treatments-xml/data/03/AF/0B/03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E.xml

<document id="4363D152DD04BB1CCA3B9D1049141E26" ID-CLB-Dataset="299055" ID-DOI="10.5281/zenodo.11067355" ID-GBIF-Dataset="67ec2de4-1816-4969-8132-f6796d3afa06" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067355" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052343267" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2021" docId="03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E" docLanguage="en" docName="ProcCalifAcadSci.67.4.85-182.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (4)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Broscodera (Sinobrosculus) chukuai Kavanaugh and Liang 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="103" masterDocId="FF967374FFDFFFB5FFE4FFCFA106FFEE" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." masterLastPageNumber="182" masterPageNumber="85" pageNumber="99" updateTime="1719540814554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="664329B6F236655D01EE5011A2EDEE0D">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae).</mods:title>
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<mods:namePart id="AE4F9CF1DB0B5706B18EEEBE21AB11D5">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="E5327EF100F1EFBC54B9D98C4B7F753A">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="6DB17FA4BCED2700E7DE8F4C987A8B59">Liang, Hongbin</mods:namePart>
<mods:affiliation id="4CF13DD6BB63133C25C8CD80D100F3FC">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title id="702D966C4A30CF04AD3C781343C2DF73">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="1D5A11DB1CD9CE4F9A84FC75BE61208C">
<mods:date id="245BE88DD7B9146F329D0526458E5BF4">2021</mods:date>
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<treatment id="03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E" ID-DOI="http://doi.org/10.5281/zenodo.12575554" ID-GBIF-Taxon="233814492" ID-Zenodo-Dep="12575554" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E" httpUri="http://treatment.plazi.org/id/03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E" lastPageId="18" lastPageNumber="103" pageId="14" pageNumber="99">
<subSubSection id="C31CE991FFD1FFBBFF9FFBC4A29EFBCD" box="[123,920,1035,1059]" pageId="14" pageNumber="99" type="nomenclature">
<paragraph id="8BB9BA1AFFD1FFBBFF9FFBC4A29EFBCD" blockId="14.[123,920,1035,1088]" box="[123,920,1035,1059]" pageId="14" pageNumber="99">
<heading id="D0F10D76FFD1FFBBFF9FFBC4A29EFBCD" box="[123,920,1035,1059]" fontSize="10" level="2" pageId="14" pageNumber="99" reason="7">
<emphasis id="B9726608FFD1FFBBFF9FFBC4A29EFBCD" bold="true" box="[123,920,1035,1059]" pageId="14" pageNumber="99">
1. 
<taxonomicName id="4C06C199FFD1FFBBFF72FBC4A232FBCD" authority="Kavanaugh and Liang" authorityName="Kavanaugh and Liang" authorityYear="2021" box="[150,820,1035,1059]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="99" phylum="Arthropoda" rank="species" species="chukuai" status="sp. nov." subGenus="Sinobrosculus">
<emphasis id="B9726608FFD1FFBBFF72FBC4A012FBCD" bold="true" box="[150,276,1035,1059]" italics="true" pageId="14" pageNumber="99">Broscodera</emphasis>
(
<emphasis id="B9726608FFD1FFBBFEC0FBC4A0C7FBCD" bold="true" box="[292,449,1035,1059]" italics="true" pageId="14" pageNumber="99">Sinobrosculus</emphasis>
) 
<emphasis id="B9726608FFD1FFBBFE35FBC4A32DFBCD" bold="true" box="[465,555,1035,1059]" italics="true" pageId="14" pageNumber="99">chukuai</emphasis>
Kavanaugh and Liang
</taxonomicName>
, 
<taxonomicNameLabel id="A241DB73FFD1FFBBFCA5FBC4A29EFBCD" box="[833,920,1035,1059]" pageId="14" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFD1FFBBFF9FFBE3A095FBD1" box="[123,403,1068,1088]" pageId="14" pageNumber="99" type="description">
<paragraph id="8BB9BA1AFFD1FFBBFF9FFBE3A095FBD1" blockId="14.[123,920,1035,1088]" box="[123,403,1068,1088]" pageId="14" pageNumber="99">
<figureCitation id="133DA69FFFD1FFBBFF9FFBE3A1D2FBAE" box="[123,212,1068,1088]" captionStart="FIGURE" captionStartId="8.[171,183,1513,1532]" captionTargetBox="[139,1176,224,1495]" captionTargetId="figure-55@8.[120,1176,216,1495]" captionTargetPageId="8" captionText="FIGURE 5. Forebody of broscine carabids, dorsal aspect. a. Eoboscus bhutanensis Morvan (CASENT1019340; Lishadi Township, Fugong County, Yunnan, China); b. Broscus punctatus (Dejean) (CASENT1011066; Heiwadi, Cikai Township, Gongshan County, Yunnan, China); c. Broscodera chukuai sp. nov. (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo Township, Gongshan County, Yunnan, China); d. Broscodera gaoligongensis sp. nov. (CASENT1020104; South Fork of Yamu He, Lumadeng Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067361" httpUri="https://zenodo.org/record/11067361/files/figure.png" pageId="14" pageNumber="99">Figures 5c</figureCitation>
, 
<figureCitation id="133DA69FFFD1FFBBFF04FBE3A1F0FBD1" box="[224,246,1068,1087]" captionStart="FIGURE" captionStartId="11.[171,183,1003,1022]" captionTargetBox="[120,1176,216,984]" captionTargetId="figure-67@11.[120,1176,216,985]" captionTargetPageId="11" captionText="FIGURE 8. Left metatrochanter, broscine species, ventral aspect. a. Broscodera gaoligongensis sp. nov. (Yaojiaping, Lushui County, Yunnan, China); b. Broscodera chukuai sp. nov. (CASENT1026771; Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1034187; Qiqi-Dulong Yakou, Cikai Township, Gongshan County, Yunnan, China); d. B. ribbei Putzeys (CASENT1000578; Nankang Yakou, Longyang County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067367" httpUri="https://zenodo.org/record/11067367/files/figure.png" pageId="14" pageNumber="99">8b</figureCitation>
, 
<figureCitation id="133DA69FFFD1FFBBFEE4FBE3A013FBD1" box="[256,277,1068,1087]" captionStart="Figure 11" captionStartId="15.[174,230,1616,1635]" captionTargetBox="[139,1176,216,1591]" captionTargetId="figure-67@15.[120,1176,216,1591]" captionTargetPageId="15" captionText="Figure 11. Broscodera chukuai sp. nov. a. Habitus (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067373" httpUri="https://zenodo.org/record/11067373/files/figure.png" pageId="14" pageNumber="99">11</figureCitation>
, 
<figureCitation id="133DA69FFFD1FFBBFEC4FBE3A033FBD1" box="[288,309,1068,1087]" captionStart="FIGURE" captionStartId="16.[171,183,933,952]" captionTargetBox="[147,1135,222,918]" captionTargetId="figure-1483@16.[120,1176,210,928]" captionTargetPageId="16" captionText="FIGURE 12. Male genitalia, Broscodera chukuai sp. nov. (CASENT1026169; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067375" httpUri="https://zenodo.org/record/11067375/files/figure.png" pageId="14" pageNumber="99">12</figureCitation>
, 
<figureCitation id="133DA69FFFD1FFBBFEA5FBE3A050FBD1" box="[321,342,1068,1087]" captionStart="FIGURE" captionStartId="80.[171,183,1579,1598]" captionTargetBox="[127,1170,216,1566]" captionTargetId="figure-57@80.[127,1170,216,1566]" captionTargetPageId="80" captionText="FIGURE 40. Photographs of habitats for broscine species in the Gaoligong Shan region. a. SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, 3950 m, 27.982°/098.480°, Bingzhongluo Township, Gongshan County, Yunnan (habitat for Brosocodera chukuai sp. nov.); b. Second cirque S of Shibali Yakou at border post “31”, 3710 m, 27.203°/098.693°, Lumadeng Township, Fugong County, Yunnan (habitat for Brosocodera chukuai sp. nov., Broscosoma furvum sp. nov., and Broscosoma viridicollare sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067439" httpUri="https://zenodo.org/record/11067439/files/figure.png" pageId="14" pageNumber="99">40</figureCitation>
, 
<figureCitation id="133DA69FFFD1FFBBFE85FBE3A095FBD1" box="[353,403,1068,1088]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="88.[171,183,995,1014]" captionStartId-1="88.[184,196,1682,1701]" captionStartId-2="89.[171,183,1461,1480]" captionTargetBox-0="[130,1164,233,969]" captionTargetBox-1="[130,1162,1063,1660]" captionTargetBox-2="[140,1154,249,1428]" captionTargetId-0="figure-57@88.[120,1176,215,989]" captionTargetId-1="figure-332@88.[120,1176,1054,1668]" captionTargetId-2="figure-68@89.[120,1176,216,1448]" captionTargetPageId-0="88" captionTargetPageId-1="88" captionTargetPageId-2="89" captionText-0="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." captionText-2="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi-0="http://doi.org/10.5281/zenodo.11067455" figureDoi-1="http://doi.org/10.5281/zenodo.11067457" figureDoi-2="http://doi.org/10.5281/zenodo.11067459" httpUri-0="https://zenodo.org/record/11067455/files/figure.png" httpUri-1="https://zenodo.org/record/11067457/files/figure.png" httpUri-2="https://zenodo.org/record/11067459/files/figure.png" pageId="14" pageNumber="99">48-50</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="8BB9BA1AFFD1FFA5FF4FFB92A1B8F9F6" blockId="14.[123,1174,1116,1726]" lastBlockId="16.[123,1174,1043,1726]" lastPageId="16" lastPageNumber="101" pageId="14" pageNumber="99">
<emphasis id="B9726608FFD1FFBBFF4FFB92A054FB9B" bold="true" box="[171,338,1116,1141]" pageId="14" pageNumber="99">Type material.</emphasis>
<materialsCitation id="3B6EB047FFD1FFBBFEBDFB93A244FB18" ID-GBIF-Occurrence="4899905439" collectingDate="2006-08-19" collectionCode="IOZ" collectorName="D. H. Kavanaugh &amp; J. A. Miller &amp; D. Z. Dong" country="China" county="Gongshan County" elevation="3745" latitude="27.98393" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" pageId="14" pageNumber="99" specimenCode="CASENT 1026170" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54BD04B8FFD1FFBBFEBDFB93A0BAFB9A" box="[345,444,1116,1140]" pageId="14" pageNumber="99" type="holotype">Holotype</typeStatus>
, a 
<specimenCount id="9D007193FFD1FFBBFE3AFB93A314FB9A" box="[478,530,1116,1140]" pageId="14" pageNumber="99" type="male">male</specimenCount>
, in 
<collectionCode id="ED1722DFFFD1FFBBFDD9FB92A36FFB9B" box="[573,617,1117,1141]" pageId="14" pageNumber="99">IOZ</collectionCode>
, labeled: “ 
<specimenCode id="DBA01261FFD1FFBBFD0CFB92A2B2FB9A" box="[744,948,1116,1141]" pageId="14" pageNumber="99">CASENT 1026170</specimenCode>
”/ “ 
<collectingCountry id="F311FA8AFFD1FFBBFC3FFB92A529FB9B" box="[987,1071,1117,1141]" name="China" pageId="14" pageNumber="99">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFD1FFBBFBD9FB92A588FB9A" box="[1085,1166,1117,1140]" country="China" name="Yunnan" pageId="14" pageNumber="99">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFD1FFBBFF9FFBB2A046FB7B" box="[123,320,1149,1173]" pageId="14" pageNumber="99">Gongshan County</collectingCounty>
, 
<collectingMunicipality id="6BDD2060FFD1FFBBFEB0FBB2A366FB7B" box="[340,608,1149,1173]" pageId="14" pageNumber="99">Bingzhongluo Township</collectingMunicipality>
, SW slope of 
<location id="8ED9ECC1FFD1FFBBFCEBFBB2A2CDFB7B" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFD1FFBBFCEBFBB2A2CDFB7B" box="[783,971,1149,1173]" country="China" county="Gongshan County" latitude="27.98393" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="14" pageNumber="99" stateProvince="Yunnan">Kawakarpu Shan</location>
, 
<locationDeviation id="9A0BA338FFD1FFBBFC3BFBB2A017FB5B" location="Chukuai Lake" pageId="14" pageNumber="99">
<quantity id="4CFE17FFFFD1FFBBFC3BFBB2A529FB7B" box="[991,1071,1149,1173]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="99" unit="km" value="0.3">0.3 km</quantity>
NNE of Chukuai Lake
</locationDeviation>
, 
<geoCoordinate id="EE32DCDDFFD1FFBBFEFAFB51A0A6FB58" box="[286,416,1182,1206]" degrees="27. 98393" direction="north" orientation="latitude" pageId="14" pageNumber="99" precision="1" value="27.98393">N27.98393°</geoCoordinate>
/ 
<geoCoordinate id="EE32DCDDFFD1FFBBFE43FB51A332FB58" box="[423,564,1182,1206]" degrees="098. 47491" direction="east" orientation="longitude" pageId="14" pageNumber="99" precision="1" value="98.47491">E098.47491°</geoCoordinate>
”/ “ 
<quantity id="4CFE17FFFFD1FFBBFDBDFB52A3ACFB5B" box="[601,682,1181,1205]" metricMagnitude="3" metricUnit="m" metricValue="3.745" pageId="14" pageNumber="99" unit="m" value="3745.0">
<elevation id="002B5D29FFD1FFBBFDBDFB52A3ACFB5B" box="[601,682,1181,1205]" metricMagnitude="3" metricUnit="m" metricValue="3.745" pageId="14" pageNumber="99" unit="m" value="3745.0">3745 m</elevation>
</quantity>
, 
<date id="FFB89CDAFFD1FFBBFD53FB51A266FB5B" box="[695,864,1181,1206]" pageId="14" pageNumber="99" value="2006-08-19">
<collectingDate id="EFFC6532FFD1FFBBFD53FB51A266FB5B" box="[695,864,1181,1206]" pageId="14" pageNumber="99" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-081 
<collectorName id="26F3DFCCFFD1FFBBFB85FB51A1F2FB38" pageId="14" pageNumber="99">D.H. Kavanaugh</collectorName>
, 
<collectorName id="26F3DFCCFFD1FFBBFEECFB71A085FB38" box="[264,387,1214,1238]" pageId="14" pageNumber="99">J.A. Miller</collectorName>
&amp; 
<collectorName id="26F3DFCCFFD1FFBBFE55FB71A32CFB38" box="[433,554,1214,1238]" pageId="14" pageNumber="99">D.Z. Dong</collectorName>
collectors”/ “HOLOTYPE 
<taxonomicName id="4C06C199FFD1FFBBFC81FB71A0B7FB18" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="99" phylum="Arthropoda" rank="species" species="chukuai" status="sp. nov." subGenus="Sinobrosculus">
<emphasis id="B9726608FFD1FFBBFC81FB71A2E6FB38" box="[869,992,1214,1238]" italics="true" pageId="14" pageNumber="99">Broscodera</emphasis>
(
<emphasis id="B9726608FFD1FFBBFC12FB71A58AFB38" box="[1014,1164,1214,1238]" italics="true" pageId="14" pageNumber="99">Sinobrosculus</emphasis>
) 
<emphasis id="B9726608FFD1FFBBFF9FFB11A1C9FB18" box="[123,207,1246,1270]" italics="true" pageId="14" pageNumber="99">chukuai</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD1FFBBFE5CFB10A30DFB18" box="[440,523,1247,1270]" pageId="14" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [red label]
</materialsCitation>
. 
<materialsCitation id="3B6EB047FFD1FFBBFCABFB10A0EBFAD6" ID-GBIF-Occurrence="4899905482" collectingDate="2006-08-19" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; J. A. Miller &amp; D. Z. Dong" country="China" county="Gongshan County" elevation="3745" latitude="27.98393" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" pageId="14" pageNumber="99" specimenCode="CASENT 1026167, CASENT 1026169" specimenCount="4" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54BD04B8FFD1FFBBFCABFB10A2B1FB19" box="[847,951,1247,1271]" pageId="14" pageNumber="99" type="paratype">Paratypes</typeStatus>
(a total of 31): 
<specimenCount id="9D007193FFD1FFBBFB84FB11A1BFFAF9" count="3" pageId="14" pageNumber="99" type="male">three males</specimenCount>
and 
<specimenCount id="9D007193FFD1FFBBFF1EFB30A07DFAF9" box="[250,379,1279,1303]" pageId="14" pageNumber="99">ten females</specimenCount>
(
<collectionCode id="ED1722DFFFD1FFBBFE75FB30A0C3FAF9" box="[401,453,1279,1303]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="99" type="Museum">CAS</collectionCode>
, 
<collectionCode id="ED1722DFFFD1FFBBFE3DFB30A303FAF9" box="[473,517,1279,1303]" pageId="14" pageNumber="99">IOZ</collectionCode>
) labeled: same as holotype except first label: “ 
<specimenCode id="DBA01261FFD1FFBBFBCAFB30A1DEFAD9" pageId="14" pageNumber="99">CASENT 1026167</specimenCode>
” to “ 
<specimenCode id="DBA01261FFD1FFBBFEF1FAEFA0E7FAD9" box="[277,481,1311,1336]" pageId="14" pageNumber="99">CASENT 1026169</specimenCode>
”
</materialsCitation>
<materialsCitation id="3B6EB047FFD1FFBBFE11FAD0A196FAB6" ID-GBIF-Occurrence="4899905557" collectionCode="CAS, IOZ" country="China" pageId="14" pageNumber="99" specimenCode="CASENT 1026171, CASENT 1026180" specimenCount="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
and “ 
<specimenCode id="DBA01261FFD1FFBBFDD4FAEFA3FAFAD9" box="[560,764,1311,1336]" pageId="14" pageNumber="99">CASENT 1026171</specimenCode>
” to “ 
<specimenCode id="DBA01261FFD1FFBBFCDDFAEFA503FAD9" box="[825,1029,1311,1336]" pageId="14" pageNumber="99">CASENT 1026180</specimenCode>
”, respectively
</materialsCitation>
; 
<materialsCitation id="3B6EB047FFD1FFBBFF79FA8FA1A4FA96" ID-GBIF-Occurrence="4899905336" collectingDate="2006-08-18" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; J. A. Miller &amp; D. Z. Dong &amp; Y. Liu" country="China" county="Gongshan County" elevation="3720" latitude="27.98121" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" pageId="14" pageNumber="99" specimenCode="CASENT 1026771, CASENT 1026773" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFD1FFBBFF79FA8FA010FAB6" box="[157,278,1344,1368]" count="3" pageId="14" pageNumber="99" type="male">three males</specimenCount>
and 
<specimenCount id="9D007193FFD1FFBBFEACFA8FA0CBFAB6" box="[328,461,1344,1368]" count="4" pageId="14" pageNumber="99" type="female">four females</specimenCount>
(
<collectionCode id="ED1722DFFFD1FFBBFE38FA8FA316FAB6" box="[476,528,1344,1368]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="99" type="Museum">CAS</collectionCode>
, 
<collectionCode id="ED1722DFFFD1FFBBFDF8FA8FA34EFAB6" box="[540,584,1344,1368]" pageId="14" pageNumber="99">IOZ</collectionCode>
) labeled: “ 
<specimenCode id="DBA01261FFD1FFBBFD59FA8FA281FAB6" box="[701,903,1344,1368]" pageId="14" pageNumber="99">CASENT 1026771</specimenCode>
” to “ 
<specimenCode id="DBA01261FFD1FFBBFC5BFA8FA58FFAB6" box="[959,1161,1344,1368]" pageId="14" pageNumber="99">CASENT 1026773</specimenCode>
” and
</materialsCitation>
<materialsCitation id="3B6EB047FFD1FFBBFF49FAAEA0EBFA34" ID-GBIF-Occurrence="4899905512" collectingDate="2006-08-18" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; J. A. Miller &amp; D. Z. Dong &amp; Y. Liu" country="China" county="Gongshan County" elevation="3720" latitude="27.98121" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" pageId="14" pageNumber="99" specimenCode="CASENT 1026774, CASENT 1026777" specimenCount="1" specimenCount-female="4" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
“ 
<specimenCode id="DBA01261FFD1FFBBFF5DFAAEA08CFA96" box="[185,394,1376,1401]" pageId="14" pageNumber="99">CASENT 1026774</specimenCode>
” to “ 
<specimenCode id="DBA01261FFD1FFBBFE2BFAAEA399FA96" box="[463,671,1376,1401]" pageId="14" pageNumber="99">CASENT 1026777</specimenCode>
”, respectively/ “ 
<collectingCountry id="F311FA8AFFD1FFBBFCBBFAAEA2B5FA97" box="[863,947,1377,1401]" name="China" pageId="14" pageNumber="99">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFD1FFBBFC21FAAEA510FA96" box="[965,1046,1377,1400]" country="China" name="Yunnan" pageId="14" pageNumber="99">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFD1FFBBFBCDFAAFA1CEFA77" pageId="14" pageNumber="99">Gongshan County</collectingCounty>
, 
<collectingMunicipality id="6BDD2060FFD1FFBBFF3CFA4EA0E4FA77" box="[216,482,1409,1433]" pageId="14" pageNumber="99">Bingzhongluo Township</collectingMunicipality>
, SW slope of 
<location id="8ED9ECC1FFD1FFBBFD62FA4EA246FA77" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFD1FFBBFD62FA4EA246FA77" box="[646,832,1409,1433]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="14" pageNumber="99" stateProvince="Yunnan">Kawakarpu Shan</location>
at 
<location id="8ED9ECC1FFD1FFBBFC83FA4EA507FA77" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFD1FFBBFC83FA4EA507FA77" box="[871,1025,1409,1433]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Chukuai Lake" pageId="14" pageNumber="99" stateProvince="Yunnan">Chukuai Lake</location>
, 
<quantity id="4CFE17FFFFD1FFBBFBF6FA4EA560FA76" box="[1042,1126,1409,1433]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="14" pageNumber="99" unit="m" value="3720.0">
<elevation id="002B5D29FFD1FFBBFBF6FA4EA560FA76" box="[1042,1126,1409,1433]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="14" pageNumber="99" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,”/ “ 
<geoCoordinate id="EE32DCDDFFD1FFBBFF9FFA6DA1FBFA54" box="[123,253,1442,1466]" degrees="27. 98121" direction="north" orientation="latitude" pageId="14" pageNumber="99" precision="1" value="27.98121">N27.98121°</geoCoordinate>
/ 
<geoCoordinate id="EE32DCDDFFD1FFBBFEE0FA6DA097FA54" box="[260,401,1441,1466]" degrees="098. 47580" direction="east" orientation="longitude" pageId="14" pageNumber="99" precision="1" value="98.4758">E098.47580°</geoCoordinate>
, 
<date id="FFB89CDAFFD1FFBBFE44FA6DA34AFA57" box="[416,588,1441,1466]" pageId="14" pageNumber="99" value="2006-08-18">
<collectingDate id="EFFC6532FFD1FFBBFE44FA6DA34AFA57" box="[416,588,1441,1466]" pageId="14" pageNumber="99" value="2006-08-18">18 August 2006</collectingDate>
</date>
, Stop #DHK-2006-079 
<collectorName id="26F3DFCCFFD1FFBBFCB0FA6DA50CFA57" box="[852,1034,1441,1465]" pageId="14" pageNumber="99">D.H. Kavanaugh</collectorName>
, 
<collectorName id="26F3DFCCFFD1FFBBFBFDFA6DA589FA57" box="[1049,1167,1441,1466]" pageId="14" pageNumber="99">J.A. Miller</collectorName>
, 
<collectorName id="26F3DFCCFFD1FFBBFF9FFA0DA1F2FA34" box="[123,244,1474,1498]" pageId="14" pageNumber="99">D.Z. Dong</collectorName>
&amp; 
<collectorName id="26F3DFCCFFD1FFBBFEC7FA0DA06AFA34" box="[291,364,1474,1498]" pageId="14" pageNumber="99">Y. Liu</collectorName>
collectors”
</materialsCitation>
; 
<materialsCitation id="3B6EB047FFD1FFBBFDE5FA0DA564FA34" ID-GBIF-Occurrence="4899905500" box="[513,1122,1474,1498]" collectingDate="2006-08-20" collectionCode="CAS, IOZ" collectorName="Y. Liu &amp; P. Hu &amp; D. Z. Dong &amp; J. Wang" country="China" county="Gongshan County" elevation="3950" latitude="27.98206" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" pageId="14" pageNumber="99" specimenCode="CASENT 1024860" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFD1FFBBFDE5FA0DA373FA34" box="[513,629,1474,1498]" count="2" pageId="14" pageNumber="99" type="male">two males</specimenCount>
(
<collectionCode id="ED1722DFFFD1FFBBFD6FFA0DA3B9FA34" box="[651,703,1474,1498]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="99" type="Museum">CAS</collectionCode>
, 
<collectionCode id="ED1722DFFFD1FFBBFD37FA0DA3F9FA34" box="[723,767,1474,1498]" pageId="14" pageNumber="99">IOZ</collectionCode>
) labeled: “ 
<specimenCode id="DBA01261FFD1FFBBFC60FA0DA553FA34" box="[900,1109,1474,1498]" pageId="14" pageNumber="99">CASENT 1024860</specimenCode>
”
</materialsCitation>
<materialsCitation id="3B6EB047FFD1FFBBFB8BFA0DA03EF9B2" ID-GBIF-Occurrence="4899905532" collectingDate="2006-08-20" collectionCode="CAS, IOZ" collectorName="Y. Liu &amp; P. Hu &amp; D. Z. Dong &amp; J. Wang" country="China" county="Gongshan County" elevation="3950" latitude="27.98206" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" pageId="14" pageNumber="99" specimenCode="CASENT 1024861" specimenCount="1" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
and “ 
<specimenCode id="DBA01261FFD1FFBBFF63FA2CA053FA14" box="[135,341,1506,1531]" pageId="14" pageNumber="99">CASENT 1024861</specimenCode>
”, respectively/ “ 
<collectingCountry id="F311FA8AFFD1FFBBFDF6FA2CA360FA15" box="[530,614,1507,1531]" name="China" pageId="14" pageNumber="99">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFD1FFBBFD93FA2CA3CEFA14" box="[631,712,1507,1530]" country="China" name="Yunnan" pageId="14" pageNumber="99">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFD1FFBBFD3EFA2DA298FA15" box="[730,926,1506,1531]" pageId="14" pageNumber="99">Gongshan County</collectingCounty>
, 
<collectingMunicipality id="6BDD2060FFD1FFBBFC4BFA2DA1AFF9F5" pageId="14" pageNumber="99">Bingzhongluo Township</collectingMunicipality>
, SW slope of 
<location id="8ED9ECC1FFD1FFBBFE93F9CCA346F9F5" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFD1FFBBFE93F9CCA346F9F5" box="[375,576,1539,1563]" country="China" county="Gongshan County" latitude="27.98206" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="14" pageNumber="99" stateProvince="Yunnan">Kawakarpu Shan</location>
, on slope 
<locationDeviation id="9A0BA338FFD1FFBBFD33F9CCA2E7F9F5" box="[727,993,1539,1563]" location="Chukuai Lake" pageId="14" pageNumber="99">NE of Chukuai Lake</locationDeviation>
, 
<quantity id="4CFE17FFFFD1FFBBFC1FF9CCA55DF9F4" box="[1019,1115,1539,1563]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="14" pageNumber="99" unit="m" value="3950.0">
<elevation id="002B5D29FFD1FFBBFC1FF9CCA55DF9F4" box="[1019,1115,1539,1563]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="14" pageNumber="99" unit="m" value="3950.0">3950 m</elevation>
</quantity>
,”/ “ 
<geoCoordinate id="EE32DCDDFFD1FFBBFF9FF9EBA1FBF9D2" box="[123,253,1571,1596]" degrees="27. 98206" direction="north" orientation="latitude" pageId="14" pageNumber="99" precision="1" value="27.98206">N27.98206°</geoCoordinate>
/ 
<geoCoordinate id="EE32DCDDFFD1FFBBFEE0F9EBA097F9D2" box="[260,401,1572,1596]" degrees="098. 48027" direction="east" orientation="longitude" pageId="14" pageNumber="99" precision="1" value="98.48027">E098.48027°</geoCoordinate>
, 
<date id="FFB89CDAFFD1FFBBFE7BF9EBA34CF9D5" box="[415,586,1571,1596]" pageId="14" pageNumber="99" value="2006-08-20">
<collectingDate id="EFFC6532FFD1FFBBFE7BF9EBA34CF9D5" box="[415,586,1571,1596]" pageId="14" pageNumber="99" value="2006-08-20">20 August 2006</collectingDate>
</date>
, Stop #DHK-2006-086 
<collectorName id="26F3DFCCFFD1FFBBFCABF9EBA295F9D5" box="[847,915,1571,1595]" pageId="14" pageNumber="99">Y. Liu</collectorName>
, 
<collectorName id="26F3DFCCFFD1FFBBFC46F9EBA2DBF9D5" box="[930,989,1572,1595]" pageId="14" pageNumber="99">P. Hu</collectorName>
, 
<collectorName id="26F3DFCCFFD1FFBBFC0FF9EBA559F9D2" box="[1003,1119,1572,1596]" pageId="14" pageNumber="99">D.Z. Dong</collectorName>
&amp; 
<collectorName id="26F3DFCCFFD1FFBBFB60F9EBA1BFF9B2" pageId="14" pageNumber="99">J. Wang</collectorName>
collectors”
</materialsCitation>
; 
<materialsCitation id="3B6EB047FFD1FFBBFEA8F98BA590F9B2" ID-GBIF-Occurrence="4899905436" box="[332,1174,1604,1628]" collectingDate="2006-08-20" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; J. A. Miller &amp; J. Xiong &amp; C. H. Li" country="China" county="Gongshan County" elevation="3920" latitude="27.98785" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47322" municipality="Bingzhongluo Township" pageId="14" pageNumber="99" specimenCode="CASENT 1024904" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFD1FFBBFEA8F98BA0B5F9B2" box="[332,435,1604,1628]" count="1" pageId="14" pageNumber="99" type="male">one male</specimenCount>
and 
<specimenCount id="9D007193FFD1FFBBFE17F98BA37CF9B2" box="[499,634,1604,1628]" count="2" pageId="14" pageNumber="99" type="female">two females</specimenCount>
(
<collectionCode id="ED1722DFFFD1FFBBFD74F98BA3C2F9B2" box="[656,708,1604,1628]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="99" type="Museum">CAS</collectionCode>
, 
<collectionCode id="ED1722DFFFD1FFBBFD33F98BA205F9B2" box="[727,771,1604,1628]" pageId="14" pageNumber="99">IOZ</collectionCode>
) labeled: “ 
<specimenCode id="DBA01261FFD1FFBBFC61F98BA550F9B2" box="[901,1110,1604,1628]" pageId="14" pageNumber="99">CASENT 1024904</specimenCode>
” and
</materialsCitation>
<materialsCitation id="3B6EB047FFD1FFA5FF9FF9AAA368FBC2" ID-GBIF-Occurrence="4899905485" collectingDate="2006-08-20" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; J. A. Miller &amp; J. Xiong &amp; C. H. Li" country="China" county="Gongshan County" elevation="3920" lastPageId="16" lastPageNumber="101" latitude="27.98785" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47322" municipality="Bingzhongluo Township" pageId="14" pageNumber="99" specimenCode="CASENT 1024905, CASENT 1024906" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
“ 
<specimenCode id="DBA01261FFD1FFBBFF63F9AAA057F992" box="[135,337,1636,1661]" pageId="14" pageNumber="99">CASENT 1024905</specimenCode>
” to “ 
<specimenCode id="DBA01261FFD1FFBBFE6FF9AAA350F992" box="[395,598,1636,1661]" pageId="14" pageNumber="99">CASENT 1024906</specimenCode>
”, respectively/ “ 
<collectingCountry id="F311FA8AFFD1FFBBFCF4F9AAA262F993" box="[784,868,1637,1661]" name="China" pageId="14" pageNumber="99">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFD1FFBBFC95F9AAA2C4F992" box="[881,962,1637,1660]" country="China" name="Yunnan" pageId="14" pageNumber="99">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFD1FFBBFC2BF9ABA589F993" box="[975,1167,1636,1661]" pageId="14" pageNumber="99">Gongshan County</collectingCounty>
, 
<collectingMunicipality id="6BDD2060FFD1FFBBFF9FF94AA095F973" box="[123,403,1669,1693]" pageId="14" pageNumber="99">Bingzhongluo Township</collectingMunicipality>
, SW slope of 
<location id="8ED9ECC1FFD1FFBBFDBCF94AA218F973" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFD1FFBBFDBCF94AA218F973" box="[600,798,1669,1693]" country="China" county="Gongshan County" latitude="27.98785" longLatPrecision="1" longitude="98.47322" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="14" pageNumber="99" stateProvince="Yunnan">Kawakarpu Shan</location>
, 
<locationDeviation id="9A0BA338FFD1FFBBFCD2F94AA589F973" box="[822,1167,1669,1693]" location="Chukuai Lake" pageId="14" pageNumber="99">
<quantity id="4CFE17FFFFD1FFBBFCD2F94AA28BF973" box="[822,909,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="99" unit="km" value="0.8">0.8 km</quantity>
N of Chukuai Lake
</locationDeviation>
, 
<geoCoordinate id="EE32DCDDFFD1FFBBFF9FF969A1FBF950" box="[123,253,1701,1726]" degrees="27. 98785" direction="north" orientation="latitude" pageId="14" pageNumber="99" precision="1" value="27.98785">N27.98785°</geoCoordinate>
/ 
<geoCoordinate id="EE32DCDDFFD1FFBBFEE0F969A097F950" box="[260,401,1702,1726]" degrees="098. 47322" direction="east" orientation="longitude" pageId="14" pageNumber="99" precision="1" value="98.47322">E098.47322°</geoCoordinate>
”/ “ 
<quantity id="4CFE17FFFFD1FFBBFE5DF969A30BF953" box="[441,525,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="3.92" pageId="14" pageNumber="99" unit="m" value="3920.0">
<elevation id="002B5D29FFD1FFBBFE5DF969A30BF953" box="[441,525,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="3.92" pageId="14" pageNumber="99" unit="m" value="3920.0">3920 m</elevation>
</quantity>
, 
<date id="FFB89CDAFFD1FFBBFDF9F969A3CAF953" box="[541,716,1701,1726]" pageId="14" pageNumber="99" value="2006-08-20">
<collectingDate id="EFFC6532FFD1FFBBFDF9F969A3CAF953" box="[541,716,1701,1726]" pageId="14" pageNumber="99" value="2006-08-20">20 August 2006</collectingDate>
</date>
, Stop #DHK-2006-088 
<collectorName id="26F3DFCCFFD1FFBBFC3CF969A588F953" box="[984,1166,1701,1725]" pageId="14" pageNumber="99">D.H. Kavanaugh</collectorName>
, 
<collectorName id="26F3DFCCFFCFFFA5FF9FFBDBA1F2FBC5" box="[123,244,1043,1068]" pageId="16" pageNumber="101">J.A. Miller</collectorName>
, 
<collectorName id="26F3DFCCFFCFFFA5FEE2FBDBA060FBC5" box="[262,358,1043,1067]" pageId="16" pageNumber="101">J. Xiong</collectorName>
, &amp; 
<collectorName id="26F3DFCCFFCFFFA5FE7DFBDBA0E9FBC5" box="[409,495,1043,1068]" pageId="16" pageNumber="101">C.H. Li</collectorName>
collectors”
</materialsCitation>
; 
<materialsCitation id="3B6EB047FFCFFFA5FD66FBDBA0E4FB60" ID-GBIF-Occurrence="4899905489" collectingDate="2006-08-21" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="China" county="Gongshan County" elevation="4035" location="Kawakarpu Shan" longitude="98.47518" municipality="Bingzhongluo Township" pageId="16" pageNumber="101" specimenCode="CASENT 1025034" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFCFFFA5FD66FBDBA3EEFBC5" box="[642,744,1043,1067]" count="1" pageId="16" pageNumber="101" type="male">one male</specimenCount>
(
<collectionCode id="ED1722DFFFCFFFA5FD19FBDBA237FBC2" box="[765,817,1044,1068]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="101" type="Museum">CAS</collectionCode>
) labeled: “ 
<specimenCode id="DBA01261FFCFFFA5FC56FBDBA584FBC5" box="[946,1154,1043,1068]" pageId="16" pageNumber="101">CASENT 1025034</specimenCode>
”/ “ 
<collectingCountry id="F311FA8AFFCFFFA5FF63FBFAA1DDFBA3" box="[135,219,1077,1101]" name="China" pageId="16" pageNumber="101">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFCFFFA5FF03FBFAA03EFBA2" box="[231,312,1077,1100]" country="China" name="Yunnan" pageId="16" pageNumber="101">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFCFFFA5FEA2FBFBA300FBA3" box="[326,518,1076,1101]" pageId="16" pageNumber="101">Gongshan County</collectingCounty>
, 
<collectingMunicipality id="6BDD2060FFCFFFA5FDF7FBFBA21FFBA2" box="[531,793,1076,1100]" pageId="16" pageNumber="101">Bingzhongluo Township</collectingMunicipality>
, SW slope of 
<location id="8ED9ECC1FFCFFFA5FC54FBFBA560FBA2" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFCFFFA5FC54FBFBA560FBA2" box="[944,1126,1076,1100]" country="China" county="Gongshan County" longitude="98.47518" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="16" pageNumber="101" stateProvince="Yunnan">Kawakarpu Shan</location>
, 
<locationDeviation id="9A0BA338FFCFFFA5FB90FBFAA07CFB83" location="Chukuai Lake" pageId="16" pageNumber="101">
<quantity id="4CFE17FFFFCFFFA5FB90FBFAA19BFB83" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="101" unit="km" value="0.9">0.9 km</quantity>
N of Chukuai Lake
</locationDeviation>
,”/ “ 
<geoCoordinate id="EE32DCDDFFCFFFA5FE4EFB9AA319FB83" box="[426,543,1109,1133]" degrees="098. 47518" direction="east" orientation="longitude" pageId="16" pageNumber="101" precision="1" value="98.47518">27.99005°/</geoCoordinate>
<geoCoordinate id="EE32DCDDFFCFFFA5FDC4FB99A3AAFB83" box="[544,684,1109,1133]" degrees="098. 47518" direction="east" orientation="longitude" pageId="16" pageNumber="101" precision="1" value="98.47518">E098.47518°</geoCoordinate>
, 
<quantity id="4CFE17FFFFCFFFA5FD57FB9AA200FB83" box="[691,774,1109,1133]" metricMagnitude="3" metricUnit="m" metricValue="4.035" pageId="16" pageNumber="101" unit="m" value="4035.0">
<elevation id="002B5D29FFCFFFA5FD57FB9AA200FB83" box="[691,774,1109,1133]" metricMagnitude="3" metricUnit="m" metricValue="4.035" pageId="16" pageNumber="101" unit="m" value="4035.0">4035 m</elevation>
</quantity>
, 
<date id="FFB89CDAFFCFFFA5FCF2FB9AA2C5FB83" box="[790,963,1109,1134]" pageId="16" pageNumber="101" value="2006-08-21">
<collectingDate id="EFFC6532FFCFFFA5FCF2FB9AA2C5FB83" box="[790,963,1109,1134]" pageId="16" pageNumber="101" value="2006-08-21">21 August 2006</collectingDate>
</date>
Stop #DHK-2006- 090 
<collectorName id="26F3DFCCFFCFFFA5FF54FBB8A06DFB60" box="[176,363,1142,1166]" pageId="16" pageNumber="101">D.H. Kavanaugh</collectorName>
collector”
</materialsCitation>
; 
<materialsCitation id="3B6EB047FFCFFFA5FE13FBB8A019FB41" ID-GBIF-Occurrence="4899905471" collectingDate="2005-08-17" collectingDateMax="2006-08-19" collectingDateMin="2005-08-17" collectionCode="CAS, IOZ" collectorName="Y. Liu &amp; H. B. Liang &amp; D. Z. Dong &amp; J. F. Zhang" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="16" pageNumber="101" specimenCode="CASENT 1025823" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFCFFFA5FE13FBB8A366FB60" box="[503,608,1142,1166]" count="1" pageId="16" pageNumber="101" type="male">one male</specimenCount>
and 
<specimenCount id="9D007193FFCFFFA5FD45FBB8A219FB60" box="[673,799,1142,1166]" count="1" pageId="16" pageNumber="101" type="female">one female</specimenCount>
(
<collectionCode id="ED1722DFFFCFFFA5FCD1FBB9A26FFB60" box="[821,873,1142,1166]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="101" type="Museum">CAS</collectionCode>
, 
<collectionCode id="ED1722DFFFCFFFA5FC99FBB9A2AFFB60" box="[893,937,1142,1166]" pageId="16" pageNumber="101">IOZ</collectionCode>
) labeled: “ 
<specimenCode id="DBA01261FFCFFFA5FBCAFBB9A1DDFB41" pageId="16" pageNumber="101">CASENT 1025823</specimenCode>
” and
</materialsCitation>
<materialsCitation id="3B6EB047FFCFFFA5FED4FB58A2D7FB1E" ID-GBIF-Occurrence="4899905304" collectingDate="2006-08-19" collectingDateMax="2006-08-19" collectingDateMin="2005-08-17" collectionCode="CAS, IOZ" collectorName="Y. Liu &amp; H. B. Liang &amp; D. Z. Dong &amp; J. F. Zhang" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="16" pageNumber="101" specimenCode="CASENT 1025824" specimenCount="1" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
“ 
<specimenCode id="DBA01261FFCFFFA5FED9FB58A311FB41" box="[317,535,1175,1199]" pageId="16" pageNumber="101">CASENT 1025824</specimenCode>
”, respectively/ “ 
<collectingCountry id="F311FA8AFFCFFFA5FD02FB58A23AFB41" box="[742,828,1175,1199]" name="China" pageId="16" pageNumber="101">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFCFFFA5FCB7FB57A2A1FB41" box="[851,935,1176,1199]" country="China" name="Yunnan" pageId="16" pageNumber="101">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFCFFFA5FC5BFB58A588FB41" box="[959,1166,1175,1199]" pageId="16" pageNumber="101">Gongshan County</collectingCounty>
, 
<collectingMunicipality id="6BDD2060FFCFFFA5FF9FFB77A087FB3E" box="[123,385,1208,1232]" pageId="16" pageNumber="101">Bingzhongluo Township</collectingMunicipality>
, SW slope of 
<location id="8ED9ECC1FFCFFFA5FDFCFB77A3C9FB3E" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFCFFFA5FDFCFB77A3C9FB3E" box="[536,719,1208,1232]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="16" pageNumber="101" stateProvince="Yunnan">Kawakarpu Shan</location>
, 
<locationDeviation id="9A0BA338FFCFFFA5FD39FB77A509FB3E" box="[733,1039,1208,1232]" location="Chukuai Lake" pageId="16" pageNumber="101">
<quantity id="4CFE17FFFFCFFFA5FD39FB77A221FB3E" box="[733,807,1208,1232]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="101" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite, 
<geoCoordinate id="EE32DCDDFFCFFFA5FF9FFB16A1FBFB1F" box="[123,253,1241,1265]" degrees="27. 97686" direction="north" orientation="latitude" pageId="16" pageNumber="101" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/ 
<geoCoordinate id="EE32DCDDFFCFFFA5FEE0FB16A097FB1F" box="[260,401,1241,1265]" degrees="098. 47799" direction="east" orientation="longitude" pageId="16" pageNumber="101" precision="1" value="98.47799">E098.47799°</geoCoordinate>
”/ “ 
<quantity id="4CFE17FFFFCFFFA5FE52FB16A300FB1E" box="[438,518,1241,1265]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="16" pageNumber="101" unit="m" value="3750.0">
<elevation id="002B5D29FFCFFFA5FE52FB16A300FB1E" box="[438,518,1241,1265]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="16" pageNumber="101" unit="m" value="3750.0">3750 m</elevation>
</quantity>
, 
<date id="FFB89CDAFFCFFFA5FDF7FB16A3BDFB1F" box="[531,699,1241,1265]" pageId="16" pageNumber="101" value="2006-08-19">
<collectingDate id="EFFC6532FFCFFFA5FDF7FB16A3BDFB1F" box="[531,699,1241,1265]" pageId="16" pageNumber="101" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-082 
<collectorName id="26F3DFCCFFCFFFA5FC5EFB16A2FAFB1F" box="[954,1020,1241,1265]" pageId="16" pageNumber="101">
Y. 
<materialsCitation id="3B6EB047FFCFFFA5FC33FB16A3FAFAFC" ID-GBIF-Occurrence="4899905305" collectingDate="2005-08-17" collectingDateMax="2006-08-19" collectingDateMin="2005-08-17" collectionCode="CAS, IOZ" collectorName="Y. Liu &amp; H. B. Liang &amp; D. Z. Dong &amp; J. F. Zhang" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="16" pageNumber="101" specimenCount="2" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
Liu collector; 
<specimenCount id="9D007193FFCFFFA5FB89FB16A1C8FAFC" count="2" pageId="16" pageNumber="101" type="female">two females</specimenCount>
(
<collectionCode id="ED1722DFFFCFFFA5FF02FB35A01CFAFC" box="[230,282,1274,1298]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="101" type="Museum">CAS</collectionCode>
, 
<collectionCode id="ED1722DFFFCFFFA5FED4FB35A05AFAFC" box="[304,348,1274,1298]" pageId="16" pageNumber="101">IOZ</collectionCode>
) labeled: “CASENT 1017826” and
</materialsCitation>
</collectorName>
</materialsCitation>
<materialsCitation id="3B6EB047FFCFFFA5FCEFFB35A366FA9B" ID-GBIF-Occurrence="4899905395" collectingDate="2005-08-17" collectingDateMax="2006-08-19" collectingDateMin="2005-08-17" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; D. Z. Dong &amp; J. F. Zhang" country="China" county="Gongshan County" elevation="3710" latitude="27.97686" location="Fugong County" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="16" pageNumber="101" specimenCode="CASENT 1017827" specimenCount="1" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
“ 
<specimenCode id="DBA01261FFCFFFA5FCF3FB35A2EDFAFC" box="[791,1003,1274,1298]" pageId="16" pageNumber="101">CASENT 1017827</specimenCode>
”, respectively/ “ 
<collectingCountry id="F311FA8AFFCFFFA5FF63FAD4A1DDFADD" box="[135,219,1307,1331]" name="China" pageId="16" pageNumber="101">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFCFFFA5FF03FAD4A03EFADC" box="[231,312,1307,1330]" country="China" name="Yunnan" pageId="16" pageNumber="101">Yunnan</collectingRegion>
, 
<location id="8ED9ECC1FFCFFFA5FEA2FAD4A0EDFADD" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFCFFFA5FEA2FAD4A0EDFADD" box="[326,491,1307,1331]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Fugong County" pageId="16" pageNumber="101" stateProvince="Yunnan">Fugong County</location>
, 
<location id="8ED9ECC1FFCFFFA5FE1CFAD4A3DFFADD" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFCFFFA5FE1CFAD4A3DFFADD" box="[504,729,1307,1331]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Lumadeng Township" pageId="16" pageNumber="101" stateProvince="Yunnan">Lumadeng Township</location>
, second cirque 
<locationDeviation id="9A0BA338FFCFFFA5FC64FAD4A54EFADD" box="[896,1096,1307,1331]" location="Shibali Yakou" pageId="16" pageNumber="101">S of Shibali Yakou</locationDeviation>
at border post “31”, 
<geoCoordinate id="EE32DCDDFFCFFFA5FEF3FAF3A09FFABA" box="[279,409,1340,1364]" degrees="27. 20333" direction="north" orientation="latitude" pageId="16" pageNumber="101" precision="1" value="27.20333">N27.20333°</geoCoordinate>
/ 
<geoCoordinate id="EE32DCDDFFCFFFA5FE44FAF3A32BFABA" box="[416,557,1339,1364]" degrees="098. 69303" direction="east" orientation="longitude" pageId="16" pageNumber="101" precision="1" value="98.69303">E098.69303°</geoCoordinate>
”/ “ 
<quantity id="4CFE17FFFFCFFFA5FDB6FAF3A3A5FABD" box="[594,675,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="3.71" pageId="16" pageNumber="101" unit="m" value="3710.0">
<elevation id="002B5D29FFCFFFA5FDB6FAF3A3A5FABD" box="[594,675,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="3.71" pageId="16" pageNumber="101" unit="m" value="3710.0">3710 m</elevation>
</quantity>
, 
<date id="FFB89CDAFFCFFFA5FD55FAF3A25CFABD" box="[689,858,1339,1364]" pageId="16" pageNumber="101" value="2005-08-17">
<collectingDate id="EFFC6532FFCFFFA5FD55FAF3A25CFABD" box="[689,858,1339,1364]" pageId="16" pageNumber="101" value="2005-08-17">17 August 2005</collectingDate>
</date>
, Stop #DHK-2005-095, 
<collectorName id="26F3DFCCFFCFFFA5FB87FAF3A1BFFA9A" pageId="16" pageNumber="101">H.B. Liang</collectorName>
, 
<collectorName id="26F3DFCCFFCFFFA5FF28FA92A042FA9B" box="[204,324,1373,1397]" pageId="16" pageNumber="101">D.Z. Dong</collectorName>
, &amp; 
<collectorName id="26F3DFCCFFCFFFA5FE9CFA92A0E6FA9A" box="[376,480,1372,1396]" pageId="16" pageNumber="101">J.F.Zhang</collectorName>
collectors”
</materialsCitation>
; 
<materialsCitation id="3B6EB047FFCFFFA5FD97FA92A0DFFA39" ID-GBIF-Occurrence="4899905404" collectingDate="2005-08-07" collectionCode="IOZ" collectorName="H. B. Liang &amp; J. F. Zhang" country="China" county="Fugong County" elevation="3450" latitude="27.21034" location="North Fork Yamu He" longLatPrecision="1" longitude="98.70141" municipality="Lishadi Township" pageId="16" pageNumber="101" specimenCode="CASENT 1018367" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFCFFFA5FD97FA92A3E9FA9A" box="[627,751,1372,1396]" count="1" pageId="16" pageNumber="101" type="female">one female</specimenCount>
(
<collectionCode id="ED1722DFFFCFFFA5FCE0FA92A236FA9B" box="[772,816,1373,1397]" pageId="16" pageNumber="101">IOZ</collectionCode>
) labeled: “ 
<specimenCode id="DBA01261FFCFFFA5FC55FA92A584FA9A" box="[945,1154,1372,1397]" pageId="16" pageNumber="101">CASENT 1018367</specimenCode>
”/ “ 
<collectingCountry id="F311FA8AFFCFFFA5FF63FAB2A1DDFA7B" box="[135,219,1405,1429]" name="China" pageId="16" pageNumber="101">CHINA</collectingCountry>
, 
<collectingRegion id="49C274F8FFCFFFA5FF0CFAB1A03FFA7B" box="[232,313,1406,1429]" country="China" name="Yunnan" pageId="16" pageNumber="101">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFCFFFA5FEADFAB1A0E9FA7B" box="[329,495,1405,1430]" pageId="16" pageNumber="101">Fugong County</collectingCounty>
, 
<collectingMunicipality id="6BDD2060FFCFFFA5FE19FAB2A3B8FA7B" box="[509,702,1405,1429]" pageId="16" pageNumber="101">Lishadi Township</collectingMunicipality>
, headwaters of 
<location id="8ED9ECC1FFCFFFA5FC8EFAB2A549FA7B" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFCFFFA5FC8EFAB2A549FA7B" box="[874,1103,1405,1429]" country="China" county="Fugong County" latitude="27.21034" longLatPrecision="1" longitude="98.70141" municipality="Lishadi Township" name="North Fork Yamu He" pageId="16" pageNumber="101" stateProvince="Yunnan">North Fork Yamu He</location>
just 
<locationDeviation id="9A0BA338FFCFFFA5FB61FAB1A036FA58" location="Shibali Yakou" pageId="16" pageNumber="101">E of Shibali Yakou</locationDeviation>
, 
<quantity id="4CFE17FFFFCFFFA5FEDBFA51A094FA58" box="[319,402,1438,1462]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="16" pageNumber="101" unit="m" value="3450.0">
<elevation id="002B5D29FFCFFFA5FEDBFA51A094FA58" box="[319,402,1438,1462]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="16" pageNumber="101" unit="m" value="3450.0">3450 m</elevation>
</quantity>
,”/ “ 
<geoCoordinate id="EE32DCDDFFCFFFA5FE5BFA50A344FA58" box="[447,578,1438,1462]" degrees="27. 21034" direction="north" orientation="latitude" pageId="16" pageNumber="101" precision="1" value="27.21034">N27.21034°</geoCoordinate>
/ 
<geoCoordinate id="EE32DCDDFFCFFFA5FDADFA50A3D3FA58" box="[585,725,1438,1462]" degrees="098. 70141" direction="east" orientation="longitude" pageId="16" pageNumber="101" precision="1" value="98.70141">E098.70141°</geoCoordinate>
, 
<date id="FFB89CDAFFCFFFA5FD00FA50A284FA58" box="[740,898,1438,1462]" pageId="16" pageNumber="101" value="2005-08-07">
<collectingDate id="EFFC6532FFCFFFA5FD00FA50A284FA58" box="[740,898,1438,1462]" pageId="16" pageNumber="101" value="2005-08-07">7 August 2005</collectingDate>
</date>
, Stop# LHB-05-52, 
<collectorName id="26F3DFCCFFCFFFA5FB87FA50A1BCFA39" pageId="16" pageNumber="101">H.B. Liang</collectorName>
&amp; 
<collectorName id="26F3DFCCFFCFFFA5FF0BFA0FA05EFA39" box="[239,344,1471,1495]" pageId="16" pageNumber="101">J.F.Zhang</collectorName>
collectors.
</materialsCitation>
All paratypes also bear the following label: “PARATYPE 
<taxonomicName id="4C06C199FFCFFFA5FF9FFA2FA3F9FA16" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[123,767,1504,1528]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="101" phylum="Arthropoda" rank="species" species="chukuai" status="sp. nov." subGenus="Sinobrosculus">
<emphasis id="B9726608FFCFFFA5FF9FFA2FA1F0FA16" box="[123,246,1504,1528]" italics="true" pageId="16" pageNumber="101">Broscodera</emphasis>
(
<emphasis id="B9726608FFCFFFA5FEE8FA2FA0A4FA16" box="[268,418,1504,1528]" italics="true" pageId="16" pageNumber="101">Sinobrosculus</emphasis>
) 
<emphasis id="B9726608FFCFFFA5FE5CFA2FA30AFA16" box="[440,524,1504,1528]" italics="true" pageId="16" pageNumber="101">chukuai</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
, 
<taxonomicNameLabel id="A241DB73FFCFFFA5FCF6FA2EA26DFA16" box="[786,875,1505,1528]" pageId="16" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [yellow label].
</paragraph>
<caption id="DF79EA92FFD0FFBAFF4AF99FA37EF978" ID-DOI="http://doi.org/10.5281/zenodo.11067373" ID-Zenodo-Dep="11067373" httpUri="https://zenodo.org/record/11067373/files/figure.png" pageId="15" pageNumber="100" startId="15.[174,230,1616,1635]" targetBox="[139,1176,216,1591]" targetPageId="15" targetType="figure">
<paragraph id="8BB9BA1AFFD0FFBAFF4AF99FA37EF978" blockId="15.[126,1177,1616,1686]" pageId="15" pageNumber="100">
Figure 11. 
<taxonomicName id="4C06C199FFD0FFBAFEEFF99FA0B1F98D" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[267,439,1616,1635]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="100" phylum="Arthropoda" rank="species" species="chukuai" status="sp. nov.">
<emphasis id="B9726608FFD0FFBAFEEFF99FA0B1F98D" box="[267,439,1616,1635]" italics="true" pageId="15" pageNumber="100">Broscodera chukuai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFD0FFBAFE58F99FA0F8F98D" box="[444,510,1616,1635]" pageId="15" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
a. Habitus (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<caption id="DF79EA92FFCFFFA5FF4FFC6AA3B3FC05" ID-DOI="http://doi.org/10.5281/zenodo.11067375" ID-Zenodo-Dep="11067375" httpUri="https://zenodo.org/record/11067375/files/figure.png" pageId="16" pageNumber="101" startId="16.[171,183,933,952]" targetBox="[147,1135,222,918]" targetPageId="16" targetType="figure">
<paragraph id="8BB9BA1AFFCFFFA5FF4FFC6AA3B3FC05" blockId="16.[123,1173,933,1003]" pageId="16" pageNumber="101">
<smallCapsWord id="8D5F2CC6FFCFFFA5FF4FFC6AA1EEFC59" baselines="947,947" box="[171,232,933,952]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="16" pageNumber="101">FIGURE</smallCapsWord>
12. Male genitalia, 
<taxonomicName id="4C06C199FFCFFFA5FE7CFC6AA343FC56" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[408,581,933,952]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="101" phylum="Arthropoda" rank="species" species="chukuai" status="sp. nov.">
<emphasis id="B9726608FFCFFFA5FE7CFC6AA343FC56" box="[408,581,933,952]" italics="true" pageId="16" pageNumber="101">Broscodera chukuai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFCFFFA5FDAFFC6AA388FC56" box="[587,654,933,952]" pageId="16" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
(CASENT1026169; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB9BA1AFFCFFFA5FF4FF9EDA314F9B4" blockId="16.[123,1174,1043,1726]" pageId="16" pageNumber="101">
<materialsCitation id="3B6EB047FFCFFFA5FF4FF9EDA30DF9B4" ID-GBIF-Occurrence="4899905458" collectorName="Bingzhongluo Township, SW" country="China" county="Gongshan County" location="Kawakarpu Shan" municipality="Bingzhongluo Township" pageId="16" pageNumber="101" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B9726608FFCFFFA5FF4FF9EDA044F9D4" bold="true" box="[171,322,1570,1594]" pageId="16" pageNumber="101">Type locality</emphasis>
. 
<collectingCountry id="F311FA8AFFCFFFA5FEB2F9EDA090F9D4" box="[342,406,1570,1594]" name="China" pageId="16" pageNumber="101">China</collectingCountry>
, 
<collectingRegion id="49C274F8FFCFFFA5FE4CF9EDA0FFF9D7" box="[424,505,1570,1593]" country="China" name="Yunnan" pageId="16" pageNumber="101">Yunnan</collectingRegion>
, 
<collectingCounty id="62D8C296FFCFFFA5FDE9F9EDA3D5F9D4" box="[525,723,1570,1594]" pageId="16" pageNumber="101">Gongshan County</collectingCounty>
, 
<collectorName id="26F3DFCCFFCFFFA5FD02F9EDA528F9D4" box="[742,1070,1570,1594]" pageId="16" pageNumber="101">Bingzhongluo Township, SW</collectorName>
slope of 
<location id="8ED9ECC1FFCFFFA5FF9FF98CA034F9B5" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFCFFFA5FF9FF98CA034F9B5" box="[123,306,1603,1627]" country="China" county="Gongshan County" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="16" pageNumber="101" stateProvince="Yunnan">Kawakarpu Shan</location>
, 
<location id="8ED9ECC1FFCFFFA5FEA4F98CA30DF9B4" LSID="urn:lsid:plazi:treatment:03AF0B0CFFD1FFA7FF9FFBC4A1E6F99E:8ED9ECC1FFCFFFA5FEA4F98CA30DF9B4" box="[320,523,1603,1627]" country="China" county="Gongshan County" municipality="Bingzhongluo Township" name="Chukuai Lake area" pageId="16" pageNumber="101" stateProvince="Yunnan">Chukuai Lake area</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFCFFFA5FF4FF9ACA3B7F953" pageId="16" pageNumber="101" type="etymology">
<paragraph id="8BB9BA1AFFCFFFA5FF4FF9ACA3B7F953" blockId="16.[123,1174,1043,1726]" pageId="16" pageNumber="101">
<emphasis id="B9726608FFCFFFA5FF4FF9ACA0E7F992" bold="true" box="[171,481,1635,1660]" pageId="16" pageNumber="101">Derivation of species name.</emphasis>
The species epithet, 
<taxonomicName id="4C06C199FFCFFFA5FD24F9ABA212F992" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[704,788,1636,1660]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="101" phylum="Arthropoda" rank="species" species="chukuai">
<emphasis id="B9726608FFCFFFA5FD24F9ABA212F992" box="[704,788,1636,1660]" italics="true" pageId="16" pageNumber="101">chukuai</emphasis>
</taxonomicName>
, is a noun in apposition and derived from the name of a lake on the southwestern slope of Kawakarpu 
<collectingRegion id="49C274F8FFCFFFA5FCB4F94BA283F972" box="[848,901,1668,1692]" country="Myanmar" name="Shan" pageId="16" pageNumber="101">Shan</collectingRegion>
, in the vicinity of which most of the specimens of the 
<typeStatus id="54BD04B8FFCFFFA5FE53F969A0E3F950" box="[439,485,1702,1726]" pageId="16" pageNumber="101">type</typeStatus>
series were found.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFCEFFA4FF4FFF14A1F8FD7B" pageId="17" pageNumber="102" type="diagnosis">
<paragraph id="8BB9BA1AFFCEFFA4FF4FFF14A1F8FD7B" blockId="17.[123,1174,219,1726]" pageId="17" pageNumber="102">
<emphasis id="B9726608FFCEFFA4FF4FFF14A019FF1D" bold="true" box="[171,287,219,243]" pageId="17" pageNumber="102">Diagnosis.</emphasis>
Adults of this species (
<figureCitation id="133DA69FFFCEFFA4FDC5FF14A37FFF1C" box="[545,633,219,243]" captionStart="Figure 11" captionStartId="15.[174,230,1616,1635]" captionTargetBox="[139,1176,216,1591]" captionTargetId="figure-67@15.[120,1176,216,1591]" captionTargetPageId="15" captionText="Figure 11. Broscodera chukuai sp. nov. a. Habitus (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067373" httpUri="https://zenodo.org/record/11067373/files/figure.png" pageId="17" pageNumber="102">Fig. 11a</figureCitation>
) can be distinguished from those of other species in the region by the following combination of character states: size small, BL male = 8.0–8. 
<quantity id="4CFE17FFFFCEFFA4FBB5FF34A588FEFD" box="[1105,1166,251,275]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="102" unit="mm" value="8.0">8 mm</quantity>
, female = 7.8–9. 
<quantity id="4CFE17FFFFCEFFA4FEC5FED4A058FEDD" box="[289,350,283,307]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="102" unit="mm" value="5.0">5 mm</quantity>
; body color reddish-brown; head (
<figureCitation id="133DA69FFFCEFFA4FD2BFED4A21FFEDD" box="[719,793,283,307]" captionStart="FIGURE" captionStartId="8.[171,183,1513,1532]" captionTargetBox="[139,1176,224,1495]" captionTargetId="figure-55@8.[120,1176,216,1495]" captionTargetPageId="8" captionText="FIGURE 5. Forebody of broscine carabids, dorsal aspect. a. Eoboscus bhutanensis Morvan (CASENT1019340; Lishadi Township, Fugong County, Yunnan, China); b. Broscus punctatus (Dejean) (CASENT1011066; Heiwadi, Cikai Township, Gongshan County, Yunnan, China); c. Broscodera chukuai sp. nov. (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo Township, Gongshan County, Yunnan, China); d. Broscodera gaoligongensis sp. nov. (CASENT1020104; South Fork of Yamu He, Lumadeng Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067361" httpUri="https://zenodo.org/record/11067361/files/figure.png" pageId="17" pageNumber="102">Fig. 5c</figureCitation>
) with eyes small, only slightly convex, their diameter slightly less than length of tempora; frontal furrows deeply impressed, smooth and impunctate; vertex with post-temporal transverse sulcus deep, sharply impressed and only narrowly punctate along bottom of sulcus; gula smooth, without transverse grooves; antennomere 3 with pubescence on apical half; pronotum with posterior transverse impression shallow and vaguely delineated; elytra with lateral margins slightly arcuate, microsculpture comprised of distinct and deeply impressed isodiametric meshes; hindwings slightly reduced; prosternum, proepisternum, metasternum, and metepisternum smooth or faintly punctulate; hind trochanter unisetose; abdominal sternite VI of female with two pairs of apicoparamedial setae. Male genitalia as in 
<figureCitation id="133DA69FFFCEFFA4FBA4FDD2A588FDDB" box="[1088,1166,541,565]" captionStart="FIGURE" captionStartId="16.[171,183,933,952]" captionTargetBox="[147,1135,222,918]" captionTargetId="figure-1483@16.[120,1176,210,928]" captionTargetPageId="16" captionText="FIGURE 12. Male genitalia, Broscodera chukuai sp. nov. (CASENT1026169; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067375" httpUri="https://zenodo.org/record/11067375/files/figure.png" pageId="17" pageNumber="102">Fig. 12</figureCitation>
; median lobe with ventral surface moderately concave with ventrolateral edges elevated and distinctly flanged near mid-length, especially on right edge; apex of apical lamella evenly rounded in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFCEFFA7FF4FFD51A0E3FEFD" lastPageId="18" lastPageNumber="103" pageId="17" pageNumber="102" type="description">
<paragraph id="8BB9BA1AFFCEFFA4FF4FFD51A25EFD38" blockId="17.[123,1174,219,1726]" pageId="17" pageNumber="102">
<emphasis id="B9726608FFCEFFA4FF4FFD51A030FD58" bold="true" box="[171,310,670,694]" pageId="17" pageNumber="102">Description.</emphasis>
<figureCitation id="133DA69FFFCEFFA4FED8FD51A095FD58" box="[316,403,670,694]" captionStart="Figure 11" captionStartId="15.[174,230,1616,1635]" captionTargetBox="[139,1176,216,1591]" captionTargetId="figure-67@15.[120,1176,216,1591]" captionTargetPageId="15" captionText="Figure 11. Broscodera chukuai sp. nov. a. Habitus (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067373" httpUri="https://zenodo.org/record/11067373/files/figure.png" pageId="17" pageNumber="102">Fig. 11a</figureCitation>
. Size small, BL male = 8.0–8. 
<quantity id="4CFE17FFFFCEFFA4FD3CFD51A210FD58" box="[728,790,670,694]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="102" unit="mm" value="8.0">8 mm</quantity>
, female = 7.8–9. 
<quantity id="4CFE17FFFFCEFFA4FC28FD51A50FFD58" box="[972,1033,670,694]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="102" unit="mm" value="5.0">5 mm</quantity>
, ratio EL/PL = 2.2-2.4. Body color reddish brown, all appendages reddish brown.
</paragraph>
<paragraph id="8BB9BA1AFFCEFFA4FF4FFD10A3FAFCB9" blockId="17.[123,1174,219,1726]" pageId="17" pageNumber="102">
Head. 
<figureCitation id="133DA69FFFCEFFA4FF14FD10A03CFD18" box="[240,314,734,759]" captionStart="FIGURE" captionStartId="8.[171,183,1513,1532]" captionTargetBox="[139,1176,224,1495]" captionTargetId="figure-55@8.[120,1176,216,1495]" captionTargetPageId="8" captionText="FIGURE 5. Forebody of broscine carabids, dorsal aspect. a. Eoboscus bhutanensis Morvan (CASENT1019340; Lishadi Township, Fugong County, Yunnan, China); b. Broscus punctatus (Dejean) (CASENT1011066; Heiwadi, Cikai Township, Gongshan County, Yunnan, China); c. Broscodera chukuai sp. nov. (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo Township, Gongshan County, Yunnan, China); d. Broscodera gaoligongensis sp. nov. (CASENT1020104; South Fork of Yamu He, Lumadeng Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067361" httpUri="https://zenodo.org/record/11067361/files/figure.png" pageId="17" pageNumber="102">Fig. 5c</figureCitation>
. Eyes small, only slightly convex, their diameter slightly less than length of tempora. Frontal furrows deeply impressed, narrow, smooth, impunctate. Vertex with post-temporal transverse sulcus deep, sharply impressed and only narrowly punctate along bottom of sulcus. Tempora roundly subangulate. Gula without transverse grooves.
</paragraph>
<paragraph id="8BB9BA1AFFCEFFA4FF4FFCAFA0D4FBB7" blockId="17.[123,1174,219,1726]" pageId="17" pageNumber="102">Pronotum. Markedly cordate, moderately convex, narrowed posteriorly, widest distinctly anterior to middle of discal region; apical margination absent; lateral margination present, narrow, distinctly delineated throughout pronotal length; basal margination absent; anterior transverse impression absent or shallow and indistinct; median longitudinal impression moderately impressed; posterior transverse impression shallow and vaguely delineated; anterior region smooth, or faintly and sparsely rugulose; pronotal base coarsely punctate and rugulose; one pair of midlateral pronotal setae present at anterior one-third to one-fourth, one pair of basolateral pronotal setae present slightly anterior to basal angles.</paragraph>
<paragraph id="8BB9BA1AFFCEFFA4FF4FFBAEA0FEFA35" blockId="17.[123,1174,219,1726]" pageId="17" pageNumber="102">Elytra. Elytral silhouette symmetrically subovoid, widest at or just behind middle, ratio EL/EW = 1.5–1.7, greatest elevation above lateral margin (in lateral view) posterior to middle; humeri distinctly present, roundly obtuse; lateral margins slightly arcuate; parascutellar striole present, short, distally merged in a distinct jog with free base of stria 1. Nine striae present; stria 1 moderately deeply and sharply impressed; striae 2 to 3, 4, or 5 moderately impressed in basal two-thirds and successively more shallowly impressed in apical third, striae 3 to 8 successively more shallowly impressed laterally and apically; stria 9 divergent medially from lateral groove in apical half; stria 1 moderately or finely punctate at least in basal half; striae 2 to 8 shallowly and sparsely punctate in basal two-thirds and successively more shallowly and sparsely punctate in apical third. Parascutellar setiferous pore absent, discal setae absent, umbilicate series comprised of one subhumeral and three subapical setae. Elytral microsculpture comprised of distinct and deeply impressed isodiametric sculpticells.</paragraph>
<paragraph id="8BB9BA1AFFCEFFA4FF4FFA2BA237FA12" blockId="17.[123,1174,219,1726]" box="[171,817,1508,1532]" pageId="17" pageNumber="102">Hindwings. Slightly reduced, incapable of supporting flight.</paragraph>
<paragraph id="8BB9BA1AFFCEFFA4FF4FF9CBA3FCF9B2" blockId="17.[123,1174,219,1726]" pageId="17" pageNumber="102">Thoracic venter. Prosternum and proepisternum smooth or faintly punctulate, mesosternum and mesepisternum coarsely but sparsely punctate, metasternum and metepisternum smooth or faintly punctulate. Metepisternum ratio ML/MW = 2.0–2.1.</paragraph>
<paragraph id="8BB9BA1AFFCEFFA4FF4FF9AAA0DBF953" blockId="17.[123,1174,219,1726]" pageId="17" pageNumber="102">Legs. Hind trochanter unisetose. Tarsomeres with sparse long setae dorsally; tarsomere 5 with a single subapical pair of setae ventrally; male pro- and mesotarsi each with small pads of adhesive setae on tarsomeres 1 and 2.</paragraph>
<paragraph id="8BB9BA1AFFCDFFA7FF4FFF14A590FF1C" blockId="18.[123,1174,219,1649]" box="[171,1174,219,243]" pageId="18" pageNumber="103">Abdomen. Sternite VI of male with one pair of apicoparamedial setae, female with two pairs.</paragraph>
<paragraph id="8BB9BA1AFFCDFFA7FF4FFF33A0E3FEFD" blockId="18.[123,1174,219,1649]" box="[171,485,252,276]" pageId="18" pageNumber="103">
Male genitalia. As in 
<figureCitation id="133DA69FFFCDFFA7FE76FF33A0D9FEFA" box="[402,479,252,276]" captionStart="FIGURE" captionStartId="16.[171,183,933,952]" captionTargetBox="[147,1135,222,918]" captionTargetId="figure-1483@16.[120,1176,210,928]" captionTargetPageId="16" captionText="FIGURE 12. Male genitalia, Broscodera chukuai sp. nov. (CASENT1026169; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067375" httpUri="https://zenodo.org/record/11067375/files/figure.png" pageId="18" pageNumber="103">Fig. 12</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFCDFFA7FF4FFED3A215FB06" pageId="18" pageNumber="103" type="discussion">
<paragraph id="8BB9BA1AFFCDFFA7FF4FFED3A2A5FBED" blockId="18.[123,1174,219,1649]" pageId="18" pageNumber="103">
<emphasis id="B9726608FFCDFFA7FF4FFED3A02BFEDB" bold="true" box="[171,301,284,309]" pageId="18" pageNumber="103">Comments.</emphasis>
Members of this species can be distinguished from those of 
<taxonomicName id="4C06C199FFCDFFA7FC51FED2A575FEDA" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[949,1139,284,308]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="gaoligongensis">
<emphasis id="B9726608FFCDFFA7FC51FED2A575FEDA" box="[949,1139,284,308]" italics="true" pageId="18" pageNumber="103">B. gaoligongensis</emphasis>
</taxonomicName>
by the features listed above in the key and diagnosis. They differ from members of all three subspecies of 
<taxonomicName id="4C06C199FFCDFFA7FF7EFE90A1FBFE98" authorityName="Deuve" authorityYear="1990" box="[154,253,350,374]" class="Insecta" family="Carabidae" genus="Sinobrosculus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="dreuxi">
<emphasis id="B9726608FFCDFFA7FF7EFE90A1FBFE98" box="[154,253,350,374]" italics="true" pageId="18" pageNumber="103">B. dreuxi</emphasis>
</taxonomicName>
from 
<collectingRegion id="49C274F8FFCDFFA7FEA7FE91A307FE98" box="[323,513,350,374]" country="China" name="Sichuan" pageId="18" pageNumber="103">Sichuan Province</collectingRegion>
, 
<collectingCountry id="F311FA8AFFCDFFA7FDF4FE91A356FE98" box="[528,592,350,374]" name="China" pageId="18" pageNumber="103">China</collectingCountry>
in having a lighter body color (reddish-brown rather than dark brown to piceous), head with impunctate (rather than punctate) frontal furrows and the post-temporal transverse sulcus only narrowly (rather than more broadly) punctate, the elytral striae only finely (rather than more coarsely) punctate, and the median lobe of the male genitalia shorter, thicker, and slightly more arcuate in lateral view compared with the illustration provided by 
<bibRefCitation id="EF97C7EBFFCDFFA7FF9FFDCDA046FDF7" author="Deuve, T." box="[123,320,513,537]" pageId="18" pageNumber="103" pagination="227 - 228" refId="ref52833" refString="Deuve, T. 1998. Note sur le genre Broscodera Lindroth, repandu depuis l'Himalaya jusqu'a l'Amerique du Nord (Coleotera, Broscidae). Nouvelle Revue d'Entomologie (N. S.) 14: 227 - 228." type="journal article" year="1998">Deuve (1998:228)</bibRefCitation>
. They differ from members of 
<taxonomicName id="4C06C199FFCDFFA7FD7EFDCEA50DFDF7" authority="Deuve, 2004" authorityName="Deuve" authorityYear="2004" box="[666,1035,513,537]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="morvani">
<emphasis id="B9726608FFCDFFA7FD7EFDCEA271FDF7" box="[666,887,513,537]" italics="true" pageId="18" pageNumber="103">Broscodera morvani</emphasis>
<bibRefCitation id="EF97C7EBFFCDFFA7FC64FDCDA50DFDF7" author="Deuve, T." box="[896,1035,513,537]" pageId="18" pageNumber="103" pagination="141 - 142" refId="ref52874" refString="Deuve, T. 2004. Nouveaux Broscidae de la region Himalayano-Tibetaine [Coleoptera, Caraboidea]. Revue Francaise d'Entomologie (N. S.) 26: 141 - 142." type="journal article" year="2004">Deuve, 2004</bibRefCitation>
</taxonomicName>
from 
<collectingRegion id="49C274F8FFCDFFA7FBB5FDCEA1DCFDD4" country="China" name="Gansu" pageId="18" pageNumber="103">Gansu Province</collectingRegion>
, 
<collectingCountry id="F311FA8AFFCDFFA7FF0FFDEDA02DFDD4" box="[235,299,546,570]" name="China" pageId="18" pageNumber="103">China</collectingCountry>
in having the eyes smaller, their diameter less than the length of the tempora (rather than greater than the temporal length), the antennae with antennomere 2 (the pedicel) longer, antennomere 3 with pubescence in its apical half (rather than without pubescence in addition to the apical setal whorl), the pronotum relatively longer and with a single midlateral seta on each side (rather with than two or three such setae), the elytral silhouette elongate-subovid with the lateral margins less rounded (rather than shorter, more ovoid, and with more markedly rounded lateral margins), and the median lobe of the male genitalia shorter, thicker, and more arcuate in lateral view (compared with the illustration provided for 
<taxonomicName id="4C06C199FFCDFFA7FD4EFCC7A227FCF1" authorityName="Deuve" authorityYear="2004" box="[682,801,776,799]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="morvani">
<emphasis id="B9726608FFCDFFA7FD4EFCC7A227FCF1" box="[682,801,776,799]" italics="true" pageId="18" pageNumber="103">B. morvani</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF97C7EBFFCDFFA7FCD4FCC8A2B4FCF1" author="Deuve, T." box="[816,946,775,799]" pageId="18" pageNumber="103" pagination="141 - 142" refId="ref52874" refString="Deuve, T. 2004. Nouveaux Broscidae de la region Himalayano-Tibetaine [Coleoptera, Caraboidea]. Revue Francaise d'Entomologie (N. S.) 26: 141 - 142." type="journal article" year="2004">Deuve 2004</bibRefCitation>
, 
<figureCitation id="133DA69FFFCDFFA7FC5BFCC8A2FBFCF1" box="[959,1021,775,799]" captionStart="FIGURE" captionStartId="4.[171,183,1161,1180]" captionTargetBox="[120,1176,216,1144]" captionTargetId="figure-1025@4.[120,1176,216,1144]" captionTargetPageId="4" captionText="FIGURE 2–4. Fig. 2. Map showing natural extent of study area, colored in green (however, sampling was permitted only in those portions in Yunnan Province. Fig. 3. Map showing location of core sampling areas. Fig. 4. Map showing locations of all entomological sampling sites. Scale lines = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067359" httpUri="https://zenodo.org/record/11067359/files/figure.png" pageId="18" pageNumber="103">Fig. 3</figureCitation>
). Members of 
<taxonomicName id="4C06C199FFCDFFA7FF9FFCE7A1EBFCD1" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[123,237,807,831]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="chukuai">
<emphasis id="B9726608FFCDFFA7FF9FFCE7A1EBFCD1" box="[123,237,807,831]" italics="true" pageId="18" pageNumber="103">B. chukuai</emphasis>
</taxonomicName>
differ from those of 
<taxonomicName id="4C06C199FFCDFFA7FE2BFCE8A257FCD1" authority="Wrase, 1995" authorityName="Wrase" authorityYear="1995" box="[463,849,807,831]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="holzschuhi">
<emphasis id="B9726608FFCDFFA7FE2BFCE8A3C2FCD1" box="[463,708,807,831]" italics="true" pageId="18" pageNumber="103">Broscodera holzschuhi</emphasis>
<bibRefCitation id="EF97C7EBFFCDFFA7FD2FFCE7A257FCD1" author="Wrase, D. W." box="[715,849,807,831]" pageId="18" pageNumber="103" pagination="337 - 366" refId="ref54880" refString="Wrase, D. W. 1995. Taxonomische und faunistische Bemerkungen uber einige palaarktische Carabiden-Arten (Coleoptera, Carabidae). Linzer Biologische Beitrage 27: 337 - 366." type="journal article" year="1995">Wrase, 1995</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F311FA8AFFCDFFA7FC70FCE8A2D2FCD1" box="[916,980,807,831]" name="Nepal" pageId="18" pageNumber="103">Nepal</collectingCountry>
in having smaller eyes, their diameter less than the length of the tempora (rather than greater than the temporal length), the pronotum relatively shorter and broader and widest distinctly anterior to mid-length (rather than closer to mid-length), and the elytral silhouette relatively shorter, broader, laterally more evenly curved throughout, and widest at or closer to midlength (rather than laterally nearly straight in basal half and more curved posteriorly and widest near two-thirds elytral length). We have not examined a male of 
<taxonomicName id="4C06C199FFCDFFA7FE5DFC23A34FFBEA" authorityName="Wrase" authorityYear="1995" box="[441,585,1004,1028]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="holzschuhi">
<emphasis id="B9726608FFCDFFA7FE5DFC23A34FFBEA" box="[441,585,1004,1028]" italics="true" pageId="18" pageNumber="103">B. holzschuhi</emphasis>
</taxonomicName>
for comparison of the genitalia.
</paragraph>
<paragraph id="8BB9BA1AFFCDFFA7FF4FFBC3A215FB06" blockId="18.[123,1174,219,1649]" pageId="18" pageNumber="103">
<emphasis id="B9726608FFCDFFA7FF4FFBC3A09EFBCA" bold="true" box="[171,408,1036,1060]" pageId="18" pageNumber="103">Habitat distribution.</emphasis>
Members of this species have been found under stones on open slopes above treeline, under deeply embedded stones on organic substrate in alpine tundra areas and near snow patches (
<figureCitation id="133DA69FFFCDFFA7FEF8FB81A070FB8B" box="[284,374,1102,1126]" captionStart="FIGURE" captionStartId="80.[171,183,1579,1598]" captionTargetBox="[127,1170,216,1566]" captionTargetId="figure-57@80.[127,1170,216,1566]" captionTargetPageId="80" captionText="FIGURE 40. Photographs of habitats for broscine species in the Gaoligong Shan region. a. SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, 3950 m, 27.982°/098.480°, Bingzhongluo Township, Gongshan County, Yunnan (habitat for Brosocodera chukuai sp. nov.); b. Second cirque S of Shibali Yakou at border post “31”, 3710 m, 27.203°/098.693°, Lumadeng Township, Fugong County, Yunnan (habitat for Brosocodera chukuai sp. nov., Broscosoma furvum sp. nov., and Broscosoma viridicollare sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067439" httpUri="https://zenodo.org/record/11067439/files/figure.png" pageId="18" pageNumber="103">Fig. 40a</figureCitation>
), and under stones on the upper banks of meltwater streams and on adjacent tundra flats. Specimens collected in Fugong County near Shibali Yakou were found under stones on organic soil along a small stream and in bamboo thickets (
<figureCitation id="133DA69FFFCDFFA7FC47FB40A504FB49" box="[931,1026,1167,1191]" captionStart="FIGURE" captionStartId="80.[171,183,1579,1598]" captionTargetBox="[127,1170,216,1566]" captionTargetId="figure-57@80.[127,1170,216,1566]" captionTargetPageId="80" captionText="FIGURE 40. Photographs of habitats for broscine species in the Gaoligong Shan region. a. SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, 3950 m, 27.982°/098.480°, Bingzhongluo Township, Gongshan County, Yunnan (habitat for Brosocodera chukuai sp. nov.); b. Second cirque S of Shibali Yakou at border post “31”, 3710 m, 27.203°/098.693°, Lumadeng Township, Fugong County, Yunnan (habitat for Brosocodera chukuai sp. nov., Broscosoma furvum sp. nov., and Broscosoma viridicollare sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067439" httpUri="https://zenodo.org/record/11067439/files/figure.png" pageId="18" pageNumber="103">Fig. 40b</figureCitation>
). Within the Gaoligong 
<collectingRegion id="49C274F8FFCDFFA7FF11FB7FA071FB26" box="[245,375,1200,1224]" country="Myanmar" name="Shan" pageId="18" pageNumber="103">Shan region</collectingRegion>
, this species occurs at relatively high elevations, with our records documenting its occurrence in the 
<quantity id="4CFE17FFFFCDFFA7FE5EFB1FA363FB06" box="[442,613,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="3.7424999999999997" metricValueMax="4.035" metricValueMin="3.45" pageId="18" pageNumber="103" unit="m" value="3742.5" valueMax="4035.0" valueMin="3450.0">3450 to 4035 m</quantity>
range (
<figureCitation id="133DA69FFFCDFFA7FD53FB1FA202FB07" box="[695,772,1232,1257]" captionStart="FIGURE" captionStartId="88.[184,196,1682,1701]" captionTargetBox="[130,1162,1063,1660]" captionTargetId="figure-332@88.[120,1176,1054,1668]" captionTargetPageId="88" captionText="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." figureDoi="http://doi.org/10.5281/zenodo.11067457" httpUri="https://zenodo.org/record/11067457/files/figure.png" pageId="18" pageNumber="103">Fig. 49</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFCDFFA7FF4FFB3EA1E6F99E" pageId="18" pageNumber="103" type="distribution">
<paragraph id="8BB9BA1AFFCDFFA7FF4FFB3EA08BFA65" blockId="18.[123,1174,219,1649]" pageId="18" pageNumber="103">
<emphasis id="B9726608FFCDFFA7FF4FFB3EA216FAE7" bold="true" box="[171,784,1265,1289]" pageId="18" pageNumber="103">
Geographical distribution within the Gaoligong 
<collectingRegion id="49C274F8FFCDFFA7FD2BFB3EA20FFAE7" box="[719,777,1265,1289]" country="Myanmar" name="Shan" pageId="18" pageNumber="103">Shan</collectingRegion>
.
</emphasis>
<figureCitation id="133DA69FFFCDFFA7FCF3FB3EA269FAE7" box="[791,879,1265,1289]" captionStart="Figure 11" captionStartId="15.[174,230,1616,1635]" captionTargetBox="[139,1176,216,1591]" captionTargetId="figure-67@15.[120,1176,216,1591]" captionTargetPageId="15" captionText="Figure 11. Broscodera chukuai sp. nov. a. Habitus (CASENT1026168; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067373" httpUri="https://zenodo.org/record/11067373/files/figure.png" pageId="18" pageNumber="103">Fig. 11b</figureCitation>
. We examined a total of 
<specimenCount id="9D007193FFCDFFA7FB9FFB3EA1ECFAC4" count="32" pageId="18" pageNumber="103" type="generic">32 specimens</specimenCount>
(
<specimenCount id="9D007193FFCDFFA7FF1FFADDA05DFAC4" box="[251,347,1298,1322]" count="12" pageId="18" pageNumber="103" type="male">12 males</specimenCount>
and 
<specimenCount id="9D007193FFCDFFA7FE76FADDA301FAC4" box="[402,519,1298,1322]" count="20" pageId="18" pageNumber="103" type="female">20 females</specimenCount>
) from the northern to the northcentral part of the Gaoligong 
<collectingRegion id="49C274F8FFCDFFA7FF9FFAFCA1B6FAA5" box="[123,176,1331,1355]" country="Myanmar" name="Shan" pageId="18" pageNumber="103">Shan</collectingRegion>
in Fugong and Gongshan Counties. Our records from Gongshan and Fugong Counties are on the eastern side of the range (in Core Areas 2 and 3, respectively (
<figureCitation id="133DA69FFFCDFFA7FCD7FA9CA278FA85" box="[819,894,1363,1387]" captionStart="FIGURE" captionStartId="88.[171,183,995,1014]" captionTargetBox="[130,1164,233,969]" captionTargetId="figure-57@88.[120,1176,215,989]" captionTargetPageId="88" captionText="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." figureDoi="http://doi.org/10.5281/zenodo.11067455" httpUri="https://zenodo.org/record/11067455/files/figure.png" pageId="18" pageNumber="103">Fig. 48</figureCitation>
) (see 
<typeStatus id="54BD04B8FFCDFFA7FC59FA9BA2F4FA82" box="[957,1010,1364,1388]" pageId="18" pageNumber="103">Type</typeStatus>
material above for exact collection data).
</paragraph>
<paragraph id="8BB9BA1AFFCDFFA7FF4FFA5BA2B0FA23" blockId="18.[123,1174,219,1649]" pageId="18" pageNumber="103">
<emphasis id="B9726608FFCDFFA7FF4FFA5BA328FA43" bold="true" box="[171,558,1428,1453]" pageId="18" pageNumber="103">Overall geographical distribution.</emphasis>
This species currently is known only from the northern to northcentral part of the Gaoligong Shan, in western 
<collectingRegion id="49C274F8FFCDFFA7FD4EFA79A267FA23" box="[682,865,1461,1485]" country="China" name="Yunnan" pageId="18" pageNumber="103">Yunnan Province</collectingRegion>
, 
<collectingCountry id="F311FA8AFFCDFFA7FC8BFA7AA2A9FA23" box="[879,943,1461,1485]" name="China" pageId="18" pageNumber="103">China</collectingCountry>
.
</paragraph>
<paragraph id="8BB9BA1AFFCDFFA7FF4FFA19A1E6F99E" blockId="18.[123,1174,219,1649]" pageId="18" pageNumber="103">
<emphasis id="B9726608FFCDFFA7FF4FFA19A277FA00" bold="true" box="[171,881,1494,1518]" pageId="18" pageNumber="103">
Geographical relationships with other 
<taxonomicName id="4C06C199FFCDFFA7FD6FFA19A277FA00" box="[651,881,1494,1518]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9726608FFCDFFA7FD6FFA19A20CFA00" bold="true" box="[651,778,1494,1518]" italics="true" pageId="18" pageNumber="103">Broscodera</emphasis>
species.
</taxonomicName>
</emphasis>
As noted below for 
<taxonomicName id="4C06C199FFCDFFA7FB9AFA18A01AF9E1" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="gaoligongensis">
<emphasis id="B9726608FFCDFFA7FB9AFA18A01AF9E1" italics="true" pageId="18" pageNumber="103">B. gaoligongensis</emphasis>
</taxonomicName>
, the known geographical range of that species partially overlaps with that of 
<taxonomicName id="4C06C199FFCDFFA7FB9AFA38A1C9F9C1" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="chukuai">
<emphasis id="B9726608FFCDFFA7FB9AFA38A1C9F9C1" italics="true" pageId="18" pageNumber="103">B. chukuai</emphasis>
</taxonomicName>
, but they have not been found together (
<figureCitation id="133DA69FFFCDFFA7FD76F9D8A3E7F9C1" box="[658,737,1559,1583]" captionStart="FIGURE" captionStartId="89.[171,183,1461,1480]" captionTargetBox="[140,1154,249,1428]" captionTargetId="figure-68@89.[120,1176,216,1448]" captionTargetPageId="89" captionText="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi="http://doi.org/10.5281/zenodo.11067459" httpUri="https://zenodo.org/record/11067459/files/figure.png" pageId="18" pageNumber="103">Fig. 50</figureCitation>
) and appear to occupy non-overlapping altitudinal ranges. No other described species of 
<taxonomicName id="4C06C199FFCDFFA7FD43F9F7A224F9BE" authorityName="Lindroth" authorityYear="1961" box="[679,802,1592,1616]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9726608FFCDFFA7FD43F9F7A224F9BE" box="[679,802,1592,1616]" italics="true" pageId="18" pageNumber="103">Broscodera</emphasis>
</taxonomicName>
has been recorded from 
<collectingRegion id="49C274F8FFCDFFA7FBA0F9F6A1DCF99F" country="China" name="Yunnan" pageId="18" pageNumber="103">Yunnan Province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>