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<document id="5CB826E65E671FC7287DA43665FDD133" ID-DOI="10.1649/0010-065X-67.3.335" ID-ISSN="1938-4394" ID-Zenodo-Dep="10100754" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630003365932" checkinUser="diego" docAuthor="Park, J. - S. &amp; Carlton, C. E." docDate="2013" docId="03933A62FFD1761F75F7A2B457156A67" docLanguage="en" docName="0010-065X-67.3.335.pdf" docOrigin="The Coleopterists Bulletin 67 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-67.3.335" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Stenosagola butcheri Park and Carlton, new species" docType="treatment" docVersion="4" lastPageNumber="357" masterDocId="FFAA421AFFC476097722A66C5656695D" masterDocTitle="A Revision of the New Zealand GenusStenosagolaBroun, 1921 (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)" masterLastPageNumber="359" masterPageNumber="335" pageNumber="356" updateTime="1699605239367" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8FDAFF5504DB85E076F0693FCFAEFA6B">A Revision of the New Zealand GenusStenosagolaBroun, 1921 (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)</mods:title>
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<mods:namePart id="C9CBD3FFB8BA77BC3E9D96A6F431F4C4">Park, J. - S.</mods:namePart>
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<mods:namePart id="D057E17A398ED890D29AFF2FA2E575F9">Carlton, C. E.</mods:namePart>
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<mods:title id="264A4DCB5A5A261DC730CDF2EDBB3EDC">The Coleopterists Bulletin</mods:title>
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<mods:date id="8B707F365352E0ED2FC6846D5F48F67B">2013</mods:date>
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<paragraph id="8B858B74FFD1761C75F7A2B4558D6C55" blockId="21.[725,1132,1240,1288]" pageId="21" pageNumber="356">
<heading id="D0CD3C18FFD1761C75F7A2B4558D6C55" centered="true" fontSize="10" level="2" pageId="21" pageNumber="356" reason="2">
<emphasis id="B94E5766FFD1761C75F7A2B4558D6C55" bold="true" pageId="21" pageNumber="356">
<taxonomicName id="4C3AF0F7FFD1761C75F7A2B452316DB3" ID-CoL="524SZ" authority="Park and Carlton" authorityName="Park and Carlton" box="[725,1127,1240,1262]" class="Insecta" family="Staphylinidae" genus="Stenosagola" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="356" phylum="Arthropoda" rank="species" species="butcheri" status="sp. nov.">
<emphasis id="B94E5766FFD1761C75F7A2B455F36DB3" bold="true" box="[725,933,1240,1262]" italics="true" pageId="21" pageNumber="356">Stenosagola butcheri</emphasis>
Park and Carlton
</taxonomicName>
,
<taxonomicNameLabel id="A27DEA1DFFD1761C7447A29F558D6C55" box="[869,987,1267,1288]" pageId="21" pageNumber="356" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C320D8FFFFD1761C75E7A37255E96FD1" pageId="21" pageNumber="356" type="materials_examined">
<paragraph id="8B858B74FFD1761C75E7A37255E96FD1" blockId="21.[685,1173,1310,1729]" pageId="21" pageNumber="356">
<emphasis id="B94E5766FFD1761C75E7A372550B6C6E" bold="true" box="[709,861,1310,1331]" pageId="21" pageNumber="356">Type Material.</emphasis>
<materialsCitation id="3B528129FFD1761C7448A37255986C8E" collectingDate="1981-10-11" collectionCode="WO, NZAC" collectorName="C. F. Butcher Litter &amp; N. Z. Arthropod Collection" country="New Zealand" location="Mt Karioi" pageId="21" pageNumber="356" specimenCount="1" specimenCount-male="1" stateProvince="Waikato" typeStatus="holotype">
<emphasis id="B94E5766FFD1761C7448A37252C56C6E" bold="true" box="[874,1171,1310,1331]" pageId="21" pageNumber="356">
<typeStatus id="548135D6FFD1761C7448A372559F6C6E" box="[874,969,1310,1331]" pageId="21" pageNumber="356" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F32DCBE4FFD1761C74F7A37252D96C6E" box="[981,1167,1310,1331]" name="New Zealand" pageId="21" pageNumber="356">NEW ZEALAND</collectingCountry>
:
</emphasis>
<collectingRegion id="49FE4596FFD1761C758FA35555766C13" box="[685,800,1337,1358]" country="New Zealand" name="Waikato" pageId="21" pageNumber="356">WAIKATO</collectingRegion>
(
<collectionCode id="ED2B13B1FFD1761C7411A35555356C13" box="[819,867,1337,1358]" pageId="21" pageNumber="356">WO</collectionCode>
):
<specimenCount id="9D3C40FDFFD1761C745BA35555CE6C13" box="[889,920,1336,1358]" pageId="21" pageNumber="356" type="male">1♂</specimenCount>
(winged male), “
<collectingCountry id="F32DCBE4FFD1761C7379A35555726C35" name="New Zealand" pageId="21" pageNumber="356">NEW ZEALAND</collectingCountry>
WO
<location id="8EE5DDAFFFD1761C7441A33F559B6C35" LSID="urn:lsid:plazi:treatment:03933A62FFD1761F75F7A2B457156A67:8EE5DDAFFFD1761C7441A33F559B6C35" box="[867,973,1363,1384]" country="New Zealand" name="Mt Karioi" pageId="21" pageNumber="356" stateProvince="Waikato">Mt Karioi</location>
<date id="FF84ADB4FFD1761C74FAA33F52086C35" box="[984,1118,1363,1384]" pageId="21" pageNumber="356" value="1981-10-11">
<collectingDate id="EFC0545CFFD1761C74FAA33F52086C35" box="[984,1118,1363,1384]" pageId="21" pageNumber="356" value="1981-10-11">11 Oct 1981</collectingDate>
</date>
<collectorName id="26CFEEA2FFD1761C734BA33F556D6CDE" pageId="21" pageNumber="356">C.F. Butcher Litter</collectorName>
81/95”, “
<collectorName id="26CFEEA2FFD1761C7486A302548E6CC0" pageId="21" pageNumber="356">N.Z. Arthropod Collection</collectorName>
,
<collectionCode id="ED2B13B1FFD1761C75C5A3E4557D6CC0" box="[743,811,1416,1437]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="21" pageNumber="356" type="Other institutional type">NZAC</collectionCode>
Private Bag 92170
<collectingRegion id="49FE4596FFD1761C7325A3E452C26CC0" box="[1031,1172,1416,1437]" country="New Zealand" name="Auckland" pageId="21" pageNumber="356">AUCKLAND</collectingRegion>
New Zealand”, “
<typeStatus id="548135D6FFD1761C746EA3CF55916CE5" box="[844,967,1443,1464]" pageId="21" pageNumber="356" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C3AF0F7FFD1761C74ECA3CE550C6C8F" authority="Park and Carlton" authorityName="Park and Carlton" class="Insecta" family="Staphylinidae" genus="Stenosagola" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="356" phylum="Arthropoda" rank="species" species="butcheri">
<emphasis id="B94E5766FFD1761C74ECA3CE52C26CEA" box="[974,1172,1442,1463]" italics="true" pageId="21" pageNumber="356">Stenosagola butcheri</emphasis>
Park and Carlton
</taxonomicName>
des. 2013”
</materialsCitation>
.
<materialsCitation id="3B528129FFD1761C74FCA3D152D96F60" collectingDate="2010-03-19" collectionCode="DSC" collectorName="D. S. Chandler" country="New Zealand" elevation="20" location="Clevedon" pageId="21" pageNumber="356" specimenCount="3" specimenCount-male="3" stateProvince="Auckland" typeStatus="paratype">
<emphasis id="B94E5766FFD1761C74FCA3D1523E6C8F" bold="true" box="[990,1128,1468,1490]" pageId="21" pageNumber="356">
<typeStatus id="548135D6FFD1761C74FCA3D152116C8F" box="[990,1095,1469,1490]" pageId="21" pageNumber="356" type="paratype">Paratypes</typeStatus>
(
<emphasis id="B94E5766FFD1761C7379A3D0523E6C8F" bold="true" box="[1115,1128,1468,1490]" italics="true" pageId="21" pageNumber="356">n</emphasis>
</emphasis>
=
<emphasis id="B94E5766FFD1761C73AAA3D155956CB0" bold="true" pageId="21" pageNumber="356">
<specimenCount id="9D3C40FDFFD1761C73AAA3D154BB6CB0" pageId="21" pageNumber="356" type="male">2 males</specimenCount>
).
<collectingCountry id="F32DCBE4FFD1761C7420A3B455E96CB0" box="[770,959,1496,1517]" name="New Zealand" pageId="21" pageNumber="356">NEW ZEALAND</collectingCountry>
:
</emphasis>
<collectingRegion id="49FE4596FFD1761C74EEA3B452046CB0" box="[972,1106,1496,1517]" country="New Zealand" name="Auckland" pageId="21" pageNumber="356">AUCKLAND</collectingRegion>
(AK):
<specimenCount id="9D3C40FDFFD1761C758FA39E549A6F55" box="[685,716,1522,1544]" pageId="21" pageNumber="356" type="male">1♂</specimenCount>
(slide-mounted),
<location id="8EE5DDAFFFD1761C74B1A39E55A36F5A" LSID="urn:lsid:plazi:treatment:03933A62FFD1761F75F7A2B457156A67:8EE5DDAFFFD1761C74B1A39E55A36F5A" box="[915,1013,1522,1543]" country="New Zealand" name="Clevedon" pageId="21" pageNumber="356" stateProvince="Auckland">Clevedon</location>
Scenic Reser.
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<quantity id="4CC22691FFD1761C758FA06154B96F7F" box="[685,751,1549,1570]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="356" unit="km" value="0.5">0.5km</quantity>
N Clevedon
</locationDeviation>
,
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<elevation id="00176C47FFD1761C74B5A061559E6F7F" box="[919,968,1549,1570]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="356" unit="m" value="20.0">20m</elevation>
</quantity>
,
<date id="FF84ADB4FFD1761C74F8A06152016F7F" box="[986,1111,1549,1570]" pageId="21" pageNumber="356" value="2010-03-19">
<collectingDate id="EFC0545CFFD1761C74F8A06152016F7F" box="[986,1111,1549,1570]" pageId="21" pageNumber="356" value="2010-03-19">III-19-2010</collectingDate>
</date>
<collectorName id="26CFEEA2FFD1761C7347A06155586F60" pageId="21" pageNumber="356">D.S. Chandler</collectorName>
, sift forest litter by stream (
<collectionCode id="ED2B13B1FFD1761C7370A04452DE6F60" box="[1106,1160,1576,1597]" country="USA" httpUri="http://grbio.org/cool/s9x3-awht" name="Dicty Stock Center" pageId="21" pageNumber="356">DSC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B528129FFD1761C758FA02E55ED6FD1" collectingDate="1976-12-15" collectionCode="NZAC" collectorName="N. M. May" country="New Zealand" elevation="400" location="Mangamuka" pageId="21" pageNumber="356" specimenCount="1" specimenCount-male="1" stateProvince="Northland" typeStatus="paratype">
<collectingRegion id="49FE4596FFD1761C758FA02E55196F0A" box="[685,847,1602,1623]" country="New Zealand" name="Northland" pageId="21" pageNumber="356">NORTHLAND</collectingRegion>
(ND):
<specimenCount id="9D3C40FDFFD1761C748EA02E559A6F05" box="[940,972,1602,1624]" pageId="21" pageNumber="356" type="male">1♂</specimenCount>
(slide-mounted),
<location id="8EE5DDAFFFD1761C758FA031557E6F2F" LSID="urn:lsid:plazi:treatment:03933A62FFD1761F75F7A2B457156A67:8EE5DDAFFFD1761C758FA031557E6F2F" box="[685,808,1629,1650]" country="New Zealand" name="Mangamuka" pageId="21" pageNumber="356" stateProvince="Northland">Mangamuka</location>
summit,
<quantity id="4CC22691FFD1761C74B2A031559D6F2F" box="[912,971,1629,1650]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="356" unit="m" value="400.0">
<elevation id="00176C47FFD1761C74B2A031559D6F2F" box="[912,971,1629,1650]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="356" unit="m" value="400.0">400m</elevation>
</quantity>
,
<date id="FF84ADB4FFD1761C74F4A03152026F2F" box="[982,1108,1629,1650]" pageId="21" pageNumber="356" value="1976-12-15">
<collectingDate id="EFC0545CFFD1761C74F4A03152026F2F" box="[982,1108,1629,1650]" pageId="21" pageNumber="356" value="1976-12-15">15 XII 1976</collectingDate>
</date>
,
<collectorName id="26CFEEA2FFD1761C7342A031548D6FD1" pageId="21" pageNumber="356">N.M. May</collectorName>
, litter 76/108 (
<collectionCode id="ED2B13B1FFD1761C744FA01B55E36FD1" box="[877,949,1655,1676]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="21" pageNumber="356" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320D8FFFFD1761C75E7A0FE52436F9C" pageId="21" pageNumber="356" type="etymology">
<paragraph id="8B858B74FFD1761C75E7A0FE52436F9C" blockId="21.[685,1173,1310,1729]" pageId="21" pageNumber="356">
<emphasis id="B94E5766FFD1761C75E7A0FE556E6FFA" bold="true" box="[709,824,1682,1703]" pageId="21" pageNumber="356">Etymology.</emphasis>
This species is named for the collector of the
<typeStatus id="548135D6FFD1761C7409A0C055D76F9C" box="[811,897,1708,1729]" pageId="21" pageNumber="356" type="holotype">holotype</typeStatus>
, C. F. Butcher.
</paragraph>
</subSubSection>
<subSubSection id="C320D8FFFFD2761F77E7A6AE5413687A" pageId="22" pageNumber="357" type="diagnosis">
<paragraph id="8B858B74FFD2761F77E7A6AE5413687A" blockId="22.[173,660,194,826]" pageId="22" pageNumber="357">
<emphasis id="B94E5766FFD2761F77E7A6AE577B698A" bold="true" box="[197,301,194,215]" pageId="22" pageNumber="357">Diagnosis.</emphasis>
Specimens of
<taxonomicName id="4C3AF0F7FFD2761F76EAA6AE5465698A" box="[456,563,194,215]" class="Insecta" family="Staphylinidae" genus="Stenosagola" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="357" phylum="Arthropoda" rank="species" species="butcheri">
<emphasis id="B94E5766FFD2761F76EAA6AE5465698A" box="[456,563,194,215]" italics="true" pageId="22" pageNumber="357">S. butcheri</emphasis>
</taxonomicName>
are similar in appearance to those of other members of the
<taxonomicName id="4C3AF0F7FFD2761F77F7A69B57736851" baseAuthorityName="Broun" baseAuthorityYear="1911" box="[213,293,247,268]" class="Insecta" family="Staphylinidae" genus="Stenosagola" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="357" phylum="Arthropoda" rank="species" species="connata">
<emphasis id="B94E5766FFD2761F77F7A69B57736851" box="[213,293,247,268]" italics="true" pageId="22" pageNumber="357">connata</emphasis>
</taxonomicName>
group but can be distinguished by the shape of the male genitalia (
<figureCitation id="130197F1FFD2761F76C4A77E546C687A" box="[486,570,274,295]" captionStart="Fig" captionStartId="19.[175,207,746,765]" captionTargetBox="[153,633,196,723]" captionTargetId="figure-642@19.[151,638,194,725]" captionTargetPageId="19" captionText="Fig. 12. Stenosagola species. a) S. pseudoconnata, aedeagus, dorsal view, b) S. chandleri, aedeagus, dorsal view, c) S. nunni, aedeagus, dorsal view, d) S. newtoni, aedeagus, dorsal view, e) S. butcheri, aedeagus, dorsal view, f) S. fiordlandensis, aedeagus, dorsal view, g) S. fiordlandensis, wingless male head, dorsal view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10100781" httpUri="https://zenodo.org/record/10100781/files/figure.png" pageId="22" pageNumber="357">Fig. 12e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320D8FFFFD2761F77E7A74057126B8D" pageId="22" pageNumber="357" type="description">
<paragraph id="8B858B74FFD2761F77E7A74057126B8D" blockId="22.[173,660,194,826]" pageId="22" pageNumber="357">
<emphasis id="B94E5766FFD2761F77E7A7405717681C" bold="true" box="[197,321,300,321]" pageId="22" pageNumber="357">Description.</emphasis>
Length
<quantity id="4CC22691FFD2761F76BEA7405445681C" box="[412,531,300,322]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="22" pageNumber="357" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
(
<figureCitation id="130197F1FFD2761F7501A74054DE681C" box="[547,648,300,322]" captionStart="Fig" captionStartId="12.[197,229,773,792]" captionTargetBox="[173,1193,194,750]" captionTargetId="figure-443@12.[173,1193,194,750]" captionTargetPageId="12" captionText="Fig. 7. Habitus of Stenosagola species, dorsal view. a) Wingless male of S. tararuaensis, b) Female of S. tararuaensis, c) Wingless male of S. haunuiensis, d) Winged male of S. pseudoconnata, e) Wingless male of S. pseudoconnata, f) Wingless male of S. chandleri, g) Female of S. chandleri, h) Winged male of S. nunni, i) Wingless male of S. nunni, j) Female of S. nunni, k) Wingless male of S. newtoni, l) Winged male of S. butcheri, m) Wingless male of S. butcheri, n) Wingless male of S. fiordlandensis, o) Female of S. fiordlandensis. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10100768" httpUri="https://zenodo.org/record/10100768/files/figure.png" pageId="22" pageNumber="357">Fig. 7lm</figureCitation>
).
<emphasis id="B94E5766FFD2761F778FA72B56B46801" bold="true" box="[173,226,327,348]" pageId="22" pageNumber="357">Head</emphasis>
: Winged male head approximately 1.52 times, wingless male 1.50 times longer than wide. Winged male eye approximately 1.59 times, wingless male 0.9 times longer than temple. Winged male antennomeres 38 subquadrate, 910 transverse. Wingless male antennomeres 37 subquadrate, 8 slightly transverse, 910 transverse.
<emphasis id="B94E5766FFD2761F769DA78A545C68A6" bold="true" box="[447,522,486,507]" pageId="22" pageNumber="357">Thorax</emphasis>
: Pronotum as long as wide. Winged male elytra approximately 1.89, wingless male 1.29 times longer than pronotum. Winged male meso- and metaventrites trapezoidal, approximately1.65 times, wingless male 1.19 times longer than pronotum.
<emphasis id="B94E5766FFD2761F76B3A40757A36BDD" bold="true" box="[401,501,619,640]" pageId="22" pageNumber="357">Aedeagus</emphasis>
: Apical lobe of male genitalia emarginate and bent to right (
<figureCitation id="130197F1FFD2761F7519A4EA54DE6BC6" box="[571,648,646,667]" captionStart="Fig" captionStartId="19.[175,207,746,765]" captionTargetBox="[153,633,196,723]" captionTargetId="figure-642@19.[151,638,194,725]" captionTargetPageId="19" captionText="Fig. 12. Stenosagola species. a) S. pseudoconnata, aedeagus, dorsal view, b) S. chandleri, aedeagus, dorsal view, c) S. nunni, aedeagus, dorsal view, d) S. newtoni, aedeagus, dorsal view, e) S. butcheri, aedeagus, dorsal view, f) S. fiordlandensis, aedeagus, dorsal view, g) S. fiordlandensis, wingless male head, dorsal view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10100781" httpUri="https://zenodo.org/record/10100781/files/figure.png" pageId="22" pageNumber="357">Fig. 12e</figureCitation>
). Left paramere longer than right, bearing setae apically (
<figureCitation id="130197F1FFD2761F77C5A4D7576F6B8D" box="[231,313,699,720]" captionStart="Fig" captionStartId="19.[175,207,746,765]" captionTargetBox="[153,633,196,723]" captionTargetId="figure-642@19.[151,638,194,725]" captionTargetPageId="19" captionText="Fig. 12. Stenosagola species. a) S. pseudoconnata, aedeagus, dorsal view, b) S. chandleri, aedeagus, dorsal view, c) S. nunni, aedeagus, dorsal view, d) S. newtoni, aedeagus, dorsal view, e) S. butcheri, aedeagus, dorsal view, f) S. fiordlandensis, aedeagus, dorsal view, g) S. fiordlandensis, wingless male head, dorsal view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10100781" httpUri="https://zenodo.org/record/10100781/files/figure.png" pageId="22" pageNumber="357">Fig. 12e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320D8FFFFD2761F77E7A4B957156A67" pageId="22" pageNumber="357" type="distribution">
<paragraph id="8B858B74FFD2761F77E7A4B954746A58" blockId="22.[173,660,194,826]" pageId="22" pageNumber="357">
<emphasis id="B94E5766FFD2761F77E7A4B957116BB7" bold="true" box="[197,327,725,746]" pageId="22" pageNumber="357">Distribution.</emphasis>
Northern part of North Island,
<collectingCountry id="F32DCBE4FFD2761F778FA49C577B6A58" box="[173,301,752,773]" name="New Zealand" pageId="22" pageNumber="357">New Zealand</collectingCountry>
(
<figureCitation id="130197F1FFD2761F761FA49C57D36A58" box="[317,389,752,773]" captionStart="Fig" captionStartId="21.[175,207,909,928]" captionTargetBox="[153,637,194,888]" captionTargetId="figure-552@21.[151,638,189,890]" captionTargetPageId="21" captionText="Fig. 14. Locations where Stenosagola nunni (triangles), Stenosagola newtoni (circle), Stenosagola butcheri (black squares), and Stenosagola fiordlandensis (white square) have been collected in New Zealand." figureDoi="http://doi.org/10.5281/zenodo.10100785" httpUri="https://zenodo.org/record/10100785/files/figure.png" pageId="22" pageNumber="357">Fig. 14</figureCitation>
, black squares).
</paragraph>
<paragraph id="8B858B74FFD2761F77E7A56757156A67" blockId="22.[173,660,194,826]" pageId="22" pageNumber="357">
<emphasis id="B94E5766FFD2761F77E7A567574F6A7D" bold="true" box="[197,281,779,800]" pageId="22" pageNumber="357">Habitat.</emphasis>
All specimens were collected by sifting forest litter.
</paragraph>
</subSubSection>
</treatment>
</document>