225 lines
20 KiB
XML
225 lines
20 KiB
XML
<document ID-DOI="10.3853/j.2201-4349.74.2022.1775" ID-ISSN="2201-4349" ID-Zenodo-Dep="7177922" approvalRequired="7" approvalRequired_for_textStreams="5" approvalRequired_for_treatments="2" checkinTime="1665305657841" checkinUser="felipe" docAuthor="Billingham, Zacariah D. & Theischinger, Gunther" docDate="2022" docId="0392DE4AFC05FFF9FF510BA2FA57A889" docLanguage="en" docName="RecAustMus.74.1.19–40.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 74 (1)" docSource="http://dx.doi.org/10.3853/j.2201-4349.74.2022.1775" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.1:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="1" docTitle="Molophilus (Molophilus) globostylus Billingham & Theischinger 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="FFABA632FC01FFFDFF990B35FFFEAA2C" masterDocTitle="New and Poorly Known Species of Crane Flies (Diptera: Limoniidae) from New South Wales, Australia" masterLastPageNumber="40" masterPageNumber="19" pageNumber="23" updateTime="1665324244475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New and Poorly Known Species of Crane Flies (Diptera: Limoniidae) from New South Wales, Australia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Billingham, Zacariah D.</mods:namePart>
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<mods:affiliation>Water Sciences, Natural Resources and Heritage, GHD, 180 Lonsdale Street, Melbourne VIC 3000, Australia & PhD candidate, Department of Ecology, Environment and Evolution, La Trobe University, Bundoora VIC 3086, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Theischinger, Gunther</mods:namePart>
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<mods:affiliation>Research Associate, Australian Museum Research Institute, Australian Museum, 6 College Street, Sydney NSW 2010, Australia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="series">
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<mods:title>Rec. Aust. Mus.</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2022-03-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>74</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>40</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3853/j.2201-4349.74.2022.1775</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3853/j.2201-4349.74.2022.1775</mods:identifier>
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<mods:identifier type="ISSN">2201-4349</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7177922</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7175112" ID-Zenodo-Dep="7175112" LSID="urn:lsid:plazi:treatment:0392DE4AFC05FFF9FF510BA2FA57A889" httpUri="http://treatment.plazi.org/id/0392DE4AFC05FFF9FF510BA2FA57A889" lastPageNumber="23" pageId="4" pageNumber="23">
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<subSubSection pageId="4" pageNumber="23" type="nomenclature">
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<paragraph blockId="4.[200,760,151,178]" box="[200,760,151,178]" pageId="4" pageNumber="23">
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<heading bold="true" box="[200,760,151,178]" centered="true" fontSize="12" level="2" pageId="4" pageNumber="23" reason="2">
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<emphasis bold="true" box="[200,760,151,178]" pageId="4" pageNumber="23">
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<taxonomicName authority="Billingham & Theischinger, 2022" authorityName="Billingham & Theischinger" authorityYear="2022" box="[200,656,151,178]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="globostylus" status="sp. nov." subGenus="Molophilus">
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<emphasis bold="true" box="[200,656,151,178]" italics="true" pageId="4" pageNumber="23">Molophilus (Molophilus) globostylus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[665,760,151,177]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="4.[181,779,200,218]" box="[181,779,200,218]" pageId="4" pageNumber="23">
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<uri box="[181,779,200,218]" pageId="4" pageNumber="23">
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection box="[428,532,247,273]" pageId="4" pageNumber="23" type="description">
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<paragraph blockId="4.[428,532,247,273]" box="[428,532,247,273]" pageId="4" pageNumber="23">
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<figureCitation box="[428,532,247,273]" captionStart="Figures 7–8" captionStartId="4.[167,244,1971,1993]" captionTargetBox="[275,1382,1405,1955]" captionTargetId="figure-281@4.[167,1467,1405,1955]" captionTargetPageId="4" captionText="Figures 7–8. Molophilus (Molophilus) globostylus sp. nov., male. (7) head colouration, dorsal; (8) hypopygium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7177932" httpUri="https://zenodo.org/record/7177932/files/figure.png" pageId="4" pageNumber="23">Figs 7–8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="23" type="materials_examined">
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<paragraph blockId="4.[166,794,294,494]" pageId="4" pageNumber="23">
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<materialsCitation collectingDate="2018-11-11" collectionCode="AM" collectorName="Z. Billingham & G. Theischinger" country="Australia" county="Gloucester Tops" latitude="-32.098" location="Gloucester Falls Walking Track" longLatPrecision="72" longitude="151.597" municipality="Tributary of Gloucester River" pageId="4" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales" typeStatus="holotype">
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<emphasis bold="true" box="[166,275,294,318]" pageId="4" pageNumber="23">
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<typeStatus box="[166,275,294,318]" pageId="4" pageNumber="23">Holotype</typeStatus>
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</emphasis>
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<specimenCount box="[286,308,295,318]" pageId="4" pageNumber="23" type="male">♂</specimenCount>
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,
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<collectingCountry box="[324,431,294,318]" name="Australia" pageId="4" pageNumber="23">Australia</collectingCountry>
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,
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<collectingRegion box="[446,656,294,318]" country="Australia" name="New South Wales" pageId="4" pageNumber="23">New South Wales</collectingRegion>
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,
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<collectingCounty pageId="4" pageNumber="23">Gloucester Tops</collectingCounty>
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,
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<collectingMunicipality box="[235,565,324,348]" pageId="4" pageNumber="23">Tributary of Gloucester River</collectingMunicipality>
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||
off
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<location LSID="urn:lsid:plazi:treatment:0392DE4AFC05FFF9FF510BA2FA57A889:8EE43987FC05FFF9FDFE0A71FEB4AB55" country="Australia" county="Gloucester Tops" latitude="-32.098" longLatPrecision="72" longitude="151.597" municipality="Tributary of Gloucester River" name="Gloucester Falls Walking Track" pageId="4" pageNumber="23" stateProvince="New South Wales">
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<location LSID="urn:lsid:plazi:treatment:0392DE4AFC05FFF9FF510BA2FA57A889:8EE43987FC05FFF9FDFE0A71FCE4AB70" box="[615,794,324,348]" country="Australia" county="Gloucester Tops" latitude="-32.098" longLatPrecision="72" longitude="151.597" municipality="Tributary of Gloucester River" name="Gloucester Falls" pageId="4" pageNumber="23" stateProvince="New South Wales">Gloucester Falls</location>
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Walking Track
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</location>
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||
(
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||
<geoCoordinate box="[349,452,353,377]" degrees="32.098" direction="south" orientation="latitude" pageId="4" pageNumber="23" precision="55" value="-32.098">32.098°S</geoCoordinate>
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<geoCoordinate box="[462,583,353,377]" degrees="151.597" direction="east" orientation="longitude" pageId="4" pageNumber="23" precision="55" value="151.597">151.597°E</geoCoordinate>
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||
),
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||
<date box="[605,756,353,377]" pageId="4" pageNumber="23" value="2018-11-11">
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<collectingDate box="[605,756,353,377]" pageId="4" pageNumber="23" value="2018-11-11">11 Nov 2018</collectingDate>
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</date>
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,
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<collectorName pageId="4" pageNumber="23">Z. Billingham</collectorName>
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&
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<collectorName box="[321,494,382,406]" pageId="4" pageNumber="23">G. Theischinger</collectorName>
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,
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<collectionCode box="[503,546,382,406]" country="Australia" name="Australian Museum" pageId="4" pageNumber="23" type="Museum">AM</collectionCode>
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||
(K.394957)
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</materialsCitation>
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.
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<materialsCitation collectingDate="2018-11-12" collectionCode="AM" collectorName="Z. Billingham & G. Theischinger" country="Australia" county="Tomalla" latitude="-31.884" location="Pheasant Creek Rd" longLatPrecision="72" longitude="151.477" municipality="Pheasant Creek" pageId="4" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales" typeStatus="paratype">
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<emphasis bold="true" box="[690,794,382,406]" pageId="4" pageNumber="23">
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||
<typeStatus box="[690,794,382,406]" pageId="4" pageNumber="23">Paratype</typeStatus>
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||
</emphasis>
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||
<specimenCount box="[166,201,412,436]" pageId="4" pageNumber="23" type="male">1♂</specimenCount>
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,
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||
<collectingCountry box="[212,313,412,436]" name="Australia" pageId="4" pageNumber="23">Australia</collectingCountry>
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||
,
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||
<collectingRegion box="[323,517,412,436]" country="Australia" name="New South Wales" pageId="4" pageNumber="23">New South Wales</collectingRegion>
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,
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||
<collectingCounty box="[527,617,412,436]" pageId="4" pageNumber="23">Tomalla</collectingCounty>
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,
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<collectingMunicipality box="[628,794,412,436]" pageId="4" pageNumber="23">Pheasant Creek</collectingMunicipality>
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||
off
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||
<location LSID="urn:lsid:plazi:treatment:0392DE4AFC05FFF9FF510BA2FA57A889:8EE43987FC05FFF9FF500A8CFE6EABFD" box="[201,400,441,465]" country="Australia" county="Tomalla" latitude="-31.884" longLatPrecision="72" longitude="151.477" municipality="Pheasant Creek" name="Pheasant Creek Rd" pageId="4" pageNumber="23" stateProvince="New South Wales">
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||
<location LSID="urn:lsid:plazi:treatment:0392DE4AFC05FFF9FF510BA2FA57A889:8EE43987FC05FFF9FF500A8CFE92ABFD" box="[201,364,441,465]" country="Australia" county="Tomalla" latitude="-31.884" longLatPrecision="72" longitude="151.477" municipality="Pheasant Creek" name="Pheasant Creek" pageId="4" pageNumber="23" stateProvince="New South Wales">Pheasant Creek</location>
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Rd
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</location>
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(
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||
<geoCoordinate box="[412,513,441,465]" degrees="31.884" direction="south" orientation="latitude" pageId="4" pageNumber="23" precision="55" value="-31.884">31.884°S</geoCoordinate>
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<geoCoordinate box="[517,634,441,465]" degrees="151.477" direction="east" orientation="longitude" pageId="4" pageNumber="23" precision="55" value="151.477">151.477°E</geoCoordinate>
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),
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||
<date box="[651,790,441,465]" pageId="4" pageNumber="23" value="2018-11-12">
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<collectingDate box="[651,790,441,465]" pageId="4" pageNumber="23" value="2018-11-12">12 Nov 2018</collectingDate>
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||
</date>
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,
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||
<collectorName box="[166,316,470,494]" pageId="4" pageNumber="23">Z. Billingham</collectorName>
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&
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<collectorName box="[350,523,470,494]" pageId="4" pageNumber="23">G. Theischinger</collectorName>
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,
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<collectionCode box="[532,575,470,494]" country="Australia" name="Australian Museum" pageId="4" pageNumber="23" type="Museum">AM</collectionCode>
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||
</materialsCitation>
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||
(K.394958).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="4" pageNumber="23" type="description">
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<paragraph blockId="4.[166,795,515,1302]" box="[166,483,515,540]" pageId="4" pageNumber="23">
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<emphasis bold="true" box="[166,298,515,539]" pageId="4" pageNumber="23">Description</emphasis>
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<specimenCount box="[305,325,516,539]" pageId="4" pageNumber="23" type="male">♂</specimenCount>
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(
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<specimenCount box="[342,360,516,540]" pageId="4" pageNumber="23" type="female">♀</specimenCount>
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unknown).
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</paragraph>
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<paragraph blockId="4.[166,795,515,1302]" pageId="4" pageNumber="23">
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<emphasis box="[197,255,545,569]" italics="true" pageId="4" pageNumber="23">Head</emphasis>
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(
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<figureCitation box="[267,331,545,569]" captionStart="Figures 7–8" captionStartId="4.[167,244,1971,1993]" captionTargetBox="[275,1382,1405,1955]" captionTargetId="figure-281@4.[167,1467,1405,1955]" captionTargetPageId="4" captionText="Figures 7–8. Molophilus (Molophilus) globostylus sp. nov., male. (7) head colouration, dorsal; (8) hypopygium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7177932" httpUri="https://zenodo.org/record/7177932/files/figure.png" pageId="4" pageNumber="23">Fig. 7</figureCitation>
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): golden yellow, vertex with prominent dark greyish brown triangular marking, antennal pedicel and scape pale whitish yellow, antennal flagellum with basal segments pale whitish yellow, terminal segments darkening to greyish brown, palpus dark greyish brown. Antenna of moderate length, extending back to the transverse suture.
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</paragraph>
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<paragraph blockId="4.[166,795,515,1302]" pageId="4" pageNumber="23">
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<emphasis box="[197,280,721,745]" italics="true" pageId="4" pageNumber="23">Thorax.</emphasis>
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Pronotum with dorsal angle pale yellow to white, otherwise golden yellow. Prescutum and scutum golden brown, the posterior margins of scutal lobes pale yellowish brown. Scutellum and mediotergite largely golden yellow, each with darker golden brown central area. Paratergite bright white, in combination with pale dorsal angle of pronotum forming a distinct pale stripe between terga and pleura. Cervical sclerite, propleuron and pleurites golden yellow. Forecoxa golden yellow, meso- and metacoxa and trochanters pale whitish yellow. Basal ⅓ of femora pale golden yellow, remainder darkening to greyish brown. Tibiae and tarsi dark greyish brown. Wing pale grey tinged with yellow, veins yellowish brown. Halter with stem and knob whitish yellow.
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</paragraph>
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<paragraph blockId="4.[166,795,515,1302]" box="[197,472,1132,1156]" pageId="4" pageNumber="23">
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<emphasis box="[197,298,1132,1156]" italics="true" pageId="4" pageNumber="23">Abdomen</emphasis>
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: greyish brown.
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</paragraph>
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<paragraph blockId="4.[166,795,515,1302]" lastBlockId="4.[839,1467,151,352]" pageId="4" pageNumber="23">
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<taxonomicName box="[197,313,1161,1185]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="23" phylum="Tracheophyta" rank="genus">
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<emphasis box="[197,313,1161,1185]" italics="true" pageId="4" pageNumber="23">Terminalia</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[327,391,1161,1185]" captionStart="Figures 7–8" captionStartId="4.[167,244,1971,1993]" captionTargetBox="[275,1382,1405,1955]" captionTargetId="figure-281@4.[167,1467,1405,1955]" captionTargetPageId="4" captionText="Figures 7–8. Molophilus (Molophilus) globostylus sp. nov., male. (7) head colouration, dorsal; (8) hypopygium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7177932" httpUri="https://zenodo.org/record/7177932/files/figure.png" pageId="4" pageNumber="23">Fig. 8</figureCitation>
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). Hypopygium golden yellow, lobe of tergite 9 rather long, narrowly trapezoidal. Gonocoxite with moderately sized, apically bluntly rounded, ventral lobe, dorsal and medial lobes rather small. Outer gonostylus a short rod terminating in bifurcate subequal spines, bearing on its dorsal surface a large swollen globe-shaped appendage. Inner gonostylus long, heavily sclerotized and terminating in an acute spine, strongly bent at about half its length to project caudad behind the globe-like appendage of the outer gonostylus. Aedeagus long and narrow. Parameres fused, poorly developed.
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</paragraph>
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<paragraph blockId="4.[839,1467,151,352]" box="[869,1463,327,352]" pageId="4" pageNumber="23">
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<emphasis box="[869,1003,328,351]" italics="true" pageId="4" pageNumber="23">Dimensions.</emphasis>
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Body length
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<quantity box="[1146,1230,328,352]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="23" unit="mm" value="4.7">4.7 mm</quantity>
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, wing length
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<quantity box="[1374,1459,328,352]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="23" unit="mm" value="4.9">4.9 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="23" type="etymology">
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<paragraph blockId="4.[839,1468,373,426]" pageId="4" pageNumber="23">
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<emphasis bold="true" box="[839,963,373,397]" pageId="4" pageNumber="23">Etymology</emphasis>
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. This species is named with reference to the prominent globe-shaped appendage of the outer gonostylus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="23" type="discussion">
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<paragraph blockId="4.[839,1467,447,677]" pageId="4" pageNumber="23">
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<emphasis bold="true" box="[839,964,447,471]" pageId="4" pageNumber="23">Discussion</emphasis>
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.
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<taxonomicName authorityName="Billingham & Theischinger" authorityYear="2022" box="[983,1307,447,471]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="globostylus" subGenus="Molophilus">
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<emphasis box="[983,1307,447,471]" italics="true" pageId="4" pageNumber="23">Molophilus (M.) globostylus</emphasis>
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</taxonomicName>
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is allied to a large group of morphologically similar species around
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<taxonomicName authority="Alexander, 1927" authorityName="Alexander" authorityYear="1927" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="capitatus" subGenus="Molophilus">
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<emphasis italics="true" pageId="4" pageNumber="23">M. (M.) capitatus</emphasis>
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<bibRefCitation author="Alexander, C. P." box="[1007,1191,506,530]" pageId="4" pageNumber="23" pagination="42 - 72" refId="ref12190" refString="Alexander, C. P. 1927. The interpretation of the radial field of the wing in the Nematocerous Diptera, with special reference to the Tipulidae. Proceedings of the Linnaean Society of New South Wales 52: 42 - 72." type="journal article" year="1927">Alexander, 1927</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName authority="Theischinger, 1988" authorityName="Theischinger" authorityYear="1988" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="auriculifer" subGenus="Molophilus">
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<emphasis box="[1252,1467,506,530]" italics="true" pageId="4" pageNumber="23">M. (M.) auriculifer</emphasis>
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Theischinger, 1988
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</taxonomicName>
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(see
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<bibRefCitation author="Theischinger, G." box="[1096,1294,535,560]" pageId="4" pageNumber="23" pagination="5 - 150" refId="ref12776" refString="Theischinger, G. 1992. The Limoniinae (Diptera: Tipulidae) of Australia. II. The genus Molophilus Curtis. Stapfia 27: 5 - 150." type="journal article" year="1992">Theischinger 1992</bibRefCitation>
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for more details on these species), and most closely resembles
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<taxonomicName authority="Alexander, 1927" authorityName="Alexander" authorityYear="1927" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="sigma" subGenus="Molophilus">
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<emphasis box="[1318,1466,565,589]" italics="true" pageId="4" pageNumber="23">M. (M.) sigma</emphasis>
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<bibRefCitation author="Alexander, C. P." box="[839,1025,594,618]" pageId="4" pageNumber="23" pagination="42 - 72" refId="ref12190" refString="Alexander, C. P. 1927. The interpretation of the radial field of the wing in the Nematocerous Diptera, with special reference to the Tipulidae. Proceedings of the Linnaean Society of New South Wales 52: 42 - 72." type="journal article" year="1927">Alexander, 1927</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName authorityName="Billingham & Theischinger" authorityYear="2022" box="[1039,1353,594,618]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="globostylus" subGenus="Molophilus">
|
||
<emphasis box="[1039,1353,594,618]" italics="true" pageId="4" pageNumber="23">Molophilus (M.) globostylus</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished from all similar species by the bifurcate outer gonostylus with its large globe-shaped dorsal appendage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
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||
</document> |