123,1174,219,1726]" pageId="46" pageNumber="131">
<emphasis id="B9726608FFF1FF9BFF4FFC4FA02BFC76" bold="true" box="[171,301,896,920]" pageId="46" pageNumber="131">Comments.</emphasis>
This is apparently the only species in genus
<taxonomicName id="4C06C199FFF1FF9BFCE9FC4EA296FC76" box="[781,912,897,920]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B9726608FFF1FF9BFCE9FC4EA296FC76" box="[781,912,897,920]" italics="true" pageId="46" pageNumber="131">Broscosoma</emphasis>
</taxonomicName>
in which members have the pronotum with complete and well-defined lateral margination (Fig, 9a). Lateral margination is also present
<taxonomicName id="4C06C199FFF1FF9BFEE4FC0EA071FC36" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[256,375,960,984]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="131" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFF1FF9BFEE4FC0EA071FC36" box="[256,375,960,984]" italics="true" pageId="46" pageNumber="131">B. resbecqi</emphasis>
</taxonomicName>
, in which it is nearly complete but interrupted in the basal one-third of the discal region (Fig, 9b), and in most specimens of
<taxonomicName id="4C06C199FFF1FF9BFD66FC2EA3D8FC16" authorityName="Putzeys. We" authorityYear="1877" box="[642,734,992,1016]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="131" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFF1FF9BFD66FC2EA3D8FC16" box="[642,734,992,1016]" italics="true" pageId="46" pageNumber="131">B. ribbei</emphasis>
</taxonomicName>
, in which it is present in the area of insertion of the midlateral seta and extended slightly anteriorly and posteriorly in a some specimens (Fig, 9b). Members of both of these species have the dorsum with vivid green metallic reflection, whereas
<taxonomicName id="4C06C199FFF1FF9BFF3EFB8DA0ACFBB7" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[218,426,1089,1113]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="131" phylum="Arthropoda" rank="species" species="holomarginatum">
<emphasis id="B9726608FFF1FF9BFF3EFB8DA0ACFBB7" box="[218,426,1089,1113]" italics="true" pageId="46" pageNumber="131">B. holomarginatum</emphasis>
</taxonomicName>
members are distinctly blue or greenish-blue dorsally. The left paramere of the male genitalia (
<figureCitation id="133DA69FFFF1FF9BFE41FBAEA0F8FB97" box="[421,510,1121,1145]" captionStart="FIGURE" captionStartId="45.[171,183,939,958]" captionTargetBox="[137,1161,228,920]" captionTargetId="figure-1604@45.[120,1176,215,935]" captionTargetPageId="45" captionText="FIGURE 25. Male genitalia, Broscosoma holomarginatum sp. nov. (CASENT1016743; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067405" httpUri="https://zenodo.org/record/11067405/files/figure.png" pageId="46" pageNumber="131">Fig. 25b</figureCitation>
) is unusual for a
<taxonomicName id="4C06C199FFF1FF9BFD58FBADA295FB97" box="[700,915,1121,1145]" pageId="46" pageNumber="131">
<emphasis id="B9726608FFF1FF9BFD58FBADA239FB97" box="[700,831,1122,1145]" italics="true" pageId="46" pageNumber="131">Broscosoma</emphasis>
species
</taxonomicName>
and similar only to that of
<taxonomicName id="4C06C199FFF1FF9BFF7CFB4DA016FB74" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[152,272,1154,1178]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="131" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFF1FF9BFF7CFB4DA016FB74" box="[152,272,1154,1178]" italics="true" pageId="46" pageNumber="131">B. resbecqi</emphasis>
</taxonomicName>
males among species of the region.
</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FFB6DA52CFA55" blockId="46.[123,1174,219,1726]" pageId="46" pageNumber="131">
<emphasis id="B9726608FFF1FF9BFF4FFB6DA09EFB54" bold="true" box="[171,408,1186,1210]" pageId="46" pageNumber="131">Habitat distribution.</emphasis>
Within the study area, members of this species have been found under stones on the upper sandy beaches of rivers (
<figureCitation id="133DA69FFFF1FF9BFD80FB0DA3B8FB34" box="[612,702,1218,1242]" captionStart="FIGURE" captionStartId="84.[171,183,1585,1604]" captionTargetBox="[147,1149,216,1572]" captionTargetId="figure-57@84.[147,1149,216,1572]" captionTargetPageId="84" captionText="FIGURE 44. Photographs of habitats for Broscosoma species in the Gaoligong Shan region. a. 0.5 km E of Fengxue Yakou, 3150 m, 25.972º/098.683 º, Luzhang Township, Lushui County, Yunnan (habitat for Broscosoma gaoligongense sp. nov.); b. Xiao Jiang at Gangfang Sancha Lukou, 1500 m, 26.122º/98.573º, Lushui County, Yunnan (habitat for Broscosoma holomarginatum sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067447" httpUri="https://zenodo.org/record/11067447/files/figure.png" pageId="46" pageNumber="131">Fig. 44b</figureCitation>
) and along roadcuts with sandy soil during daytime. Adults are active on the surface at night, along roadcuts on moist sandy substrate stabilized by mosses and on open, sandy river beaches, where they congregate near or on boulders surrounded by sand, especially on boulders covered with mosses. Most of these sandy flats have little cover under which these beetles can hide during the day, so it seem likely that many spend daylight hours burrowed in the sand itself. Within the Gaoligong
<collectingRegion id="49C274F8FFF1FF9BFD01FAACA261FA95" box="[741,871,1379,1403]" country="Myanmar" name="Shan" pageId="46" pageNumber="131">Shan region</collectingRegion>
, this species occurs at relatively low elevations, with our records documenting its occurrence in the
<quantity id="4CFE17FFFFF1FF9BFC47FA4CA554FA75" box="[931,1106,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.5574999999999999" metricValueMax="1.6149999999999998" metricValueMin="1.5" pageId="46" pageNumber="131" unit="m" value="1557.5" valueMax="1615.0" valueMin="1500.0">1500 to 1615 m</quantity>
range (
<figureCitation id="133DA69FFFF1FF9BFF60FA6BA1D6FA52" box="[132,208,1444,1468]" captionStart="FIGURE" captionStartId="88.[184,196,1682,1701]" captionTargetBox="[130,1162,1063,1660]" captionTargetId="figure-332@88.[120,1176,1054,1668]" captionTargetPageId="88" captionText="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." figureDoi="http://doi.org/10.5281/zenodo.11067457" httpUri="https://zenodo.org/record/11067457/files/figure.png" pageId="46" pageNumber="131">Fig. 49</figureCitation>
). The record from Medog (
<collectingRegion id="49C274F8FFF1FF9BFE1DFA6BA336FA52" box="[505,560,1444,1468]" country="China" name="Xizang" pageId="46" pageNumber="131">Tibet</collectingRegion>
) is from a slightly higher elevation of
<quantity id="4CFE17FFFFF1FF9BFC36FA6BA525FA55" box="[978,1059,1444,1468]" metricMagnitude="3" metricUnit="m" metricValue="2.023" pageId="46" pageNumber="131" unit="m" value="2023.0">2023 m</quantity>
.
</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FFA0BA1B1F9B2" blockId="46.[123,1174,219,1726]" pageId="46" pageNumber="131">
<emphasis id="B9726608FFF1FF9BFF4FFA0BA216FA32" bold="true" box="[171,784,1476,1500]" pageId="46" pageNumber="131">
Geographical distribution within the Gaoligong
<collectingRegion id="49C274F8FFF1FF9BFD2BFA0BA20FFA32" box="[719,777,1476,1500]" country="Myanmar" name="Shan" pageId="46" pageNumber="131">Shan</collectingRegion>
.
</emphasis>
<figureCitation id="133DA69FFFF1FF9BFCF2FA0BA269FA32" box="[790,879,1476,1500]" captionStart="FIGURE" captionStartId="44.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1568]" captionTargetId="figure-56@44.[120,1176,216,1568]" captionTargetPageId="44" captionText="FIGURE 24. Broscosoma holomarginatum sp. nov. a. Habitus (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067401" httpUri="https://zenodo.org/record/11067401/files/figure.png" pageId="46" pageNumber="131">Fig. 24b</figureCitation>
. We examined a total of
<specimenCount id="9D007193FFF1FF9BFB9FFA0BA1ECFA12" count="20" pageId="46" pageNumber="131" type="generic">20 specimens</specimenCount>
(ten males and ten females) from the northern and southcentral parts of the Gaoligong
<collectingRegion id="49C274F8FFF1FF9BFF9FF9CBA1B6F9F2" box="[123,176,1540,1564]" country="Myanmar" name="Shan" pageId="46" pageNumber="131">Shan</collectingRegion>
in Gongshan and Lushui Counties. Our records from both areas are on the western side of the range (in Core Areas 1 and 4, respectively (
<figureCitation id="133DA69FFFF1FF9BFDB7F9EBA3A6F9D3" box="[595,672,1572,1597]" captionStart="FIGURE" captionStartId="88.[171,183,995,1014]" captionTargetBox="[130,1164,233,969]" captionTargetId="figure-57@88.[120,1176,215,989]" captionTargetPageId="88" captionText="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." figureDoi="http://doi.org/10.5281/zenodo.11067455" httpUri="https://zenodo.org/record/11067455/files/figure.png" pageId="46" pageNumber="131">Fig. 48</figureCitation>
) (see
<typeStatus id="54BD04B8FFF1FF9BFD06F9EAA211F9D3" box="[738,791,1573,1597]" pageId="46" pageNumber="131">Type</typeStatus>
material above for exact collection data).
</paragraph>
<paragraph id="8BB9BA1AFFF1FF9BFF4FF9AAA215F953" blockId="46.[123,1174,219,1726]" pageId="46" pageNumber="131">
<emphasis id="B9726608FFF1FF9BFF4FF9AAA329F993" bold="true" box="[171,559,1637,1661]" pageId="46" pageNumber="131">Overall geographical distribution.</emphasis>
<figureCitation id="133DA69FFFF1FF9BFDD1F9AAA387F993" box="[565,641,1637,1661]" captionStart="FIGURE" captionStartId="87.[171,183,970,989]" captionTargetBox="[120,1180,216,954]" captionTargetId="figure-67@87.[120,1180,216,954]" captionTargetPageId="87" captionText="FIGURE 47. Map showing approximate known overall geographical distributions of broscine species occurring in the Gaoligong Shan region as well as outside the study area. a. Broscus punctatus (Dejean); b. Eobroscus bhutanensis Morvan; c. Broscosoma ribbei Putzeys; d. Broscosoma holomarginatum sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 k" figureDoi="http://doi.org/10.5281/zenodo.11067453" httpUri="https://zenodo.org/record/11067453/files/figure.png" pageId="46" pageNumber="131">Fig. 47</figureCitation>
. This species has been recorded only from Gongshan and Lushui Counties in
<collectingRegion id="49C274F8FFF1FF9BFE50F949A303F973" box="[436,517,1670,1693]" country="China" name="Yunnan" pageId="46" pageNumber="131">Yunnan</collectingRegion>
and Medog County in Xizhang (
<collectingRegion id="49C274F8FFF1FF9BFC8FF94AA2A4F973" box="[875,930,1669,1693]" country="China" name="Xizang" pageId="46" pageNumber="131">Tibet</collectingRegion>
). Its occurrence in the study area represents the southern part its geographical range.
</paragraph>
<paragraph id="8BB9BA1AFFF0FF9AFF4FFF14A2FAFEBD" blockId="47.[123,1174,219,339]" pageId="47" pageNumber="132">
<emphasis id="B9726608FFF0FF9AFF4FFF14A246FF1D" bold="true" box="[171,832,219,243]" pageId="47" pageNumber="132">
Geographical relationships with other
<taxonomicName id="4C06C199FFF0FF9AFDB9FF14A246FF1D" box="[605,832,219,243]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9726608FFF0FF9AFDB9FF14A3E2FF1C" bold="true" box="[605,740,219,242]" italics="true" pageId="47" pageNumber="132">Broscosoma</emphasis>
species.
</taxonomicName>
</emphasis>
Within the study area, members of this species have been found together with those of
<taxonomicName id="4C06C199FFF0FF9AFD28FF33A2CBFEFD" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[716,973,252,275]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="purpureum">
<emphasis id="B9726608FFF0FF9AFD28FF33A2CBFEFD" box="[716,973,252,275]" italics="true" pageId="47" pageNumber="132">Broscosoma purpureum</emphasis>
</taxonomicName>
(described below) and
<taxonomicName id="4C06C199FFF0FF9AFF43FED3A002FEDD" authorityName="Putzeys. We" authorityYear="1877" box="[167,260,283,307]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFF0FF9AFF43FED3A002FEDD" box="[167,260,283,307]" italics="true" pageId="47" pageNumber="132">B. ribbei</emphasis>
</taxonomicName>
at one or more sites (
<figureCitation id="133DA69FFFF0FF9AFE01FED4A336FEDD" box="[485,560,283,307]" captionStart="FIGURE" captionStartId="89.[171,183,1461,1480]" captionTargetBox="[140,1154,249,1428]" captionTargetId="figure-68@89.[120,1176,216,1448]" captionTargetPageId="89" captionText="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi="http://doi.org/10.5281/zenodo.11067459" httpUri="https://zenodo.org/record/11067459/files/figure.png" pageId="47" pageNumber="132">Fig. 50</figureCitation>
). In Medog County,
<collectingRegion id="49C274F8FFF0FF9AFCEFFED4A25EFEDD" box="[779,856,283,307]" country="China" name="Xizang" pageId="47" pageNumber="132">Xizang</collectingRegion>
(
<collectingRegion id="49C274F8FFF0FF9AFC83FED4A298FEDD" box="[871,926,283,307]" country="China" name="Xizang" pageId="47" pageNumber="132">Tibet</collectingRegion>
), they have been found syntopic with members of
<taxonomicName id="4C06C199FFF0FF9AFE7EFEF3A0FEFEBD" authorityName="Putzeys. We" authorityYear="1877" box="[410,504,315,339]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFF0FF9AFE7EFEF3A0FEFEBD" box="[410,504,315,339]" italics="true" pageId="47" pageNumber="132">B. ribbei</emphasis>
</taxonomicName>
and another (undescribed)
<taxonomicName id="4C06C199FFF0FF9AFCFAFEF3A2FAFEBD" box="[798,1020,315,339]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="undefined-9">
<emphasis id="B9726608FFF0FF9AFCFAFEF3A2A7FEBD" box="[798,929,316,339]" italics="true" pageId="47" pageNumber="132">Broscosoma</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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