293 lines
32 KiB
XML
293 lines
32 KiB
XML
<document ID-DOI="10.11646/zootaxa.5263.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7804268" ID-ZooBank="20CB3CA0-BEF9-474C-8931-6A7948B9CA61" approvalRequired="5" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="4" checkinTime="1680760231104" checkinUser="plazi" docAuthor="Řezáč, Milan, Cardoso, Pedro & Řezáčová, Veronika" docDate="2023" docId="039CE028E879FFCBFF12FF36FAF46205" docLanguage="en" docName="zootaxa.5263.3.2.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpactea crespoi Rezac 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="344" masterDocId="FFA59850E870FFC2FF85FFA1FFD2667D" masterDocTitle="Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal" masterLastPageNumber="364" masterPageNumber="335" pageNumber="344" updateTime="1680760785065" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Řezáč, Milan</mods:namePart>
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<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cardoso, Pedro</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-8119-9960</mods:nameIdentifier>
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<mods:affiliation>Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History Luomus, University of Helsinki, P. O. Box 17, FI- 00014 Helsinki, Finland. pedro. cardoso @ helsinki. fi; https: // orcid. org / 0000 - 0001 - 8119 - 9960</mods:affiliation>
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<mods:nameIdentifier type="email">pedro.cardoso@helsinki.fi</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Řezáčová, Veronika</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-1749-0355</mods:nameIdentifier>
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<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia & strver @ seznam. cz; https: // orcid. org / 0000 - 0002 - 1749 - 0355</mods:affiliation>
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<mods:nameIdentifier type="email">strver@seznam.cz</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5263</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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<mods:extent unit="page">
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<mods:start>335</mods:start>
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<mods:end>364</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.3.2</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5263.3.2</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7804268</mods:identifier>
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<mods:identifier type="ZooBank">20CB3CA0-BEF9-474C-8931-6A7948B9CA61</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7804280" ID-Zenodo-Dep="7804280" LSID="urn:lsid:plazi:treatment:039CE028E879FFCBFF12FF36FAF46205" httpUri="http://treatment.plazi.org/id/039CE028E879FFCBFF12FF36FAF46205" lastPageNumber="344" pageId="9" pageNumber="344">
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<subSubSection box="[151,555,151,181]" pageId="9" pageNumber="344" type="nomenclature">
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<heading bold="true" box="[151,555,151,181]" fontSize="11" level="1" pageId="9" pageNumber="344" reason="1">
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<emphasis bold="true" box="[151,555,151,181]" pageId="9" pageNumber="344">
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<taxonomicName authority="Rezac" authorityName="Rezac" authorityYear="2023" box="[151,454,151,181]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="crespoi" status="sp. nov.">
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<emphasis bold="true" box="[151,372,151,177]" italics="true" pageId="9" pageNumber="344">Harpactea crespoi</emphasis>
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Řezáč
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</taxonomicName>
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<taxonomicNameLabel box="[462,555,151,177]" pageId="9" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,472,187,214]" pageId="9" pageNumber="344" type="description">
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<paragraph blockId="9.[151,555,151,214]" box="[151,472,187,214]" pageId="9" pageNumber="344">
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(
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<figureCitation box="[159,252,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="9" pageNumber="344">Figs 1N</figureCitation>
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,
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<figureCitation box="[264,300,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[250,1344,181,1799]" captionTargetId="figure-24@4.[244,1344,181,1799]" captionTargetPageId="4" captionText="FIGURE 2. Portuguese Harpactea spiders. Right male chelicera.Above anterior (G, J, L, M) or posterior view (A, B, C, D, E, F, H, I, K, N); below median view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804274" httpUri="https://zenodo.org/record/7804274/files/figure.png" pageId="9" pageNumber="344">2N</figureCitation>
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,
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<figureCitation box="[312,348,187,213]" captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="9" pageNumber="344">3N</figureCitation>
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,
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<figureCitation box="[360,390,187,214]" captionStart="FIGURE 5" captionStartId="10.[151,250,1545,1569]" captionTargetBox="[151,1436,236,1521]" captionTargetId="figure-42@10.[151,1436,236,1521]" captionTargetPageId="10" captionText="FIGURE 5. Portuguese Harpactea spiders. Right male bulbus. A. H. adicensis sp. nov. Portugal: Sobral da Adiça; B. H. algarvensis. Portugal: Barranco do Velho; C. H. krejcii sp. nov. Portugal:Ameixial; D. H. dolanskyi sp. nov. Portugal: Mertola; E. H. tavirensis. Portugal: Barranco do Velho; F. H. crespoi sp. nov. Portugal: Sobral da Adiça. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804284" httpUri="https://zenodo.org/record/7804284/files/figure.png" pageId="9" pageNumber="344">5F</figureCitation>
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,
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<figureCitation box="[402,436,187,213]" captionStart="FIGURE 8" captionStartId="16.[151,250,1822,1846]" captionTargetBox="[169,1418,182,1795]" captionTargetId="figure-24@16.[169,1418,181,1798]" captionTargetPageId="16" captionText="FIGURE 8. Portuguese Harpactea spiders. Vulva. A. H. magnibulbi. Portugal: Fóia; B. H. korenkoi sp. nov. Portugal: Carrapateira; C. H. crespoi sp. nov. Portugal: Sobral da Adiça; D. H. algarvensis. Portugal: Barranco do Velho; E. H. tavirensis. Portugal: Barranco do Velho. Views: dor.—dorsal, fro.—anterior, hin.—posterior, ven.—ventral. Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue." figureDoi="http://doi.org/10.5281/zenodo.7804292" httpUri="https://zenodo.org/record/7804292/files/figure.png" pageId="9" pageNumber="344">8C</figureCitation>
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,
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<figureCitation box="[448,463,187,213]" captionStart="FIGURE 9" captionStartId="22.[151,250,1892,1916]" captionTargetBox="[279,1342,195,1818]" captionTargetId="figure-24@22.[231,1356,181,1869]" captionTargetPageId="22" captionText="FIGURE 9. Known distribution of Portuguese Harpactea species. In each map the first species is represented by a transparent circle and the second species by a black diamond:A) H. adicensis & H. algarvensis; B) H. crespoi & H. dolanskyi; C) H. fageli & H. gaditana; D) H. henriquesi & H. hombergii; E) H. korenkoi & H. krejcii; F) H. magnibulbi & H. minoccii; G) H. pekari & H. proxima; H) H. stalitoides & H. subiasi; I) H. tavirensis." figureDoi="http://doi.org/10.5281/zenodo.7804296" httpUri="https://zenodo.org/record/7804296/files/figure.png" pageId="9" pageNumber="344">9</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection pageId="9" pageNumber="344" type="materials_examined">
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<paragraph blockId="9.[151,1437,253,1144]" pageId="9" pageNumber="344">
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<emphasis bold="true" box="[151,325,253,280]" pageId="9" pageNumber="344">Type material.</emphasis>
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<materialsCitation collectingDate="2005-12-27" collectorName="da Adica & L. Crespo & National Natural History Museum" latitude="37.978" location="Serra da Adica" longLatPrecision="70" longitude="-7.294" municipality="Moura" pageId="9" pageNumber="344" specimenCount="1" stateProvince="Praha" typeStatus="holotype">
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<emphasis box="[332,442,254,280]" italics="true" pageId="9" pageNumber="344">
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<typeStatus box="[332,438,254,280]" pageId="9" pageNumber="344">Holotype</typeStatus>
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.
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</emphasis>
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♁,
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<collectingMunicipality box="[483,561,254,280]" pageId="9" pageNumber="344">Moura</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:039CE028E879FFCBFF12FF36FAF46205:8EEA07E5E879FFCBFDB9FF5FFD366765" box="[572,740,254,280]" latitude="37.978" longLatPrecision="70" longitude="-7.294" municipality="Moura" name="Serra da Adica" pageId="9" pageNumber="344" stateProvince="Praha">
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||
Serra
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<collectorName box="[638,740,254,280]" pageId="9" pageNumber="344">da Adiça</collectorName>
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</location>
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,
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<geoCoordinate box="[751,831,254,280]" degrees="37.978" direction="north" orientation="latitude" pageId="9" pageNumber="344" precision="55" value="37.978">37.978</geoCoordinate>
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,
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<geoCoordinate box="[842,916,254,280]" degrees="7.294" direction="west" orientation="longitude" pageId="9" pageNumber="344" precision="55" value="-7.294">-7.294</geoCoordinate>
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,
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<taxonomicName authorityName="Brignoli" authorityYear="1984" box="[928,1133,254,280]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="344" phylum="Tracheophyta" rank="species" species="coccifera">
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<emphasis box="[928,1133,254,280]" italics="true" pageId="9" pageNumber="344">Quercus coccifera</emphasis>
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</taxonomicName>
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wood,
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<date box="[1216,1432,254,281]" pageId="9" pageNumber="344" value="2005-12-27">
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<collectingDate box="[1216,1432,254,281]" pageId="9" pageNumber="344" value="2005-12-27">27 December 2005</collectingDate>
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</date>
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, leg.
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<collectorName box="[199,313,290,316]" pageId="9" pageNumber="344">L. Crespo</collectorName>
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, coll.
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<collectorName box="[380,766,290,316]" pageId="9" pageNumber="344">National Natural History Museum</collectorName>
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,
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<collectingRegion box="[777,858,290,316]" country="Czech Republic" name="Praha" pageId="9" pageNumber="344">Prague</collectingRegion>
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, code P6A 7385.
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</paragraph>
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<paragraph blockId="9.[151,1437,253,1144]" pageId="9" pageNumber="344">
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<materialsCitation box="[199,1108,326,352]" collectorName="Moura" latitude="37.978" location="Serra da Adica" longLatPrecision="70" longitude="-7.294" pageId="9" pageNumber="344" specimenCount="1" typeStatus="paratype">
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<emphasis box="[199,320,327,352]" italics="true" pageId="9" pageNumber="344">
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<typeStatus box="[199,316,327,352]" pageId="9" pageNumber="344">Paratypes</typeStatus>
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.
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</emphasis>
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<collectorName box="[330,408,326,352]" pageId="9" pageNumber="344">Moura</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:039CE028E879FFCBFF12FF36FAF46205:8EEA07E5E879FFCBFE23FEE7FD87671D" box="[422,597,326,352]" latitude="37.978" longLatPrecision="70" longitude="-7.294" name="Serra da Adica" pageId="9" pageNumber="344">Serra da Adiça</location>
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,
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,
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<geoCoordinate box="[706,780,326,352]" degrees="7.294" direction="west" orientation="longitude" pageId="9" pageNumber="344" precision="55" value="-7.294">-7.294</geoCoordinate>
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,
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<taxonomicName authorityName="Brignoli" authorityYear="1984" box="[796,1004,326,352]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="344" phylum="Tracheophyta" rank="species" species="coccifera">
|
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<emphasis box="[796,1004,326,352]" italics="true" pageId="9" pageNumber="344">Quercus coccifera</emphasis>
|
||
</taxonomicName>
|
||
wood, 1
|
||
</materialsCitation>
|
||
♁,
|
||
<materialsCitation collectingDate="2005-11-07" collectorName="M. Rezac & Crop Research Institute" location="Prague" pageId="9" pageNumber="344" specimenCount="1" specimenCount-female="1" stateProvince="Praha" typeStatus="paratype">
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<specimenCount box="[1157,1203,326,352]" pageId="9" pageNumber="344" type="female">1 ♀</specimenCount>
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,
|
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<date box="[1220,1432,326,353]" pageId="9" pageNumber="344" value="2005-11-07">
|
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<collectingDate box="[1220,1432,326,353]" pageId="9" pageNumber="344" value="2005-11-07">7 November 2005</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[200,311,362,390]" pageId="9" pageNumber="344">M. Řezáč</collectorName>
|
||
, coll.
|
||
<collectorName box="[379,648,362,388]" pageId="9" pageNumber="344">Crop Research Institute</collectorName>
|
||
,
|
||
<collectingRegion box="[659,740,362,388]" country="Czech Republic" name="Praha" pageId="9" pageNumber="344">Prague</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2005-12-27" collectorName="L. Crespo & Crop Research Institute" location="Prague" pageId="9" pageNumber="344" specimenCount="1" specimenCount-female="1" stateProvince="Praha" typeStatus="paratype">
|
||
<specimenCount box="[753,795,362,388]" pageId="9" pageNumber="344" type="female">1 ♀</specimenCount>
|
||
,
|
||
<date box="[810,1027,362,389]" pageId="9" pageNumber="344" value="2005-12-27">
|
||
<collectingDate box="[810,1027,362,389]" pageId="9" pageNumber="344" value="2005-12-27">27 December 2005</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[1088,1201,362,388]" pageId="9" pageNumber="344">L. Crespo</collectorName>
|
||
, coll.
|
||
<collectorName pageId="9" pageNumber="344">Crop Research Institute</collectorName>
|
||
,
|
||
<collectingRegion box="[257,338,398,424]" country="Czech Republic" name="Praha" pageId="9" pageNumber="344">Prague</collectingRegion>
|
||
</materialsCitation>
|
||
; 6 ♁♁,
|
||
<materialsCitation box="[427,1288,398,426]" collectingDate="2013-03-27" collectorName="M. Rezac & Crop Research Institute" location="Prague" pageId="9" pageNumber="344" specimenCount="13" specimenCount-female="13" stateProvince="Praha" typeStatus="paratype">
|
||
<specimenCount box="[427,505,398,424]" pageId="9" pageNumber="344" type="female">13 ♀♀</specimenCount>
|
||
,
|
||
<date box="[518,692,398,424]" pageId="9" pageNumber="344" value="2013-03-27">
|
||
<collectingDate box="[518,692,398,424]" pageId="9" pageNumber="344" value="2013-03-27">27 March 2013</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[751,861,398,426]" pageId="9" pageNumber="344">M. Řezáč</collectorName>
|
||
, coll.
|
||
<collectorName box="[928,1196,398,424]" pageId="9" pageNumber="344">Crop Research Institute</collectorName>
|
||
,
|
||
<collectingRegion box="[1207,1288,398,424]" country="Czech Republic" name="Praha" pageId="9" pageNumber="344">Prague</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="344" type="etymology">
|
||
<paragraph blockId="9.[151,1437,253,1144]" pageId="9" pageNumber="344">
|
||
<emphasis bold="true" box="[199,334,434,460]" pageId="9" pageNumber="344">Etymology.</emphasis>
|
||
Named after the Portuguese arachnologist Luis Crespo, our friend, who helped to discover this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="344" type="diagnosis">
|
||
<paragraph blockId="9.[151,1437,253,1144]" pageId="9" pageNumber="344">
|
||
<emphasis bold="true" box="[199,322,505,531]" pageId="9" pageNumber="344">Diagnosis.</emphasis>
|
||
The species is characteristic by the cymbium bent around tegulum (
|
||
<figureCitation box="[1102,1194,506,532]" captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="9" pageNumber="344">Fig. 3N</figureCitation>
|
||
), tegulum wider than long and extremely long embolus (
|
||
<figureCitation box="[535,620,542,569]" captionStart="FIGURE 5" captionStartId="10.[151,250,1545,1569]" captionTargetBox="[151,1436,236,1521]" captionTargetId="figure-42@10.[151,1436,236,1521]" captionTargetPageId="10" captionText="FIGURE 5. Portuguese Harpactea spiders. Right male bulbus. A. H. adicensis sp. nov. Portugal: Sobral da Adiça; B. H. algarvensis. Portugal: Barranco do Velho; C. H. krejcii sp. nov. Portugal:Ameixial; D. H. dolanskyi sp. nov. Portugal: Mertola; E. H. tavirensis. Portugal: Barranco do Velho; F. H. crespoi sp. nov. Portugal: Sobral da Adiça. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804284" httpUri="https://zenodo.org/record/7804284/files/figure.png" pageId="9" pageNumber="344">Fig. 5F</figureCitation>
|
||
). The embolus is very long also in
|
||
<taxonomicName authority="Řezáč & Cardoso & Řezáčová, 2023" authorityName="Řezáč & Cardoso & Řezáčová" authorityYear="2023" box="[1002,1142,542,568]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="dolanskyi" status="sp. nov.">
|
||
<emphasis box="[1002,1142,542,568]" italics="true" pageId="9" pageNumber="344">H. dolanskyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1148,1236,542,568]" pageId="9" pageNumber="344">
|
||
<taxonomicNameLabel box="[1148,1236,542,568]" pageId="9" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authorityName="Wunderlich" authorityYear="2020" box="[1288,1429,543,568]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="tavirensis">
|
||
<emphasis box="[1288,1429,543,568]" italics="true" pageId="9" pageNumber="344">H. tavirensis</emphasis>
|
||
</taxonomicName>
|
||
, but these species possess conductor, in
|
||
<taxonomicName authority="Řezáč & Cardoso & Řezáčová, 2023" authorityName="Řezáč & Cardoso & Řezáčová" authorityYear="2023" box="[586,701,579,604]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="crespoi" status="sp. nov.">
|
||
<emphasis box="[586,701,579,604]" italics="true" pageId="9" pageNumber="344">H. crespoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[708,796,578,604]" pageId="9" pageNumber="344">
|
||
<taxonomicNameLabel box="[708,796,578,604]" pageId="9" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it is absent. The vulva is similar to that of
|
||
<taxonomicName authorityName="Ferrandez" authorityYear="1990" box="[1267,1429,578,604]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="algarvensis">
|
||
<emphasis box="[1267,1429,578,604]" italics="true" pageId="9" pageNumber="344">H. algarvensis</emphasis>
|
||
</taxonomicName>
|
||
, but it lacks two pockets of whitish tissue in front of the anterior arc, and is less sclerotised, the only obviously sclerotised part of the anterior arc is the median rod (
|
||
<figureCitation box="[744,833,650,676]" captionStart="FIGURE 8" captionStartId="16.[151,250,1822,1846]" captionTargetBox="[169,1418,182,1795]" captionTargetId="figure-24@16.[169,1418,181,1798]" captionTargetPageId="16" captionText="FIGURE 8. Portuguese Harpactea spiders. Vulva. A. H. magnibulbi. Portugal: Fóia; B. H. korenkoi sp. nov. Portugal: Carrapateira; C. H. crespoi sp. nov. Portugal: Sobral da Adiça; D. H. algarvensis. Portugal: Barranco do Velho; E. H. tavirensis. Portugal: Barranco do Velho. Views: dor.—dorsal, fro.—anterior, hin.—posterior, ven.—ventral. Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue." figureDoi="http://doi.org/10.5281/zenodo.7804292" httpUri="https://zenodo.org/record/7804292/files/figure.png" pageId="9" pageNumber="344">Fig. 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="344" type="description">
|
||
<paragraph blockId="9.[151,1437,253,1144]" pageId="9" pageNumber="344">
|
||
<emphasis bold="true" box="[199,345,685,711]" pageId="9" pageNumber="344">Description.</emphasis>
|
||
<emphasis box="[358,551,686,712]" italics="true" pageId="9" pageNumber="344">
|
||
Male (
|
||
<typeStatus box="[437,539,686,712]" pageId="9" pageNumber="344">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Carapace pale brown, matting (
|
||
<figureCitation box="[939,1034,686,712]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="9" pageNumber="344">Fig. 1N</figureCitation>
|
||
). Sternum brown-yellow, matting. Chelicerae pale brown. Legs are pale yellow, pedipalps darker. For measurements and leg spination see the
|
||
<tableCitation captionStart="TABLE 1" captionStartId="17.[152,244,151,175]" captionText="TABLE 1. Measurements and spination of holotypes of the new Portuguese Harpactea species and Harpactea tavirensis." pageId="9" pageNumber="344">Table 1</tableCitation>
|
||
. Relative leg length: IV>I>II>III. Cymbium slightly bent around the bulbus, with elongated distal part (
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="9" pageNumber="344">Fig. 3N</figureCitation>
|
||
), and slightly concave basal half of the prolateral side. Tegulum wider than long (
|
||
<figureCitation box="[1123,1210,794,821]" captionStart="FIGURE 5" captionStartId="10.[151,250,1545,1569]" captionTargetBox="[151,1436,236,1521]" captionTargetId="figure-42@10.[151,1436,236,1521]" captionTargetPageId="10" captionText="FIGURE 5. Portuguese Harpactea spiders. Right male bulbus. A. H. adicensis sp. nov. Portugal: Sobral da Adiça; B. H. algarvensis. Portugal: Barranco do Velho; C. H. krejcii sp. nov. Portugal:Ameixial; D. H. dolanskyi sp. nov. Portugal: Mertola; E. H. tavirensis. Portugal: Barranco do Velho; F. H. crespoi sp. nov. Portugal: Sobral da Adiça. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804284" httpUri="https://zenodo.org/record/7804284/files/figure.png" pageId="9" pageNumber="344">Fig. 5F</figureCitation>
|
||
). Conductor absent. Embolus dark, extremely long, slightly undulated, directing to the side, the base is bent in 180° angle. Opisthosoma cylindrical, whitish.
|
||
</paragraph>
|
||
<paragraph blockId="9.[151,1437,253,1144]" pageId="9" pageNumber="344">
|
||
<emphasis box="[199,427,902,928]" italics="true" pageId="9" pageNumber="344">
|
||
Female (
|
||
<typeStatus box="[298,414,902,928]" pageId="9" pageNumber="344">paratypes</typeStatus>
|
||
).
|
||
</emphasis>
|
||
All somatic characters as described for the male. The vulva is of the type
|
||
<taxonomicName authorityName="Ferrandez" authorityYear="1990" box="[1255,1384,902,928]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="algarvensis">
|
||
<emphasis box="[1255,1384,902,928]" italics="true" pageId="9" pageNumber="344">algarvensis</emphasis>
|
||
</taxonomicName>
|
||
(see Materials and Methods) (
|
||
<figureCitation box="[432,515,938,964]" captionStart="FIGURE 8" captionStartId="16.[151,250,1822,1846]" captionTargetBox="[169,1418,182,1795]" captionTargetId="figure-24@16.[169,1418,181,1798]" captionTargetPageId="16" captionText="FIGURE 8. Portuguese Harpactea spiders. Vulva. A. H. magnibulbi. Portugal: Fóia; B. H. korenkoi sp. nov. Portugal: Carrapateira; C. H. crespoi sp. nov. Portugal: Sobral da Adiça; D. H. algarvensis. Portugal: Barranco do Velho; E. H. tavirensis. Portugal: Barranco do Velho. Views: dor.—dorsal, fro.—anterior, hin.—posterior, ven.—ventral. Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue." figureDoi="http://doi.org/10.5281/zenodo.7804292" httpUri="https://zenodo.org/record/7804292/files/figure.png" pageId="9" pageNumber="344">Fig. 8C</figureCitation>
|
||
).The vulva is relatively small and only slightly sclerotised. The median rod is very weak. The posterior diverticle is a rudimentary spherical sclerotised spot.
|
||
</paragraph>
|
||
<paragraph blockId="9.[151,1437,253,1144]" pageId="9" pageNumber="344">
|
||
<emphasis bold="true" box="[199,329,1009,1036]" pageId="9" pageNumber="344">Size range</emphasis>
|
||
. Male carapace length
|
||
<quantity box="[617,759,1010,1036]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="9" pageNumber="344" unit="mm" value="1.95" valueMax="2.3" valueMin="1.6">1.6–2.3 mm</quantity>
|
||
(1.9±0.18, N=8), female carapace length
|
||
<quantity box="[1267,1437,1010,1037]" metricMagnitude="-3" metricUnit="m" metricValue="1.695" metricValueMax="1.86" metricValueMin="1.53" pageId="9" pageNumber="344" unit="mm" value="1.695" valueMax="1.86" valueMin="1.53">1.53–1.86 mm</quantity>
|
||
(1.72±0.17, N=3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1410,1082,1108]" pageId="9" pageNumber="344" type="biology_ecology">
|
||
<paragraph blockId="9.[151,1437,253,1144]" box="[199,1410,1082,1108]" pageId="9" pageNumber="344">
|
||
<emphasis bold="true" box="[199,300,1082,1108]" pageId="9" pageNumber="344">Ecology.</emphasis>
|
||
Adults were found in early spring in leaf litter of
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1984" box="[859,1063,1082,1108]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="344" phylum="Tracheophyta" rank="species" species="coccifera">
|
||
<emphasis box="[859,1063,1082,1108]" italics="true" pageId="9" pageNumber="344">Quercus coccifera</emphasis>
|
||
</taxonomicName>
|
||
forests on northeastern slopes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1318,1117,1144]" pageId="9" pageNumber="344" type="distribution">
|
||
<paragraph blockId="9.[151,1437,253,1144]" box="[199,1318,1117,1144]" pageId="9" pageNumber="344">
|
||
<emphasis bold="true" box="[199,353,1117,1143]" pageId="9" pageNumber="344">Distribution.</emphasis>
|
||
So far known only from Serra da Adiça, a small mountain range in southern
|
||
<collectingCountry box="[1216,1315,1118,1144]" name="Portugal" pageId="9" pageNumber="344">Portugal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |