481 lines
56 KiB
XML
481 lines
56 KiB
XML
<document id="A528DF361B1ABFFE58929FCE5109177A" ID="10.11646/zootaxa.4269.4.9SLASH11131" ID-CLB-Dataset="32980" ID-DOI="10.11646/zootaxa.4269.4.9" ID-GBIF-Dataset="ea96b06d-bfc2-45d7-8e60-68e24c23ccf8" ID-ISSN="1175-5326" ID-Zenodo-Dep="582788" ID-ZooBank="DDEF97BF-EE79-468C-A3B1-1F8BFCB2DA17" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495615120791" checkinUser="plazi" docAuthor="Winterbottom, Richard" docDate="2017" docId="03ABCE693556A37D6DC4F92D1D97FD16" docLanguage="en" docName="zootaxa.4269.4.9SLASH11131.pdf" docOrigin="Zootaxa 4269 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trimma bathum Winterbottom, 2017, new species" docType="treatment" docUuid="FC0DD9A7-1CE1-4982-9DFA-1DFF5C050D3E" docUuidSource="ZooBank" docVersion="9" lastPageNumber="564" masterDocId="FF92B6113557A3786D53FFC11F08FFE6" masterDocTitle="Two new species of Trimma (Pisces; Gobiidae) from Fiji, south-western Pacific Ocean" masterLastPageNumber="570" masterPageNumber="559" pageNumber="560" updateTime="1698439880072" updateUser="ExternalLinkService">
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<mods:title id="D4A6C5E07C98312048BD1070CDC9E552">Two new species of Trimma (Pisces; Gobiidae) from Fiji, south-western Pacific Ocean</mods:title>
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<mods:name id="D41C40509E07AD929B72A4862F6347C9" type="personal">
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<mods:namePart id="E9C51BE2C132C483E6CB5BCCF0D6E0A0">Winterbottom, Richard</mods:namePart>
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<mods:title id="A74DDFFD3954F46643F13FE14E8CCBA0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="92E93362CB4D60475F8DCC368B3B3AD0">
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<mods:date id="F7F866C59E71CC5F845999784948FABC">2017</mods:date>
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<mods:number id="2157CD78D9E4F43B702E72EAD1305229">2017-05-24</mods:number>
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<mods:identifier id="3051207C180DF664D505963D05120B4E" type="DOI">10.11646/zootaxa.4269.4.9</mods:identifier>
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<treatment id="03ABCE693556A37D6DC4F92D1D97FD16" ID-DOI="http://doi.org/10.5281/zenodo.6048827" ID-GBIF-Taxon="130844706" ID-Zenodo-Dep="6048827" LSID="urn:lsid:zoobank.org:act:FC0DD9A7-1CE1-4982-9DFA-1DFF5C050D3E" httpUri="http://treatment.plazi.org/id/03ABCE693556A37D6DC4F92D1D97FD16" lastPageId="5" lastPageNumber="564" pageId="1" pageNumber="560">
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<subSubSection id="C3182CF43556A3796DC4F92D1E0EF8AD" pageId="1" pageNumber="560" type="nomenclature">
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<paragraph id="8BBD7F7F3556A3796DC4F92D1EFBF8E0" blockId="1.[151,499,1772,1867]" box="[151,499,1772,1798]" pageId="1" pageNumber="560">
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<heading id="D0F5C8133556A3796DC4F92D1EFBF8E0" bold="true" box="[151,499,1772,1798]" fontSize="11" level="1" pageId="1" pageNumber="560" reason="1">
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<emphasis id="B976A36D3556A3796DC4F92D1EFBF8E0" bold="true" box="[151,499,1772,1798]" pageId="1" pageNumber="560">
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<taxonomicName id="4C0204FC3556A3796DC4F92D1E54F8E0" ID-CoL="58V4Q" box="[151,348,1772,1798]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="560" phylum="Chordata" rank="species" species="bathum" status="sp. nov.">
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<emphasis id="B976A36D3556A3796DC4F92D1E54F8E0" bold="true" box="[151,348,1772,1798]" italics="true" pageId="1" pageNumber="560">Trimma bathum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2451E163556A3796C30F92D1EFBF8E0" box="[355,499,1772,1798]" pageId="1" pageNumber="560" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBD7F7F3556A3796DC4F8CF1E0EF8AD" blockId="1.[151,499,1772,1867]" pageId="1" pageNumber="560">
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Deep-reef Pygmygoby
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<figureCitation id="133963FA3556A3796DC4F8F31E0AF8AD" box="[151,258,1842,1867]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1935,1957]" captionStartId-1="4.[151,250,863,885]" captionTargetBox-0="[176,1402,193,1887]" captionTargetBox-1="[193,1394,193,841]" captionTargetId-0="figure@3.[166,1402,193,1888]" captionTargetId-1="figure@4.[193,1394,193,841]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Trimma bathum, 16.4 mm SL male holotype, Suva, Fiji. Lateral (A, image reversed) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin yellow lines. Photos by R. Winterbottom." captionText-1="FIGURE 2. Trimma bathum, 17.6 (UK T 4454) & 16.4 (ROM T 02301) mm SL, Viti Levu, Fiji. Photo by R. Pyle." httpUri-0="https://zenodo.org/record/582789/files/figure.png" httpUri-1="https://zenodo.org/record/582790/files/figure.png" pageId="1" pageNumber="560">Figs. 1–2</figureCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3182CF43556A3796DC4F8BA1D9DF876" box="[151,661,1914,1936]" pageId="1" pageNumber="560" type="description">
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<paragraph id="8BBD7F7F3556A3796DC4F8BA1D9DF876" blockId="1.[151,661,1914,1936]" box="[151,661,1914,1936]" pageId="1" pageNumber="560">
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<treatmentCitationGroup id="AB1258513556A3796DC4F8BA1D9DF876" box="[151,661,1914,1936]" pageId="1" pageNumber="560">
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<taxonomicName id="4C0204FC3556A3796DC4F8BA1FEEF876" ID-CoL="827C" box="[151,230,1915,1936]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="560" phylum="Chordata" rank="genus">
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||
<emphasis id="B976A36D3556A3796DC4F8BA1FEEF876" box="[151,230,1915,1936]" italics="true" pageId="1" pageNumber="560">Trimma</emphasis>
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</taxonomicName>
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RW sp. 97—
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<treatmentCitation id="0AA3596E3556A3796C3DF8BB1D9DF876" author="Winterbottom" box="[366,661,1914,1936]" page="83" pageId="1" pageNumber="560" year="2014">
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<bibRefCitation id="EF93028E3556A3796C3DF8BB1D71F876" author="Winterbottom" box="[366,633,1914,1936]" pageId="1" pageNumber="560" refString="Winterbottom, R., Hanner, R. H., Burridge, M. & Zur, M. (2014) A cornucopia of cryptic species-a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys, 381, 79 - 111." type="journal article" year="2014">
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||
Winterbottom
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<emphasis id="B976A36D3556A3796F50F8BA1D27F876" box="[515,559,1914,1936]" italics="true" pageId="1" pageNumber="560">et al</emphasis>
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., 2014
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||
</bibRefCitation>
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||
:83
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||
</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3182CF43556A37A6DC4F87C1D21FE4B" lastPageId="2" lastPageNumber="561" pageId="1" pageNumber="560" type="materials_examined">
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<paragraph id="8BBD7F7F3556A37A6DC4F87C1D92FF33" blockId="1.[151,1436,1981,2006]" lastBlockId="2.[151,1437,151,2015]" lastPageId="2" lastPageNumber="561" pageId="1" pageNumber="560">
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<materialsCitation id="3B6A75223556A37A6DC4F87C1BEDFF49" ID-GBIF-Occurrence="1503738028" collectionCode="ROM" country="Fiji" county="Fish Patch" elevation="53" lastPageId="2" lastPageNumber="561" latitude="-18.16" location="Viti Levu" longLatPrecision="765" longitude="178.4" municipality="Suva" pageId="1" pageNumber="560" specimenCode="ROM 101384" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B976A36D3556A3796DC4F87C1EF6F830" bold="true" box="[151,510,1981,2006]" pageId="1" pageNumber="560">
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||
Material examined.
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||
<typeStatus id="54B9C1DD3556A3796CD9F87C1EF1F830" box="[394,505,1981,2006]" pageId="1" pageNumber="560" type="holotype">Holotype</typeStatus>
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.
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||
</emphasis>
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||
<specimenCode id="DBA4D7043556A3796F55F87F1DAAF830" box="[518,674,1982,2006]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="560">
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<collectionCode id="ED13E7BA3556A3796F55F87F1D4EF830" box="[518,582,1982,2006]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="560">ROM</collectionCode>
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<date id="FFBC59BF3556A3796F1DF87F1DAAF830" box="[590,674,1982,2006]" pageId="1" pageNumber="560" value="84-10-13">101384</date>
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</specimenCode>
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(ex-BPBM 40009),
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<quantity id="4CFAD29A3556A3796EDEF87C1CFAF833" box="[909,1010,1981,2006]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="1" pageNumber="560" unit="mm" value="16.4">16.4 mm</quantity>
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||
SL male,
|
||
<collectingCountry id="F3153FEF3556A379693AF87C1B9BF830" box="[1129,1171,1981,2006]" name="Fiji" pageId="1" pageNumber="560">Fiji</collectingCountry>
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,
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<location id="8EDD29A43556A37969F3F87C1A06F833" LSID="urn:lsid:plazi:treatment:03ABCE693556A37D6DC4F92D1D97FD16:8EDD29A43556A37969F3F87C1A06F833" box="[1184,1294,1981,2006]" country="Fiji" county="Fish Patch" latitude="-18.16" longLatPrecision="765" longitude="178.4" municipality="Suva" name="Viti Levu" pageId="1" pageNumber="560">Viti Levu</location>
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,
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<collectingMunicipality id="6BD9E5053556A3796848F87F1A5EF830" box="[1307,1366,1982,2006]" pageId="1" pageNumber="560">Suva</collectingMunicipality>
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, outer reef, S of "
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<collectingCounty id="62DC07F33555A37A6C5DFF561E81FF56" box="[270,393,151,176]" pageId="2" pageNumber="561">Fish Patch</collectingCounty>
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" southern wall, vertical reef drop-off,
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<geoCoordinate id="EE3619B83555A37A6E64FF561C91FF56" box="[823,921,151,176]" direction="south" orientation="latitude" pageId="2" pageNumber="561" precision="555" value="-18.16">18.16° S</geoCoordinate>
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,
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<geoCoordinate id="EE3619B83555A37A6EF5FF591B11FF49" box="[934,1049,152,176]" direction="east" orientation="longitude" pageId="2" pageNumber="561" precision="555" value="178.4">178.40° E</geoCoordinate>
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,
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<quantity id="4CFAD29A3555A37A6977FF561B7CFF49" box="[1060,1140,151,176]" metricMagnitude="1" metricUnit="m" metricValue="5.25" pageId="2" pageNumber="561" unit="m" value="52.5">
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<elevation id="002F984C3555A37A6977FF561B7CFF49" box="[1060,1140,151,176]" metricMagnitude="1" metricUnit="m" metricValue="5.25" pageId="2" pageNumber="561" unit="m" value="52.5">52.5 m</elevation>
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||
</quantity>
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, rotenone
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</materialsCitation>
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,
|
||
<date id="FFBC59BF3555A37A69A3FF591A31FF56" box="[1264,1337,151,176]" pageId="2" pageNumber="561">3 Feb.</date>
|
||
, 2002, J. L. Earle & J. Dituri, Field # RLP-FIJI02–015.
|
||
</paragraph>
|
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<paragraph id="8BBD7F7F3555A37A6D94FF1E1D21FE4B" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="561">
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<materialsCitation id="3B6A75223555A37A6D94FF1E1B62FEA6" ID-GBIF-Occurrence="1503738027" collectingDate="1931-01" collectionCode="BPBM" collectorName="J. L. Earle & J. Dituri & Field" country="Fiji" county="Fish Patch" elevation="87" latitude="-18.1642" location="Viti Levu" longLatPrecision="527" longitude="178.4" municipality="Suva Harbour" pageId="2" pageNumber="561" specimenCode="BPBM 39900, 15.5" specimenCount="1" typeStatus="paratype">
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<emphasis id="B976A36D3555A37A6D94FF1E1E4EFF1E" bold="true" box="[199,326,223,248]" pageId="2" pageNumber="561">
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<typeStatus id="54B9C1DD3555A37A6D94FF1E1E49FF1E" box="[199,321,223,248]" pageId="2" pageNumber="561" type="paratype">Paratypes</typeStatus>
|
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.
|
||
</emphasis>
|
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<specimenCode id="DBA4D7043555A37A6C1FFF211D2CFF1E" box="[332,548,223,248]" pageId="2" pageNumber="561">
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<collectionCode id="ED13E7BA3555A37A6C1FFF211E92FF11" box="[332,410,224,247]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="561">BPBM</collectionCode>
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39900,
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<quantity id="4CFAD29A3555A37A6CA0FF1E1D2CFF1E" box="[499,548,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="2" pageNumber="561" unit="mm" value="15.5">15.5</quantity>
|
||
</specimenCode>
|
||
mm SL,
|
||
<collectingCountry id="F3153FEF3555A37A6FD8FF1E1DBDFF1E" box="[651,693,223,248]" name="Fiji" pageId="2" pageNumber="561">Fiji</collectingCountry>
|
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,
|
||
<location id="8EDD29A43555A37A6FECFF1E1C23FF11" LSID="urn:lsid:plazi:treatment:03ABCE693556A37D6DC4F92D1D97FD16:8EDD29A43555A37A6FECFF1E1C23FF11" box="[703,811,223,248]" country="Fiji" county="Fish Patch" latitude="-18.1642" longLatPrecision="527" longitude="178.4" municipality="Suva Harbour" name="Viti Levu" pageId="2" pageNumber="561">Viti Levu</location>
|
||
, outside of
|
||
<collectingMunicipality id="6BD9E5053555A37A6EF8FF211B42FF1E" box="[939,1098,223,248]" pageId="2" pageNumber="561">Suva Harbour</collectingMunicipality>
|
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; beyond "
|
||
<collectingCounty id="62DC07F33555A37A69E5FF1E1A3CFF1E" box="[1206,1332,223,248]" pageId="2" pageNumber="561">Fish Patch</collectingCounty>
|
||
", directly off bow of old shipwreck on top of reef, vertical reef drop-off with small ledges and holes,
|
||
<geoCoordinate id="EE3619B83555A37A69CAFEC51A10FEFA" box="[1177,1304,260,285]" direction="south" orientation="latitude" pageId="2" pageNumber="561" precision="5" value="-18.1642">18.1642° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3619B83555A37A6877FEC51A9FFEFA" box="[1316,1431,260,284]" direction="east" orientation="longitude" pageId="2" pageNumber="561" precision="555" value="178.4">178.40° E</geoCoordinate>
|
||
,
|
||
<quantity id="4CFAD29A3555A37A6DC4FEE91E08FED9" box="[151,256,296,320]" metricMagnitude="1" metricUnit="m" metricValue="8.7" metricValueMax="9.1" metricValueMin="8.3" pageId="2" pageNumber="561" unit="m" value="87.0" valueMax="91.0" valueMin="83.0">
|
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<elevation id="002F984C3555A37A6DC4FEE91E08FED9" box="[151,256,296,320]" metricMagnitude="1" metricUnit="m" metricValue="8.7" metricValueMax="9.1" metricValueMin="8.3" pageId="2" pageNumber="561" unit="m" value="87.0" valueMax="91.0" valueMin="83.0">83–91 m</elevation>
|
||
</quantity>
|
||
, rotenone,
|
||
<date id="FFBC59BF3555A37A6CD0FEE91ED1FED9" box="[387,473,296,320]" pageId="2" pageNumber="561" value="1931-01">
|
||
<collectingDate id="EFF8A0573555A37A6CD0FEE91ED1FED9" box="[387,473,296,320]" pageId="2" pageNumber="561" value="1931-01">31 Jan.</collectingDate>
|
||
</date>
|
||
, 2002,
|
||
<collectorName id="26F71AA93555A37A6F61FEE91DA5FEA6" box="[562,685,295,320]" pageId="2" pageNumber="561">J. L. Earle</collectorName>
|
||
&
|
||
<collectorName id="26F71AA93555A37A6F8AFEE91C34FEA6" box="[729,828,295,320]" pageId="2" pageNumber="561">J. Dituri</collectorName>
|
||
,
|
||
<collectorName id="26F71AA93555A37A6E19FEE61C8CFEA6" box="[842,900,295,320]" pageId="2" pageNumber="561">Field</collectorName>
|
||
# RLP-FIJI02–009
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6A75223555A37A692BFEE91C7BFE6E" ID-GBIF-Occurrence="1503738033" collectingDate="1928-01" collectionCode="BPBM" collectorName="R. L. Pyle & J. L. Earle & Field" country="Fiji" county="Fish Patch" elevation="62" latitude="-18.15983" location="Viti Levu" longLatPrecision="5" longitude="178.3993" municipality="Suva" pageId="2" pageNumber="561" specimenCode="BPBM 41320" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBA4D7043555A37A692BFEE91A1EFEA6" box="[1144,1302,296,320]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="561">
|
||
<collectionCode id="ED13E7BA3555A37A692BFEE91BCEFED9" box="[1144,1222,296,319]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="561">BPBM</collectionCode>
|
||
41320
|
||
</specimenCode>
|
||
(ex-BPBM 39712),
|
||
<quantity id="4CFAD29A3555A37A6DA7FE8D1E50FE82" box="[244,344,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="561" unit="mm" value="12.5">12.5 mm</quantity>
|
||
SL,
|
||
<collectingCountry id="F3153FEF3555A37A6CDFFE8D1EBEFE83" box="[396,438,332,357]" name="Fiji" pageId="2" pageNumber="561">Fiji</collectingCountry>
|
||
,
|
||
<location id="8EDD29A43555A37A6C92FE8D1D25FE82" LSID="urn:lsid:plazi:treatment:03ABCE693556A37D6DC4F92D1D97FD16:8EDD29A43555A37A6C92FE8D1D25FE82" box="[449,557,332,357]" country="Fiji" county="Fish Patch" latitude="-18.15983" longLatPrecision="5" longitude="178.3993" municipality="Suva" name="Viti Levu" pageId="2" pageNumber="561">Viti Levu</location>
|
||
,
|
||
<collectingMunicipality id="6BD9E5053555A37A6F6BFE8D1D7BFE82" box="[568,627,332,356]" pageId="2" pageNumber="561">Suva</collectingMunicipality>
|
||
, outer reef, "
|
||
<collectingCounty id="62DC07F33555A37A6E53FE8D1C75FE83" box="[768,893,332,357]" pageId="2" pageNumber="561">Fish Patch</collectingCounty>
|
||
",
|
||
<geoCoordinate id="EE3619B83555A37A6EC2FE8D1B15FE82" box="[913,1053,332,357]" direction="south" orientation="latitude" pageId="2" pageNumber="561" precision="1" value="-18.15983">18.15983° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3619B83555A37A697AFE8D1BBFFE82" box="[1065,1207,332,356]" direction="east" orientation="longitude" pageId="2" pageNumber="561" precision="5" value="178.3993">178.3993° E</geoCoordinate>
|
||
,
|
||
<quantity id="4CFAD29A3555A37A6990FE8D1A20FE82" box="[1219,1320,332,357]" metricMagnitude="1" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="2" pageNumber="561" unit="m" value="62.0" valueMax="67.0" valueMin="57.0">
|
||
<elevation id="002F984C3555A37A6990FE8D1A20FE82" box="[1219,1320,332,357]" metricMagnitude="1" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="2" pageNumber="561" unit="m" value="62.0" valueMax="67.0" valueMin="57.0">57–67 m</elevation>
|
||
</quantity>
|
||
, rotenone,
|
||
<date id="FFBC59BF3555A37A6DC4FEB11FE3FE61" box="[151,235,368,392]" pageId="2" pageNumber="561" value="1928-01">
|
||
<collectingDate id="EFF8A0573555A37A6DC4FEB11FE3FE61" box="[151,235,368,392]" pageId="2" pageNumber="561" value="1928-01">28 Jan.</collectingDate>
|
||
</date>
|
||
, 2002,
|
||
<collectorName id="26F71AA93555A37A6C11FEB11EA7FE6E" box="[322,431,367,392]" pageId="2" pageNumber="561">R.L. Pyle</collectorName>
|
||
&
|
||
<collectorName id="26F71AA93555A37A6C8BFEB11D42FE6E" box="[472,586,367,392]" pageId="2" pageNumber="561">J.L. Earle</collectorName>
|
||
,
|
||
<collectorName id="26F71AA93555A37A6F04FEAE1D99FE6E" box="[599,657,367,392]" pageId="2" pageNumber="561">Field</collectorName>
|
||
# RLP-FIJI02–001
|
||
</materialsCitation>
|
||
. ROM
|
||
<date id="FFBC59BF3555A37A6E99FEAE1B16FE6E" box="[970,1054,367,392]" pageId="2" pageNumber="561" value="85-10-13">101385</date>
|
||
(ex-BPBM 40009),
|
||
<quantity id="4CFAD29A3555A37A685FFEAE1A7AFE61" box="[1292,1394,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="2" pageNumber="561" unit="mm" value="15.8">15.8 mm</quantity>
|
||
SL female, collected with the holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3182CF43555A37A6D94FE761D58FD86" pageId="2" pageNumber="561" type="description">
|
||
<paragraph id="8BBD7F7F3555A37A6D94FE761D58FD86" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="561">
|
||
<emphasis id="B976A36D3555A37A6D94FE761EBFFE36" bold="true" box="[199,439,439,464]" pageId="2" pageNumber="561">Tissues (non-types):</emphasis>
|
||
KU:KUI T4454 (ex-ROM T02300; ex-BPBM 39849) and ROM T02301 (ex-KU:KUI T4458, ex-BPBM 39849), 17.6 and
|
||
<quantity id="4CFAD29A3555A37A6F6CFE1D1DAFFE12" box="[575,679,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="2" pageNumber="561" unit="mm" value="16.4">16.4 mm</quantity>
|
||
SL respectively,
|
||
<collectingCountry id="F3153FEF3555A37A6E26FE1D1C97FE13" box="[885,927,476,501]" name="Fiji" pageId="2" pageNumber="561">Fiji</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3153FEF3555A37A6EFDFE1D1CD0FE13" box="[942,984,476,501]" name="Fiji" pageId="2" pageNumber="561">Viti</collectingCountry>
|
||
Levu, Suva, outside of Suva Harbour, beyond "Fish patch", directly off bow of old shipwreck on top of reef, vertical reef drop-off with a large diagonal crack and over-hang, with some sea fans,
|
||
<geoCoordinate id="EE3619B83555A37A6F22FDE51DF5FDDA" box="[625,765,548,573]" direction="south" orientation="latitude" pageId="2" pageNumber="561" precision="1" value="-18.16422">18.16422° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3619B83555A37A6E58FDE51C92FDDA" box="[779,922,548,572]" direction="east" orientation="longitude" pageId="2" pageNumber="561" precision="5" value="178.4003">178.4003° E</geoCoordinate>
|
||
,
|
||
<quantity id="4CFAD29A3555A37A6EF4FDE51B04FDDA" box="[935,1036,548,573]" metricMagnitude="1" metricUnit="m" metricValue="6.85" metricValueMax="7.0" metricValueMin="6.7" pageId="2" pageNumber="561" unit="m" value="68.5" valueMax="70.0" valueMin="67.0">67–70 m</quantity>
|
||
, rotenone,
|
||
<date id="FFBC59BF3555A37A69D9FDE51BD5FDDA" box="[1162,1245,548,572]" pageId="2" pageNumber="561" value="1931-01">31 Jan.</date>
|
||
, 2002, J.L. Earle & D.F. Pence, Field # RLP-Fiji02–007.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3182CF43555A37A6D94FDAD1B11FCF1" pageId="2" pageNumber="561" type="diagnosis">
|
||
<paragraph id="8BBD7F7F3555A37A6D94FDAD1B11FCF1" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="561">
|
||
<emphasis id="B976A36D3555A37A6D94FDAD1E49FD63" bold="true" box="[199,321,620,645]" pageId="2" pageNumber="561">Diagnosis.</emphasis>
|
||
A species of
|
||
<taxonomicName id="4C0204FC3555A37A6CB5FDAC1D35FD62" box="[486,573,621,644]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="561" phylum="Chordata" rank="genus">
|
||
<emphasis id="B976A36D3555A37A6CB5FDAC1D35FD62" box="[486,573,621,644]" italics="true" pageId="2" pageNumber="561">Trimma</emphasis>
|
||
</taxonomicName>
|
||
with scales absent from the cheeks, opercle and predorsal midline, 18–19 unbranched pectoral fin rays, an unbranched 5th pelvic fin ray that is 40–56% the length of the 4th ray, 17–18 total gill rakers, a U-shaped interorbital and a narrow slit-like postorbital trench, a low, median fleshy ridge extending half-way towards the orbit from the origin of the first dorsal fin, and, when freshly collected, a pink head and body with most body scales having an orange-brown spot or short bar at their centres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3182CF43555A37C6D94FCE31CFFFA68" lastPageId="4" lastPageNumber="563" pageId="2" pageNumber="561" type="description">
|
||
<paragraph id="8BBD7F7F3555A37A6D94FCE31E8FF839" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="561">
|
||
<emphasis id="B976A36D3555A37A6D94FCE31E51FCDD" bold="true" box="[199,345,802,827]" pageId="2" pageNumber="561">Description.</emphasis>
|
||
The description is based on
|
||
<specimenCount id="9D04B4F63555A37A6FF3FCE21C26FCDD" box="[672,814,802,827]" pageId="2" pageNumber="561" type="generic">4 specimens</specimenCount>
|
||
,
|
||
<quantity id="4CFAD29A3555A37A6E68FCE31CD7FCDC" box="[827,991,802,827]" metricMagnitude="-2" metricUnit="m" metricValue="1.4449999999999998" metricValueMax="1.64" metricValueMin="1.25" pageId="2" pageNumber="561" unit="mm" value="14.45" valueMax="16.4" valueMin="12.5">12.5–16.4 mm</quantity>
|
||
SL (mean =15.1) from 3 lots collected off Suva Harbour,
|
||
<collectingCountry id="F3153FEF3555A37A6C3AFC871E9BFCB9" box="[361,403,838,863]" name="Fiji" pageId="2" pageNumber="561">Viti</collectingCountry>
|
||
Levu,
|
||
<collectingCountry id="F3153FEF3555A37A6CB7FC871D03FCB9" box="[484,523,838,863]" name="Fiji" pageId="2" pageNumber="561">Fiji</collectingCountry>
|
||
in 2002. An additional two specimens of this species were collected for analysis of the COI gene (see Discussion). Dorsal fin VI +
|
||
<date id="FFBC59BF3555A37A6FB8FCAA1C21FC65" box="[747,809,874,899]" pageId="2" pageNumber="561">
|
||
I 8–
|
||
<emphasis id="B976A36D3555A37A6E48FCAB1C21FC65" bold="true" box="[795,809,874,899]" pageId="2" pageNumber="561">9</emphasis>
|
||
</date>
|
||
(mean = 8.8, n = 4), second spine slightly elongated, reaching to base of spine of second dorsal fin, first ray of second dorsal fin
|
||
<emphasis id="B976A36D3555A37A6E89FC4F1B43FC41" bold="true" box="[986,1099,910,935]" pageId="2" pageNumber="561">branched</emphasis>
|
||
(n = 2) or unbranched (n = 2), remaining fin rays branched except for posterior element of last ray, which reaches posteriorly 68–
|
||
<emphasis id="B976A36D3555A37A6852FC731A15FC2D" bold="true" box="[1281,1309,946,971]" pageId="2" pageNumber="561">79</emphasis>
|
||
% (mean = 73.7%, n = 2) distance between its base and first exposed dorsal procurrent caudal fin ray; anal fin
|
||
<date id="FFBC59BF3555A37A69A8FC191A3EFC09" box="[1275,1334,982,1007]" pageId="2" pageNumber="561">
|
||
I 8–
|
||
<emphasis id="B976A36D3555A37A687BFC171A3EFC09" bold="true" box="[1320,1334,982,1007]" pageId="2" pageNumber="561">9</emphasis>
|
||
</date>
|
||
(mean = 8.8, n = 4), first ray
|
||
<emphasis id="B976A36D3555A37A6C2FFC3B1EE5FBF5" bold="true" box="[380,493,1018,1043]" pageId="2" pageNumber="561">branched</emphasis>
|
||
(n = 2) or unbranched (n = 2), last ray reaches posteriorly 65–
|
||
<emphasis id="B976A36D3555A37A69E3FC3B1BC4FBF5" bold="true" box="[1200,1228,1018,1043]" pageId="2" pageNumber="561">66</emphasis>
|
||
% (n = 2) distance between its base and first exposed ventral procurrent caudal fin ray; pectoral fin 18–
|
||
<emphasis id="B976A36D3555A37A6902FBDF1B65FBD1" bold="true" box="[1105,1133,1054,1079]" pageId="2" pageNumber="561">19</emphasis>
|
||
(mean = 18.8, n = 4), rays unbranched, fin reaching posteriorly to region above urogenital papilla to anal spine; pelvic fin
|
||
<date id="FFBC59BF3555A37A6854FB821A24FBBD" box="[1287,1324,1090,1115]" pageId="2" pageNumber="561">I 5</date>
|
||
, fifth ray unbranched and
|
||
<emphasis id="B976A36D3555A37A6C01FBA71E66FB99" bold="true" box="[338,366,1126,1151]" pageId="2" pageNumber="561">40</emphasis>
|
||
–56% (mean = 49.5, n = 3) length of fourth ray, which reaches posteriorly to between bases of second to fourth anal rays, pelvic rays 1–4 with a single sequential branch point; basal membrane apparently absent; no fraenum. Lateral scales 23 (n = 3); anterior transverse scales 9–
|
||
<emphasis id="B976A36D3555A37A6E86FB6F1CF9FB21" bold="true" box="[981,1009,1198,1223]" pageId="2" pageNumber="561">10</emphasis>
|
||
(n = 2); posterior transverse scales 8–
|
||
<emphasis id="B976A36D3555A37A6DC4FB131FADFB0D" bold="true" box="[151,165,1234,1259]" pageId="2" pageNumber="561">9</emphasis>
|
||
(n = 2); cheek, opercle and midline of predorsal without scales; anteriormost scale on side of nape reaching anteriorly to one scale width posterior to upper margin of eye; 3 vertical rows of cycloid scales on pectoral fin base with
|
||
<quantity id="4CFAD29A3555A37A6D82FADA1E12FAD5" box="[209,282,1306,1331]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="2" pageNumber="561" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">
|
||
1–
|
||
<emphasis id="B976A36D3555A37A6DBEFADB1FF3FAD5" bold="true" box="[237,251,1306,1331]" pageId="2" pageNumber="561">2</emphasis>
|
||
in
|
||
</quantity>
|
||
anteriormost row,
|
||
<quantity id="4CFAD29A3555A37A6CA1FADA1D16FAD5" box="[498,542,1306,1331]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="561" unit="in" value="4.0">4 in</quantity>
|
||
second row and
|
||
<quantity id="4CFAD29A3555A37A6FB1FADB1C06FAD5" box="[738,782,1306,1331]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="561" unit="in" value="5.0">5 in</quantity>
|
||
outer row (n = 2); 6 cycloid scales in midline anterior to pelvic fin base (n = 2); area between pelvic spine and ventral margin of pectoral fin base, midline of belly and sometimes anteriormost row of body scales beneath axil of pectoral fin base with cycloid scales. Circumpeduncular scales 12, scales rows in midline between base of last anal ray and first ventral procurrent caudal fin ray 7 (n = 2). Upper jaw teeth: outer row of enlarged, curved, spaced canines, decreasing in size posteriorly to distal end of premaxilla, several irregular inner rows of small, short conical teeth near symphysis, number of rows decreasing to a single row at bend of jaw. Lower jaw teeth: outer row of enlarged, curved, spaced canines reaching to bend of dentary, several irregular rows of small conical teeth at symphysis, decreasing to single row of somewhat enlarged (½ height of outer teeth), slightly curved teeth reaching to coronoid process. Tongue truncate with rounded edges. Gill opening extending anteroventrally to below mid-pupil; gill rakers
|
||
<emphasis id="B976A36D3555A37A6EE1F99F1CC8F991" bold="true" box="[946,960,1630,1655]" pageId="2" pageNumber="561">3</emphasis>
|
||
–4 + 13–
|
||
<emphasis id="B976A36D3555A37A6976F99F1B49F991" bold="true" box="[1061,1089,1630,1655]" pageId="2" pageNumber="561">14</emphasis>
|
||
=
|
||
<emphasis id="B976A36D3555A37A6933F99F1B74F991" bold="true" box="[1120,1148,1630,1655]" pageId="2" pageNumber="561">17</emphasis>
|
||
–18 (mean = 3.5 + 13.8 = 17.3, n = 4). Anterior nares in short tube reaching anteriorly to above anterior margin of upper lip, posterior opening pore-like with raised rim, separated from bony front of orbit by 4 times its diameter, nasal sac raised and on anterior one-third of snout. Bony interorbital width 36.0–
|
||
<emphasis id="B976A36D3555A37A6E05F90B1C8FF905" bold="true" box="[854,903,1738,1763]" pageId="2" pageNumber="561">39.5</emphasis>
|
||
% (n = 2) pupil diameter; moderate U-shaped interorbital trench and narrow, groove-like postorbital slit; epaxialis extending anteriorly to point above middle of pupil (
|
||
<figureCitation id="133963FA3555A37A6DB3F8D31E2BF8CD" box="[224,291,1810,1835]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1957]" captionTargetBox="[176,1402,193,1887]" captionTargetId="figure@3.[166,1402,193,1888]" captionTargetPageId="3" captionText="FIGURE 1. Trimma bathum, 16.4 mm SL male holotype, Suva, Fiji. Lateral (A, image reversed) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin yellow lines. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/582789/files/figure.png" pageId="2" pageNumber="561">Fig. 1</figureCitation>
|
||
B); narrow ridge of skin from origin of first dorsal fin extending along anterior midline of nape halfway to orbit. Caudal peduncle depth as percentage caudal peduncle length
|
||
<emphasis id="B976A36D3555A37A6E84F8F71B00F8A9" bold="true" box="[983,1032,1846,1871]" pageId="2" pageNumber="561">52.3</emphasis>
|
||
–57.4; head length as percentage SL
|
||
<emphasis id="B976A36D3555A37A6DC4F89B1FC0F895" bold="true" box="[151,200,1882,1907]" pageId="2" pageNumber="561">32.1</emphasis>
|
||
–32.9; as percentage head length: horizontal eye diameter 37.4–
|
||
<emphasis id="B976A36D3555A37A6EE0F89B1CECF895" bold="true" box="[947,996,1882,1907]" pageId="2" pageNumber="561">39.4</emphasis>
|
||
; snout length 19.5–
|
||
<emphasis id="B976A36D3555A37A699DF89B1BF7F895" bold="true" box="[1230,1279,1882,1907]" pageId="2" pageNumber="561">22.3</emphasis>
|
||
; cheek depth 20.2–
|
||
<emphasis id="B976A36D3555A37A6D85F8BF1E0FF871" bold="true" box="[214,263,1918,1943]" pageId="2" pageNumber="561">25.1</emphasis>
|
||
. Cephalic sensory papillae as in
|
||
<figureCitation id="133963FA3555A37A6FD6F8BF1DC0F871" box="[645,712,1918,1943]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1957]" captionTargetBox="[176,1402,193,1887]" captionTargetId="figure@3.[166,1402,193,1888]" captionTargetPageId="3" captionText="FIGURE 1. Trimma bathum, 16.4 mm SL male holotype, Suva, Fiji. Lateral (A, image reversed) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin yellow lines. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/582789/files/figure.png" pageId="2" pageNumber="561">Fig. 1</figureCitation>
|
||
, number of papillae in each row given in
|
||
<tableCitation id="C6804AC43555A37A69E2F8BF1A0FF871" box="[1201,1287,1918,1943]" captionStart="TABLE 1" captionStartId="1.[151,239,548,571]" captionText="TABLE 1. Counts of cephalic sensory papillae in the different rows of the two new species of Trimma. Data given as minimum to maximum values, with the mean, Standard Deviation and number of specimens examined respectively given in parentheses, except where the value was invariant, where only the number of specimens is given. Terminology follows that used in Winterbottom et al. (2015)." pageId="2" pageNumber="561">Table 1</tableCitation>
|
||
. Abdominal/ caudal vertebral transition presumed to be
|
||
<typeStatus id="54B9C1DD3555A37A6FD3F8621DB0F85D" box="[640,696,1955,1979]" pageId="2" pageNumber="561">Type</typeStatus>
|
||
B (inferred from the narrow bony interorbital width of less than 80% pupil-diameter).
|
||
</paragraph>
|
||
<caption id="DF7D2FF73554A37B6DC4F84E1E23F804" httpUri="https://zenodo.org/record/582789/files/figure.png" pageId="3" pageNumber="562" targetBox="[176,1402,193,1887]" targetPageId="3">
|
||
<paragraph id="8BBD7F7F3554A37B6DC4F84E1E23F804" blockId="3.[151,1436,1935,2018]" pageId="3" pageNumber="562">
|
||
<emphasis id="B976A36D3554A37B6DC4F84E1E1BF842" bold="true" box="[151,275,1935,1957]" pageId="3" pageNumber="562">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C0204FC3554A37B6C4AF8511EB2F843" box="[281,442,1935,1957]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="562" phylum="Chordata" rank="species" species="bathum">
|
||
<emphasis id="B976A36D3554A37B6C4AF8511EB2F843" box="[281,442,1935,1957]" italics="true" pageId="3" pageNumber="562">Trimma bathum</emphasis>
|
||
</taxonomicName>
|
||
, 16.4 mm SL male holotype, Suva, Fiji. Lateral (A, image reversed) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin yellow lines. Photos by R. Winterbottom.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7D2FF73553A37C6DC4FC9E1A1FFC93" box="[151,1303,863,885]" httpUri="https://zenodo.org/record/582790/files/figure.png" pageId="4" pageNumber="563" targetBox="[193,1394,193,841]" targetPageId="4">
|
||
<paragraph id="8BBD7F7F3553A37C6DC4FC9E1A1FFC93" blockId="4.[151,1303,863,885]" box="[151,1303,863,885]" pageId="4" pageNumber="563">
|
||
<emphasis id="B976A36D3553A37C6DC4FC9E1E1AFC92" bold="true" box="[151,274,863,885]" pageId="4" pageNumber="563">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C0204FC3553A37C6C4AFCA11EB2FC93" box="[281,442,863,885]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="bathum">
|
||
<emphasis id="B976A36D3553A37C6C4AFCA11EB2FC93" box="[281,442,863,885]" italics="true" pageId="4" pageNumber="563">Trimma bathum</emphasis>
|
||
</taxonomicName>
|
||
, 17.6 (UK T4454) & 16.4 (ROM T02301) mm SL, Viti Levu, Fiji. Photo by R. Pyle.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD7F7F3553A37C6D94FC601E2DFB50" blockId="4.[151,1436,929,2037]" pageId="4" pageNumber="563">
|
||
<emphasis id="B976A36D3553A37C6D94FC601D48FC5C" box="[199,576,929,954]" italics="true" pageId="4" pageNumber="563">Colour pattern. Freshly collected</emphasis>
|
||
, based on photograph of two specimens saved as tissues (
|
||
<figureCitation id="133963FA3553A37C6987FC601A13FC5C" box="[1236,1307,929,954]" captionStart="FIGURE 2" captionStartId="4.[151,250,863,885]" captionTargetBox="[193,1394,193,841]" captionTargetId="figure@4.[193,1394,193,841]" captionTargetPageId="4" captionText="FIGURE 2. Trimma bathum, 17.6 (UK T 4454) & 16.4 (ROM T 02301) mm SL, Viti Levu, Fiji. Photo by R. Pyle." httpUri="https://zenodo.org/record/582790/files/figure.png" pageId="4" pageNumber="563">Fig. 2</figureCitation>
|
||
). Head and anterior body pink, fading gradually posteriorly until almost translucent at end of caudal peduncle. Centres of most body scales with orange-brown spot or short bar, spots/bars may join each other, becoming a diffuse, slightly lighter wash dorsally and ventrally on peduncle. Second dorsal fin (and possibly anal fin) with a basal orangebrown stripe, posterior half of fin with similarly-coloured mottling. Caudal fin with ill-defined scattered lighter spots and blotches, about one-third pupil-diameter in size. Iris with narrow white ring around pupil, yellowishblack below pupil, and broad, very diffuse dark, pupil-width oblique stripe from anterodorsal to posteroventral across pupil.
|
||
</paragraph>
|
||
<paragraph id="8BBD7F7F3553A37C6D94FB001CFFFA68" blockId="4.[151,1436,929,2037]" pageId="4" pageNumber="563">
|
||
<emphasis id="B976A36D3553A37C6D94FB001E3FFB3C" box="[199,311,1217,1242]" italics="true" pageId="4" pageNumber="563">Preserved</emphasis>
|
||
.
|
||
<typeStatus id="54B9C1DD3553A37C6C15FB001EA6FB3C" box="[326,430,1217,1242]" pageId="4" pageNumber="563" type="holotype">Holotype</typeStatus>
|
||
straw-yellow and faded, with fairly even scattering of very small melanophores on head and anterior nape (fewer on cheeks), none on ventral surface of head and abdomen. Melanophores also tend be concentrated along anterior margins of scale pockets on body so that they show through at middle of scale in front (note: most body scales have been abraded off specimens), melanophores decreasing in number posteriorly. A few slightly larger melanophores along base of second dorsal fin which may indicate a basal stripe in life.
|
||
<typeStatus id="54B9C1DD3553A37C687CFA931A94FA8C" box="[1327,1436,1362,1386]" pageId="4" pageNumber="563" type="paratype">Paratypes</typeStatus>
|
||
similar, but with few small melanophores on lips and ventral surface of head.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3182CF43553A37C6D94FA581A8EFA30" pageId="4" pageNumber="563" type="etymology">
|
||
<paragraph id="8BBD7F7F3553A37C6D94FA581A8EFA30" blockId="4.[151,1436,929,2037]" pageId="4" pageNumber="563">
|
||
<emphasis id="B976A36D3553A37C6D94FA581E46FA54" bold="true" box="[199,334,1433,1458]" pageId="4" pageNumber="563">Etymology.</emphasis>
|
||
From the Greek word ‘bathos’ (βάΘος), meaning deep or deep water, in reference to the depth at which these specimens were collected (
|
||
<quantity id="4CFAD29A3553A37C6F03FA7C1DB0FA30" box="[592,696,1469,1494]" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="4" pageNumber="563" unit="m" value="70.0" valueMax="90.0" valueMin="50.0">50–90 m</quantity>
|
||
). The name is treated as an adjective in the nominative singular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3182CF43553A37D6D94FA201EA2FDDB" lastPageId="5" lastPageNumber="564" pageId="4" pageNumber="563" type="distribution">
|
||
<paragraph id="8BBD7F7F3553A37C6D94FA201BD7FA1F" blockId="4.[151,1436,929,2037]" box="[199,1247,1505,1530]" pageId="4" pageNumber="563">
|
||
<emphasis id="B976A36D3553A37C6D94FA201E68FA1C" bold="true" box="[199,352,1505,1530]" pageId="4" pageNumber="563">Distribution.</emphasis>
|
||
Currently recorded only from the
|
||
<collectingCountry id="F3153FEF3553A37C6FB1FA201C57FA1C" box="[738,863,1505,1530]" name="Fiji" pageId="4" pageNumber="563">Fiji Islands</collectingCountry>
|
||
in the vicinity of Suva,
|
||
<collectingCountry id="F3153FEF3553A37C693EFA201B9FFA1C" box="[1133,1175,1505,1530]" name="Fiji" pageId="4" pageNumber="563">Viti</collectingCountry>
|
||
Levu.
|
||
</paragraph>
|
||
<paragraph id="8BBD7F7F3553A37D6D94F9C41A99FEFA" blockId="4.[151,1436,929,2037]" lastBlockId="5.[151,1436,151,752]" lastPageId="5" lastPageNumber="564" pageId="4" pageNumber="563">
|
||
<emphasis id="B976A36D3553A37C6D94F9C41E64F9F8" bold="true" box="[199,364,1541,1566]" pageId="4" pageNumber="563">Comparisons.</emphasis>
|
||
The colour pattern combination of a pink head and body with orange-brown spots/short bars in the centres of the scale rows on the body is unique to
|
||
<taxonomicName id="4C0204FC3553A37C6FBFF9EA1C51F9A4" box="[748,857,1577,1602]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="bathum">
|
||
<emphasis id="B976A36D3553A37C6FBFF9EA1C51F9A4" box="[748,857,1577,1602]" italics="true" pageId="4" pageNumber="563">T. bathum</emphasis>
|
||
</taxonomicName>
|
||
. Four other valid species of
|
||
<taxonomicName id="4C0204FC3553A37C69CAF9EA1BF8F9A4" box="[1177,1264,1579,1602]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="genus">
|
||
<emphasis id="B976A36D3553A37C69CAF9EA1BF8F9A4" box="[1177,1264,1579,1602]" italics="true" pageId="4" pageNumber="563">Trimma</emphasis>
|
||
</taxonomicName>
|
||
lack predorsal, cheek and opercular scales, and have unbranched pectoral and 5th pelvic fin rays [
|
||
<taxonomicName id="4C0204FC3553A37C6969F98E1A90F980" authority="Winterbottom, 2000" authorityName="Winterbottom" authorityYear="2000" box="[1082,1432,1613,1638]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="anaima">
|
||
<emphasis id="B976A36D3553A37C6969F98E1BA2F983" box="[1082,1194,1614,1638]" italics="true" pageId="4" pageNumber="563">T. anaima</emphasis>
|
||
<bibRefCitation id="EF93028E3553A37C69E2F98C1A90F980" author="Winterbottom" box="[1201,1432,1613,1638]" pageId="4" pageNumber="563" refString="Winterbottom, R. (2000) Four new species of Trimma (Gobiidae), from the Indian and western Pacific oceans. aqua, Journal of Ichthyology and aquatic Biology, 4 (2), 57 - 66." type="journal article" year="2000">Winterbottom, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C0204FC3553A37C6DC4F9B21D1BF96C" authority="Tomiyama, 1936" authorityName="Tomiyama" authorityYear="1936" box="[151,531,1649,1674]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="grammistes">
|
||
<emphasis id="B976A36D3553A37C6DC4F9B21E3DF96C" box="[151,309,1651,1674]" italics="true" pageId="4" pageNumber="563">T. grammistes</emphasis>
|
||
(
|
||
<bibRefCitation id="EF93028E3553A37C6C15F9B01D03F96C" author="Tomiyama" box="[326,523,1649,1674]" pageId="4" pageNumber="563" refString="Tomiyama, I. (1936) Gobiidae of Japan. Japanese Journal of Zoology, 7, 37 - 112." type="journal article" year="1936">Tomiyama, 1936</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C0204FC3553A37C6F72F9B21B58F96C" authority="Winterbottom & Southcott, 2007" authorityName="Winterbottom & Southcott" authorityYear="2007" box="[545,1104,1649,1674]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="sanguinellus">
|
||
<emphasis id="B976A36D3553A37C6F72F9B21DC3F96C" box="[545,715,1649,1674]" italics="true" pageId="4" pageNumber="563">T. sanguinellus</emphasis>
|
||
<bibRefCitation id="EF93028E3553A37C6F86F9B01B58F96C" author="Winterbottom" box="[725,1104,1649,1674]" pageId="4" pageNumber="563" refString="Winterbottom, R. & Southcott, L. (2007) Two new species of the genus Trimma from western Thailand. aqua, Journal of Ichthyology and aquatic Biology, 13 (2), 69 - 76." type="journal article" year="2007">Winterbottom & Southcott, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C0204FC3553A37C69DDF9B21FDDF948" authority="Winterbottom, 2004" authorityName="Winterbottom" authorityYear="2004" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="sostra">
|
||
<emphasis id="B976A36D3553A37C69DDF9B21BF8F96C" box="[1166,1264,1651,1674]" italics="true" pageId="4" pageNumber="563">T. sostra</emphasis>
|
||
<bibRefCitation id="EF93028E3553A37C69AAF9B01FDDF948" author="Winterbottom" pageId="4" pageNumber="563" refString="Winterbottom, R. (2004) Three new species of Trimma (Pisces; Gobiidae) from the central, western and south Pacific. aqua, Journal of Ichthyology and aquatic Biology, 9 (1), 7 - 16." type="journal article" year="2004">Winterbottom, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
]. This species differs further from
|
||
<taxonomicName id="4C0204FC3553A37C6F3DF9561DEAF94B" box="[622,738,1686,1710]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="anaima">
|
||
<emphasis id="B976A36D3553A37C6F3DF9561DEAF94B" box="[622,738,1686,1710]" italics="true" pageId="4" pageNumber="563">T. anaima</emphasis>
|
||
</taxonomicName>
|
||
in usually having 9 dorsal and anal fin rays (vs. 8), 18–19 pectoral rays (vs. 15–17), 13–14 lower gill rakers (vs. 11–12), a narrower bony interorbital (about 40% pupil-width vs. 60%), a U-shaped trench between the eyes with a narrow slit-like post-orbital trench (vs. both absent), and a dermal fold in the dorsal midline anterior to the first dorsal spine (vs. absent).
|
||
<taxonomicName id="4C0204FC3553A37C697BF8C21A01F8FC" box="[1064,1289,1795,1818]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="grammistes">
|
||
<emphasis id="B976A36D3553A37C697BF8C21A01F8FC" box="[1064,1289,1795,1818]" italics="true" pageId="4" pageNumber="563">Trimma grammistes</emphasis>
|
||
</taxonomicName>
|
||
differs most prominently in colour pattern, with a brownish body (darker above) and a pupil diameter black stripe from the upper jaw through the eye and just above the pectoral fin base which curves gently down to end at the middle of the caudal peduncle and fades into the base of the caudal fin (vs. no such stripe); it also has higher mean values for number of dorsal and anal fin rays (9.8 and 9.5 vs. 8.8 and 8.8 respectively), a longer fifth pelvic fin ray (mean = 80 % length of fourth ray vs. 49.5%), and fewer total gill rakers (13–16 vs. 17–18).
|
||
<taxonomicName id="4C0204FC3553A37C696FF8771A22F828" box="[1084,1322,1973,1998]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="563" phylum="Chordata" rank="species" species="sanguinellus">
|
||
<emphasis id="B976A36D3553A37C696FF8771A22F828" box="[1084,1322,1973,1998]" italics="true" pageId="4" pageNumber="563">Trimma sanguinellus</emphasis>
|
||
</taxonomicName>
|
||
is a plain orange-red species with a somewhat elongate second spine in the first dorsal fin (to bases of 2nd to 4th rays of second dorsal fin vs. base of spine), and 15–17 total gill rakers (vs. 17–18).
|
||
<taxonomicName id="4C0204FC3552A37D6EC2FF581B3BFF56" box="[913,1075,153,176]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="sostra">
|
||
<emphasis id="B976A36D3552A37D6EC2FF581B3BFF56" box="[913,1075,153,176]" italics="true" pageId="5" pageNumber="564">Trimma sostra</emphasis>
|
||
</taxonomicName>
|
||
has a white body with six large, red midlateral blotches on the body with another over the opercular region (vs. blotches absent), a very elongate second spine of the first dorsal fin (vs. spine not elongate), usually (in 30 of
|
||
<specimenCount id="9D04B4F63552A37D6EAEFF211B90FF1E" box="[1021,1176,223,248]" pageId="5" pageNumber="564" type="generic">32 specimens</specimenCount>
|
||
) at least some pectoral fin rays branched (vs. all unbranched) and no fleshy ridge in midline anterior to first dorsal fin (vs. ridge present).
|
||
</paragraph>
|
||
<paragraph id="8BBD7F7F3552A37D6D94FEE61EA2FDDB" blockId="5.[151,1436,151,752]" pageId="5" pageNumber="564">
|
||
The phenetic relationships from an analysis of the COI gene (see below) suggest that
|
||
<taxonomicName id="4C0204FC3552A37D69D4FEE81FC7FE82" authority="Winterbottom, 2001" authorityName="Winterbottom" authorityYear="2001" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="stobbsi">
|
||
<emphasis id="B976A36D3552A37D69D4FEE81BFAFEA6" box="[1159,1266,295,320]" italics="true" pageId="5" pageNumber="564">T. stobbsi</emphasis>
|
||
<bibRefCitation id="EF93028E3552A37D69AAFEE61FC7FE82" author="Winterbottom" pageId="5" pageNumber="564" refString="Winterbottom, R. (2001) Two new gobiid fish species in Trimma and Trimmatom (Teleostei: Gobiidae) from the Indian and Western Pacific Oceans. aqua, Journal of Ichthyology and aquatic Biology, 5 (1), 19 - 24." type="journal article" year="2001">Winterbottom, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0204FC3552A37D6C56FE8C1DABFE83" authority="Winterbottom et al., 2015" authorityName="Winterbottom et al." authorityYear="2015" box="[261,675,332,357]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="kardium">
|
||
<emphasis id="B976A36D3552A37D6C56FE8C1E72FE83" box="[261,378,332,357]" italics="true" pageId="5" pageNumber="564">T. kardium</emphasis>
|
||
<bibRefCitation id="EF93028E3552A37D6CD0FE8D1DABFE83" author="Winterbottom" box="[387,675,332,357]" pageId="5" pageNumber="564" refString="Winterbottom, R., Erdmann, M. V. & Cahyani, N. K D. (2015) New species of Trimma (Actinopterygii, Gobiidae) from Indonesia, with comments on head papillae nomenclature. Zootaxa, 3973 (2), 201 - 226." type="journal article" year="2015">
|
||
Winterbottom
|
||
<emphasis id="B976A36D3552A37D6F75FE8C1D5FFE83" box="[550,599,332,357]" italics="true" pageId="5" pageNumber="564">et al</emphasis>
|
||
., 2015
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
are most similar to the new species. The colour pattern of
|
||
<taxonomicName id="4C0204FC3552A37D6862FE8C1A93FE83" box="[1329,1435,332,357]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="stobbsi">
|
||
<emphasis id="B976A36D3552A37D6862FE8C1A93FE83" box="[1329,1435,332,357]" italics="true" pageId="5" pageNumber="564">T. stobbsi</emphasis>
|
||
</taxonomicName>
|
||
is overall brownish background (vs. pink) with a prominent, half-pupil diameter black spot over the posterodorsal region of the opercle (vs. black spot absent), and branched rays are present (vs. absent) in the pectoral fin.
|
||
<taxonomicName id="4C0204FC3552A37D6816FE541FFBFE36" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="kardium">
|
||
<emphasis id="B976A36D3552A37D6816FE541FFBFE36" italics="true" pageId="5" pageNumber="564">Trimma kardium</emphasis>
|
||
</taxonomicName>
|
||
is overall yellow with a pinkish head, and a prominent, heart-shaped red blotch over the hyoid arch (vs. not as above). In addition, the fifth pelvic fin ray of
|
||
<taxonomicName id="4C0204FC3552A37D6FB8FE1C1C6AFE13" box="[747,866,476,501]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="kardium">
|
||
<emphasis id="B976A36D3552A37D6FB8FE1C1C6AFE13" box="[747,866,476,501]" italics="true" pageId="5" pageNumber="564">T. kardium</emphasis>
|
||
</taxonomicName>
|
||
may be branched (vs. unbranched), and it usually has at least some scales in the predorsal midline (vs. scales absent), and a row of scales on the opercle (vs. opercular scales absent).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3182CF43552A37D6D94FD861D97FD16" pageId="5" pageNumber="564" type="discussion">
|
||
<paragraph id="8BBD7F7F3552A37D6D94FD861D97FD16" blockId="5.[151,1436,151,752]" pageId="5" pageNumber="564">
|
||
<emphasis id="B976A36D3552A37D6D94FD861E44FD86" bold="true" box="[199,332,583,608]" pageId="5" pageNumber="564">Discussion.</emphasis>
|
||
A Neighbour-Joining analysis of available COI data for
|
||
<taxonomicName id="4C0204FC3552A37D6E91FD881B11FD86" box="[962,1049,585,608]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="genus">
|
||
<emphasis id="B976A36D3552A37D6E91FD881B11FD86" box="[962,1049,585,608]" italics="true" pageId="5" pageNumber="564">Trimma</emphasis>
|
||
</taxonomicName>
|
||
places the specimens of
|
||
<taxonomicName id="4C0204FC3552A37D687EFD881A94FD86" box="[1325,1436,583,608]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="bathum">
|
||
<emphasis id="B976A36D3552A37D687EFD881A94FD86" box="[1325,1436,583,608]" italics="true" pageId="5" pageNumber="564">T. bathum</emphasis>
|
||
</taxonomicName>
|
||
sequenced (listed as
|
||
<emphasis id="B976A36D3552A37D6CD7FDAC1E99FD62" box="[388,401,621,644]" italics="true" pageId="5" pageNumber="564">T</emphasis>
|
||
. RW sp 97) as being phenetically closest to
|
||
<taxonomicName id="4C0204FC3552A37D6EC7FDAC1B03FD63" box="[916,1035,620,645]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="kardium">
|
||
<emphasis id="B976A36D3552A37D6EC7FDAC1B03FD63" box="[916,1035,620,645]" italics="true" pageId="5" pageNumber="564">T. kardium</emphasis>
|
||
</taxonomicName>
|
||
(listed as
|
||
<emphasis id="B976A36D3552A37D69D5FDAC1B9CFD62" box="[1158,1172,621,644]" italics="true" pageId="5" pageNumber="564">T</emphasis>
|
||
. RW sp 98), these two species together being phenetically closest to
|
||
<taxonomicName id="4C0204FC3552A37D6FC6FD501DF7FD4E" box="[661,767,655,680]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="stobbsi">
|
||
<emphasis id="B976A36D3552A37D6FC6FD501DF7FD4E" box="[661,767,655,680]" italics="true" pageId="5" pageNumber="564">T. stobbsi</emphasis>
|
||
</taxonomicName>
|
||
(Box J of
|
||
<figureCitation id="133963FA3552A37D6E27FD4E1CBEFD4E" box="[884,950,655,680]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1957]" captionTargetBox="[176,1402,193,1887]" captionTargetId="figure@3.[166,1402,193,1888]" captionTargetPageId="3" captionText="FIGURE 1. Trimma bathum, 16.4 mm SL male holotype, Suva, Fiji. Lateral (A, image reversed) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin yellow lines. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/582789/files/figure.png" pageId="5" pageNumber="564">
|
||
Fig.
|
||
<quantity id="4CFAD29A3552A37D6EFBFD511CBEFD4E" box="[936,950,656,680]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="564" unit="in" value="1.0">1</quantity>
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||
</figureCitation>
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||
in
|
||
<bibRefCitation id="EF93028E3552A37D6E8AFD4E1BF6FD4E" author="Winterbottom" box="[985,1278,655,680]" pageId="5" pageNumber="564" refString="Winterbottom, R., Hanner, R. H., Burridge, M. & Zur, M. (2014) A cornucopia of cryptic species-a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys, 381, 79 - 111." type="journal article" year="2014">
|
||
Winterbottom
|
||
<emphasis id="B976A36D3552A37D692FFD501BA5FD4E" box="[1148,1197,655,680]" italics="true" pageId="5" pageNumber="564">et al</emphasis>
|
||
., 2014
|
||
</bibRefCitation>
|
||
: 85). The new species differs from
|
||
<taxonomicName id="4C0204FC3552A37D6C29FD741EF8FD2B" box="[378,496,692,717]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="kardium">
|
||
<emphasis id="B976A36D3552A37D6C29FD741EF8FD2B" box="[378,496,692,717]" italics="true" pageId="5" pageNumber="564">T. kardium</emphasis>
|
||
</taxonomicName>
|
||
by almost 10% of the COI base pairs, and these two species together differ from the
|
||
<taxonomicName id="4C0204FC3552A37D6DC4FD181E09FD16" box="[151,257,727,752]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="564" phylum="Chordata" rank="species" species="stobbsi">
|
||
<emphasis id="B976A36D3552A37D6DC4FD181E09FD16" box="[151,257,727,752]" italics="true" pageId="5" pageNumber="564">T. stobbsi</emphasis>
|
||
</taxonomicName>
|
||
complex by slightly more than 11%.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |