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<document id="34BBE06128568668123FAD96B4FF6DE1" ID-CLB-Dataset="299055" ID-DOI="10.5281/zenodo.11067355" ID-GBIF-Dataset="67ec2de4-1816-4969-8132-f6796d3afa06" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067355" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052343267" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2021" docId="03AF0B0CFFDAFFB2FF58FD4DA597FB18" docLanguage="en" docName="ProcCalifAcadSci.67.4.85-182.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (4)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." docType="key" docVersion="4" lastPageNumber="92" masterDocId="FF967374FFDFFFB5FFE4FFCFA106FFEE" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." masterLastPageNumber="182" masterPageNumber="85" pageNumber="90" updateTime="1719538714492" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8BF0438F085641EDE7EA80FBDB9B7DA5">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae).</mods:title>
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<mods:name id="1EBC2E1529B5AA7E35DEFB9A1D96FC6A" type="personal">
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<mods:namePart id="2CF63080520B90E40784F902EFEEE7AA">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5F80B5C6324BFAFC5B744E51AEF4F6A8">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="E95FCEE3C4F8BCEA859AE01BD127EA35">Liang, Hongbin</mods:namePart>
<mods:affiliation id="B6ED70FDB65E19974C3AAB213A748B11">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title id="B92A9DA490A67E46CDF771DAD5678B38">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="E3B9C168AAEF81201F097D488FA7F12E">
<mods:date id="8BB5184DB9115C88BBA7FF010FC7C3DA">2021</mods:date>
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<mods:number id="311A2F88455F5F156DA437A10BCFFABA">2021-09-30</mods:number>
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<mods:number id="08886F4E521F447A586E42E208DDF610">67</mods:number>
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<mods:classification id="C27C7E3C27EC9D1D23DDD72AD8E077DE">journal article</mods:classification>
<mods:identifier id="EA73C9D977B8C212BB3E4666A354A688" type="CLB-Dataset">299055</mods:identifier>
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<treatment id="03AF0B0CFFDAFFB2FF58FD4DA597FB18" ID-DOI="http://doi.org/10.5281/zenodo.12576227" ID-Zenodo-Dep="12576227" LSID="urn:lsid:plazi:treatment:03AF0B0CFFDAFFB2FF58FD4DA597FB18" httpUri="http://treatment.plazi.org/id/03AF0B0CFFDAFFB2FF58FD4DA597FB18" lastPageId="7" lastPageNumber="92" pageId="5" pageNumber="90">
<subSubSection id="C31CE991FFDAFFB0FF58FD4DA552FD74" box="[188,1108,641,666]" pageId="5" pageNumber="90" type="nomenclature">
<paragraph id="8BB9BA1AFFDAFFB0FF58FD4DA552FD74" blockId="5.[188,1108,641,666]" box="[188,1108,641,666]" pageId="5" pageNumber="90">
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<emphasis id="B9726608FFDAFFB0FF58FD4DA552FD74" bold="true" box="[188,1108,641,666]" pageId="5" pageNumber="90">
Key for identification of adults of broscine species of the Gaoligong
<collectingRegion id="49C274F8FFDAFFB0FC27FD4EA552FD74" box="[963,1108,641,666]" country="Myanmar" name="Shan" pageId="5" pageNumber="90">Shan Region</collectingRegion>
</emphasis>
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<subSubSection id="C31CE991FFDAFFB2FF9FFD62A597FB18" lastPageId="7" lastPageNumber="92" pageId="5" pageNumber="90" type="key">
<key id="FF97ED2BFFDAFFB3FF9FFD62A597FAAE" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
<keyStep id="30F2A2BFFFDAFFB0FF9FFD62A597FD0B" pageId="5" pageNumber="90">
<paragraph id="8BB9BA1AFFDAFFB0FF9FFD62A597FD2B" blockId="5.[123,1169,684,741]" box="[123,1169,684,709]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFD62A597FD2B" box="[123,1169,684,709]" pageId="5" pageNumber="90">1 Size large, BL of male or female greater than 15.0 mm............................ 2</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFDAFFB0FF9FFD02A597FD0B" blockId="5.[123,1169,684,741]" box="[123,1169,717,741]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFD02A597FD0B" box="[123,1169,717,741]" pageId="5" pageNumber="90">1’ Size moderate, BL of male or female less than 13.0 mm........................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F2A2BFFFDAFFB0FF9FFD37A597FC5E" pageId="5" pageNumber="90">
<paragraph id="8BB9BA1AFFDAFFB0FF9FFD37A597FCA1" blockId="5.[123,1174,759,944]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFD37A597FCA1" pageId="5" pageNumber="90">
2(1) Head (
<figureCitation id="133DA69FFFDAFFB0FF1BFD38A04CFCE1" box="[255,330,759,783]" captionStart="FIGURE" captionStartId="8.[171,183,1513,1532]" captionTargetBox="[139,1176,224,1495]" captionTargetId="figure-55@8.[120,1176,216,1495]" captionTargetPageId="8" captionText="FIGURE 5. Forebody of broscine carabids, dorsal aspect. a. Eoboscus bhutanensis Morvan (CASENT1019340; Lishadi Township, Fugong County, Yunnan, China); b. Broscus punctatus (Dejean) (CASENT1011066; Heiwadi, Cikai Township, Gongshan County, Yunnan, China); c. Broscodera chukuai sp. nov. (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo Township, Gongshan County, Yunnan, China); d. Broscodera gaoligongensis sp. nov. (CASENT1020104; South Fork of Yamu He, Lumadeng Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067361" httpUri="https://zenodo.org/record/11067361/files/figure.png" pageId="5" pageNumber="90">Fig. 5a</figureCitation>
) with very deep, sharply defined and impunctate cleft on vertex between tempora; male with protarsomeres 1 and 2 and mesotarsomeres 1 and 2 with ventral pads of adhesive setae … Genus
<taxonomicName id="4C06C199FFDAFFB0FE90FCF7A39FFCA1" authority="Kryzhanovskij." authorityName="Kryzhanovskij." box="[372,665,824,848]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9726608FFDAFFB0FE90FCF7A0E2FCBE" box="[372,484,824,848]" italics="true" pageId="5" pageNumber="90">Eobroscus</emphasis>
Kryzhanovskij.
</taxonomicName>
............
<taxonomicName id="4C06C199FFDAFFB0FCDBFCF7A597FCA1" authority="Morvan" authorityName="Morvan" box="[831,1169,824,848]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="bhutanensis">
<emphasis id="B9726608FFDAFFB0FCDBFCF7A533FCBE" box="[831,1077,824,848]" italics="true" pageId="5" pageNumber="90">Eobroscus bhutanensis</emphasis>
Morvan
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFDAFFB0FF9FFC97A597FC5E" blockId="5.[123,1174,759,944]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFC97A597FC5E" pageId="5" pageNumber="90">
2’ Head (
<figureCitation id="133DA69FFFDAFFB0FF14FC97A03DFC9E" box="[240,315,856,880]" captionStart="FIGURE" captionStartId="8.[171,183,1513,1532]" captionTargetBox="[139,1176,224,1495]" captionTargetId="figure-55@8.[120,1176,216,1495]" captionTargetPageId="8" captionText="FIGURE 5. Forebody of broscine carabids, dorsal aspect. a. Eoboscus bhutanensis Morvan (CASENT1019340; Lishadi Township, Fugong County, Yunnan, China); b. Broscus punctatus (Dejean) (CASENT1011066; Heiwadi, Cikai Township, Gongshan County, Yunnan, China); c. Broscodera chukuai sp. nov. (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo Township, Gongshan County, Yunnan, China); d. Broscodera gaoligongensis sp. nov. (CASENT1020104; South Fork of Yamu He, Lumadeng Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067361" httpUri="https://zenodo.org/record/11067361/files/figure.png" pageId="5" pageNumber="90">Fig. 5b</figureCitation>
) with shallower, broader, and distinctly punctate transverse sulcus between tempora; male protarsomeres 1 to 3 with ventral pads of adhesive setae, all male mesotarsomeres lacking ventral pads of adhesive setae … Genus
<taxonomicName id="4C06C199FFDAFFB0FD47FC56A25CFC5E" authority="Panzer." authorityName="Panzer." box="[675,858,921,944]" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9726608FFDAFFB0FD47FC56A3FEFC5E" box="[675,760,921,944]" italics="true" pageId="5" pageNumber="90">Broscus</emphasis>
Panzer.
</taxonomicName>
<taxonomicName id="4C06C199FFDAFFB0FC8CFC56A597FC5E" authority="(Dejean)" authorityName="" authorityYear="1828" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[872,1169,920,944]" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="B9726608FFDAFFB0FC8CFC56A52AFC5E" box="[872,1068,921,944]" italics="true" pageId="5" pageNumber="90">Broscus punctatus</emphasis>
(Dejean)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F2A2BFFFDAFFB0FF9FFC0CA597FB75" pageId="5" pageNumber="90">
<paragraph id="8BB9BA1AFFDAFFB0FF9FFC0CA597FBD5" blockId="5.[123,1174,963,1180]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFC0CA597FBD5" pageId="5" pageNumber="90">
3(1’) Pronotum (
<figureCitation id="133DA69FFFDAFFB0FEDFFC0CA084FC35" box="[315,386,963,987]" captionStart="FIGURE" captionStartId="8.[171,183,1513,1532]" captionTargetBox="[139,1176,224,1495]" captionTargetId="figure-55@8.[120,1176,216,1495]" captionTargetPageId="8" captionText="FIGURE 5. Forebody of broscine carabids, dorsal aspect. a. Eoboscus bhutanensis Morvan (CASENT1019340; Lishadi Township, Fugong County, Yunnan, China); b. Broscus punctatus (Dejean) (CASENT1011066; Heiwadi, Cikai Township, Gongshan County, Yunnan, China); c. Broscodera chukuai sp. nov. (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo Township, Gongshan County, Yunnan, China); d. Broscodera gaoligongensis sp. nov. (CASENT1020104; South Fork of Yamu He, Lumadeng Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067361" httpUri="https://zenodo.org/record/11067361/files/figure.png" pageId="5" pageNumber="90">Figs. 5</figureCitation>
c-d) with basolateral setae present; antennomere 3 with sparse pubescence in apical one-third to one half, antennomere 4 pubescent; male protarsomeres and mesotarsomeres 1 and 2 with small ventral pads of adhesive setae … Genus
<taxonomicName id="4C06C199FFDAFFB0FC9BFBCCA559FBF5" authority="Lindroth" authorityName="Lindroth" authorityYear="1961" box="[895,1119,1027,1051]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9726608FFDAFFB0FC9BFBCCA2FCFBF5" box="[895,1018,1027,1051]" italics="true" pageId="5" pageNumber="90">Broscodera</emphasis>
Lindroth
</taxonomicName>
subgenus
<taxonomicName id="4C06C199FFDAFFB0FF39FBECA0DDFBD5" authority="Deuve." authorityName="Deuve." box="[221,475,1059,1083]" pageId="5" pageNumber="90" rank="subGenus" subGenus="Sinobrosculus">
<emphasis id="B9726608FFDAFFB0FF39FBECA075FBD5" box="[221,371,1059,1083]" italics="true" pageId="5" pageNumber="90">Sinobrosculus</emphasis>
Deuve.
</taxonomicName>
.................................................. 4
</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFDAFFB0FF9FFB8CA597FB75" blockId="5.[123,1174,963,1180]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFB8CA597FB75" pageId="5" pageNumber="90">
3’ Pronotum (
<figureCitation id="133DA69FFFDAFFB0FEC7FB8CA06AFBB5" box="[291,364,1091,1115]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="5" pageNumber="90">Figs. 6</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FE9EFB8BA081FBB5" box="[378,391,1092,1115]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="5" pageNumber="90">7</figureCitation>
) with basolateral setae absent; antennomeres 3 and 4 glabrous except for apical whorl of setae; male protarsomeres 1 to 3 with larger ventral pads of adhesive setae, mesotarsomeres 1 and 2 with larger ventral pads … Genus
<taxonomicName id="4C06C199FFDAFFB0FCF1FB4BA535FB75" authority="Rosenhauer." authorityName="Rosenhauer." box="[789,1075,1156,1180]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9726608FFDAFFB0FCF1FB4BA29EFB75" box="[789,920,1156,1179]" italics="true" pageId="5" pageNumber="90">Broscosoma</emphasis>
Rosenhauer.
</taxonomicName>
..... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F2A2BFFFDAFFB0FF9FFB60A597FA09" pageId="5" pageNumber="90">
<paragraph id="8BB9BA1AFFDAFFB0FF9FFB60A597FAA9" blockId="5.[123,1174,1198,1511]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFB60A597FAA9" pageId="5" pageNumber="90">
4 (3) Size larger, BL = 10.0 mm or more; body color piceous or dark brown; head (
<figureCitation id="133DA69FFFDAFFB0FBEEFB61A553FB28" box="[1034,1109,1198,1222]" captionStart="FIGURE" captionStartId="8.[171,183,1513,1532]" captionTargetBox="[139,1176,224,1495]" captionTargetId="figure-55@8.[120,1176,216,1495]" captionTargetPageId="8" captionText="FIGURE 5. Forebody of broscine carabids, dorsal aspect. a. Eoboscus bhutanensis Morvan (CASENT1019340; Lishadi Township, Fugong County, Yunnan, China); b. Broscus punctatus (Dejean) (CASENT1011066; Heiwadi, Cikai Township, Gongshan County, Yunnan, China); c. Broscodera chukuai sp. nov. (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo Township, Gongshan County, Yunnan, China); d. Broscodera gaoligongensis sp. nov. (CASENT1020104; South Fork of Yamu He, Lumadeng Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067361" httpUri="https://zenodo.org/record/11067361/files/figure.png" pageId="5" pageNumber="90">Fig. 5d</figureCitation>
) with eyes more convex and slightly larger, their diameter equal to length of tempora, vertex with post-temporal transverse sulcus densely, coarsely and, more widely punctate; hindwings full-sized, functional for flight; hind trochanter asetose (
<figureCitation id="133DA69FFFDAFFB0FD65FAC0A3CAFAC8" box="[641,716,1295,1319]" captionStart="FIGURE" captionStartId="11.[171,183,1003,1022]" captionTargetBox="[120,1176,216,984]" captionTargetId="figure-67@11.[120,1176,216,985]" captionTargetPageId="11" captionText="FIGURE 8. Left metatrochanter, broscine species, ventral aspect. a. Broscodera gaoligongensis sp. nov. (Yaojiaping, Lushui County, Yunnan, China); b. Broscodera chukuai sp. nov. (CASENT1026771; Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1034187; Qiqi-Dulong Yakou, Cikai Township, Gongshan County, Yunnan, China); d. B. ribbei Putzeys (CASENT1000578; Nankang Yakou, Longyang County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067367" httpUri="https://zenodo.org/record/11067367/files/figure.png" pageId="5" pageNumber="90">Fig. 8a</figureCitation>
); male genitalia as in
<figureCitation id="133DA69FFFDAFFB0FC58FAC0A50EFAC9" box="[956,1032,1295,1319]" captionStart="FIGURE" captionStartId="21.[171,183,1024,1043]" captionTargetBox="[169,1140,222,993]" captionTargetId="figure-1454@21.[120,1173,216,1006]" captionTargetPageId="21" captionText="FIGURE 14. Male genitalia, Broscodera gaoligongensis sp. nov. (Yaojiaping, Lushui County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067379" httpUri="https://zenodo.org/record/11067379/files/figure.png" pageId="5" pageNumber="90">Fig. 14</figureCitation>
.......................................................
<emphasis id="B9726608FFDAFFB0FCEEFAE1A597FAA9" bold="true" box="[778,1169,1326,1351]" pageId="5" pageNumber="90">
<taxonomicName id="4C06C199FFDAFFB0FCEEFAE1A535FAA8" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[778,1075,1326,1350]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="gaoligongensis" status="sp. nov.">
<emphasis id="B9726608FFDAFFB0FCEEFAE1A535FAA8" bold="true" box="[778,1075,1326,1350]" italics="true" pageId="5" pageNumber="90">Broscodera gaoligongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFDAFFB0FBDFFAE0A597FAA9" box="[1083,1169,1327,1351]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFDAFFB0FF9FFA80A597FA09" blockId="5.[123,1174,1198,1511]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFA80A597FA09" pageId="5" pageNumber="90">
4’ Size smaller, BL =
<quantity id="4CFE17FFFFDAFFB0FE9FFA80A0CAFA88" box="[379,460,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="90" unit="mm" value="9.5">9.5 mm</quantity>
or less; body color reddish brown; head (
<figureCitation id="133DA69FFFDAFFB0FC6AFA80A2DEFA88" box="[910,984,1359,1383]" captionStart="FIGURE" captionStartId="8.[171,183,1513,1532]" captionTargetBox="[139,1176,224,1495]" captionTargetId="figure-55@8.[120,1176,216,1495]" captionTargetPageId="8" captionText="FIGURE 5. Forebody of broscine carabids, dorsal aspect. a. Eoboscus bhutanensis Morvan (CASENT1019340; Lishadi Township, Fugong County, Yunnan, China); b. Broscus punctatus (Dejean) (CASENT1011066; Heiwadi, Cikai Township, Gongshan County, Yunnan, China); c. Broscodera chukuai sp. nov. (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo Township, Gongshan County, Yunnan, China); d. Broscodera gaoligongensis sp. nov. (CASENT1020104; South Fork of Yamu He, Lumadeng Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067361" httpUri="https://zenodo.org/record/11067361/files/figure.png" pageId="5" pageNumber="90">Fig. 5c</figureCitation>
) with eyes flatter and smaller, their diameter less than length of tempora, vertex with post-temporal transverse sulcus narrowly punctate only along bottom of sulcus; hindwings slightly reduced, non-functional for flight; hind trochanter unisetose (
<figureCitation id="133DA69FFFDAFFB0FDC0FA60A376FA29" box="[548,624,1455,1479]" captionStart="FIGURE" captionStartId="11.[171,183,1003,1022]" captionTargetBox="[120,1176,216,984]" captionTargetId="figure-67@11.[120,1176,216,985]" captionTargetPageId="11" captionText="FIGURE 8. Left metatrochanter, broscine species, ventral aspect. a. Broscodera gaoligongensis sp. nov. (Yaojiaping, Lushui County, Yunnan, China); b. Broscodera chukuai sp. nov. (CASENT1026771; Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1034187; Qiqi-Dulong Yakou, Cikai Township, Gongshan County, Yunnan, China); d. B. ribbei Putzeys (CASENT1000578; Nankang Yakou, Longyang County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067367" httpUri="https://zenodo.org/record/11067367/files/figure.png" pageId="5" pageNumber="90">Fig. 8b</figureCitation>
); male genitalia as in
<figureCitation id="133DA69FFFDAFFB0FC84FA60A2AAFA29" box="[864,940,1455,1479]" captionStart="FIGURE" captionStartId="16.[171,183,933,952]" captionTargetBox="[147,1135,222,918]" captionTargetId="figure-1483@16.[120,1176,210,928]" captionTargetPageId="16" captionText="FIGURE 12. Male genitalia, Broscodera chukuai sp. nov. (CASENT1026169; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067375" httpUri="https://zenodo.org/record/11067375/files/figure.png" pageId="5" pageNumber="90">Fig. 12</figureCitation>
....................................................................
<emphasis id="B9726608FFDAFFB0FCB0FA00A597FA09" bold="true" box="[852,1169,1487,1511]" pageId="5" pageNumber="90">
<taxonomicName id="4C06C199FFDAFFB0FCB0FA00A535FA09" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[852,1075,1487,1511]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="chukuai" status="sp. nov.">
<emphasis id="B9726608FFDAFFB0FCB0FA00A535FA09" bold="true" box="[852,1075,1487,1511]" italics="true" pageId="5" pageNumber="90">Broscodera chukuai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFDAFFB0FBDFFA00A597FA09" box="[1083,1169,1487,1511]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F2A2BFFFDAFFB0FF9FFA35A597F9BD" pageId="5" pageNumber="90">
<paragraph id="8BB9BA1AFFDAFFB0FF9FFA35A597F9DC" blockId="5.[123,1173,1530,1619]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FFA35A597F9DC" pageId="5" pageNumber="90">
5 Elytral silhouette subparallel to subovoid, with humeri (shoulders) more (
<figureCitation id="133DA69FFFDAFFB0FC25FA35A522F9FC" box="[961,1060,1530,1554]" captionStart="FIGURE" captionStartId="44.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1568]" captionTargetId="figure-56@44.[120,1176,216,1568]" captionTargetPageId="44" captionText="FIGURE 24. Broscosoma holomarginatum sp. nov. a. Habitus (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067401" httpUri="https://zenodo.org/record/11067401/files/figure.png" pageId="5" pageNumber="90">Figs. 24a</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FBD6FA35A55EF9FC" box="[1074,1112,1530,1554]" captionStart="FIGURE" captionStartId="52.[171,183,1586,1605]" captionTargetBox="[119,1175,216,1566]" captionTargetId="figure-56@52.[119,1175,216,1566]" captionTargetPageId="52" captionText="FIGURE 28. Broscosoma purpureum sp. nov. a. Habitus (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067411" httpUri="https://zenodo.org/record/11067411/files/figure.png" pageId="5" pageNumber="90">28a</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FB82FA35A58BF9FC" box="[1126,1165,1530,1554]" captionStart="FIGURE" captionStartId="61.[171,183,1593,1612]" captionTargetBox="[120,1174,216,1575]" captionTargetId="figure-68@61.[120,1175,216,1577]" captionTargetPageId="61" captionText="FIGURE 32. Broscosoma ribbei Putzeys. a. Habitus (CASENT1021037; 2.8 km above Shibali, Lishadi Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067421" httpUri="https://zenodo.org/record/11067421/files/figure.png" pageId="5" pageNumber="90">32a</figureCitation>
) or less (
<figureCitation id="133DA69FFFDAFFB0FF09F9D5A056F9DC" box="[237,336,1562,1586]" captionStart="FIGURE" captionStartId="29.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1596]" captionTargetId="figure-68@29.[120,1176,216,1598]" captionTargetPageId="29" captionText="FIGURE 17. Broscosoma danzhuense sp. nov. a. Habitus (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067385" httpUri="https://zenodo.org/record/11067385/files/figure.png" pageId="5" pageNumber="90">Figs. 17a</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FEBAF9D5A082F9DC" box="[350,388,1562,1586]" captionStart="FIGURE" captionStartId="48.[171,183,1587,1606]" captionTargetBox="[132,1176,216,1571]" captionTargetId="figure-56@48.[120,1176,216,1571]" captionTargetPageId="48" captionText="FIGURE 26. Broscosoma parvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067407" httpUri="https://zenodo.org/record/11067407/files/figure.png" pageId="5" pageNumber="90">26a</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FE76F9D5A0BFF9DC" box="[402,441,1562,1586]" captionStart="FIGURE" captionStartId="57.[171,183,1577,1596]" captionTargetBox="[135,1176,216,1562]" captionTargetId="figure-68@57.[120,1176,216,1563]" captionTargetPageId="57" captionText="FIGURE 30. Broscosoma resbecqi sp. nov. a. Habitus (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067417" httpUri="https://zenodo.org/record/11067417/files/figure.png" pageId="5" pageNumber="90">30a</figureCitation>
) distinct............................................. 6
</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFDAFFB0FF9FF9F5A597F9BD" blockId="5.[123,1173,1530,1619]" box="[123,1169,1594,1619]" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB0FF9FF9F5A597F9BD" box="[123,1169,1594,1619]" pageId="5" pageNumber="90">
5’ Elytral silhouette ovoid, with humeri indistinct (
<figureCitation id="133DA69FFFDAFFB0FD57F9F5A210F9BC" box="[691,790,1594,1618]" captionStart="FIGURE" captionStartId="25.[171,183,1609,1628]" captionTargetBox="[139,1176,216,1587]" captionTargetId="figure-67@25.[120,1176,216,1587]" captionTargetPageId="25" captionText="FIGURE 15. Broscosoma bicoloratum sp. nov. a. Habitus (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067381" httpUri="https://zenodo.org/record/11067381/files/figure.png" pageId="5" pageNumber="90">Figs. 15a</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FCC7F9F4A24CF9BC" box="[803,842,1595,1619]" captionStart="FIGURE" captionStartId="33.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1586]" captionTargetId="figure-67@33.[120,1176,216,1586]" captionTargetPageId="33" captionText="FIGURE 18. Broscosoma furvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 k." figureDoi="http://doi.org/10.5281/zenodo.11067389" httpUri="https://zenodo.org/record/11067389/files/figure.png" pageId="5" pageNumber="90">18a</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FCBCF9F4A279F9BC" box="[856,895,1595,1619]" captionStart="FIGURE" captionStartId="37.[171,183,1600,1619]" captionTargetBox="[125,1176,216,1581]" captionTargetId="figure-67@37.[120,1176,216,1586]" captionTargetPageId="37" captionText="FIGURE 20. Broscosoma gaoligongense Deuve and Wrase. a. Habitus (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067393" httpUri="https://zenodo.org/record/11067393/files/figure.png" pageId="5" pageNumber="90">20a</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FC68F9F4A2B5F9BC" box="[908,947,1595,1619]" captionStart="FIGURE" captionStartId="40.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1566]" captionTargetId="figure-56@40.[120,1176,216,1566]" captionTargetPageId="40" captionText="FIGURE 22. Broscosoma gongshanense sp. nov. a. Habitus (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067397" httpUri="https://zenodo.org/record/11067397/files/figure.png" pageId="5" pageNumber="90">22a</figureCitation>
,
<figureCitation id="133DA69FFFDAFFB0FC25F9F4A2EEF9BC" box="[961,1000,1595,1619]" captionStart="FIGURE" captionStartId="68.[171,183,1584,1603]" captionTargetBox="[120,1176,216,1563]" captionTargetId="figure-56@68.[120,1176,216,1563]" captionTargetPageId="68" captionText="FIGURE 34. Broscosoma viridicollare sp. nov. a. Habitus (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067427" httpUri="https://zenodo.org/record/11067427/files/figure.png" pageId="5" pageNumber="90">34a</figureCitation>
)......... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F2A2BFFFDAFFB3FF9FF9AAA597FEDD" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
<paragraph id="8BB9BA1AFFDAFFB3FF9FF9AAA597FEFD" blockId="5.[123,1174,1637,1725]" lastBlockId="6.[123,1169,219,308]" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
<keyLead id="30F7192FFFDAFFB3FF9FF9AAA597FEFD" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
6 (5’) Pronotum with lateral margination (lateral bead) complete (
<figureCitation id="133DA69FFFDAFFB0FC80F9AAA2B7F993" box="[868,945,1637,1661]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="5" pageNumber="90">Fig. 9a</figureCitation>
) between apical and basal margins; dorsal surface with distinct blue or blue-green luster, legs pale rufous; elytral microsculpture distinct, comprised of isodiametric to slightly longitudinally stretched sculpticells, elytral striae distinctly punctate; male genitalia as in
<figureCitation id="133DA69FFFD9FFB3FCEAFF14A25DFF1D" box="[782,859,219,243]" captionStart="FIGURE" captionStartId="45.[171,183,939,958]" captionTargetBox="[137,1161,228,920]" captionTargetId="figure-1604@45.[120,1176,215,935]" captionTargetPageId="45" captionText="FIGURE 25. Male genitalia, Broscosoma holomarginatum sp. nov. (CASENT1016743; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067405" httpUri="https://zenodo.org/record/11067405/files/figure.png" pageId="6" pageNumber="91">Fig. 25</figureCitation>
..................................................................
<emphasis id="B9726608FFD9FFB3FD08FF33A597FEFD" bold="true" box="[748,1169,251,275]" pageId="6" pageNumber="91">
<taxonomicName id="4C06C199FFD9FFB3FD08FF33A532FEFD" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[748,1076,251,275]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="holomarginatum" status="sp. nov.">
<emphasis id="B9726608FFD9FFB3FD08FF33A532FEFD" bold="true" box="[748,1076,251,275]" italics="true" pageId="6" pageNumber="91">Broscosoma holomarginatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD9FFB3FBDFFF34A597FEFD" box="[1083,1169,251,275]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFD9FFB3FF9FFED3A597FEDD" blockId="6.[123,1169,219,308]" box="[123,1169,284,308]" pageId="6" pageNumber="91">
<keyLead id="30F7192FFFD9FFB3FF9FFED3A597FEDD" box="[123,1169,284,308]" pageId="6" pageNumber="91">
6’ Pronotum with lateral margination absent or incomplete (
<figureCitation id="133DA69FFFD9FFB3FCF1FED3A258FEDA" box="[789,862,284,308]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="6" pageNumber="91">Figs. 9</figureCitation>
b-d)................. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F2A2BFFFD9FFB3FF9FFE87A596FD8C" pageId="6" pageNumber="91">
<paragraph id="8BB9BA1AFFD9FFB3FF9FFE87A597FDEF" blockId="6.[123,1174,327,610]" pageId="6" pageNumber="91">
<keyLead id="30F7192FFFD9FFB3FF9FFE87A597FDEF" pageId="6" pageNumber="91">
7 (6’) Pronotum with lateral margination complete anterior to midlateral seta, interrupted briefly in basal half (
<figureCitation id="133DA69FFFD9FFB3FEC9FEA8A07CFE91" box="[301,378,359,384]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="6" pageNumber="91">Fig. 9b</figureCitation>
); elytral humeri as in
<figureCitation id="133DA69FFFD9FFB3FD89FEA8A3BCFE6E" box="[621,698,359,384]" captionStart="FIGURE" captionStartId="57.[171,183,1577,1596]" captionTargetBox="[135,1176,216,1562]" captionTargetId="figure-68@57.[120,1176,216,1563]" captionTargetPageId="57" captionText="FIGURE 30. Broscosoma resbecqi sp. nov. a. Habitus (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067417" httpUri="https://zenodo.org/record/11067417/files/figure.png" pageId="6" pageNumber="91">Fig. 30</figureCitation>
; dorsal surface of head and pronotum vivid metallic green, elytra darker blue-green, legs pale rufous; vertex with post-temporal transverse sulcus relatively shallow and broadly and densely punctate (
<figureCitation id="133DA69FFFD9FFB3FCD8FE67A28DFE2E" box="[828,907,424,448]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="6" pageNumber="91">Fig. 6d</figureCitation>
); elytral microsculpture effaced; metatrochanter asetose (
<figureCitation id="133DA69FFFD9FFB3FE1DFE06A342FE0E" box="[505,580,457,481]" captionStart="FIGURE" captionStartId="11.[171,183,1003,1022]" captionTargetBox="[120,1176,216,984]" captionTargetId="figure-67@11.[120,1176,216,985]" captionTargetPageId="11" captionText="FIGURE 8. Left metatrochanter, broscine species, ventral aspect. a. Broscodera gaoligongensis sp. nov. (Yaojiaping, Lushui County, Yunnan, China); b. Broscodera chukuai sp. nov. (CASENT1026771; Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1034187; Qiqi-Dulong Yakou, Cikai Township, Gongshan County, Yunnan, China); d. B. ribbei Putzeys (CASENT1000578; Nankang Yakou, Longyang County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067367" httpUri="https://zenodo.org/record/11067367/files/figure.png" pageId="6" pageNumber="91">Fig. 8c</figureCitation>
); male genitalia as in
<figureCitation id="133DA69FFFD9FFB3FCD7FE06A279FE0F" box="[819,895,457,481]" captionStart="FIGURE" captionStartId="53.[171,183,939,958]" captionTargetBox="[151,1168,221,932]" captionTargetId="figure-1570@53.[120,1176,216,938]" captionTargetPageId="53" captionText="FIGURE 29. Male genitalia, Broscosoma purpureum sp. nov. (CASENT1015869; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral Maspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067415" httpUri="https://zenodo.org/record/11067415/files/figure.png" pageId="6" pageNumber="91">Fig. 29</figureCitation>
......................................................................
<emphasis id="B9726608FFD9FFB3FCAFFE26A597FDEF" bold="true" box="[843,1169,488,513]" pageId="6" pageNumber="91">
<taxonomicName id="4C06C199FFD9FFB3FCAFFE26A535FDEE" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[843,1075,488,512]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="resbecqi" status="sp. nov.">
<emphasis id="B9726608FFD9FFB3FCAFFE26A535FDEE" bold="true" box="[843,1075,488,512]" italics="true" pageId="6" pageNumber="91">Broscosoma resbecqi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD9FFB3FBDEFE26A597FDEF" box="[1082,1169,489,513]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFD9FFB3FF9FFDC5A596FD8C" blockId="6.[123,1174,327,610]" pageId="6" pageNumber="91">
<keyLead id="30F7192FFFD9FFB3FF9FFDC5A596FD8C" pageId="6" pageNumber="91">
7’ Pronotum with lateral margination absent, present only posterior to sub-basal constriction, or also present in area of midlateral seta (in a few specimens of
<taxonomicName id="4C06C199FFD9FFB3FCCDFDE5A281FDAC" authorityName="Putzeys. We" authorityYear="1877" box="[809,903,554,578]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFD9FFB3FCCDFDE5A281FDAC" box="[809,903,554,578]" italics="true" pageId="6" pageNumber="91">B. ribbei</emphasis>
</taxonomicName>
extended almost to anterior margin (
<figureCitation id="133DA69FFFD9FFB3FEC5FD85A06DFD8C" box="[289,363,586,610]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="6" pageNumber="91">Fig. 9c</figureCitation>
), but these specimens have the metatrochanter unisetose)........... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F2A2BFFFD9FFB3FF9FFDB9A597FBBD" pageId="6" pageNumber="91">
<paragraph id="8BB9BA1AFFD9FFB3FF9FFDB9A597FC7F" blockId="6.[123,1174,629,1107]" pageId="6" pageNumber="91">
<keyLead id="30F7192FFFD9FFB3FF9FFDB9A597FC7F" pageId="6" pageNumber="91">
8(7’) Elytral microsculpture distinct, comprised of moderately impressed isodiametric sculpticells; metatrochanters unisetose (
<figureCitation id="133DA69FFFD9FFB3FE59FD59A30FFD40" box="[445,521,662,686]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="6" pageNumber="91">Fig. 9d</figureCitation>
) (except asetose unilaterally in some specimens); elytral silhouette as in
<figureCitation id="133DA69FFFD9FFB3FEC7FD79A07DFD20" box="[291,379,694,718]" captionStart="FIGURE" captionStartId="61.[171,183,1593,1612]" captionTargetBox="[120,1174,216,1575]" captionTargetId="figure-68@61.[120,1175,216,1577]" captionTargetPageId="61" captionText="FIGURE 32. Broscosoma ribbei Putzeys. a. Habitus (CASENT1021037; 2.8 km above Shibali, Lishadi Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067421" httpUri="https://zenodo.org/record/11067421/files/figure.png" pageId="6" pageNumber="91">Fig. 32a</figureCitation>
; size larger, BL =
<quantity id="4CFE17FFFFD9FFB3FDDAFD79A388FD20" box="[574,654,694,718]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="6" pageNumber="91" unit="mm" value="8.9">8.9 mm</quantity>
or more; pronotum globose anterior to sub-basal constriction as in
<figureCitation id="133DA69FFFD9FFB3FEB3FD18A0A5FD01" box="[343,419,726,751]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="6" pageNumber="91">Fig. 6b</figureCitation>
, in most specimens with lateral margination present in region of midlateral seta (
<figureCitation id="133DA69FFFD9FFB3FEFFFD38A060FCE0" box="[283,358,759,783]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="6" pageNumber="91">Fig. 9c</figureCitation>
) and also basal to sub-basal constriction and/or extended further anterior to midlateral seta in some specimens (absent entirely from a few specimens); vertex with post-temporal transverse sulcus relatively shallow and narrowly and sparsely punctate (
<figureCitation id="133DA69FFFD9FFB3FC16FCF7A539FCBE" box="[1010,1087,824,848]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="6" pageNumber="91">Fig. 6b</figureCitation>
); entire dorsum of body bright metallic golden-green, green (
<figureCitation id="133DA69FFFD9FFB3FD37FC97A22AFC9E" box="[723,812,856,880]" captionStart="FIGURE" captionStartId="61.[171,183,1593,1612]" captionTargetBox="[120,1174,216,1575]" captionTargetId="figure-68@61.[120,1175,216,1577]" captionTargetPageId="61" captionText="FIGURE 32. Broscosoma ribbei Putzeys. a. Habitus (CASENT1021037; 2.8 km above Shibali, Lishadi Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067421" httpUri="https://zenodo.org/record/11067421/files/figure.png" pageId="6" pageNumber="91">Fig. 32a</figureCitation>
), or blue-green; male genitalia as in
<figureCitation id="133DA69FFFD9FFB3FF50FCB7A006FC7F" box="[180,256,888,913]" captionStart="FIGURE" captionStartId="62.[171,183,976,995]" captionTargetBox="[137,1162,224,952]" captionTargetId="figure-1498@62.[120,1176,216,972]" captionTargetPageId="62" captionText="FIGURE 33. Male genitalia, Broscosoma ribbei Putzeys (CASENT1025422; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067425" httpUri="https://zenodo.org/record/11067425/files/figure.png" pageId="6" pageNumber="91">Fig. 33</figureCitation>
.............................................
<taxonomicName id="4C06C199FFD9FFB3FC89FCB6A597FC7F" authority="Putzeys" authorityName="Putzeys. We" authorityYear="1877" box="[877,1169,888,913]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFD9FFB3FC89FCB6A531FC7E" box="[877,1079,888,912]" italics="true" pageId="6" pageNumber="91">Broscosoma ribbei</emphasis>
Putzeys
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFD9FFB3FF9FFC56A597FBBD" blockId="6.[123,1174,629,1107]" pageId="6" pageNumber="91">
<keyLead id="30F7192FFFD9FFB3FF9FFC56A597FBBD" pageId="6" pageNumber="91">
8’ Elytral microsculpture faintly impressed or effaced; metatrochanters asetose (
<figureCitation id="133DA69FFFD9FFB3FC33FC56A526FC5E" box="[983,1056,921,945]" captionStart="FIGURE" captionStartId="11.[171,183,1003,1022]" captionTargetBox="[120,1176,216,984]" captionTargetId="figure-67@11.[120,1176,216,985]" captionTargetPageId="11" captionText="FIGURE 8. Left metatrochanter, broscine species, ventral aspect. a. Broscodera gaoligongensis sp. nov. (Yaojiaping, Lushui County, Yunnan, China); b. Broscodera chukuai sp. nov. (CASENT1026771; Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1034187; Qiqi-Dulong Yakou, Cikai Township, Gongshan County, Yunnan, China); d. B. ribbei Putzeys (CASENT1000578; Nankang Yakou, Longyang County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067367" httpUri="https://zenodo.org/record/11067367/files/figure.png" pageId="6" pageNumber="91">Fig. 8c</figureCitation>
) (although a few specimens of
<taxonomicName id="4C06C199FFD9FFB3FE97FC75A30EFC3F" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[371,520,954,977]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="purpureum">
<emphasis id="B9726608FFD9FFB3FE97FC75A30EFC3F" box="[371,520,954,977]" italics="true" pageId="6" pageNumber="91">B. purpureum</emphasis>
</taxonomicName>
have one or both metatrochanters unisetose); size generally smaller, BL =
<quantity id="4CFE17FFFFD9FFB3FED1FC15A08EFC1F" box="[309,392,985,1010]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="91" unit="mm" value="8.5">8.5 mm</quantity>
or less (except BL of
<taxonomicName id="4C06C199FFD9FFB3FD64FC15A21AFC1C" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[640,796,986,1010]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="danzhuense">
<emphasis id="B9726608FFD9FFB3FD64FC15A21AFC1C" box="[640,796,986,1010]" italics="true" pageId="6" pageNumber="91">B. danzhuense</emphasis>
</taxonomicName>
<typeStatus id="54BD04B8FFD9FFB3FCC1FC15A284FC1C" box="[805,898,986,1010]" pageId="6" pageNumber="91" type="holotype">holotype</typeStatus>
=
<quantity id="4CFE17FFFFD9FFB3FC47FC15A2F3FC1F" box="[931,1013,986,1010]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="6" pageNumber="91" unit="mm" value="9.3">9.3 mm</quantity>
); pronotum of different shape (
<figureCitation id="133DA69FFFD9FFB3FEADFC35A0C4FBFC" box="[329,450,1018,1042]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="6" pageNumber="91">Figs. 6c,e,f</figureCitation>
), with lateral margination absent or present only at or posterior to sub-basal constriction (
<figureCitation id="133DA69FFFD9FFB3FE75FBD5A0DBFBDC" box="[401,477,1050,1074]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="6" pageNumber="91">Fig. 9d</figureCitation>
); dorsum of body bright metallic blue, blue-green, or darker and duller green................................................................ 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F2A2BFFFD9FFB3FF9FFBA9A597FAAE" pageId="6" pageNumber="91">
<paragraph id="8BB9BA1AFFD9FFB3FF9FFBA9A596FB31" blockId="6.[123,1174,1126,1344]" pageId="6" pageNumber="91">
<keyLead id="30F7192FFFD9FFB3FF9FFBA9A596FB31" pageId="6" pageNumber="91">
9 (8’) Dorsum of body bright metallic blue or bluish-purple, head with greenish hue in some specimens, legs (including femora) pale rufous; elytral humeri distinct (
<figureCitation id="133DA69FFFD9FFB3FC92FB49A2D6FB70" box="[886,976,1158,1183]" captionStart="FIGURE" captionStartId="52.[171,183,1586,1605]" captionTargetBox="[119,1175,216,1566]" captionTargetId="figure-56@52.[119,1175,216,1566]" captionTargetPageId="52" captionText="FIGURE 28. Broscosoma purpureum sp. nov. a. Habitus (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067411" httpUri="https://zenodo.org/record/11067411/files/figure.png" pageId="6" pageNumber="91">Fig. 28a</figureCitation>
), angulate, elytral striae coarsely punctate, elytral microsculpture effaced; shape of pronotum as in
<figureCitation id="133DA69FFFD9FFB3FBE2FB68A554FB50" box="[1030,1106,1191,1215]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="6" pageNumber="91">Fig. 6c</figureCitation>
; male genitalia as in
<figureCitation id="133DA69FFFD9FFB3FED0FB08A087FB31" box="[308,385,1223,1247]" captionStart="FIGURE" captionStartId="53.[171,183,939,958]" captionTargetBox="[151,1168,221,932]" captionTargetId="figure-1570@53.[120,1176,216,938]" captionTargetPageId="53" captionText="FIGURE 29. Male genitalia, Broscosoma purpureum sp. nov. (CASENT1015869; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral Maspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067415" httpUri="https://zenodo.org/record/11067415/files/figure.png" pageId="6" pageNumber="91">Fig. 29</figureCitation>
..............................
<emphasis id="B9726608FFD9FFB3FCCCFB08A596FB31" bold="true" box="[808,1168,1223,1247]" pageId="6" pageNumber="91">
<taxonomicName id="4C06C199FFD9FFB3FCCCFB08A534FB30" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[808,1074,1223,1246]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="purpureum" status="sp. nov.">
<emphasis id="B9726608FFD9FFB3FCCCFB08A534FB30" bold="true" box="[808,1074,1223,1246]" italics="true" pageId="6" pageNumber="91">Broscosoma purpureum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD9FFB3FBDEFB08A596FB31" box="[1082,1168,1223,1247]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB9BA1AFFD9FFB3FF9FFB27A597FAAE" blockId="6.[123,1174,1126,1344]" pageId="6" pageNumber="91">
<keyLead id="30F7192FFFD9FFB3FF9FFB27A597FAAE" pageId="6" pageNumber="91">
9’ Dorsum of body green or blue-green, legs with femora darker, reddish-brown to piceus; elytral humeri (
<figureCitation id="133DA69FFFD9FFB3FEFAFAC7A086FAF1" box="[286,384,1288,1312]" captionStart="FIGURE" captionStartId="29.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1596]" captionTargetId="figure-68@29.[120,1176,216,1598]" captionTargetPageId="29" captionText="FIGURE 17. Broscosoma danzhuense sp. nov. a. Habitus (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067385" httpUri="https://zenodo.org/record/11067385/files/figure.png" pageId="6" pageNumber="91">Figs. 17a</figureCitation>
,
<figureCitation id="133DA69FFFD9FFB3FE6AFAC7A0B3FAF1" box="[398,437,1288,1312]" captionStart="FIGURE" captionStartId="48.[171,183,1587,1606]" captionTargetBox="[132,1176,216,1571]" captionTargetId="figure-56@48.[120,1176,216,1571]" captionTargetPageId="48" captionText="FIGURE 26. Broscosoma parvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067407" httpUri="https://zenodo.org/record/11067407/files/figure.png" pageId="6" pageNumber="91">26a</figureCitation>
) less distinct, more rounded, elytral striae more finely punctate, elytral microsculpture faint but evident; shape of pronotum as in
<figureCitation id="133DA69FFFD9FFB3FCD7FAE7A291FAAE" box="[819,919,1320,1344]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="6" pageNumber="91">Figs. 6e,f</figureCitation>
...............
<figureCitation id="133DA69FFFD9FFB3FB92FAE7A597FAAE" box="[1142,1169,1320,1344]" captionStart="FIGURE" captionStartId="13.[171,183,1441,1460]" captionTargetBox="[404,932,216,1422]" captionTargetId="figure-66@13.[404,932,216,1422]" captionTargetPageId="13" captionText="FIGURE 10. Head, Broscosoma species, lateral aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote endpoints of genal ridge." figureDoi="http://doi.org/10.5281/zenodo.11067371" httpUri="https://zenodo.org/record/11067371/files/figure.png" pageId="6" pageNumber="91">10</figureCitation>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8BB9BA1AFFD9FFB3FF9FFA9BA597FA03" blockId="6.[123,1174,1364,1725]" pageId="6" pageNumber="91">
10 (9’) Body larger, BL =
<quantity id="4CFE17FFFFD9FFB3FE79FA9BA0E8FA85" box="[413,494,1364,1388]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="6" pageNumber="91" unit="mm" value="9.3">9.3 mm</quantity>
; elytral silhouette (
<figureCitation id="133DA69FFFD9FFB3FD24FA9BA21EFA85" box="[704,792,1364,1388]" captionStart="FIGURE" captionStartId="29.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1596]" captionTargetId="figure-68@29.[120,1176,216,1598]" captionTargetPageId="29" captionText="FIGURE 17. Broscosoma danzhuense sp. nov. a. Habitus (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067385" httpUri="https://zenodo.org/record/11067385/files/figure.png" pageId="6" pageNumber="91">Fig. 17a</figureCitation>
) with greatest width distinctly posterior to midlength; eyes (
<figureCitation id="133DA69FFFD9FFB3FE5CFABBA303FA65" box="[440,517,1396,1420]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="6" pageNumber="91">Fig. 6e</figureCitation>
) moderately convex; elytra with only stria 1 impressed, all other striae represented only be punctures diminishing in size and depth laterally and posteriorly, elytral intervals flat; entire dorsum of body dark green; male unknown................................................................
<emphasis id="B9726608FFD9FFB3FCC7FA1AA597FA03" bold="true" box="[803,1169,1493,1517]" pageId="6" pageNumber="91">
<taxonomicName id="4C06C199FFD9FFB3FCC7FA1AA535FA03" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[803,1075,1493,1517]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="danzhuense" status="sp. nov.">
<emphasis id="B9726608FFD9FFB3FCC7FA1AA535FA03" bold="true" box="[803,1075,1493,1517]" italics="true" pageId="6" pageNumber="91">Broscosoma danzhuense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD9FFB3FBDFFA1AA597FA03" box="[1083,1169,1493,1517]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BB9BA1AFFD9FFB3FF9FFA39A597F941" blockId="6.[123,1174,1364,1725]" pageId="6" pageNumber="91">
10’ Body smaller, BL
<quantity id="4CFE17FFFFD9FFB3FE9EFA39A0CAF9E3" box="[378,460,1525,1550]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="91" unit="mm" value="8.5">8.5 mm</quantity>
or less; elytral silhouette (
<figureCitation id="133DA69FFFD9FFB3FD0AFA3AA241F9E3" box="[750,839,1525,1549]" captionStart="FIGURE" captionStartId="48.[171,183,1587,1606]" captionTargetBox="[132,1176,216,1571]" captionTargetId="figure-56@48.[120,1176,216,1571]" captionTargetPageId="48" captionText="FIGURE 26. Broscosoma parvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067407" httpUri="https://zenodo.org/record/11067407/files/figure.png" pageId="6" pageNumber="91">Fig. 26a</figureCitation>
) with greatest width at or near midlength; eyes (
<figureCitation id="133DA69FFFD9FFB3FEB0F9D9A09AF9C0" box="[340,412,1558,1582]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="6" pageNumber="91">Fig. 6f</figureCitation>
) less convex, slightly flattened in some specimens; elytra with two or more striae impressed, all striae punctate with punctures diminishing in size and depth laterally and posteriorly, at least medial two or more elytral intervals very slightly to moderately convex; dorsum with head green, elytra blue-green and pronotum either green or blue-green; male genitalia as in
<figureCitation id="133DA69FFFD9FFB3FEEAF958A05CF941" box="[270,346,1687,1711]" captionStart="FIGURE" captionStartId="49.[171,183,963,982]" captionTargetBox="[144,1168,235,949]" captionTargetId="figure-1710@49.[120,1176,216,957]" captionTargetPageId="49" captionText="FIGURE 27. Male genitalia, Broscosoma parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067409" httpUri="https://zenodo.org/record/11067409/files/figure.png" pageId="6" pageNumber="91">Fig. 27</figureCitation>
....................................
<emphasis id="B9726608FFD9FFB3FCB5F957A597F941" bold="true" box="[849,1169,1687,1711]" pageId="6" pageNumber="91">
<taxonomicName id="4C06C199FFD9FFB3FCB5F957A535F941" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[849,1075,1688,1711]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="parvum" status="sp. nov.">
<emphasis id="B9726608FFD9FFB3FCB5F957A535F941" bold="true" box="[849,1075,1688,1711]" italics="true" pageId="6" pageNumber="91">Broscosoma parvum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD9FFB3FBDFF958A597F941" box="[1083,1169,1687,1711]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BB9BA1AFFD9FFB3FF9FF960A179F953" blockId="6.[123,1174,1364,1725]" box="[123,127,1711,1725]" pageId="6" pageNumber="91">.</paragraph>
<paragraph id="8BB9BA1AFFD8FFB2FF9FFF14A597FF1D" blockId="7.[123,1173,219,307]" box="[123,1169,219,243]" pageId="7" pageNumber="92">
11 (5’) Metatrochanter unisetose (
<figureCitation id="133DA69FFFD8FFB2FE14FF14A33AFF1D" box="[496,572,219,243]" captionStart="FIGURE" captionStartId="11.[171,183,1003,1022]" captionTargetBox="[120,1176,216,984]" captionTargetId="figure-67@11.[120,1176,216,985]" captionTargetPageId="11" captionText="FIGURE 8. Left metatrochanter, broscine species, ventral aspect. a. Broscodera gaoligongensis sp. nov. (Yaojiaping, Lushui County, Yunnan, China); b. Broscodera chukuai sp. nov. (CASENT1026771; Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1034187; Qiqi-Dulong Yakou, Cikai Township, Gongshan County, Yunnan, China); d. B. ribbei Putzeys (CASENT1000578; Nankang Yakou, Longyang County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067367" httpUri="https://zenodo.org/record/11067367/files/figure.png" pageId="7" pageNumber="92">Fig. 8d</figureCitation>
)........................................ 12
</paragraph>
<paragraph id="8BB9BA1AFFD8FFB2FF9FFF34A597FEDD" blockId="7.[123,1173,219,307]" pageId="7" pageNumber="92">
11’ Metatrochanter asetose (
<figureCitation id="133DA69FFFD8FFB2FE2EFF34A31FFEFD" box="[458,537,251,275]" captionStart="FIGURE" captionStartId="11.[171,183,1003,1022]" captionTargetBox="[120,1176,216,984]" captionTargetId="figure-67@11.[120,1176,216,985]" captionTargetPageId="11" captionText="FIGURE 8. Left metatrochanter, broscine species, ventral aspect. a. Broscodera gaoligongensis sp. nov. (Yaojiaping, Lushui County, Yunnan, China); b. Broscodera chukuai sp. nov. (CASENT1026771; Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1034187; Qiqi-Dulong Yakou, Cikai Township, Gongshan County, Yunnan, China); d. B. ribbei Putzeys (CASENT1000578; Nankang Yakou, Longyang County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067367" httpUri="https://zenodo.org/record/11067367/files/figure.png" pageId="7" pageNumber="92">Fig. 8c</figureCitation>
) (except unilaterally unisetose in very few specimens of
<taxonomicName id="4C06C199FFD8FFB2FF73FED3A03EFEDD" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[151,312,283,307]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFD8FFB2FF73FED3A03EFEDD" box="[151,312,283,307]" italics="true" pageId="7" pageNumber="92">B. bicoloratum</emphasis>
</taxonomicName>
)............................................................ 13
</paragraph>
<paragraph id="8BB9BA1AFFD8FFB2FF9FFE89A597FE10" blockId="7.[123,1174,326,703]" pageId="7" pageNumber="92">
12 (11’) Dorsum of body piceous, without any trace of metallic reflection (
<figureCitation id="133DA69FFFD8FFB2FC71FE89A2EDFEB3" box="[917,1003,326,350]" captionStart="FIGURE" captionStartId="33.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1586]" captionTargetId="figure-67@33.[120,1176,216,1586]" captionTargetPageId="33" captionText="FIGURE 18. Broscosoma furvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 k." figureDoi="http://doi.org/10.5281/zenodo.11067389" httpUri="https://zenodo.org/record/11067389/files/figure.png" pageId="7" pageNumber="92">Fig. 18a</figureCitation>
); antennomeres 3 and 4 distinctly darker than antennomeres 1 and 2; pronotum anterior to sub-basal constriction (
<figureCitation id="133DA69FFFD8FFB2FF44FE49A1EAFE70" box="[160,236,390,414]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="7" pageNumber="92">Fig. 7a</figureCitation>
) longitudinally slightly ovoid in dorsal view; genal ridge evident, extended from base of head to ventral margin of eye (
<figureCitation id="133DA69FFFD8FFB2FE1EFE69A357FE50" box="[506,593,422,446]" captionStart="FIGURE" captionStartId="13.[171,183,1441,1460]" captionTargetBox="[404,932,216,1422]" captionTargetId="figure-66@13.[404,932,216,1422]" captionTargetPageId="13" captionText="FIGURE 10. Head, Broscosoma species, lateral aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote endpoints of genal ridge." figureDoi="http://doi.org/10.5281/zenodo.11067371" httpUri="https://zenodo.org/record/11067371/files/figure.png" pageId="7" pageNumber="92">Fig. 10a</figureCitation>
); male genitalia as in
<figureCitation id="133DA69FFFD8FFB2FCDFFE69A280FE50" box="[827,902,422,446]" captionStart="FIGURE" captionStartId="34.[171,183,963,982]" captionTargetBox="[142,1161,229,948]" captionTargetId="figure-1616@34.[120,1176,216,960]" captionTargetPageId="34" captionText="FIGURE 19. Male genitalia, Broscosoma furvum sp. nov. (CASENT1023548; 1.0 km E of Shibali Yakou, Lishadi Township, Fugong County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067391" httpUri="https://zenodo.org/record/11067391/files/figure.png" pageId="7" pageNumber="92">Fig. 19</figureCitation>
, with median lobe longer and more slender and with apical lamella distinctly expanded ventrally in lateral view..........................................................
<emphasis id="B9726608FFD8FFB2FCB0FE28A597FE10" bold="true" box="[852,1169,486,510]" pageId="7" pageNumber="92">
<taxonomicName id="4C06C199FFD8FFB2FCB0FE28A532FE10" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[852,1076,486,510]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="furvum" status="sp. nov.">
<emphasis id="B9726608FFD8FFB2FCB0FE28A532FE10" bold="true" box="[852,1076,486,510]" italics="true" pageId="7" pageNumber="92">Broscosoma furvum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD8FFB2FBDFFE29A597FE10" box="[1083,1169,486,510]" pageId="7" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BB9BA1AFFD8FFB2FF9FFDC9A597FD50" blockId="7.[123,1174,326,703]" pageId="7" pageNumber="92">
12’ Dorsum of head and pronotum black or dark piceous, elytra with bright metallic green or blue-green reflection (
<figureCitation id="133DA69FFFD8FFB2FEB4FDE9A0A1FDD0" box="[336,423,550,575]" captionStart="FIGURE" captionStartId="37.[171,183,1600,1619]" captionTargetBox="[125,1176,216,1581]" captionTargetId="figure-67@37.[120,1176,216,1586]" captionTargetPageId="37" captionText="FIGURE 20. Broscosoma gaoligongense Deuve and Wrase. a. Habitus (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067393" httpUri="https://zenodo.org/record/11067393/files/figure.png" pageId="7" pageNumber="92">Fig. 20a</figureCitation>
); antennomeres 1 through 4 concolorous, pale rufous; pronotum anterior to sub-basal constriction (
<figureCitation id="133DA69FFFD8FFB2FE3DFD89A323FDB0" box="[473,549,582,606]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="7" pageNumber="92">Fig. 7d</figureCitation>
) globose in dorsal view; genal ridge evident basal to post-temporal constriction but extended only to or slightly anterior to the constriction (
<figureCitation id="133DA69FFFD8FFB2FBCDFDA8A583FD91" box="[1065,1157,615,639]" captionStart="FIGURE" captionStartId="13.[171,183,1441,1460]" captionTargetBox="[404,932,216,1422]" captionTargetId="figure-66@13.[404,932,216,1422]" captionTargetPageId="13" captionText="FIGURE 10. Head, Broscosoma species, lateral aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote endpoints of genal ridge." figureDoi="http://doi.org/10.5281/zenodo.11067371" httpUri="https://zenodo.org/record/11067371/files/figure.png" pageId="7" pageNumber="92">Fig. 10b</figureCitation>
); male genitalia as in
<figureCitation id="133DA69FFFD8FFB2FE9CFD48A0C0FD71" box="[376,454,647,671]" captionStart="FIGURE" captionStartId="38.[171,183,927,946]" captionTargetBox="[145,1143,223,907]" captionTargetId="figure-1565@38.[132,1162,218,913]" captionTargetPageId="38" captionText="FIGURE 21. Male genitalia, Broscosoma gaoligongense Deuve and Wrase. (CASENT1027154; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067395" httpUri="https://zenodo.org/record/11067395/files/figure.png" pageId="7" pageNumber="92">Fig. 21</figureCitation>
, with median lobe slightly shorter and thicker and apical lamella more symmetrically rounded in lateral view...
<taxonomicName id="4C06C199FFD8FFB2FD43FD67A597FD50" authority="Deuve and Wrase" authorityName="Deuve and Wrase" box="[679,1169,679,703]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFD8FFB2FD43FD67A2CBFD51" box="[679,973,679,703]" italics="true" pageId="7" pageNumber="92">Broscosoma gaoligongense</emphasis>
Deuve and Wrase
</taxonomicName>
</paragraph>
<paragraph id="8BB9BA1AFFD8FFB2FF9FFD1DA597FC44" blockId="7.[123,1174,721,1098]" pageId="7" pageNumber="92">
13 (11’) Head and pronotum black to piceous but with distinct metallic green band across anterior part of pronotum (
<figureCitation id="133DA69FFFD8FFB2FE6DFD3DA0DFFCE4" box="[393,473,754,778]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="7" pageNumber="92">Fig. 7b</figureCitation>
) anterior to anterior transverse impression; head with tempora slightly inflated, sharply convex or vaguely angulate; pronotum anterior to sub-basal constriction slightly to moderately longitudinally ovoid in dorsal view; elytral microsculpture distinct, comprised of moderately impressed isodiametric sculpticells; male genitalia as in
<figureCitation id="133DA69FFFD8FFB2FBEFFC9DA55EFC84" box="[1035,1112,850,874]" captionStart="FIGURE" captionStartId="69.[171,183,922,941]" captionTargetBox="[144,1155,225,900]" captionTargetId="figure-1615@69.[127,1169,215,908]" captionTargetPageId="69" captionText="FIGURE 35. Male genitalia, Broscosoma viridicollare sp. nov. (CASENT1017820; second cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067429" httpUri="https://zenodo.org/record/11067429/files/figure.png" pageId="7" pageNumber="92">Fig. 35</figureCitation>
, with median lobe slightly longer and apical lamella broader in lateral view...................................................................
<emphasis id="B9726608FFD8FFB2FCC4FC5DA597FC44" bold="true" box="[800,1169,914,938]" pageId="7" pageNumber="92">
<taxonomicName id="4C06C199FFD8FFB2FCC4FC5DA532FC44" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[800,1076,914,938]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="viridicollare" status="sp. nov.">
<emphasis id="B9726608FFD8FFB2FCC4FC5DA532FC44" bold="true" box="[800,1076,914,938]" italics="true" pageId="7" pageNumber="92">Broscosoma viridicollare</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD8FFB2FBDFFC5DA597FC44" box="[1083,1169,914,938]" pageId="7" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BB9BA1AFFD8FFB2FF9FFC7DA597FBA4" blockId="7.[123,1174,721,1098]" pageId="7" pageNumber="92">
13’ Head and pronotum black to piceous, without metallic reflection (except a few specimens of
<taxonomicName id="4C06C199FFD8FFB2FF73FC1CA03CFC04" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[151,314,978,1002]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFD8FFB2FF73FC1CA03CFC04" box="[151,314,978,1002]" italics="true" pageId="7" pageNumber="92">B. bicoloratum</emphasis>
</taxonomicName>
have faint a metallic green band across anterior part of pronotum); head with tempora only slightly (
<figureCitation id="133DA69FFFD8FFB2FE6FFC3DA0D3FBE4" box="[395,469,1010,1034]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="7" pageNumber="92">Fig. 7e</figureCitation>
) or distinctly (
<figureCitation id="133DA69FFFD8FFB2FD90FC3DA3B9FBE4" box="[628,703,1010,1034]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="7" pageNumber="92">Fig. 7c</figureCitation>
) convex; pronotum either globose or slightly longitudinally ovoid in dorsal view; elytral microsculpture indistinct, effaced or only extremely faintly evident in some areas............................................... 14
</paragraph>
<paragraph id="8BB9BA1AFFD8FFB2FF9FFB92A597FB7B" blockId="7.[123,1185,1117,1270]" pageId="7" pageNumber="92">
14 (13’) Pronotum globose anterior to sub-basal constriction as in
<figureCitation id="133DA69FFFD8FFB2FCA3FB92A294FB9B" box="[839,914,1117,1141]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="7" pageNumber="92">Fig. 7c</figureCitation>
; head with tempora distinctly convex behind eyes; male genitalia as in
<figureCitation id="133DA69FFFD8FFB2FD79FBB2A3ECFB7B" box="[669,746,1149,1173]" captionStart="FIGURE" captionStartId="26.[171,183,963,982]" captionTargetBox="[142,1152,219,938]" captionTargetId="figure-1559@26.[120,1176,215,955]" captionTargetPageId="26" captionText="FIGURE 16. Male genitalia, Broscosoma bicoloratum sp. nov. (CASENT1025709; southeast slope of Heipu Yakou, Cikai Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067383" httpUri="https://zenodo.org/record/11067383/files/figure.png" pageId="7" pageNumber="92">Fig. 16</figureCitation>
...
<emphasis id="B9726608FFD8FFB2FCC4FBB1A597FB7B" bold="true" box="[800,1169,1149,1173]" pageId="7" pageNumber="92">
<taxonomicName id="4C06C199FFD8FFB2FCC4FBB1A535FB7B" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[800,1075,1149,1173]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="bicoloratum" status="sp. nov.">
<emphasis id="B9726608FFD8FFB2FCC4FBB1A535FB7B" bold="true" box="[800,1075,1149,1173]" italics="true" pageId="7" pageNumber="92">Broscosoma bicoloratum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD8FFB2FBDFFBB2A597FB7B" box="[1083,1169,1149,1173]" pageId="7" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BB9BA1AFFD8FFB2FF9FFB52A597FB18" blockId="7.[123,1185,1117,1270]" pageId="7" pageNumber="92">
14’ Pronotum anterior to sub-basal constriction slightly longitudinally ovoid in dorsal view (
<figureCitation id="133DA69FFFD8FFB2FB8FFB52A590FB5B" box="[1131,1174,1181,1205]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="7" pageNumber="92">Fig.</figureCitation>
7e); head with tempora straight or only slightly convex behind eyes; male genitalia as in
<figureCitation id="133DA69FFFD8FFB2FBB1FB72A5A7FB3B" box="[1109,1185,1213,1237]" captionStart="FIGURE" captionStartId="41.[171,183,967,986]" captionTargetBox="[157,1154,219,934]" captionTargetId="figure-1587@41.[121,1176,216,953]" captionTargetPageId="41" captionText="FIGURE 23. Male genitalia, Broscosoma gongshanense sp. nov. (CASENT1026429; 2.0 km S of Heipu Yakou, Cikai Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Medi- an lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067399" httpUri="https://zenodo.org/record/11067399/files/figure.png" pageId="7" pageNumber="92">Fig. 23</figureCitation>
..............................................
<emphasis id="B9726608FFD8FFB2FCEDFB11A597FB18" bold="true" box="[777,1169,1245,1270]" pageId="7" pageNumber="92">
<taxonomicName id="4C06C199FFD8FFB2FCEDFB11A532FB1B" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[777,1076,1245,1269]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="gongshanense" status="sp. nov.">
<emphasis id="B9726608FFD8FFB2FCEDFB11A532FB1B" bold="true" box="[777,1076,1245,1269]" italics="true" pageId="7" pageNumber="92">Broscosoma gongshanense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD8FFB2FBDFFB11A597FB18" box="[1083,1169,1246,1270]" pageId="7" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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