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<document ID-DOI="10.3853/j.2201-4349.74.2022.1775" ID-ISSN="2201-4349" ID-Zenodo-Dep="7177922" approvalRequired="7" approvalRequired_for_textStreams="5" approvalRequired_for_treatments="2" checkinTime="1665305657841" checkinUser="felipe" docAuthor="Billingham, Zacariah D. &amp; Theischinger, Gunther" docDate="2022" docId="0392DE4AFC00FFFFFF5F08D7FE01A907" docLanguage="en" docName="RecAustMus.74.1.1940.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 74 (1)" docSource="http://dx.doi.org/10.3853/j.2201-4349.74.2022.1775" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.1:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="1" docTitle="Molophilus (Molophilus) avia Billingham &amp; Theischinger 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="20" masterDocId="FFABA632FC01FFFDFF990B35FFFEAA2C" masterDocTitle="New and Poorly Known Species of Crane Flies (Diptera: Limoniidae) from New South Wales, Australia" masterLastPageNumber="40" masterPageNumber="19" pageNumber="20" updateTime="1665324244475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New and Poorly Known Species of Crane Flies (Diptera: Limoniidae) from New South Wales, Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Billingham, Zacariah D.</mods:namePart>
<mods:affiliation>Water Sciences, Natural Resources and Heritage, GHD, 180 Lonsdale Street, Melbourne VIC 3000, Australia &amp; PhD candidate, Department of Ecology, Environment and Evolution, La Trobe University, Bundoora VIC 3086, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Theischinger, Gunther</mods:namePart>
<mods:affiliation>Research Associate, Australian Museum Research Institute, Australian Museum, 6 College Street, Sydney NSW 2010, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="series">
<mods:title>Rec. Aust. Mus.</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2022-03-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>74</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>40</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3853/j.2201-4349.74.2022.1775</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3853/j.2201-4349.74.2022.1775</mods:identifier>
<mods:identifier type="ISSN">2201-4349</mods:identifier>
<mods:identifier type="Zenodo-Dep">7177922</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7175106" ID-Zenodo-Dep="7175106" LSID="urn:lsid:plazi:treatment:0392DE4AFC00FFFFFF5F08D7FE01A907" httpUri="http://treatment.plazi.org/id/0392DE4AFC00FFFFFF5F08D7FE01A907" lastPageId="2" lastPageNumber="20" pageId="1" pageNumber="20">
<subSubSection pageId="1" pageNumber="20" type="nomenclature">
<paragraph blockId="1.[198,671,994,1021]" box="[198,671,994,1021]" pageId="1" pageNumber="20">
<heading bold="true" box="[198,671,994,1021]" centered="true" fontSize="12" level="2" pageId="1" pageNumber="20" reason="2">
<emphasis bold="true" box="[198,671,994,1021]" pageId="1" pageNumber="20">
<taxonomicName authority="Billingham &amp; Theischinger, 2022" authorityName="Billingham &amp; Theischinger" authorityYear="2022" box="[198,567,994,1021]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="avia" status="sp. nov." subGenus="Molophilus">
<emphasis bold="true" box="[198,567,994,1021]" italics="true" pageId="1" pageNumber="20">Molophilus (Molophilus) avia</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[576,671,995,1021]" pageId="1" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="1.[137,732,1044,1062]" box="[137,732,1044,1062]" pageId="1" pageNumber="20">
<uri box="[137,732,1044,1062]" pageId="1" pageNumber="20">
urn:lsid:zoobank.org:act:
<uuid box="[337,732,1044,1062]" pageId="1" pageNumber="20">0897FEEB-8893-4EAB-9001-4C582C077F4C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[383,486,1083,1109]" pageId="1" pageNumber="20" type="description">
<paragraph blockId="1.[383,486,1083,1109]" box="[383,486,1083,1109]" pageId="1" pageNumber="20">
<figureCitation box="[383,486,1083,1109]" captionStart="Figures 12" captionStartId="1.[121,198,1983,2005]" captionTargetBox="[214,1326,1397,1973]" captionTargetId="figure-582@1.[121,1421,1396,1973]" captionTargetPageId="1" captionText="Figures 12. Molophilus (Molophilus) avia sp. nov., male. (1) head colouration, dorsal; (2) hypopygium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7177924" httpUri="https://zenodo.org/record/7177924/files/figure.png" pageId="1" pageNumber="20">Figs 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="20" type="materials_examined">
<paragraph blockId="1.[121,749,1122,1381]" pageId="1" pageNumber="20">
<materialsCitation collectingDate="2018-11-09" collectionCode="AM" collectorName="Z. Billingham &amp; G. Theischinger" country="Australia" county="Upper Karuah River" latitude="-32.232" location="Frying Pan Rd" longLatPrecision="72" longitude="151.764" municipality="Burrawang Creek" pageId="1" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales" typeStatus="holotype">
<emphasis bold="true" box="[121,225,1122,1146]" pageId="1" pageNumber="20">
<typeStatus box="[121,225,1122,1146]" pageId="1" pageNumber="20">Holotype</typeStatus>
</emphasis>
<specimenCount box="[234,256,1122,1145]" pageId="1" pageNumber="20" type="male"></specimenCount>
,
<collectingCountry box="[269,370,1122,1146]" name="Australia" pageId="1" pageNumber="20">Australia</collectingCountry>
,
<collectingRegion box="[382,580,1122,1146]" country="Australia" name="New South Wales" pageId="1" pageNumber="20">New South Wales</collectingRegion>
,
<collectingCounty pageId="1" pageNumber="20">Upper Karuah River</collectingCounty>
,
<collectingMunicipality box="[200,401,1151,1176]" pageId="1" pageNumber="20">Burrawang Creek</collectingMunicipality>
off
<location LSID="urn:lsid:plazi:treatment:0392DE4AFC00FFFFFF5F08D7FE01A907:8EE43987FC00FFFCFE5E0F4AFD8FAEBB" box="[455,625,1151,1175]" country="Australia" county="Upper Karuah River" latitude="-32.232" longLatPrecision="72" longitude="151.764" municipality="Burrawang Creek" name="Frying Pan Rd" pageId="1" pageNumber="20" stateProvince="New South Wales">Frying Pan Rd</location>
(
<geoCoordinate box="[643,748,1151,1175]" degrees="32.232" direction="south" orientation="latitude" pageId="1" pageNumber="20" precision="55" value="-32.232">32.232°S</geoCoordinate>
<geoCoordinate box="[121,238,1180,1204]" degrees="151.764" direction="east" orientation="longitude" pageId="1" pageNumber="20" precision="55" value="151.764">151.764°E</geoCoordinate>
),
<date box="[256,382,1181,1205]" pageId="1" pageNumber="20" value="2018-11-09">
<collectingDate box="[256,382,1181,1205]" pageId="1" pageNumber="20" value="2018-11-09">9 Nov 2018</collectingDate>
</date>
,
<collectorName box="[392,540,1180,1204]" pageId="1" pageNumber="20">Z. Billingham</collectorName>
&amp;
<collectorName box="[572,745,1180,1205]" pageId="1" pageNumber="20">G. Theischinger</collectorName>
,
<collectionCode box="[121,164,1210,1234]" country="Australia" name="Australian Museum" pageId="1" pageNumber="20" type="Museum">AM</collectionCode>
(K.394953)
</materialsCitation>
.
<materialsCitation accessionNumber="T22391" collectingDate="2018-11-10" collectionCode="T" collectorName="Z. Billingham &amp; G. Theischinger" country="Australia" county="Berrico Rd" latitude="-32.151" location="Avon River" longLatPrecision="72" longitude="151.844" municipality="Tributary of Avon River" pageId="1" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales" typeStatus="paratype">
<emphasis bold="true" box="[311,425,1210,1234]" pageId="1" pageNumber="20">
<typeStatus box="[311,425,1210,1234]" pageId="1" pageNumber="20">Paratypes</typeStatus>
</emphasis>
<specimenCount box="[433,468,1210,1234]" pageId="1" pageNumber="20" type="male">1♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:0392DE4AFC00FFFFFF5F08D7FE01A907:8EE43987FC00FFFCFE790F8FFDA0AEFE" box="[480,606,1210,1234]" country="Australia" county="Berrico Rd" latitude="-32.151" longLatPrecision="72" longitude="151.844" municipality="Tributary of Avon River" name="Avon River" pageId="1" pageNumber="20" stateProvince="New South Wales">Avon River</location>
,
<collectingMunicipality pageId="1" pageNumber="20">Tributary of Avon River</collectingMunicipality>
off
<collectingCounty box="[290,408,1239,1263]" pageId="1" pageNumber="20">Berrico Rd</collectingCounty>
(
<geoCoordinate box="[423,524,1239,1263]" degrees="32.151" direction="south" orientation="latitude" pageId="1" pageNumber="20" precision="55" value="-32.151">32.151°S</geoCoordinate>
<geoCoordinate box="[531,649,1239,1263]" degrees="151.844" direction="east" orientation="longitude" pageId="1" pageNumber="20" precision="55" value="151.844">151.844°E</geoCoordinate>
),
<date pageId="1" pageNumber="20" value="2018-11-10">
<collectingDate pageId="1" pageNumber="20" value="2018-11-10">10 Nov 2018</collectingDate>
</date>
,
<collectorName box="[191,343,1268,1292]" pageId="1" pageNumber="20">Z. Billingham</collectorName>
&amp;
<collectorName box="[383,559,1268,1293]" pageId="1" pageNumber="20">G. Theischinger</collectorName>
, GHD (
<accessionNumber box="[648,736,1269,1293]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/T22391" pageId="1" pageNumber="20">
<collectionCode box="[648,665,1269,1293]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="20">T</collectionCode>
22391
</accessionNumber>
)
</materialsCitation>
;
<materialsCitation collectingDate="2018-11-17" collectionCode="AM" collectorName="Z. Billingham &amp; G. Theischinger" country="Australia" county="Knob Rd" latitude="-32.377" location="Bulahdelah" longLatPrecision="72" longitude="152.153" municipality="Black Camp Creek" pageId="1" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales" typeStatus="paratype">
<specimenCount box="[121,156,1298,1322]" pageId="1" pageNumber="20" type="male">1♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:0392DE4AFC00FFFFFF5F08D7FE01A907:8EE43987FC00FFFCFF3F0E27FEDCAF06" box="[166,290,1298,1322]" country="Australia" county="Knob Rd" latitude="-32.377" longLatPrecision="72" longitude="152.153" municipality="Black Camp Creek" name="Bulahdelah" pageId="1" pageNumber="20" stateProvince="New South Wales">Bulahdelah</location>
,
<collectingMunicipality box="[300,500,1298,1322]" pageId="1" pageNumber="20">Black Camp Creek</collectingMunicipality>
off
<collectingCounty box="[540,635,1298,1322]" pageId="1" pageNumber="20">Knob Rd</collectingCounty>
(
<geoCoordinate box="[648,749,1298,1322]" degrees="32.377" direction="south" orientation="latitude" pageId="1" pageNumber="20" precision="55" value="-32.377">32.377°S</geoCoordinate>
<geoCoordinate box="[121,237,1327,1351]" degrees="152.153" direction="east" orientation="longitude" pageId="1" pageNumber="20" precision="55" value="152.153">152.153°E</geoCoordinate>
),
<date box="[253,390,1327,1351]" pageId="1" pageNumber="20" value="2018-11-17">
<collectingDate box="[253,390,1327,1351]" pageId="1" pageNumber="20" value="2018-11-17">17 Nov 2018</collectingDate>
</date>
,
<collectorName box="[399,546,1327,1351]" pageId="1" pageNumber="20">Z. Billingham</collectorName>
&amp;
<collectorName box="[575,744,1327,1351]" pageId="1" pageNumber="20">G. Theischinger</collectorName>
,
<collectionCode box="[121,164,1357,1381]" country="Australia" name="Australian Museum" pageId="1" pageNumber="20" type="Museum">AM</collectionCode>
</materialsCitation>
(K.394954).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="20" type="description">
<paragraph blockId="1.[793,1422,151,1202]" box="[794,1110,151,176]" pageId="1" pageNumber="20">
<emphasis bold="true" box="[794,926,151,175]" pageId="1" pageNumber="20">Description</emphasis>
<specimenCount box="[932,952,152,175]" pageId="1" pageNumber="20" type="male"></specimenCount>
(
<specimenCount box="[970,988,152,176]" pageId="1" pageNumber="20" type="female"></specimenCount>
unknown).
</paragraph>
<paragraph blockId="1.[793,1422,151,1202]" pageId="1" pageNumber="20">
<emphasis box="[824,882,181,205]" italics="true" pageId="1" pageNumber="20">Head</emphasis>
(
<figureCitation box="[894,955,181,205]" captionStart="Figures 12" captionStartId="1.[121,198,1983,2005]" captionTargetBox="[214,1326,1397,1973]" captionTargetId="figure-582@1.[121,1421,1396,1973]" captionTargetPageId="1" captionText="Figures 12. Molophilus (Molophilus) avia sp. nov., male. (1) head colouration, dorsal; (2) hypopygium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7177924" httpUri="https://zenodo.org/record/7177924/files/figure.png" pageId="1" pageNumber="20">Fig. 1</figureCitation>
): golden yellow, vertex with prominent dark greyish brown teardrop-shaped marking, antennal pedicel and scape pale whitish yellow, antennal flagellum with basal segments pale whitish yellow, terminal segments darkening to greyish brown, palpus dark greyish brown. Antenna of moderate length, extending back to the transverse suture.
</paragraph>
<paragraph blockId="1.[793,1422,151,1202]" pageId="1" pageNumber="20">
<emphasis box="[824,908,357,381]" italics="true" pageId="1" pageNumber="20">Thorax.</emphasis>
Pronotum with lateral edges golden yellow and dorsal angle dark greyish brown, otherwise whitish yellow. Prescutum and scutum golden brown, the posterior margins of scutal lobes pale yellowish brown. Scutellum mostly golden yellow with small darker golden brown basal area. Mediotergite largely golden yellow with darker golden brown central area. Paratergite bright white, in combination with pale portions of pronotum forming a distinct pale stripe between terga and pleura. Cervical sclerite, propleuron and pleurites golden yellow. Forecoxa golden yellow, meso- and metacoxa and trochanters pale whitish yellow. Basal ⅓ of femora pale golden yellow, remainder of femora darkening to greyish brown. Tibiae and tarsi dark greyish brown. Wing pale grey tinged with yellow, veins yellowish brown. Halter with stem and knob whitish yellow.
</paragraph>
<paragraph blockId="1.[793,1422,151,1202]" box="[824,1100,797,821]" pageId="1" pageNumber="20">
<emphasis box="[824,925,797,821]" italics="true" pageId="1" pageNumber="20">Abdomen</emphasis>
: greyish brown.
</paragraph>
<paragraph blockId="1.[793,1422,151,1202]" pageId="1" pageNumber="20">
<taxonomicName box="[824,939,826,850]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis box="[824,939,826,850]" italics="true" pageId="1" pageNumber="20">Terminalia</emphasis>
</taxonomicName>
(
<figureCitation box="[954,1019,826,850]" captionStart="Figures 12" captionStartId="1.[121,198,1983,2005]" captionTargetBox="[214,1326,1397,1973]" captionTargetId="figure-582@1.[121,1421,1396,1973]" captionTargetPageId="1" captionText="Figures 12. Molophilus (Molophilus) avia sp. nov., male. (1) head colouration, dorsal; (2) hypopygium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7177924" httpUri="https://zenodo.org/record/7177924/files/figure.png" pageId="1" pageNumber="20">Fig. 2</figureCitation>
). Hypopygium golden yellow, lobe of tergite 9 rather long, rectangular, caudal margin shallowly concave. Gonocoxite with large, tapering, apically rounded, ventral lobe bearing a fringe of strong setae on the medial edge, dorsal and medial lobes of gonocoxite undeveloped. Outer gonostylus short and club-shaped, caudal margin irregular serrate with a short prominent spine, ventral face with a serrate spine. Inner gonostylus long, heavily sclerotized and terminating in an axe-head-shaped blade, strongly bent at about half its length to project caudad. Aedeagus long and narrow. Parameres fused, poorly developed.
</paragraph>
<paragraph blockId="1.[793,1422,151,1202]" pageId="1" pageNumber="20">
<emphasis box="[824,957,1150,1173]" italics="true" pageId="1" pageNumber="20">Dimensions.</emphasis>
Body length (excluding antenna)
<quantity box="[1331,1417,1149,1173]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="20" unit="mm" value="4.3">4.3 mm</quantity>
, wing length
<quantity box="[927,1012,1178,1202]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="20" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="20" type="etymology">
<paragraph blockId="1.[794,1421,1223,1277]" pageId="1" pageNumber="20">
<emphasis bold="true" box="[794,917,1224,1248]" pageId="1" pageNumber="20">Etymology</emphasis>
. Latin for grandmother, this species is named in memory of Theresa “Tess” Billingham.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="discussion">
<paragraph blockId="1.[794,1421,1298,1381]" lastBlockId="2.[166,794,670,811]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<emphasis bold="true" box="[794,914,1298,1322]" pageId="1" pageNumber="20">Discussion</emphasis>
.
<taxonomicName authorityName="Billingham &amp; Theischinger" authorityYear="2022" box="[927,1155,1298,1322]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="avia" subGenus="Molophilus">
<emphasis box="[927,1155,1298,1322]" italics="true" pageId="1" pageNumber="20">Molophilus (M.) avia</emphasis>
</taxonomicName>
is allied to a large group of morphologically similar species around
<taxonomicName authority="Alexander, 1927" authorityName="Alexander" authorityYear="1927" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="capitatus" subGenus="Molophilus">
<emphasis box="[1239,1421,1328,1351]" italics="true" pageId="1" pageNumber="20">M. (M.) capitatus</emphasis>
<bibRefCitation author="Alexander, C. P." box="[794,969,1357,1381]" pageId="1" pageNumber="20" pagination="42 - 72" refId="ref12190" refString="Alexander, C. P. 1927. The interpretation of the radial field of the wing in the Nematocerous Diptera, with special reference to the Tipulidae. Proceedings of the Linnaean Society of New South Wales 52: 42 - 72." type="journal article" year="1927">Alexander, 1927</bibRefCitation>
</taxonomicName>
and
<taxonomicName authority="Theischinger, 1988" authorityName="Theischinger" authorityYear="1988" box="[1017,1421,1357,1381]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="auriculifer" subGenus="Molophilus">
<emphasis box="[1017,1215,1357,1381]" italics="true" pageId="1" pageNumber="20">M. (M.) auriculifer</emphasis>
Theischinger, 1988
</taxonomicName>
(see
<bibRefCitation author="Theischinger, G." box="[216,421,670,694]" pageId="2" pageNumber="21" pagination="5 - 150" refId="ref12776" refString="Theischinger, G. 1992. The Limoniinae (Diptera: Tipulidae) of Australia. II. The genus Molophilus Curtis. Stapfia 27: 5 - 150." type="journal article" year="1992">Theischinger, 1992</bibRefCitation>
for more details on these species), and most closely resembles
<taxonomicName authority="Theischinger, 1992" authorityName="Theischinger" authorityYear="1992" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="gununo" subGenus="Molophilus">
<emphasis box="[470,641,699,723]" italics="true" pageId="2" pageNumber="21">M. (M.) gununo</emphasis>
<bibRefCitation author="Theischinger, G." pageId="2" pageNumber="21" pagination="5 - 150" refId="ref12776" refString="Theischinger, G. 1992. The Limoniinae (Diptera: Tipulidae) of Australia. II. The genus Molophilus Curtis. Stapfia 27: 5 - 150." type="journal article" year="1992">Theischinger, 1992</bibRefCitation>
</taxonomicName>
.
<taxonomicName authorityName="Billingham &amp; Theischinger" authorityYear="2022" box="[236,469,728,752]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="avia" subGenus="Molophilus">
<emphasis box="[236,469,728,752]" italics="true" pageId="2" pageNumber="21">Molophilus (M.) avia</emphasis>
</taxonomicName>
is readily distinguished from all similar species by the form of the inner gonostylus, with terminal axe-head shaped blade.
</paragraph>
</subSubSection>
</treatment>
</document>