treatments-xml/data/03/AF/0B/03AF0B0CFFE8FF8EFF9EF9DEA0D5FBD7.xml

<document id="2D3FD7915A2D0C57571C6F22F3B541A3" ID-CLB-Dataset="299055" ID-DOI="10.5281/zenodo.11067355" ID-GBIF-Dataset="67ec2de4-1816-4969-8132-f6796d3afa06" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067355" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052343267" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2021" docId="03AF0B0CFFE8FF8EFF9EF9DEA0D5FBD7" docLanguage="en" docName="ProcCalifAcadSci.67.4.85-182.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (4)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Broscosoma resbecqi Kavanaugh and Liang 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="144" masterDocId="FF967374FFDFFFB5FFE4FFCFA106FFEE" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." masterLastPageNumber="182" masterPageNumber="85" pageNumber="140" updateTime="1719540814554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="85319F29D479C2AC2C3E908BD5864A2A">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae).</mods:title>
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<mods:name id="6FD65EA64BC4DD4684AE3169B076B5BB" type="personal">
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<mods:namePart id="7BEAEE339F4218D8924D7E4218B2D306">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="E7BF4A6CC10A803C9447098FE6629F34">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="F0A3EC51A655C43BBE73DBB4202434F9">Liang, Hongbin</mods:namePart>
<mods:affiliation id="DA27717926F11A9429217DC761B5B7EB">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title id="A8DF01D0B01B367E3BA5FC39D61D991B">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="487C07A8756D784205CAB59040D96A0D">
<mods:date id="E825F88DD94555461F60AE1D7750ADFA">2021</mods:date>
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<treatment id="03AF0B0CFFE8FF8EFF9EF9DEA0D5FBD7" ID-DOI="http://doi.org/10.5281/zenodo.12575582" ID-GBIF-Taxon="233814478" ID-Zenodo-Dep="12575582" LSID="urn:lsid:plazi:treatment:03AF0B0CFFE8FF8EFF9EF9DEA0D5FBD7" httpUri="http://treatment.plazi.org/id/03AF0B0CFFE8FF8EFF9EF9DEA0D5FBD7" lastPageId="59" lastPageNumber="144" pageId="55" pageNumber="140">
<subSubSection id="C31CE991FFE8FF82FF9EF9DEA3F0F9C4" box="[122,758,1553,1578]" pageId="55" pageNumber="140" type="nomenclature">
<paragraph id="8BB9BA1AFFE8FF82FF9EF9DEA3F0F9C4" blockId="55.[122,758,1553,1608]" box="[122,758,1553,1578]" pageId="55" pageNumber="140">
<heading id="D0F10D76FFE8FF82FF9EF9DEA3F0F9C4" box="[122,758,1553,1578]" fontSize="10" level="2" pageId="55" pageNumber="140" reason="7">
<emphasis id="B9726608FFE8FF82FF9EF9DEA3F0F9C4" bold="true" box="[122,758,1553,1578]" pageId="55" pageNumber="140">
11. 
<taxonomicName id="4C06C199FFE8FF82FF45F9DDA394F9C7" authority="Kavanaugh and Liang" authorityName="Kavanaugh and Liang" authorityYear="2021" box="[161,658,1553,1578]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="140" phylum="Arthropoda" rank="species" species="resbecqi" status="sp. nov.">
<emphasis id="B9726608FFE8FF82FF45F9DDA08FF9C7" bold="true" box="[161,393,1553,1577]" italics="true" pageId="55" pageNumber="140">Broscosoma resbecqi</emphasis>
Kavanaugh and Liang
</taxonomicName>
, 
<taxonomicNameLabel id="A241DB73FFE8FF82FD7BF9DDA3F0F9C4" box="[671,758,1554,1578]" pageId="55" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFE8FF82FF9EF9FDA0C4F9A6" box="[122,450,1586,1608]" pageId="55" pageNumber="140" type="description">
<paragraph id="8BB9BA1AFFE8FF82FF9EF9FDA0C4F9A6" blockId="55.[122,758,1553,1608]" box="[122,450,1586,1608]" pageId="55" pageNumber="140">
<figureCitation id="133DA69FFFE8FF82FF9EF9FDA1E7F9A6" box="[122,225,1586,1608]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="55" pageNumber="140">Figures 6d</figureCitation>
, 
<figureCitation id="133DA69FFFE8FF82FF09F9FDA003F9A6" box="[237,261,1586,1608]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="55" pageNumber="140">9b</figureCitation>
, 
<figureCitation id="133DA69FFFE8FF82FEF6F9FDA02CF9A9" box="[274,298,1586,1607]" captionStart="FIGURE" captionStartId="57.[171,183,1577,1596]" captionTargetBox="[135,1176,216,1562]" captionTargetId="figure-68@57.[120,1176,216,1563]" captionTargetPageId="57" captionText="FIGURE 30. Broscosoma resbecqi sp. nov. a. Habitus (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067417" httpUri="https://zenodo.org/record/11067417/files/figure.png" pageId="55" pageNumber="140">30</figureCitation>
, 
<figureCitation id="133DA69FFFE8FF82FED2F9FDA048F9A9" box="[310,334,1586,1607]" captionStart="FIGURE" captionStartId="58.[171,183,951,970]" captionTargetBox="[133,1161,228,923]" captionTargetId="figure-1524@58.[120,1176,216,932]" captionTargetPageId="58" captionText="FIGURE 31. Male genitalia, Broscosoma resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067419" httpUri="https://zenodo.org/record/11067419/files/figure.png" pageId="55" pageNumber="140">31</figureCitation>
, 
<figureCitation id="133DA69FFFE8FF82FEBFF9FDA078F9A6" box="[347,382,1586,1608]" captionStart="FIGURE" captionStartId="86.[171,183,1592,1611]" captionTargetBox="[147,1149,216,1572]" captionTargetId="figure-57@86.[147,1149,216,1572]" captionTargetPageId="86" captionText="FIGURE 46. Photographs of habitats for broscine species in the Gaoligong Shan region. a. Siran Wang, 0.2 km above confluence with Dulong Jiang, 1720 m, 28.013°/ E098.321º, Dulongjiang Township, Gongshan County, Yunnan (habitat for Broscosoma resbecqi sp. nov.); b. Nankang Yakou, 2130 m, 24.828°/98.767°, Longyang County, Yunnan (habitat for Broscosoma ribbei Putzeys and Broscus punctatus (Dejean). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067451" httpUri="https://zenodo.org/record/11067451/files/figure.png" pageId="55" pageNumber="140">46a</figureCitation>
, 
<figureCitation id="133DA69FFFE8FF82FE6EF9FDA0C4F9A6" box="[394,450,1586,1608]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="88.[171,183,995,1014]" captionStartId-1="88.[184,196,1682,1701]" captionStartId-2="89.[171,183,1461,1480]" captionTargetBox-0="[130,1164,233,969]" captionTargetBox-1="[130,1162,1063,1660]" captionTargetBox-2="[140,1154,249,1428]" captionTargetId-0="figure-57@88.[120,1176,215,989]" captionTargetId-1="figure-332@88.[120,1176,1054,1668]" captionTargetId-2="figure-68@89.[120,1176,216,1448]" captionTargetPageId-0="88" captionTargetPageId-1="88" captionTargetPageId-2="89" captionText-0="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." captionText-2="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi-0="http://doi.org/10.5281/zenodo.11067455" figureDoi-1="http://doi.org/10.5281/zenodo.11067457" figureDoi-2="http://doi.org/10.5281/zenodo.11067459" httpUri-0="https://zenodo.org/record/11067455/files/figure.png" httpUri-1="https://zenodo.org/record/11067457/files/figure.png" httpUri-2="https://zenodo.org/record/11067459/files/figure.png" pageId="55" pageNumber="140">48-50</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFE8FF8DFF4EF9ABA065FE1A" lastPageId="56" lastPageNumber="141" pageId="55" pageNumber="140" type="materials_examined">
<paragraph id="8BB9BA1AFFE8FF8DFF4EF9ABA30AFEFD" blockId="55.[122,1174,1636,1726]" lastBlockId="56.[123,1174,219,1726]" lastPageId="56" lastPageNumber="141" pageId="55" pageNumber="140">
<emphasis id="B9726608FFE8FF82FF4EF9ABA057F992" bold="true" box="[170,337,1636,1660]" pageId="55" pageNumber="140">Type material.</emphasis>
<materialsCitation id="3B6EB047FFE8FF8DFEBCF9ABA30AFEFD" ID-GBIF-Occurrence="4899905326" collectingDate="2004-11-01" collectionCode="IOZ" collectorName="D. H. Kavanaugh &amp; D. - Z. Dong" country="China" county="Gongshan County" elevation="1720" lastPageId="56" lastPageNumber="141" latitude="28.01347" location="Siran Wang" longLatPrecision="1" longitude="98.32117" municipality="Dulongjiang Township" pageId="55" pageNumber="140" specimenCode="CASENT 1016251" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54BD04B8FFE8FF82FEBCF9ABA0BDF992" box="[344,443,1636,1660]" pageId="55" pageNumber="140" type="holotype">Holotype</typeStatus>
, a 
<specimenCount id="9D007193FFE8FF82FE3AF9ABA314F992" box="[478,530,1636,1660]" pageId="55" pageNumber="140" type="male">male</specimenCount>
, in 
<collectionCode id="ED1722DFFFE8FF82FDD9F9ABA36FF992" box="[573,617,1636,1660]" pageId="55" pageNumber="140">IOZ</collectionCode>
, labeled: “ 
<specimenCode id="DBA01261FFE8FF82FD0CF9ABA2B2F992" box="[744,948,1636,1660]" pageId="55" pageNumber="140">CASENT 1016251</specimenCode>
”/ “ 
<collectingCountry id="F311FA8AFFE8FF82FC3FF9ABA529F992" box="[987,1071,1636,1660]" name="China" pageId="55" pageNumber="140">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFE8FF82FBD9F9AAA588F992" box="[1085,1166,1637,1660]" country="China" name="Yunnan" pageId="55" pageNumber="140">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFE8FF82FF9EF94AA038F973" box="[122,318,1669,1693]" pageId="55" pageNumber="140">Gongshan County</collectingCounty>
, 
<collectingMunicipality id="6BDD2060FFE8FF82FEABF94AA341F973" box="[335,583,1669,1693]" pageId="55" pageNumber="140">Dulongjiang Township</collectingMunicipality>
, 
<location id="8ED9ECC1FFE8FF82FDBCF94AA3DEF973" LSID="urn:lsid:plazi:treatment:03AF0B0CFFE8FF8EFF9EF9DEA0D5FBD7:8ED9ECC1FFE8FF82FDBCF94AA3DEF973" box="[600,728,1669,1693]" country="China" county="Gongshan County" latitude="28.01347" longLatPrecision="1" longitude="98.32117" municipality="Dulongjiang Township" name="Siran Wang" pageId="55" pageNumber="140" stateProvince="Yunnan">Siran Wang</location>
, 
<quantity id="4CFE17FFFFE8FF82FD0DF94AA231F973" box="[745,823,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="140" unit="km" value="0.2">
<locationDeviation id="9A0BA338FFE8FF82FD0DF94AA231F973" box="[745,823,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="140" unit="km" value="0.2">0.2 km</locationDeviation>
</quantity>
above confluence with 
<location id="8ED9ECC1FFE8FF82FBA1F94AA1B4F953" LSID="urn:lsid:plazi:treatment:03AF0B0CFFE8FF8EFF9EF9DEA0D5FBD7:8ED9ECC1FFE8FF82FBA1F94AA1B4F953" country="China" county="Gongshan County" latitude="28.01347" longLatPrecision="1" longitude="98.32117" municipality="Dulongjiang Township" name="Dulong Jiang" pageId="55" pageNumber="140" stateProvince="Yunnan">Dulong Jiang</location>
, 
<geoCoordinate id="EE32DCDDFFE8FF82FF20F969A040F950" box="[196,326,1702,1726]" degrees="28. 01347" direction="north" orientation="latitude" pageId="55" pageNumber="140" precision="1" value="28.01347">N28.01347°</geoCoordinate>
/ 
<geoCoordinate id="EE32DCDDFFE8FF82FEBCF969A0E5F950" box="[344,483,1702,1726]" degrees="098. 32117" direction="east" orientation="longitude" pageId="55" pageNumber="140" precision="1" value="98.32117">E098.32117°</geoCoordinate>
,”/ “ 
<quantity id="4CFE17FFFFE8FF82FDF7F969A36EF953" box="[531,616,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="55" pageNumber="140" unit="m" value="1720.0">
<elevation id="002B5D29FFE8FF82FDF7F969A36EF953" box="[531,616,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="55" pageNumber="140" unit="m" value="1720.0">1720 m</elevation>
</quantity>
, 
<date id="FFB89CDAFFE8FF82FD9EF969A244F950" box="[634,834,1701,1726]" pageId="55" pageNumber="140" value="2004-11-01">
<collectingDate id="EFFC6532FFE8FF82FD9EF969A244F950" box="[634,834,1701,1726]" pageId="55" pageNumber="140" value="2004-11-01">1 November 2004</collectingDate>
</date>
, Stop # DHK-2004-066, 
<collectorName id="26F3DFCCFFE8FF8DFB85F969A1F2FF1D" lastPageId="56" lastPageNumber="141" pageId="55" pageNumber="140">D.H. Kavanaugh</collectorName>
&amp; 
<collectorName id="26F3DFCCFFE7FF8DFEFFFF13A09FFF1A" box="[283,409,220,244]" pageId="56" pageNumber="141">D.-Z. Dong</collectorName>
collectors”/ “HOLOTYPE 
<taxonomicName id="4C06C199FFE7FF8DFD2CFF13A593FF1D" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[712,1173,219,243]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="141" phylum="Arthropoda" rank="species" species="resbecqi" status="sp. nov.">
<emphasis id="B9726608FFE7FF8DFD2CFF13A2ABFF1D" box="[712,941,219,243]" italics="true" pageId="56" pageNumber="141">Broscosoma resbecqi</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFE7FF8DFF9FFF33A1C8FEFD" box="[123,206,252,275]" pageId="56" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [red label].
</materialsCitation>
</paragraph>
<paragraph id="8BB9BA1AFFE7FF8DFF4FFED3A36DFEBD" blockId="56.[123,1174,219,1726]" pageId="56" pageNumber="141">
<emphasis id="B9726608FFE7FF8DFF4FFED3A03DFEDD" bold="true" box="[171,315,283,308]" pageId="56" pageNumber="141">Type locality</emphasis>
. China, Yunnan, Gongshan County, Dulongjiang Township, Siran Wang, 0.2. km above confluence with Dulong Jiang, 1720 m.
</paragraph>
<paragraph id="8BB9BA1AFFE7FF8DFF4FFE93A065FE1A" blockId="56.[123,1174,219,1726]" pageId="56" pageNumber="141">
<emphasis id="B9726608FFE7FF8DFF4FFE93A0E7FE9A" bold="true" box="[171,481,348,372]" pageId="56" pageNumber="141">Derivation of species name.</emphasis>
We take pleasure in naming this new species in honor of our colleague, known professionally as Thierry Deuve, but officially as Thierry de Resbecq, in recognition of his many contributions to our knowledge of the broscine fauna of Asia and for his help with this and other research projects. The species epithet, 
<taxonomicName id="4C06C199FFE7FF8DFD48FE72A203FE3B" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[684,773,445,469]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="141" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE7FF8DFD48FE72A203FE3B" box="[684,773,445,469]" italics="true" pageId="56" pageNumber="141">resbecqi</emphasis>
</taxonomicName>
, is the Latinized form of his surname (in the genitive case).
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFE7FF8DFF4FFE32A06DFCF8" pageId="56" pageNumber="141" type="diagnosis">
<paragraph id="8BB9BA1AFFE7FF8DFF4FFE32A06DFCF8" blockId="56.[123,1174,219,1726]" pageId="56" pageNumber="141">
<emphasis id="B9726608FFE7FF8DFF4FFE32A019FDFB" bold="true" box="[171,287,509,533]" pageId="56" pageNumber="141">Diagnosis.</emphasis>
Adults of this species (
<figureCitation id="133DA69FFFE7FF8DFDC5FE32A37CFDFB" box="[545,634,509,533]" captionStart="FIGURE" captionStartId="57.[171,183,1577,1596]" captionTargetBox="[135,1176,216,1562]" captionTargetId="figure-68@57.[120,1176,216,1563]" captionTargetPageId="57" captionText="FIGURE 30. Broscosoma resbecqi sp. nov. a. Habitus (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067417" httpUri="https://zenodo.org/record/11067417/files/figure.png" pageId="56" pageNumber="141">Fig. 30a</figureCitation>
) can be distinguished from those of other species in the region by the following combination of character states: size moderate, BL = 9.0 mm; dorsal surface of head and pronotum vivid metallic green, elytra darker blue-green, legs pale rufous; vertex of head with post-temporal transverse sulcus relatively shallow and broadly and densely punctate (
<figureCitation id="133DA69FFFE7FF8DFF01FDB1A036FD78" box="[229,304,638,662]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="56" pageNumber="141">Fig. 6d</figureCitation>
); antennomeres 3 and 4 glabrous except for apical whorl of setae; pronotum with lateral margination present but interrupted briefly in basal half (
<figureCitation id="133DA69FFFE7FF8DFCC9FD51A27CFD58" box="[813,890,670,694]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="56" pageNumber="141">Fig. 9b</figureCitation>
), basolateral setae absent; elytral silhouette subovoid, with humeri rounded but evident, elytral microsculpture effaced; metatrochanter asetose; male protarsi with tarsomeres 1 to 3 with ventral pads of adhesive setae; male genitalia as in 
<figureCitation id="133DA69FFFE7FF8DFEF3FD30A062FCF9" box="[279,356,767,791]" captionStart="FIGURE" captionStartId="58.[171,183,951,970]" captionTargetBox="[133,1161,228,923]" captionTargetId="figure-1524@58.[120,1176,216,932]" captionTargetPageId="58" captionText="FIGURE 31. Male genitalia, Broscosoma resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067419" httpUri="https://zenodo.org/record/11067419/files/figure.png" pageId="56" pageNumber="141">Fig. 31</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFE7FF8FFF4FFCD0A0E3FA0B" lastPageId="58" lastPageNumber="143" pageId="56" pageNumber="141" type="description">
<paragraph id="8BB9BA1AFFE7FF8DFF4FFCD0A297FC79" blockId="56.[123,1174,219,1726]" pageId="56" pageNumber="141">
<emphasis id="B9726608FFE7FF8DFF4FFCD0A030FCD9" bold="true" box="[171,310,799,823]" pageId="56" pageNumber="141">Description.</emphasis>
<figureCitation id="133DA69FFFE7FF8DFED8FCD0A092FCD9" box="[316,404,799,823]" captionStart="FIGURE" captionStartId="57.[171,183,1577,1596]" captionTargetBox="[135,1176,216,1562]" captionTargetId="figure-68@57.[120,1176,216,1563]" captionTargetPageId="57" captionText="FIGURE 30. Broscosoma resbecqi sp. nov. a. Habitus (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067417" httpUri="https://zenodo.org/record/11067417/files/figure.png" pageId="56" pageNumber="141">Fig. 30a</figureCitation>
. Size medium. BL male = 9.0 mm, ratio EL/PL = 2.5. Body color black or dark piceous, last visible abdominal sternite dark rufous brown, all antennomeres, femora, tibiae, and tarsi pale rufous. Head and pronotum with brilliant green metallic reflection, elytra with darker bluish-green metallic reflection, venter without metallic reflection.
</paragraph>
<paragraph id="8BB9BA1AFFE7FF8DFF4FFC6FA2F6FB34" blockId="56.[123,1174,219,1726]" pageId="56" pageNumber="141">
Head. 
<figureCitation id="133DA69FFFE7FF8DFF10FC6FA042FC56" box="[244,324,928,952]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="56" pageNumber="141">Fig. 6d</figureCitation>
. Eyes medium-sized, their diameter slightly greater than length of tempora. Frontal furrows moderately impressed, narrow, arcuate, slightly convergent anteriorly and posteriorly, long, extended posteriorly beyond middle of eyes, rugulose and sparsely and coarsely punctate. Vertex with post-temporal transverse sulcus densely and coarsely punctate. Tempora straight, oblique. Genal ridges present, restricted to head region distinctly posterior to post-temporal constriction. Clypeus with one pair of setae. Supraorbital setae present, one pair. Eustipes of maxilla with two setae, the dorsobasal seta distinctly more than half as long as ventrobasal seta. 
<collectingRegion id="49C274F8FFE7FF8DFBDFFBADA593FB97" box="[1083,1173,1122,1145]" country="Myanmar" name="Chin" pageId="56" pageNumber="141">Mentum</collectingRegion>
with tooth present, simple, paramedial region shallowly foveate, one pair of mental setae present. Glosal sclerite (ligula) with one pair of setae. Submentum with two pairs of setae. Gula without transverse grooves. Antennomeres 3 and 4 without pubescence, ratio A3/A5 = 1.1.
</paragraph>
<paragraph id="8BB9BA1AFFE7FF8DFF4FFB2CA1C0F9F2" blockId="56.[123,1174,219,1726]" pageId="56" pageNumber="141">
Pronotum. 
<figureCitation id="133DA69FFFE7FF8DFEC7FB2DA077FB14" box="[291,369,1250,1274]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="56" pageNumber="141">Fig. 6d</figureCitation>
. Globose anterior to sub-basal constriction, with base distinctly pedunculate, widest at middle of discal region, PL/PW = 1.1, disk markedly convex; apical margination absent; lateral margination present, narrow, distinctly delineated from apical margin to just posterior of midlateral seta, thence slightly interrupted and then evident again at and posterior to the sub-basal constriction; basal margination absent; anterior transverse impression shallow, broad and vaguely delineated, coarsely punctate both at bottom and on slopes of depression; median longitudinal impression moderately impressed; posterior transverse impression deep, sharply delineated; anterior region sparsely punctate; pronotal base coarsely and densely punctate; one pair of midlateral pronotal setae present, inserted at mid-length of discal region; basolateral pronotal setae absent.
</paragraph>
<paragraph id="8BB9BA1AFFE7FF8FFF4FF9EBA3A3FAEB" blockId="56.[123,1174,219,1726]" lastBlockId="58.[123,1174,1070,1702]" lastPageId="58" lastPageNumber="143" pageId="56" pageNumber="141">Elytra. Elytral silhouette subovoid, slightly narrowed in basal half, widest at middle, ratio EL/EW = 1.6, greatest elevation above lateral margin (in lateral view) posterior to middle; humeri distinctly present, roundly obtuse; lateral margins moderately arcuate and nearly straight in anterior half sub-basally; parascutellar striole present, short, distally merged in a distinct jog with free base of stria 1. Eight striae present; stria 1 moderately deeply and sharply impressed, striae 2 to 8 indistinct, obliterated or represented by extremely fine and shallow linear depressions, successively less evident laterally and apically, stria 8 merged with lateral groove throughout its length or nearly so; stria 1 moderately punctate at least in basal half, stria 2 moderately punctate in basal two-thirds and successively more shallowly and sparsely punctate in apical third, striae 3 to 8 successively more shallowly and sparsely punctate laterally and apically. Elytral microsculpture effaced. Parascutellar seta present at base of stria 2, discal setae absent, umbilicate series of elytral comprised of one post-humeral and two preapical setae.</paragraph>
<caption id="DF79EA92FFE6FF8CFF4FF9E6A0FDF99E" ID-DOI="http://doi.org/10.5281/zenodo.11067417" ID-Zenodo-Dep="11067417" httpUri="https://zenodo.org/record/11067417/files/figure.png" pageId="57" pageNumber="142" startId="57.[171,183,1577,1596]" targetBox="[135,1176,216,1562]" targetPageId="57" targetType="figure">
<paragraph id="8BB9BA1AFFE6FF8CFF4FF9E6A0FDF99E" blockId="57.[123,1173,1577,1648]" pageId="57" pageNumber="142">
<smallCapsWord id="8D5F2CC6FFE6FF8CFF4FF9E6A1EEF9D5" baselines="1591,1591" box="[171,232,1577,1596]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="57" pageNumber="142">FIGURE</smallCapsWord>
30. 
<taxonomicName id="4C06C199FFE6FF8CFEE8F9E5A0C6F9D2" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[268,448,1577,1596]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="142" phylum="Arthropoda" rank="species" species="resbecqi" status="sp. nov.">
<emphasis id="B9726608FFE6FF8CFEE8F9E5A0C6F9D2" box="[268,448,1577,1596]" italics="true" pageId="57" pageNumber="142">Broscosoma resbecqi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFE6FF8CFE21F9E6A300F9D2" box="[453,518,1577,1596]" pageId="57" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
a. Habitus (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<caption id="DF79EA92FFE5FF8FFF4FFC78A37CFC10" ID-DOI="http://doi.org/10.5281/zenodo.11067419" ID-Zenodo-Dep="11067419" httpUri="https://zenodo.org/record/11067419/files/figure.png" pageId="58" pageNumber="143" startId="58.[171,183,951,970]" targetBox="[133,1161,228,923]" targetPageId="58" targetType="figure">
<paragraph id="8BB9BA1AFFE5FF8FFF4FFC78A37CFC10" blockId="58.[123,1174,951,1022]" pageId="58" pageNumber="143">
<smallCapsWord id="8D5F2CC6FFE5FF8FFF4FFC78A1EEFC27" baselines="965,965" box="[171,232,951,970]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="58" pageNumber="143">FIGURE</smallCapsWord>
31. Male genitalia, 
<taxonomicName id="4C06C199FFE5FF8FFE72FC77A34AFC24" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[406,588,951,970]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="143" phylum="Arthropoda" rank="species" species="resbecqi" status="sp. nov.">
<emphasis id="B9726608FFE5FF8FFE72FC77A34AFC24" box="[406,588,951,970]" italics="true" pageId="58" pageNumber="143">Broscosoma resbecqi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFE5FF8FFDB7FC78A393FC24" box="[595,661,951,970]" pageId="58" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
(Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB9BA1AFFE5FF8FFF4FFAC1A237FACB" blockId="58.[123,1174,1070,1702]" box="[171,817,1294,1318]" pageId="58" pageNumber="143">Hindwings. Slightly reduced, incapable of supporting flight.</paragraph>
<paragraph id="8BB9BA1AFFE5FF8FFF4FFAE1A3E1FAAB" blockId="58.[123,1174,1070,1702]" box="[171,743,1326,1350]" pageId="58" pageNumber="143">Thoracic venter. Metepisternum ratio ML/MW = 1.8.</paragraph>
<paragraph id="8BB9BA1AFFE5FF8FFF4FFA81A2E8FA4B" blockId="58.[123,1174,1070,1702]" pageId="58" pageNumber="143">Legs. Hind trochanter asetose. Tarsomeres without dorsal pubescence; tarsomere 5 with two or more pairs of setae ventrally; male protarsi with pads of adhesive setae on tarsomeres 1 to 3 ventrally, male mesotarsi with pads of adhesive setae on tarsomeres 1 and 2 ventrally.</paragraph>
<paragraph id="8BB9BA1AFFE5FF8FFF4FFA61A515FA2B" blockId="58.[123,1174,1070,1702]" box="[171,1043,1454,1478]" pageId="58" pageNumber="143">Abdomen. Male sternite VI with one pair of apicolateral setae. Female unknown.</paragraph>
<paragraph id="8BB9BA1AFFE5FF8FFF4FFA01A0E3FA0B" blockId="58.[123,1174,1070,1702]" box="[171,485,1486,1510]" pageId="58" pageNumber="143">
Male genitalia. As in 
<figureCitation id="133DA69FFFE5FF8FFE76FA01A0D9FA08" box="[402,479,1486,1510]" captionStart="FIGURE" captionStartId="58.[171,183,951,970]" captionTargetBox="[133,1161,228,923]" captionTargetId="figure-1524@58.[120,1176,216,932]" captionTargetPageId="58" captionText="FIGURE 31. Male genitalia, Broscosoma resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067419" httpUri="https://zenodo.org/record/11067419/files/figure.png" pageId="58" pageNumber="143">Fig. 31</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFE5FF8EFF4FFA21A0D5FBD7" lastPageId="59" lastPageNumber="144" pageId="58" pageNumber="143" type="discussion">
<paragraph id="8BB9BA1AFFE5FF8EFF4FFA21A047FDBB" blockId="58.[123,1174,1070,1702]" lastBlockId="59.[123,1174,219,1081]" lastPageId="59" lastPageNumber="144" pageId="58" pageNumber="143">
<emphasis id="B9726608FFE5FF8FFF4FFA21A02BF9E8" bold="true" box="[171,301,1518,1542]" pageId="58" pageNumber="143">Comments.</emphasis>
As noted above, 
<taxonomicName id="4C06C199FFE5FF8FFE0FFA20A365F9E8" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[491,611,1518,1542]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="143" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE5FF8FFE0FFA20A365F9E8" box="[491,611,1518,1542]" italics="true" pageId="58" pageNumber="143">B. resbecqi</emphasis>
</taxonomicName>
is one of four species in the study area members of which have the entire dorsum of the body with metallic green reflection. Features distinguishing members of this species from those of 
<taxonomicName id="4C06C199FFE5FF8FFDF2F9E0A3B6F9A8" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[534,688,1582,1606]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="143" phylum="Arthropoda" rank="species" species="danzhuense">
<emphasis id="B9726608FFE5FF8FFDF2F9E0A3B6F9A8" box="[534,688,1582,1606]" italics="true" pageId="58" pageNumber="143">B. danzhuense</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C06C199FFE5FF8FFD00F9E0A254F9A8" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[740,850,1583,1606]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="143" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFE5FF8FFD00F9E0A254F9A8" box="[740,850,1583,1606]" italics="true" pageId="58" pageNumber="143">B. parvum</emphasis>
</taxonomicName>
have been addressed above in the Comments sections for each of those species. At first glance, 
<taxonomicName id="4C06C199FFE5FF8FFCD4F980A2A1F988" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[816,935,1614,1638]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="143" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE5FF8FFCD4F980A2A1F988" box="[816,935,1614,1638]" italics="true" pageId="58" pageNumber="143">B. resbecqi</emphasis>
</taxonomicName>
could be mistaken for a small specimen of 
<taxonomicName id="4C06C199FFE5FF8FFEB0F9A0A0B7F968" authorityName="Putzeys. We" authorityYear="1877" box="[340,433,1646,1670]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="143" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFE5FF8FFEB0F9A0A0B7F968" box="[340,433,1646,1670]" italics="true" pageId="58" pageNumber="143">B. ribbei</emphasis>
</taxonomicName>
, but several features distinguish members of these taxa. The 
<typeStatus id="54BD04B8FFE5FF8FFBDCF9A1A593F968" box="[1080,1173,1646,1670]" pageId="58" pageNumber="143" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C06C199FFE5FF8FFF7DF940A014F948" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[153,274,1678,1702]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="143" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE5FF8FFF7DF940A014F948" box="[153,274,1678,1702]" italics="true" pageId="58" pageNumber="143">B. resbecqi</emphasis>
</taxonomicName>
has the metallic reflection of the elytra a darker bluish-green, contrasting with the brighter green of the forebody, and the femora pale like the tibiae and tarsi. In 
<taxonomicName id="4C06C199FFE4FF8EFC2CFF13A520FF1D" authorityName="Putzeys. We" authorityYear="1877" box="[968,1062,219,243]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFE4FF8EFC2CFF13A520FF1D" box="[968,1062,219,243]" italics="true" pageId="59" pageNumber="144">B. ribbei</emphasis>
</taxonomicName>
members, the metallic reflection of the forebody and elytra are both a bright green, golden green, or blue-green and the femora are distinctly darker than the tibiae and tarsi. The eyes are more convex, the tempora longer and are less convex, and the post-temporal transverse sulcus more broadly and densely punctate in 
<taxonomicName id="4C06C199FFE4FF8EFEB4FE92A0C1FE9A" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[336,455,348,372]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE4FF8EFEB4FE92A0C1FE9A" box="[336,455,348,372]" italics="true" pageId="59" pageNumber="144">B. resbecqi</emphasis>
</taxonomicName>
(
<figureCitation id="133DA69FFFE4FF8EFE33FE93A325FE9A" box="[471,547,348,372]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="59" pageNumber="144">Fig. 6d</figureCitation>
) than in 
<taxonomicName id="4C06C199FFE4FF8EFD67FE92A3E7FE9A" authorityName="Putzeys. We" authorityYear="1877" box="[643,737,348,372]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFE4FF8EFD67FE92A3E7FE9A" box="[643,737,348,372]" italics="true" pageId="59" pageNumber="144">B. ribbei</emphasis>
</taxonomicName>
(
<figureCitation id="133DA69FFFE4FF8EFD15FE93A23AFE9A" box="[753,828,348,372]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="59" pageNumber="144">Fig. 6b</figureCitation>
) members. The lateral margination of the pronotum in 
<taxonomicName id="4C06C199FFE4FF8EFE9DFEB2A0E9FE7A" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[377,495,380,404]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE4FF8EFE9DFEB2A0E9FE7A" box="[377,495,380,404]" italics="true" pageId="59" pageNumber="144">B. resbecqi</emphasis>
</taxonomicName>
(
<figureCitation id="133DA69FFFE4FF8EFE1BFEB3A34DFE7A" box="[511,587,380,404]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="59" pageNumber="144">Fig. 6d</figureCitation>
) is better developed and nearly complete, whereas it is restricted to the area of the midlateral seta in most specimens 
<taxonomicName id="4C06C199FFE4FF8EFCF2FE52A272FE5A" authorityName="Putzeys. We" authorityYear="1877" box="[790,884,412,436]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFE4FF8EFCF2FE52A272FE5A" box="[790,884,412,436]" italics="true" pageId="59" pageNumber="144">B. ribbei</emphasis>
</taxonomicName>
, extended further anteriorly and posteriorly in some (
<figureCitation id="133DA69FFFE4FF8EFE4AFE72A0FAFE3B" box="[430,508,445,469]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="59" pageNumber="144">Fig. 6b</figureCitation>
), and not at all distinct in a very few specimens. The metatrochanters are asetose in the 
<taxonomicName id="4C06C199FFE4FF8EFE59FE11A333FE1B" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[445,565,477,501]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE4FF8EFE59FE11A333FE1B" box="[445,565,477,501]" italics="true" pageId="59" pageNumber="144">B. resbecqi</emphasis>
</taxonomicName>
<typeStatus id="54BD04B8FFE4FF8EFDDAFE12A39DFE1B" box="[574,667,477,501]" pageId="59" pageNumber="144" type="holotype">holotype</typeStatus>
but unisetose in most 
<taxonomicName id="4C06C199FFE4FF8EFC77FE11A2F4FE1B" authorityName="Putzeys. We" authorityYear="1877" box="[915,1010,477,501]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFE4FF8EFC77FE11A2F4FE1B" box="[915,1010,477,501]" italics="true" pageId="59" pageNumber="144">B. ribbei</emphasis>
</taxonomicName>
members, unilaterally asetose in a few. The median lobe of the male genitalia in much shorter, thicker, more arcuate and the apical lamella of very different shape in 
<taxonomicName id="4C06C199FFE4FF8EFD33FDD1A248FDDB" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[727,846,541,565]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE4FF8EFD33FDD1A248FDDB" box="[727,846,541,565]" italics="true" pageId="59" pageNumber="144">B. resbecqi</emphasis>
</taxonomicName>
(
<figureCitation id="133DA69FFFE4FF8EFCBAFDD2A2B1FDDB" box="[862,951,541,566]" captionStart="FIGURE" captionStartId="58.[171,183,951,970]" captionTargetBox="[133,1161,228,923]" captionTargetId="figure-1524@58.[120,1176,216,932]" captionTargetPageId="58" captionText="FIGURE 31. Male genitalia, Broscosoma resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067419" httpUri="https://zenodo.org/record/11067419/files/figure.png" pageId="59" pageNumber="144">Fig. 31c</figureCitation>
) compared with that in 
<taxonomicName id="4C06C199FFE4FF8EFF73FDF1A1F3FDB8" authorityName="Putzeys. We" authorityYear="1877" box="[151,245,574,598]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFE4FF8EFF73FDF1A1F3FDB8" box="[151,245,574,598]" italics="true" pageId="59" pageNumber="144">B. ribbei</emphasis>
</taxonomicName>
males.
</paragraph>
<paragraph id="8BB9BA1AFFE4FF8EFF4FFD91A58CFD78" blockId="59.[123,1174,219,1081]" pageId="59" pageNumber="144">
Among 
<taxonomicName id="4C06C199FFE4FF8EFEE6FD90A0DFFD98" box="[258,473,606,630]" pageId="59" pageNumber="144">
<emphasis id="B9726608FFE4FF8EFEE6FD90A083FD98" box="[258,389,607,630]" italics="true" pageId="59" pageNumber="144">Broscosoma</emphasis>
species
</taxonomicName>
known from outside the study area, none have members with the lateral margination of the pronotum evident, so none can be confused with 
<taxonomicName id="4C06C199FFE4FF8EFC47FDB0A51CFD78" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[931,1050,638,662]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFE4FF8EFC47FDB0A51CFD78" box="[931,1050,638,662]" italics="true" pageId="59" pageNumber="144">B. resbecqi</emphasis>
</taxonomicName>
members.
</paragraph>
<paragraph id="8BB9BA1AFFE4FF8EFF4FFD51A353FCD9" blockId="59.[123,1174,219,1081]" pageId="59" pageNumber="144">
<emphasis id="B9726608FFE4FF8EFF4FFD51A090FD59" bold="true" box="[171,406,670,695]" pageId="59" pageNumber="144">Habitat distribution.</emphasis>
The unique specimen of this species was found under a stone about 
<quantity id="4CFE17FFFFE4FF8EFB89FD50A590FD58" box="[1133,1174,671,695]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="59" pageNumber="144" unit="m" value="4.0">4 m</quantity>
back from the edge of a moderate-sized, cold, and fast-flowing stream descending through mixed coniferous/ broadleaf evergreen forest (
<figureCitation id="133DA69FFFE4FF8EFDC5FD10A37FFD18" box="[545,633,735,759]" captionStart="FIGURE" captionStartId="86.[171,183,1592,1611]" captionTargetBox="[147,1149,216,1572]" captionTargetId="figure-57@86.[147,1149,216,1572]" captionTargetPageId="86" captionText="FIGURE 46. Photographs of habitats for broscine species in the Gaoligong Shan region. a. Siran Wang, 0.2 km above confluence with Dulong Jiang, 1720 m, 28.013°/ E098.321º, Dulongjiang Township, Gongshan County, Yunnan (habitat for Broscosoma resbecqi sp. nov.); b. Nankang Yakou, 2130 m, 24.828°/98.767°, Longyang County, Yunnan (habitat for Broscosoma ribbei Putzeys and Broscus punctatus (Dejean). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067451" httpUri="https://zenodo.org/record/11067451/files/figure.png" pageId="59" pageNumber="144">Fig. 46a</figureCitation>
). The banks of the stream were largely overgrown with vegetation and shaded during most if not all of the day. The site of collection was at the relatively low elevation of 
<quantity id="4CFE17FFFFE4FF8EFE91FCEFA0C0FCD9" box="[373,454,800,824]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="59" pageNumber="144" unit="m" value="1720.0">1720 m</quantity>
(Fig. FX49).
</paragraph>
<paragraph id="8BB9BA1AFFE4FF8EFF4FFCF0A3C9FC76" blockId="59.[123,1174,219,1081]" pageId="59" pageNumber="144">
<emphasis id="B9726608FFE4FF8EFF4FFCF0A20AFCB6" bold="true" box="[171,780,831,856]" pageId="59" pageNumber="144">Geographical distribution within the Gaoligong Shan.</emphasis>
<figureCitation id="133DA69FFFE4FF8EFCF6FC8FA26CFCB6" box="[786,874,832,856]" captionStart="FIGURE" captionStartId="57.[171,183,1577,1596]" captionTargetBox="[135,1176,216,1562]" captionTargetId="figure-68@57.[120,1176,216,1563]" captionTargetPageId="57" captionText="FIGURE 30. Broscosoma resbecqi sp. nov. a. Habitus (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067417" httpUri="https://zenodo.org/record/11067417/files/figure.png" pageId="59" pageNumber="144">Fig. 30b</figureCitation>
. This species is known only from the type locality in Gongshan County (see Type material above for exact collection data) on the western slope of the range in Core Area 1 (
<figureCitation id="133DA69FFFE4FF8EFD97FC4FA3C6FC76" box="[627,704,896,920]" captionStart="FIGURE" captionStartId="88.[171,183,995,1014]" captionTargetBox="[130,1164,233,969]" captionTargetId="figure-57@88.[120,1176,215,989]" captionTargetPageId="88" captionText="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." figureDoi="http://doi.org/10.5281/zenodo.11067455" httpUri="https://zenodo.org/record/11067455/files/figure.png" pageId="59" pageNumber="144">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="8BB9BA1AFFE4FF8EFF4FFC6FA251FC36" blockId="59.[123,1174,219,1081]" pageId="59" pageNumber="144">
<emphasis id="B9726608FFE4FF8EFF4FFC6FA328FC57" bold="true" box="[171,558,928,953]" pageId="59" pageNumber="144">Overall geographical distribution.</emphasis>
This species currently is known only from the northwestern part of the Gaoligong Shan, in western 
<collectingRegion id="49C274F8FFE4FF8EFDAFFC0EA205FC37" box="[587,771,961,985]" country="China" name="Yunnan" pageId="59" pageNumber="144">Yunnan Province</collectingRegion>
, 
<collectingCountry id="F311FA8AFFE4FF8EFCF4FC0EA256FC37" box="[784,848,961,985]" name="China" pageId="59" pageNumber="144">China</collectingCountry>
.
</paragraph>
<paragraph id="8BB9BA1AFFE4FF8EFF4FFC2EA0D5FBD7" blockId="59.[123,1174,219,1081]" pageId="59" pageNumber="144">
<emphasis id="B9726608FFE4FF8EFF4FFC2EA24AFC17" bold="true" box="[171,844,993,1017]" pageId="59" pageNumber="144">
Geographical relationships with other 
<taxonomicName id="4C06C199FFE4FF8EFD8CFC2EA24AFC17" box="[616,844,993,1017]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9726608FFE4FF8EFD8CFC2EA3E9FC16" bold="true" box="[616,751,993,1016]" italics="true" pageId="59" pageNumber="144">Broscosoma</emphasis>
species.
</taxonomicName>
</emphasis>
No other 
<taxonomicName id="4C06C199FFE4FF8EFC59FC2DA593FC17" box="[957,1173,993,1017]" pageId="59" pageNumber="144">
<emphasis id="B9726608FFE4FF8EFC59FC2DA546FC17" box="[957,1088,994,1017]" italics="true" pageId="59" pageNumber="144">Broscosoma</emphasis>
species
</taxonomicName>
has been found syntopic with the unique 
<typeStatus id="54BD04B8FFE4FF8EFDD7FBCDA367FBF4" box="[563,609,1026,1050]" pageId="59" pageNumber="144">type</typeStatus>
of this species (
<figureCitation id="133DA69FFFE4FF8EFCF4FBCEA25BFBF7" box="[784,861,1025,1049]" captionStart="FIGURE" captionStartId="89.[171,183,1461,1480]" captionTargetBox="[140,1154,249,1428]" captionTargetId="figure-68@89.[120,1176,216,1448]" captionTargetPageId="89" captionText="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi="http://doi.org/10.5281/zenodo.11067459" httpUri="https://zenodo.org/record/11067459/files/figure.png" pageId="59" pageNumber="144">Fig. 50</figureCitation>
), although 
<taxonomicName id="4C06C199FFE4FF8EFC3CFBCDA590FBF7" authorityName="Putzeys" authorityYear="1877" box="[984,1174,1025,1049]" class="Insecta" family="Carabidae" genus="Brososoma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="144" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFE4FF8EFC3CFBCDA590FBF7" box="[984,1174,1025,1049]" italics="true" pageId="59" pageNumber="144">Brososoma ribbei</emphasis>
</taxonomicName>
occurs in the same general area.
</paragraph>
</subSubSection>
</treatment>
</document>