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<document ID-DOI="10.11646/zootaxa.5263.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7804268" ID-ZooBank="20CB3CA0-BEF9-474C-8931-6A7948B9CA61" approvalRequired="5" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="4" checkinTime="1680760231104" checkinUser="plazi" docAuthor="Řezáč, Milan, Cardoso, Pedro &amp; Řezáčová, Veronika" docDate="2023" docId="039CE028E87BFFC9FF12FD2EFBBF62EB" docLanguage="en" docName="zootaxa.5263.3.2.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpactea gaditana Pesarini 1988" docType="treatment" docVersion="2" lastPageNumber="346" masterDocId="FFA59850E870FFC2FF85FFA1FFD2667D" masterDocTitle="Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal" masterLastPageNumber="364" masterPageNumber="335" pageNumber="346" updateTime="1680760785065" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Řezáč, Milan</mods:namePart>
<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cardoso, Pedro</mods:namePart>
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<mods:affiliation>Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History Luomus, University of Helsinki, P. O. Box 17, FI- 00014 Helsinki, Finland. pedro. cardoso @ helsinki. fi; https: // orcid. org / 0000 - 0001 - 8119 - 9960</mods:affiliation>
<mods:nameIdentifier type="email">pedro.cardoso@helsinki.fi</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Řezáčová, Veronika</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1749-0355</mods:nameIdentifier>
<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia &amp; strver @ seznam. cz; https: // orcid. org / 0000 - 0002 - 1749 - 0355</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-06</mods:number>
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<mods:number>5263</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7804288" ID-Zenodo-Dep="7804288" LSID="urn:lsid:plazi:treatment:039CE028E87BFFC9FF12FD2EFBBF62EB" httpUri="http://treatment.plazi.org/id/039CE028E87BFFC9FF12FD2EFBBF62EB" lastPageNumber="346" pageId="11" pageNumber="346">
<subSubSection box="[151,575,654,681]" pageId="11" pageNumber="346" type="nomenclature">
<paragraph blockId="11.[151,575,654,717]" box="[151,575,654,681]" pageId="11" pageNumber="346">
<heading bold="true" box="[151,575,654,681]" fontSize="11" level="1" pageId="11" pageNumber="346" reason="1">
<taxonomicName ID-CoL="3JRST" authority="Pesarini, 1988" authorityName="Pesarini" authorityYear="1988" box="[151,575,654,681]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="gaditana">
<emphasis bold="true" box="[151,575,654,681]" pageId="11" pageNumber="346">
<emphasis bold="true" box="[151,392,654,681]" italics="true" pageId="11" pageNumber="346">Harpactea gaditana</emphasis>
<bibRefCitation author="Pesarini, C." box="[399,575,654,680]" pageId="11" pageNumber="346" pagination="179 - 184" refId="ref20734" refString="Pesarini, C. (1988) Due nuove specie di Harpactea Bristowe appartenenti alla fauna Iberica. Atti della Societ Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 129, 179 - 184." type="journal article" year="1988">Pesarini, 1988</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,422,691,717]" pageId="11" pageNumber="346" type="description">
<paragraph blockId="11.[151,575,654,717]" box="[151,422,691,717]" pageId="11" pageNumber="346">
(
<figureCitation box="[159,246,691,717]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="11" pageNumber="346">Fig. 1B</figureCitation>
,
<figureCitation box="[259,294,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[250,1344,181,1799]" captionTargetId="figure-24@4.[244,1344,181,1799]" captionTargetPageId="4" captionText="FIGURE 2. Portuguese Harpactea spiders. Right male chelicera.Above anterior (G, J, L, M) or posterior view (A, B, C, D, E, F, H, I, K, N); below median view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804274" httpUri="https://zenodo.org/record/7804274/files/figure.png" pageId="11" pageNumber="346">2B</figureCitation>
,
<figureCitation box="[305,340,691,717]" captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="11" pageNumber="346">3B</figureCitation>
,
<figureCitation box="[352,386,691,717]" captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="11" pageNumber="346">4B</figureCitation>
,
<figureCitation box="[399,414,691,717]" captionStart="FIGURE 9" captionStartId="22.[151,250,1892,1916]" captionTargetBox="[279,1342,195,1818]" captionTargetId="figure-24@22.[231,1356,181,1869]" captionTargetPageId="22" captionText="FIGURE 9. Known distribution of Portuguese Harpactea species. In each map the first species is represented by a transparent circle and the second species by a black diamond:A) H. adicensis &amp; H. algarvensis; B) H. crespoi &amp; H. dolanskyi; C) H. fageli &amp; H. gaditana; D) H. henriquesi &amp; H. hombergii; E) H. korenkoi &amp; H. krejcii; F) H. magnibulbi &amp; H. minoccii; G) H. pekari &amp; H. proxima; H) H. stalitoides &amp; H. subiasi; I) H. tavirensis." figureDoi="http://doi.org/10.5281/zenodo.7804296" httpUri="https://zenodo.org/record/7804296/files/figure.png" pageId="11" pageNumber="346">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="346" type="reference_group">
<paragraph blockId="11.[151,1437,759,814]" pageId="11" pageNumber="346">
<treatmentCitationGroup pageId="11" pageNumber="346">
<treatmentCitation author="Pesarini, C." box="[151,480,759,783]" page="180" pageId="11" pageNumber="346" year="1988">
<taxonomicName ID-CoL="3JRST" authority="Pesarini 1988: 180" authorityName="Pesarini" authorityPageNumber="180" authorityYear="1988" box="[151,480,759,783]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="gaditana">
<emphasis box="[151,275,759,783]" italics="true" pageId="11" pageNumber="346">H. gaditana</emphasis>
<bibRefCitation author="Pesarini, C." box="[284,480,759,783]" pageId="11" pageNumber="346" pagination="179 - 184" refId="ref20734" refString="Pesarini, C. (1988) Due nuove specie di Harpactea Bristowe appartenenti alla fauna Iberica. Atti della Societ Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 129, 179 - 184." type="journal article" year="1988">Pesarini 1988: 180</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 2 (description based on male);
<treatmentCitation author="Le Peru, B." box="[862,1059,759,783]" page="269" pageId="11" pageNumber="346" year="2011">
<bibRefCitation author="Le Peru, B." box="[862,1059,759,783]" pageId="11" pageNumber="346" pagination="1 - 522" refId="ref20332" refString="Le Peru, B. (2011) The spiders of Europe, a synthesis of data: Volume 1 Atypidae to Theridiidae. Memoires de la Societe Linneenne de Lyon, 2, 1 - 522." type="journal article" year="2011">
Le
<collectingCountry box="[898,944,760,783]" name="Peru" pageId="11" pageNumber="346">Peru</collectingCountry>
2011: 269
</bibRefCitation>
</treatmentCitation>
, fig. 402 (♁, redrawn from
<bibRefCitation author="Pesarini, C." pageId="11" pageNumber="346" pagination="179 - 184" refId="ref20734" refString="Pesarini, C. (1988) Due nuove specie di Harpactea Bristowe appartenenti alla fauna Iberica. Atti della Societ Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 129, 179 - 184." type="journal article" year="1988">Pesarini 1988</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="346" type="materials_examined">
<paragraph blockId="11.[151,1436,860,1175]" box="[151,640,860,887]" pageId="11" pageNumber="346">
<emphasis bold="true" box="[151,262,860,886]" pageId="11" pageNumber="346">Material.</emphasis>
<typeStatus box="[269,325,861,887]" pageId="11" pageNumber="346">Type</typeStatus>
material was not examined.
</paragraph>
<paragraph blockId="11.[151,1436,860,1175]" pageId="11" pageNumber="346">
<emphasis box="[199,358,896,922]" italics="true" pageId="11" pageNumber="346">New material.</emphasis>
<materialsCitation collectingDate="2008-06-03" collectorName="S. Korenko &amp; Crop Research Institute" latitude="40.117" location="Prague" longLatPrecision="69" longitude="-7.484" pageId="11" pageNumber="346" specimenCount="1" stateProvince="Praha">
Fundao, Alcongosta,
<geoCoordinate box="[602,681,896,922]" degrees="40.117" direction="north" orientation="latitude" pageId="11" pageNumber="346" precision="55" value="40.117">40.117</geoCoordinate>
,
<geoCoordinate box="[692,766,896,922]" degrees="7.484" direction="west" orientation="longitude" pageId="11" pageNumber="346" precision="55" value="-7.484">-7.484</geoCoordinate>
, inside house, 1 ♁,
<date box="[986,1124,896,923]" pageId="11" pageNumber="346" value="2008-06-03">
<collectingDate box="[986,1124,896,923]" pageId="11" pageNumber="346" value="2008-06-03">3 June 2008</collectingDate>
</date>
, leg.
<collectorName box="[1183,1314,896,922]" pageId="11" pageNumber="346">S. Korenko</collectorName>
, coll.
<collectorName pageId="11" pageNumber="346">Crop Research Institute</collectorName>
,
<collectingRegion box="[367,448,933,959]" country="Czech Republic" name="Praha" pageId="11" pageNumber="346">Prague</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="346" type="diagnosis">
<paragraph blockId="11.[151,1436,860,1175]" pageId="11" pageNumber="346">
<emphasis bold="true" box="[199,322,968,994]" pageId="11" pageNumber="346">Diagnosis.</emphasis>
<taxonomicName authorityName="Pesarini" authorityYear="1988" box="[332,563,968,994]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="gaditana">
<emphasis box="[332,563,968,994]" italics="true" pageId="11" pageNumber="346">Harpactea gaditana</emphasis>
</taxonomicName>
is very similar to
<taxonomicName authority="Řezáč &amp; Cardoso &amp; Řezáčová, 2023" authorityName="Řezáč &amp; Cardoso &amp; Řezáčová" authorityYear="2023" box="[784,940,968,994]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="henriquesi" status="sp. nov.">
<emphasis box="[784,940,968,994]" italics="true" pageId="11" pageNumber="346">H. henriquesi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[951,1045,968,994]" pageId="11" pageNumber="346">
<taxonomicNameLabel box="[951,1045,968,994]" pageId="11" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in thin body with relatively long legs and genitalic morphology. It differs from
<taxonomicName authority="Řezáč &amp; Cardoso &amp; Řezáčová, 2023" authorityName="Řezáč &amp; Cardoso &amp; Řezáčová" authorityYear="2023" box="[672,825,1004,1030]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="henriquesi" status="sp. nov.">
<emphasis box="[672,825,1004,1030]" italics="true" pageId="11" pageNumber="346">H. henriquesi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[832,922,1004,1030]" pageId="11" pageNumber="346">
<taxonomicNameLabel box="[832,922,1004,1030]" pageId="11" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by slightly larger body (
<figureCitation box="[1201,1288,1004,1030]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="11" pageNumber="346">Fig. 1B</figureCitation>
), less curved embolus (
<figureCitation box="[261,391,1040,1066]" captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="11" pageNumber="346">Fig. 4B—e</figureCitation>
), the base of embolus is not covered by the base of conductor.
</paragraph>
</subSubSection>
<subSubSection box="[199,810,1076,1103]" pageId="11" pageNumber="346" type="biology_ecology">
<paragraph blockId="11.[151,1436,860,1175]" box="[199,810,1076,1103]" pageId="11" pageNumber="346">
<emphasis bold="true" box="[199,300,1076,1102]" pageId="11" pageNumber="346">Ecology.</emphasis>
Adults can be found in spring (MarchJune).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="346" type="distribution">
<paragraph blockId="11.[151,1436,860,1175]" pageId="11" pageNumber="346">
<emphasis bold="true" box="[199,353,1112,1138]" pageId="11" pageNumber="346">Distribution.</emphasis>
It occurs in southern
<collectingCountry box="[604,668,1112,1138]" name="Spain" pageId="11" pageNumber="346">Spain</collectingCountry>
(near the town Vejer in Cadiz province,
<bibRefCitation author="Pesarini, C." box="[1135,1291,1112,1138]" pageId="11" pageNumber="346" pagination="179 - 184" refId="ref20734" refString="Pesarini, C. (1988) Due nuove specie di Harpactea Bristowe appartenenti alla fauna Iberica. Atti della Societ Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 129, 179 - 184." type="journal article" year="1988">Pesarini 1988</bibRefCitation>
) and eastern
<collectingCountry box="[151,250,1148,1174]" name="Portugal" pageId="11" pageNumber="346">Portugal</collectingCountry>
. In western
<collectingCountry box="[384,479,1148,1174]" name="Portugal" pageId="11" pageNumber="346">Portugal</collectingCountry>
it is replaced by the sibling species
<taxonomicName authority="Řezáč &amp; Cardoso &amp; Řezáčová, 2023" authorityName="Řezáč &amp; Cardoso &amp; Řezáčová" authorityYear="2023" box="[884,1036,1148,1174]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="henriquesi" status="sp. nov.">
<emphasis box="[884,1036,1148,1174]" italics="true" pageId="11" pageNumber="346">H. henriquesi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1043,1133,1148,1174]" pageId="11" pageNumber="346">
<taxonomicNameLabel box="[1043,1133,1148,1174]" pageId="11" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>