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<document id="97A7C9CBB2C8048EACF0D8A57460FD70" ID-CLB-Dataset="20416" ID-DOI="10.5852/ejt.2022.798.1671" ID-GBIF-Dataset="dc185c1a-2bc2-48f1-bbdd-e0ae42f852f1" ID-ISSN="2118-9773" ID-Zenodo-Dep="6323040" ID-ZooBank="F136E044-62C8-4FB3-8160-7DAE663D9600" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646225490206" checkinUser="felipe" docAuthor="Houben, Albrecht M., Monnens, Marlies, Proesmans, Willem &amp; Artois, Tom J." docDate="2022" docId="038A87DAA76FFF8207BCFE38FBEC0E7F" docLanguage="en" docName="ejt-798_70-102.pdf" docOrigin="European Journal of Taxonomy 798" docSource="http://dx.doi.org/10.5852/ejt.2022.798.1671" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Krumbachia virginiana Ruebush 1938" docType="treatment" docVersion="8" lastPageNumber="93" masterDocId="FFB3FFA2A779FF9A0626FF8AFFA50F38" masterDocTitle="Limnoterrestrial ‘ Typhloplanidae’ (Rhabdocoela, Platyhelminthes), with the description of four new species and a new genus" masterLastPageNumber="102" masterPageNumber="70" pageNumber="92" updateTime="1699316596528" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="90BD30249AA4A79866F5B53375D35BFB">Limnoterrestrial ‘ Typhloplanidae’ (Rhabdocoela, Platyhelminthes), with the description of four new species and a new genus</mods:title>
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<mods:namePart id="59E46E87DF1E6B636DFDF70623DBB814">Houben, Albrecht M.</mods:namePart>
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<mods:affiliation id="67B569D918FECD6C82005973B95A68EC">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
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<mods:namePart id="4CE7325D22E6FB4B4191408D2A5BFE51">Monnens, Marlies</mods:namePart>
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<mods:affiliation id="B1E5691E63F3E238152DCFBE0D20FA3C">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="9690F9DC594E254F5D7AFC09CEB39CA8" type="email">marlies.monnens@uhasselt.be</mods:nameIdentifier>
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<mods:namePart id="635EF9931001F8527D0751D5A3A56177">Proesmans, Willem</mods:namePart>
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<mods:affiliation id="3A7607137E2230F7CCAA5E5ECD34D87D">Agroécologie, INRAE, 17 Rue Sully, F- 21065 Dijon, France.</mods:affiliation>
<mods:nameIdentifier id="AEF9DE78CA72222F082FF2B099665DD8" type="email">willem.proesmans@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2285EFCB235655BBE06FA3E60D0E1A29">Artois, Tom J.</mods:namePart>
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<mods:affiliation id="6691CF92DF5651B9928B945D9C24D9A4">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="DF5EC3EB0C3EBAD2B0AD631EAF249D95" type="email">tom.artois@uhasselt.be</mods:nameIdentifier>
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<mods:title id="B38FDB9704FBCF18C9F4990DDAF27721">European Journal of Taxonomy</mods:title>
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<mods:date id="23BADCDE191DFF61A6983779B57BD90A">2022</mods:date>
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<mods:number id="5F6F1AE633C656B5A4138D09106C8A8B">798</mods:number>
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<treatment id="038A87DAA76FFF8207BCFE38FBEC0E7F" ID-GBIF-Taxon="193858132" LSID="urn:lsid:plazi:treatment:038A87DAA76FFF8207BCFE38FBEC0E7F" httpUri="http://treatment.plazi.org/id/038A87DAA76FFF8207BCFE38FBEC0E7F" lastPageId="24" lastPageNumber="93" pageId="22" pageNumber="92">
<subSubSection id="C3396547A76FFF8C07BCFE38FB3F0EF4" box="[410,1178,434,461]" pageId="22" pageNumber="92" type="nomenclature">
<paragraph id="8B9C36CCA76FFF8C07BCFE38FB3F0EF4" blockId="22.[410,1178,434,496]" box="[410,1178,434,461]" pageId="22" pageNumber="92">
<heading id="D0D481A0A76FFF8C07BCFE38FB3F0EF4" box="[410,1178,434,461]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="92" reason="2">
<taxonomicName id="4C234D4FA76FFF8C07BCFE38FB3F0EF4" ID-CoL="6NPD2" authority="(Kepner &amp; Carter, 1931) Ruebush, 1938" authorityName="Ruebush" authorityYear="1938" baseAuthorityName="Kepner &amp; Carter" baseAuthorityYear="1931" box="[410,1178,434,461]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="22" pageNumber="92" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76FFF8C07BCFE38FD1F0EF4" bold="true" box="[410,698,434,461]" italics="true" pageId="22" pageNumber="92">Krumbachia virginiana</emphasis>
(
<bibRefCitation id="EFB24B3DA76FFF8C04EFFE38FC7F0EF4" author="Kepner W. A. &amp; Carter J. S." box="[713,986,434,460]" pageId="22" pageNumber="92" pagination="87 - 94" refId="ref15508" refString="Kepner W. A. &amp; Carter J. S. 1931. Olisthanella virginiana nov. sp. Zoologischer Anzeiger 95: 87 - 94." type="journal article" year="1931">Kepner &amp; Carter, 1931</bibRefCitation>
)
<bibRefCitation id="EFB24B3DA76FFF8C05CFFE38FB3F0EF4" author="Ruebush T. K." box="[1001,1178,434,460]" pageId="22" pageNumber="92" pagination="260 - 265" refId="ref15981" refString="Ruebush T. K. 1938. Krumbachia minuta n. sp. (Turbellaria Rhabdocoela). Zoologischer Anzeiger 122: 260 - 265." type="journal article" year="1938">Ruebush, 1938</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396547A76FFF8C04FBFE5FFCF20EC8" box="[733,855,469,496]" pageId="22" pageNumber="92" type="description">
<paragraph id="8B9C36CCA76FFF8C04FBFE5FFCF20EC8" blockId="22.[410,1178,434,496]" box="[733,855,469,496]" pageId="22" pageNumber="92">
<figureCitation id="13182A49A76FFF8C04FBFE5FFCF20EC8" box="[733,855,469,496]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D–E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3396547A76FFF8C069BFD9DFB730D4B" pageId="22" pageNumber="92" type="reference_group">
<paragraph id="8B9C36CCA76FFF8C069BFD9DFD4B0D09" blockId="22.[189,750,535,562]" box="[189,750,535,562]" pageId="22" pageNumber="92">
<taxonomicName id="4C234D4FA76FFF8C069BFD9DFD4F0D09" ID-CoL="4987Y" authority="Kepner &amp; Carter, 1931" authorityName="Kepner &amp; Carter" authorityYear="1931" box="[189,746,535,562]" family="Typhloplanidae" genus="Olisthanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="22" pageNumber="92" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76FFF8C069BFD9DFE740D09" box="[189,465,535,561]" italics="true" pageId="22" pageNumber="92">Olisthanella virginiana</emphasis>
<bibRefCitation id="EFB24B3DA76FFF8C07FFFD92FD4F0D09" author="Kepner W. A. &amp; Carter J. S." box="[473,746,535,562]" pageId="22" pageNumber="92" pagination="87 - 94" refId="ref15508" refString="Kepner W. A. &amp; Carter J. S. 1931. Olisthanella virginiana nov. sp. Zoologischer Anzeiger 95: 87 - 94." type="journal article" year="1931">Kepner &amp; Carter, 1931</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C36CCA76FFF8C069BFDD2FB730D4B" blockId="22.[189,1238,600,627]" box="[189,1238,600,627]" pageId="22" pageNumber="92">
<emphasis id="B957EADEA76FFF8C069BFDD2FE4D0D4A" box="[189,488,600,626]" italics="true" pageId="22" pageNumber="92">
<taxonomicName id="4C234D4FA76FFF8C069BFDD2FE740D4A" ID-CoL="4987Y" authority="Kepner &amp; Carter, 1931" authorityName="Kepner &amp; Carter" authorityYear="1931" box="[189,465,600,626]" family="Typhloplanidae" genus="Olisthanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="22" pageNumber="92" phylum="Platyhelminthes" rank="species" species="virginiana">Olisthanella virginiana</taxonomicName>
–
</emphasis>
Senn 1935: 47–50.
<emphasis id="B957EADEA76FFF8C04F2FDD3FD4B0D4A" box="[724,750,601,626]" italics="true" pageId="22" pageNumber="92">—</emphasis>
Vandel 1937: 526.
<emphasis id="B957EADEA76FFF8C05F5FDD3FC480D4A" box="[979,1005,601,626]" italics="true" pageId="22" pageNumber="92">—</emphasis>
Birstein 1991: 108.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A76FFF8C069BFD10FAA60C11" pageId="22" pageNumber="92" type="materials_examined">
<paragraph id="8B9C36CCA76FFF8C069BFD10FE080D8C" blockId="22.[189,429,666,692]" box="[189,429,666,692]" pageId="22" pageNumber="92">
<heading id="D0D481A0A76FFF8C069BFD10FE080D8C" bold="true" box="[189,429,666,692]" fontSize="11" level="3" pageId="22" pageNumber="92" reason="3">
<emphasis id="B957EADEA76FFF8C069BFD10FE080D8C" bold="true" box="[189,429,666,692]" pageId="22" pageNumber="92">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA76FFF8C069BFD42FAA60C11" blockId="22.[189,1399,712,809]" pageId="22" pageNumber="92">
<materialsCitation id="3B4B3C91A76FFF8C069BFD42FB5A0C11" ID-GBIF-Occurrence="3695141302" collectedFrom="submersed roots in a puddle of mud in a forest" collectingDate="2009-05-06" collectionCode="HU" collectorName="Niels Van Steenkiste" country="Canada" latitude="43.238888" location="Dundas Valley Conservation Area" longLatPrecision="19" longitude="-79.98805" pageId="22" pageNumber="92" specimenCount="6" stateProvince="Ontario">
<collectingCountry id="F334765CA76FFF8C069BFD42FE9C0DDA" box="[189,313,712,738]" name="Canada" pageId="22" pageNumber="92">CANADA</collectingCountry>
•
<specimenCount id="9D25FD45A76FFF8C0775FD42FE0E0DDB" box="[339,427,712,739]" pageId="22" pageNumber="92">6 specs</specimenCount>
, studied alive, one of which horizontally sectioned and the rest of which sagittally sectioned;
<collectingRegion id="49E7F82EA76FFF8C071BFD61FE3F0C3D" box="[317,410,747,773]" country="Canada" name="Ontario" pageId="22" pageNumber="92">Ontario</collectingRegion>
,
<location id="8EFC6017A76FFF8C0781FD61FC980C3E" LSID="urn:lsid:plazi:treatment:038A87DAA76FFF8207BCFE38FBEC0E7F:8EFC6017A76FFF8C0781FD61FC980C3E" box="[423,829,747,774]" country="Canada" latitude="43.238888" longLatPrecision="19" longitude="-79.98805" name="Dundas Valley Conservation Area" pageId="22" pageNumber="92" stateProvince="Ontario">Dundas Valley Conservation Area</location>
;
<geoCoordinate id="EE17500BA76FFF8C056FFD60FC450C3E" box="[841,992,746,774]" degrees="43" direction="north" minutes="14" orientation="latitude" pageId="22" pageNumber="92" precision="15" seconds="20" value="43.238888">43°14′20″ N</geoCoordinate>
,
<geoCoordinate id="EE17500BA76FFF8C05CBFD60FB2F0C3E" box="[1005,1162,746,774]" degrees="79" direction="west" minutes="59" orientation="longitude" pageId="22" pageNumber="92" precision="15" seconds="17" value="-79.98805">79°59′17″ W</geoCoordinate>
;
<date id="FF9D100CA76FFF8C02B1FD61FA8E0C3E" box="[1175,1323,747,774]" pageId="22" pageNumber="92" value="2009-05-06">
<collectingDate id="EFD9E9E4A76FFF8C02B1FD61FA8E0C3E" box="[1175,1323,747,774]" pageId="22" pageNumber="92" value="2009-05-06">6 May 2009</collectingDate>
</date>
;
<collectorName id="26D6531AA76FFF8C031FFD61FECE0C11" pageId="22" pageNumber="92">Niels Van Steenkiste</collectorName>
leg.;
<collectedFrom id="383F7531A76FFF8C078BFC85FC770C11" box="[429,978,783,809]" pageId="22" pageNumber="92">submersed roots in a puddle of mud in a forest</collectedFrom>
; XIV.2.43–XIV.2.48;
<collectionCode id="ED32AE09A76FFF8C02F7FC85FB5A0C11" box="[1233,1279,783,809]" pageId="22" pageNumber="92">HU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A76FFF8C069BFCDAFC690892" pageId="22" pageNumber="92" type="description">
<paragraph id="8B9C36CCA76FFF8C069BFCDAFEEA0C52" blockId="22.[189,335,848,874]" box="[189,335,848,874]" pageId="22" pageNumber="92">
<heading id="D0D481A0A76FFF8C069BFCDAFEEA0C52" bold="true" box="[189,335,848,874]" fontSize="11" level="3" pageId="22" pageNumber="92" reason="3">
<emphasis id="B957EADEA76FFF8C069BFCDAFEEA0C52" bold="true" box="[189,335,848,874]" pageId="22" pageNumber="92">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA76FFF8C069BFCF5FED30B1E" blockId="22.[188,1401,895,1062]" pageId="22" pageNumber="92">
Specimens are very dark and around
<quantity id="4CDB9B29A76FFF8C045DFCF5FD640CA1" box="[635,705,895,921]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="92" unit="mm" value="3.0">3 mm</quantity>
long. The body is constricted at ±30% and has a rounded anterior and posterior end (
<figureCitation id="13182A49A76FFF8C0429FC28FDD70C84" box="[527,626,930,956]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D</figureCitation>
). At ±65% the body is at its widest. Rostrally, rhabdite glands (
<figureCitation id="13182A49A76FFF8C06E0FC4FFE860CE7" box="[198,291,965,991]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D</figureCitation>
: ar) are present and arranged in two groups. Dermal rhabdites are lacking. Protonephridiopores (
<figureCitation id="13182A49A76FFF8C06E3FC62FE860B3A" box="[197,291,1000,1026]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D</figureCitation>
: pp) are positioned just behind the pharynx (
<figureCitation id="13182A49A76FFF8C050EFC62FC200B3A" box="[808,901,1000,1026]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D</figureCitation>
: ph), which is situated just before the body middle (±45%).
</paragraph>
<paragraph id="8B9C36CCA76FFF8C069BFBC7FAEC0BE9" blockId="22.[189,1399,1101,1234]" pageId="22" pageNumber="92">
The gonopore (
<figureCitation id="13182A49A76FFF8C0757FBC7FE680B5F" box="[369,461,1101,1127]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: gp) is situated at ±50% of the body and connected to a genital atrium (
<figureCitation id="13182A49A76FFF8C0331FBC7FAD70B5F" box="[1303,1394,1101,1127]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: ga). The genital atrium is surrounded by an inner circular and outer longitudinal muscle layer and lined with a high, nucleated epithelium that is somewhat ruffled. Moreover, some fine-grained eosinophilic glands (
<figureCitation id="13182A49A76FFF8C073CFB3DFED00BE9" box="[282,373,1207,1233]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: gl1) are situated posterior to the atrium and open anteriorly into the genital atrium.
</paragraph>
<paragraph id="8B9C36CCA76FFF8C069BFB72FC4709E4" blockId="22.[186,1401,1272,1756]" pageId="22" pageNumber="92">
The large, elongated, paired testes (
<figureCitation id="13182A49A76FFF8C0476FB72FD090A2A" box="[592,684,1272,1298]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D</figureCitation>
: t) lie anterior to the pharynx and ventral to the paired vitellaria (
<figureCitation id="13182A49A76FFF8C06E5FA91FE800A0D" box="[195,293,1307,1333]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D</figureCitation>
: vi). The paired vasa deferentia (
<figureCitation id="13182A49A76FFF8C048DFA91FCA20A0D" box="[683,775,1307,1333]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: vde) fuse just before entering the copulatory bulbus (
<figureCitation id="13182A49A76FFF8C06E1FAB4FE810A61" box="[199,292,1342,1369]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D</figureCitation>
: co). Two layers of spiral muscles and a third, weaker, outer longitudinal muscle layer surround the 240–320 µm long, slightly bent copulatory organ (see
<figureCitation id="13182A49A76FFF8C055EFAE8FC710A44" box="[888,980,1378,1404]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
). This organ bears an intracapsular seminal vesicle (
<figureCitation id="13182A49A76FFF8C07A4FA0FFE7B0AA7" box="[386,478,1413,1439]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: vs) and a 215–250 µm long, slightly bent, ejaculatory duct (
<figureCitation id="13182A49A76FFF8C028BFA0FFAAC0AA7" box="[1197,1289,1413,1439]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: de) with a thick, sclerotised layer (5–6 µm). The entire ejaculatory duct is surrounded by a loose matrix (
<figureCitation id="13182A49A76FFF8C0331FA22FAD70AFA" box="[1303,1394,1448,1474]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: mx) and at the proximal end, it shows a constriction. Large, coarse-grained, extracapsular basophilic glands (
<figureCitation id="13182A49A76FFF8C073BFA64FE390931" box="[285,412,1518,1545]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D–E</figureCitation>
: gg) are associated with the copulatory organ. A 200–260µm long bursa (
<figureCitation id="13182A49A76FFF8C032AFA64FF760914" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D– E</figureCitation>
: bu) is situated next to the copulatory organ. This bursa is a muscular, elongated sack with a clear constriction at 20% of its length from the proximal tip (see
<figureCitation id="13182A49A76FFF8C054FF9BFFC650977" box="[873,960,1589,1615]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
), dividing it into a proximal and distal part. Both parts are lined with a very thin epithelium, which is lacking at some places. The proximal part is surrounded by two weak layers of diagonal muscles. In the distal part, these muscles divide into a strong outer circular and weaker inner longitudinal layer. This entire structure is surrounded by a matrix (
<figureCitation id="13182A49A76FFF8C06E3F948FE8409E4" box="[197,289,1730,1756]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: mx) and an additional strong layer of longitudinal muscles.
</paragraph>
<paragraph id="8B9C36CCA76FFF8C069BF889FC690892" blockId="22.[188,1400,1795,1962]" pageId="22" pageNumber="92">
The vitellaria (
<figureCitation id="13182A49A76FFF8C074EF889FE600825" box="[360,453,1795,1821]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D</figureCitation>
: vi) extend from the region of the rhabdite glands to the almost posterior end. The vitelloducts (
<figureCitation id="13182A49A76FFF8C0770F8ACFE170878" box="[342,434,1830,1856]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: vd) fuse just before opening into the female duct (
<figureCitation id="13182A49A76FFF8C0235F8ACFBD50878" box="[1043,1136,1830,1856]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: fd). At this place, the female duct also receives the oviduct (
<figureCitation id="13182A49A76FFF8C04A5F8C3FD7A085C" box="[643,735,1865,1892]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: od), a large globular seminal receptacle (
<figureCitation id="13182A49A76FFF8C02EBF8C3FAE8085C" box="[1229,1357,1865,1892]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8D–E</figureCitation>
: rs) and large, coarse-grained basophilic glands (
<figureCitation id="13182A49A76FFF8C04E3F8E7FC8508BF" box="[709,800,1901,1927]" captionStart="Fig" captionStartId="21.[189,232,1854,1880]" captionTargetBox="[270,1303,295,1804]" captionTargetId="figure-13@21.[249,1339,265,1814]" captionTargetPageId="21" captionText="Fig. 8. Two species of Krumbachia Reisinger, 1924. A–C. Krumbachia subterranea Reisinger, 1933. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 823). D–E. Krumbachia virginiana (Kepner &amp; Carter, 1931) Ruebush, 1938. D. Internal organisation of a live specimen. E. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323060" httpUri="https://zenodo.org/record/6323060/files/figure.png" pageId="22" pageNumber="92">Fig. 8E</figureCitation>
: gl2), which are situated around the receptacle. The oviduct and vitelloducts are surrounded by weak, circular muscles.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A76EFF8D069BFE82FE8A0EDC" pageId="23" pageNumber="93" type="discussion">
<paragraph id="8B9C36CCA76EFF8D069BFE82FEE70E1A" blockId="23.[189,322,264,290]" box="[189,322,264,290]" pageId="23" pageNumber="93">
<heading id="D0D481A0A76EFF8D069BFE82FEE70E1A" bold="true" box="[189,322,264,290]" fontSize="11" level="3" pageId="23" pageNumber="93" reason="3">
<emphasis id="B957EADEA76EFF8D069BFE82FEE70E1A" bold="true" box="[189,322,264,290]" pageId="23" pageNumber="93">Discussion</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA76EFF8D069BFEBDFC950E6A" blockId="23.[189,816,311,338]" box="[189,816,311,338]" pageId="23" pageNumber="93">
See the general discussion on the genus
<taxonomicName id="4C234D4FA76EFF8D04B3FEBDFC8C0E69" authorityName="Reisinger" authorityYear="1924" box="[661,809,311,337]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA76EFF8D04B3FEBDFC8C0E69" box="[661,809,311,337]" italics="true" pageId="23" pageNumber="93">Krumbachia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C36CCA76EFF8D069BFEF3FE950EAB" blockId="23.[189,304,377,403]" box="[189,304,377,403]" pageId="23" pageNumber="93">
<heading id="D0D481A0A76EFF8D069BFEF3FE950EAB" bold="true" box="[189,304,377,403]" fontSize="11" level="3" pageId="23" pageNumber="93" reason="3">
<emphasis id="B957EADEA76EFF8D069BFEF3FE950EAB" bold="true" box="[189,304,377,403]" pageId="23" pageNumber="93">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA76EFF8D069BFE2DFE8A0EDC" blockId="23.[189,1398,423,484]" pageId="23" pageNumber="93">
The bursa is filled mainly with some
<typeStatus id="5498886EA76EFF8D04ABFE22FD650EFA" box="[653,704,424,450]" pageId="23" pageNumber="93">type</typeStatus>
of secretion. The seminal receptacle contains sperm and secretion.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A76EFF8D069BFD81FED00D4F" pageId="23" pageNumber="93" type="distribution">
<paragraph id="8B9C36CCA76EFF8D069BFD81FD990D1D" blockId="23.[189,572,523,550]" box="[189,572,523,550]" pageId="23" pageNumber="93">
<heading id="D0D481A0A76EFF8D069BFD81FD990D1D" bold="true" box="[189,572,523,550]" fontSize="11" level="3" pageId="23" pageNumber="93" reason="3">
<emphasis id="B957EADEA76EFF8D069BFD81FD990D1D" bold="true" box="[189,572,523,550]" pageId="23" pageNumber="93">Previously known distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA76EFF8D069BFDB0FED00D4F" blockId="23.[188,1399,570,631]" pageId="23" pageNumber="93">
A swamp, small stream, and pond near the University of
<collectingRegion id="49E7F82EA76EFF8D056CFDB0FC080D6C" box="[842,941,570,596]" country="United States of America" name="Virginia" pageId="23" pageNumber="93">Virginia</collectingRegion>
,
<collectingRegion id="49E7F82EA76EFF8D0590FDB0FBBC0D6C" box="[950,1049,570,596]" country="United States of America" name="Virginia" pageId="23" pageNumber="93">Virginia</collectingRegion>
,
<collectingCountry id="F334765CA76EFF8D0205FDB0FBF90D6C" box="[1059,1116,570,596]" name="United States of America" pageId="23" pageNumber="93">USA</collectingCountry>
(
<bibRefCitation id="EFB24B3DA76EFF8D024CFDB0FAD70D6C" author="Kepner W. A. &amp; Carter J. S." box="[1130,1394,570,596]" pageId="23" pageNumber="93" pagination="87 - 94" refId="ref15508" refString="Kepner W. A. &amp; Carter J. S. 1931. Olisthanella virginiana nov. sp. Zoologischer Anzeiger 95: 87 - 94." type="journal article" year="1931">Kepner &amp; Carter 1931</bibRefCitation>
;
<bibRefCitation id="EFB24B3DA76EFF8D069AFDD7FECC0D4F" author="Ruebush T. K." box="[188,361,605,631]" pageId="23" pageNumber="93" pagination="260 - 265" refId="ref15981" refString="Ruebush T. K. 1938. Krumbachia minuta n. sp. (Turbellaria Rhabdocoela). Zoologischer Anzeiger 122: 260 - 265." type="journal article" year="1938">Ruebush 1938</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396547A76EFF82069BFD15FBEC0E7F" lastPageId="24" lastPageNumber="94" pageId="23" pageNumber="93" type="discussion">
<paragraph id="8B9C36CCA76EFF8D069BFD15FDD60D81" blockId="23.[189,627,670,697]" box="[189,627,670,697]" pageId="23" pageNumber="93">
<emphasis id="B957EADEA76EFF8D069BFD15FDD60D81" bold="true" box="[189,627,670,697]" pageId="23" pageNumber="93">
General discussion on
<taxonomicName id="4C234D4FA76EFF8D07FCFD14FDD60D81" authorityName="Reisinger" authorityYear="1924" box="[474,627,670,697]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA76EFF8D07FCFD14FDD60D81" bold="true" box="[474,627,670,697]" italics="true" pageId="23" pageNumber="93">Krumbachia</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B9C36CCA76EFF8D069BFD47FE190B3A" blockId="23.[188,1400,717,1026]" pageId="23" pageNumber="93">
The genus
<taxonomicName id="4C234D4FA76EFF8D076BFD47FE440DDF" authorityName="Reisinger" authorityYear="1924" box="[333,481,717,743]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA76EFF8D076BFD47FE440DDF" box="[333,481,717,743]" italics="true" pageId="23" pageNumber="93">Krumbachia</emphasis>
</taxonomicName>
is the most species-rich genus within ‘Protoplanellinae’ (
<bibRefCitation id="EFB24B3DA76EFF8D0299FD47FEE10C33" author="Van Steenkiste N. &amp; Davison P. &amp; Artois T." pageId="23" pageNumber="93" pagination="285 - 291" refId="ref16235" refString="Van Steenkiste N., Davison P. &amp; Artois T. 2010. Bryoplana xerophila n. g. n. sp., a new limnoterrestrial microturbellarian (Platyhelminthes, Typhloplanidae, Protoplanellinae) from epilithic mosses, with notes on its ecology. Zoological Science 27 (3): 285 - 291. https: // doi. org / 10.2108 / zsj. 27.285" type="journal article" year="2010">
Van Steenkiste
<emphasis id="B957EADEA76EFF8D069BFD7BFF5F0C32" box="[189,250,752,778]" italics="true" pageId="23" pageNumber="93">et al.</emphasis>
2010
</bibRefCitation>
). Members of this genus can measure from
<quantity id="4CDB9B29A76EFF8D0569FD7BFC0F0C33" box="[847,938,753,779]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="93" unit="mm" value="0.3">0.3 mm</quantity>
in the smallest species (i.e.,
<taxonomicName id="4C234D4FA76EFF8D0324FD7BFECA0C16" authority="Ruebush, 1938" authorityName="Ruebush" authorityYear="1938" family="Typhloplanidae" genus="Prorhynchella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="minuta">
<emphasis id="B957EADEA76EFF8D0324FD7BFADD0C32" box="[1282,1400,753,778]" italics="true" pageId="23" pageNumber="93">K. minuta</emphasis>
<bibRefCitation id="EFB24B3DA76EFF8D069BFC9EFECA0C16" author="Ruebush T. K." box="[189,367,788,814]" pageId="23" pageNumber="93" pagination="260 - 265" refId="ref15981" refString="Ruebush T. K. 1938. Krumbachia minuta n. sp. (Turbellaria Rhabdocoela). Zoologischer Anzeiger 122: 260 - 265." type="journal article" year="1938">Ruebush, 1938</bibRefCitation>
</taxonomicName>
) to over
<quantity id="4CDB9B29A76EFF8D07FAFC9EFD840C16" box="[476,545,788,814]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="93" unit="mm" value="3.0">3 mm</quantity>
in the largest one (i.e.,
<taxonomicName id="4C234D4FA76EFF8D051BFC9FFB380C16" authority="Schwank, 1979" authorityName="Schwank" authorityYear="1979" box="[829,1181,788,814]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="paludicola">
<emphasis id="B957EADEA76EFF8D051BFC9FFC450C16" box="[829,992,788,814]" italics="true" pageId="23" pageNumber="93">K. paludicola</emphasis>
Schwank, 1979
</taxonomicName>
and
<taxonomicName id="4C234D4FA76EFF8D02FFFC9FFD090C69" authority="(Kepner &amp; Carter, 1931) Ruebush, 1938" authorityName="Ruebush" authorityYear="1938" baseAuthorityName="Kepner &amp; Carter" baseAuthorityYear="1931" family="Typhloplanidae" genus="Olisthanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76EFF8D02FFFC9FFAD20C16" box="[1241,1399,789,814]" italics="true" pageId="23" pageNumber="93">K. virginiana</emphasis>
(
<bibRefCitation id="EFB24B3DA76EFF8D06E0FCBDFE440C69" author="Kepner W. A. &amp; Carter J. S." box="[198,481,823,849]" pageId="23" pageNumber="93" pagination="87 - 94" refId="ref15508" refString="Kepner W. A. &amp; Carter J. S. 1931. Olisthanella virginiana nov. sp. Zoologischer Anzeiger 95: 87 - 94." type="journal article" year="1931">Kepner &amp; Carter, 1931</bibRefCitation>
)
<bibRefCitation id="EFB24B3DA76EFF8D07D2FCBDFD090C69" author="Ruebush T. K." box="[500,684,823,849]" pageId="23" pageNumber="93" pagination="260 - 265" refId="ref15981" refString="Ruebush T. K. 1938. Krumbachia minuta n. sp. (Turbellaria Rhabdocoela). Zoologischer Anzeiger 122: 260 - 265." type="journal article" year="1938">Ruebush, 1938</bibRefCitation>
</taxonomicName>
). The diagnostic features are: lack of dermal rhabdites and presence of adenal rhabdites, presence of a sclerotised ejaculatory duct, and a pharynx with a median to posterior position. However, in practice, this genus is a conglomeration of animals roughly matching the diagnostic features (e.g.,
<taxonomicName id="4C234D4FA76EFF8D07CCFC2BFC9D0C83" authority="Schwank, 1980" authorityName="Schwank" authorityYear="1980" box="[490,824,928,955]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="guttulata">
<emphasis id="B957EADEA76EFF8D07CCFC2BFDDD0C82" box="[490,632,928,954]" italics="true" pageId="23" pageNumber="93">K. guttulata</emphasis>
Schwank, 1980
</taxonomicName>
possesses dermal rhabdites) and there are many similarities between the genera
<taxonomicName id="4C234D4FA76EFF8D041DFC4EFD6A0CE6" authorityName="Reisinger" authorityYear="1924" box="[571,719,964,990]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA76EFF8D041DFC4EFD6A0CE6" box="[571,719,964,990]" italics="true" pageId="23" pageNumber="93">Krumbachia</emphasis>
</taxonomicName>
and
<emphasis id="B957EADEA76EFF8D0528FC4EFC040CE6" box="[782,929,964,990]" italics="true" pageId="23" pageNumber="93">Olisthanella</emphasis>
Voigt, 1892 (see
<bibRefCitation id="EFB24B3DA76EFF8D025FFC4EFA800CE6" author="Ruebush T. K." box="[1145,1317,964,990]" pageId="23" pageNumber="93" pagination="260 - 265" refId="ref15981" refString="Ruebush T. K. 1938. Krumbachia minuta n. sp. (Turbellaria Rhabdocoela). Zoologischer Anzeiger 122: 260 - 265." type="journal article" year="1938">Ruebush 1938</bibRefCitation>
for an elaborate discussion).
</paragraph>
<paragraph id="8B9C36CCA76EFF8D069BFBA2FCD60B94" blockId="23.[188,1399,1064,1196]" pageId="23" pageNumber="93">
<taxonomicName id="4C234D4FA76EFF8D069BFBA2FE4C0B7A" authorityName="Reisinger" authorityYear="1933" box="[189,489,1064,1090]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="subterranea">
<emphasis id="B957EADEA76EFF8D069BFBA2FE4C0B7A" box="[189,489,1064,1090]" italics="true" pageId="23" pageNumber="93">Krumbachia subterranea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C234D4FA76EFF8D0400FBA3FD610B7A" authorityName="Ruebush" authorityYear="1938" baseAuthorityName="Kepner &amp; Carter" baseAuthorityYear="1931" box="[550,708,1065,1090]" family="Typhloplanidae" genus="Olisthanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76EFF8D0400FBA3FD610B7A" box="[550,708,1065,1090]" italics="true" pageId="23" pageNumber="93">K. virginiana</emphasis>
</taxonomicName>
possess the diagnostic features of the genus
<taxonomicName id="4C234D4FA76EFF8D02FAFBA2FACA0B7A" authorityName="Reisinger" authorityYear="1924" box="[1244,1391,1064,1090]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA76EFF8D02FAFBA2FACA0B7A" box="[1244,1391,1064,1090]" italics="true" pageId="23" pageNumber="93">Krumbachia</emphasis>
</taxonomicName>
. Moreover, both are quite large species, lack eyes (but see further), have testes positioned rostral to the pharynx, and possess a strongly muscular bursa. There are only two other species that correspond to this description:
<taxonomicName id="4C234D4FA76EFF8D0769FB19FE550B94" authorityName="Schwank" authorityYear="1979" box="[335,496,1170,1196]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="paludicola">
<emphasis id="B957EADEA76EFF8D0769FB19FE550B94" box="[335,496,1170,1196]" italics="true" pageId="23" pageNumber="93">K. paludicola</emphasis>
</taxonomicName>
and
<taxonomicName id="4C234D4FA76EFF8D040FFB19FCCA0B94" authority="Reisinger, 1924" authorityName="Reisinger" authorityYear="1924" box="[553,879,1170,1196]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="styriaca">
<emphasis id="B957EADEA76EFF8D040FFB19FD090B94" box="[553,684,1171,1196]" italics="true" pageId="23" pageNumber="93">K. styriaca</emphasis>
<bibRefCitation id="EFB24B3DA76EFF8D0495FB18FCCA0B94" author="Reisinger E." box="[691,879,1170,1196]" pageId="23" pageNumber="93" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger, 1924</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C36CCA76EFF8D069BFB59FC390ADC" blockId="23.[188,1400,1235,1508]" pageId="23" pageNumber="93">
<taxonomicName id="4C234D4FA76EFF8D069BFB59FE700BD5" authorityName="Ruebush" authorityYear="1938" baseAuthorityName="Kepner &amp; Carter" baseAuthorityYear="1931" box="[189,469,1235,1261]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76EFF8D069BFB59FE700BD5" box="[189,469,1235,1261]" italics="true" pageId="23" pageNumber="93">Krumbachia virginiana</emphasis>
</taxonomicName>
strongly resembles
<taxonomicName id="4C234D4FA76EFF8D04EFFB5EFCC80BD5" authorityName="Schwank" authorityYear="1979" box="[713,877,1235,1261]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="paludicola">
<emphasis id="B957EADEA76EFF8D04EFFB5EFCC80BD5" box="[713,877,1235,1261]" italics="true" pageId="23" pageNumber="93">K. paludicola</emphasis>
</taxonomicName>
because it has a bipartite bursa with strong muscles around its distal part.
<taxonomicName id="4C234D4FA76EFF8D0401FB7DFC980A28" authorityName="Ruebush" authorityYear="1938" baseAuthorityName="Kepner &amp; Carter" baseAuthorityYear="1931" box="[551,829,1271,1297]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76EFF8D0401FB7DFC980A28" box="[551,829,1271,1297]" italics="true" pageId="23" pageNumber="93">Krumbachia virginiana</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C234D4FA76EFF8D05FDFB7DFBD80A29" authorityName="Schwank" authorityYear="1979" box="[987,1149,1271,1297]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="paludicola">
<emphasis id="B957EADEA76EFF8D05FDFB7DFBD80A29" box="[987,1149,1271,1297]" italics="true" pageId="23" pageNumber="93">K. paludicola</emphasis>
</taxonomicName>
in having a joint vas deferens, lacking a blindside branch on the ejaculatory duct, and lacking a large sphincter between the genital atrium and male copulatory system. In the original description,
<bibRefCitation id="EFB24B3DA76EFF8D0277FAB4FAD20A6F" author="Kepner W. A. &amp; Carter J. S." box="[1105,1399,1341,1368]" pageId="23" pageNumber="93" pagination="87 - 94" refId="ref15508" refString="Kepner W. A. &amp; Carter J. S. 1931. Olisthanella virginiana nov. sp. Zoologischer Anzeiger 95: 87 - 94." type="journal article" year="1931">Kepner &amp; Carter (1931)</bibRefCitation>
mention the fact that the dark colouration in their specimens of
<taxonomicName id="4C234D4FA76EFF8D05F2FAEBFBD10A42" authorityName="Ruebush" authorityYear="1938" baseAuthorityName="Kepner &amp; Carter" baseAuthorityYear="1931" box="[980,1140,1377,1402]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76EFF8D05F2FAEBFBD10A42" box="[980,1140,1377,1402]" italics="true" pageId="23" pageNumber="93">K. virginiana</emphasis>
</taxonomicName>
disappears when the animals are starved for a weak, except for two tiny black spots that, according to them, may be eyes. As we did not starve the animals, this could not be confirmed.
<taxonomicName id="4C234D4FA76EFF8D0540FA2DFB250AF9" authorityName="Schwank" authorityYear="1979" box="[870,1152,1447,1473]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="paludicola">
<emphasis id="B957EADEA76EFF8D0540FA2DFB250AF9" box="[870,1152,1447,1473]" italics="true" pageId="23" pageNumber="93">Krumbachia paludicola</emphasis>
</taxonomicName>
has a wide European distribution, whereas
<taxonomicName id="4C234D4FA76EFF8D079CFA41FDF20ADC" authorityName="Ruebush" authorityYear="1938" baseAuthorityName="Kepner &amp; Carter" baseAuthorityYear="1931" box="[442,599,1483,1508]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76EFF8D079CFA41FDF20ADC" box="[442,599,1483,1508]" italics="true" pageId="23" pageNumber="93">K. virginiana</emphasis>
</taxonomicName>
has a Nearctic distribution.
</paragraph>
<paragraph id="8B9C36CCA76EFF8D069BF986FAAB09EE" blockId="23.[188,1399,1547,1750]" pageId="23" pageNumber="93">
In his extensive redescription of
<taxonomicName id="4C234D4FA76EFF8D0468F986FC64091E" authority=", Ruebush (1935)" authorityName=", Ruebush" authorityYear="1935" box="[590,961,1547,1574]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76EFF8D0468F986FD4B091D" box="[590,750,1548,1573]" italics="true" pageId="23" pageNumber="93">K. virginiana</emphasis>
,
<bibRefCitation id="EFB24B3DA76EFF8D0527F981FC64091E" author="Ruebush T. K." box="[769,961,1547,1574]" pageId="23" pageNumber="93" pagination="129 - 136" refId="ref15958" refString="Ruebush T. K. 1935. The genus Olisthanella in the United States. Zoologischer Anzeiger 112: 129 - 136." type="journal article" year="1935">Ruebush (1935)</bibRefCitation>
</taxonomicName>
mentions intraspecific variability in several features, leading to the description of a new variety:
<taxonomicName id="4C234D4FA76EFF8D05E3F9A5FA910970" baseAuthorityName="Ruebush" baseAuthorityYear="1935" box="[965,1332,1582,1609]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="variety" species="virginiana" variety="glandulosa">
<emphasis id="B957EADEA76EFF8D05E3F9A5FBCD0970" box="[965,1128,1583,1608]" italics="true" pageId="23" pageNumber="93">K. virginiana</emphasis>
var.
<emphasis id="B957EADEA76EFF8D0296F9A4FA910970" box="[1200,1332,1582,1608]" italics="true" pageId="23" pageNumber="93">glandulosa</emphasis>
</taxonomicName>
. The identification of this variety is based on the position of the pharynx, subtle differences in the construction of the adenal rhabdite tracks, and the lack of a seminal receptacle in
<taxonomicName id="4C234D4FA76EFF8D05C6F9FCFA9E09B7" baseAuthorityName="Ruebush" baseAuthorityYear="1935" box="[992,1339,1653,1679]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="variety" species="virginiana" variety="glandulosa">
<emphasis id="B957EADEA76EFF8D05C6F9FCFBD909B7" box="[992,1148,1654,1679]" italics="true" pageId="23" pageNumber="93">K. virginiana</emphasis>
var.
<emphasis id="B957EADEA76EFF8D0291F9FFFA9E09B7" box="[1207,1339,1653,1679]" italics="true" pageId="23" pageNumber="93">glandulosa</emphasis>
</taxonomicName>
. The latter character is considered a doubtful one by
<bibRefCitation id="EFB24B3DA76EFF8D04C1F912FC03098B" author="Ruebush T. K." box="[743,934,1688,1715]" pageId="23" pageNumber="93" pagination="129 - 136" refId="ref15958" refString="Ruebush T. K. 1935. The genus Olisthanella in the United States. Zoologischer Anzeiger 112: 129 - 136." type="journal article" year="1935">Ruebush (1935)</bibRefCitation>
, as this may depend on the condition of the animal. The specimens studied by us show all characteristics of the nominal
<taxonomicName id="4C234D4FA76EFF8D024CF936FAA209ED" authorityName=", Ruebush" authorityYear="1935" box="[1130,1287,1724,1749]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="virginiana">
<emphasis id="B957EADEA76EFF8D024CF936FAA209ED" box="[1130,1287,1724,1749]" italics="true" pageId="23" pageNumber="93">K. virginiana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C36CCA76EFF82069BF977FBEC0E7F" blockId="23.[188,1401,1789,1991]" lastBlockId="24.[189,1398,265,327]" lastPageId="24" lastPageNumber="94" pageId="23" pageNumber="93">
<taxonomicName id="4C234D4FA76EFF8D069BF977FE48082F" authorityName="Reisinger" authorityYear="1933" box="[189,493,1789,1815]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="subterranea">
<emphasis id="B957EADEA76EFF8D069BF977FE48082F" box="[189,493,1789,1815]" italics="true" pageId="23" pageNumber="93">Krumbachia subterranea</emphasis>
</taxonomicName>
is morphologically similar to
<emphasis id="B957EADEA76EFF8D054BF977FC58082E" box="[877,1021,1789,1814]" italics="true" pageId="23" pageNumber="93">
<taxonomicName id="4C234D4FA76EFF8D054BF977FC52082E" authorityName="Reisinger" authorityYear="1924" box="[877,1015,1789,1814]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="styriaca">K. styriaca</taxonomicName>
,
</emphasis>
and its description was based mainly on three differences (see
<bibRefCitation id="EFB24B3DA76EFF8D046EF8AAFCA40802" author="Reisinger E." box="[584,769,1824,1850]" pageId="23" pageNumber="93" pagination="155 - 160" refId="ref15862" refString="Reisinger E. 1933. Neues zur vitalen Nervenfarbung. (Gleichzeitig ein Beitrag zur Kenntnis des Protoplanelliden-Nervensystems). Verhandlungen der deutschen Zoologischen Gesellschaft: 155 - 160." type="book chapter" year="1933">Reisinger 1933</bibRefCitation>
). First,
<taxonomicName id="4C234D4FA76EFF8D0546F8ABFC400802" authorityName="Reisinger" authorityYear="1924" box="[864,997,1825,1850]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="styriaca">
<emphasis id="B957EADEA76EFF8D0546F8ABFC400802" box="[864,997,1825,1850]" italics="true" pageId="23" pageNumber="93">K. styriaca</emphasis>
</taxonomicName>
has a thin, sclerotised bursa with many bends and knicks, while it is a flattened, broad sack in
<taxonomicName id="4C234D4FA76EFF8D05A6F8CEFB940865" authorityName="Reisinger" authorityYear="1933" box="[896,1073,1859,1885]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="subterranea">
<emphasis id="B957EADEA76EFF8D05A6F8CEFB940865" box="[896,1073,1859,1885]" italics="true" pageId="23" pageNumber="93">K. subterranea</emphasis>
</taxonomicName>
. Second, the female duct in
<taxonomicName id="4C234D4FA76EFF8D069BF8EDFE9B08B8" authorityName="Reisinger" authorityYear="1924" box="[189,318,1895,1920]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="styriaca">
<emphasis id="B957EADEA76EFF8D069BF8EDFE9B08B8" box="[189,318,1895,1920]" italics="true" pageId="23" pageNumber="93">K. styriaca</emphasis>
</taxonomicName>
possesses a pronounced bend at the point where the female glands occur, while no such bend occurs in
<taxonomicName id="4C234D4FA76EFF8D0714F800FE41089B" authorityName="Reisinger" authorityYear="1933" box="[306,484,1929,1955]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="subterranea">
<emphasis id="B957EADEA76EFF8D0714F800FE41089B" box="[306,484,1929,1955]" italics="true" pageId="23" pageNumber="93">K. subterranea</emphasis>
</taxonomicName>
. Furthermore, the transition between the female duct and bursal stalk is very smooth in the latter species. Third, the vitellaria have small branches in
<taxonomicName id="4C234D4FA76EFF8D0228F827FB6508FF" authorityName="Reisinger" authorityYear="1933" box="[1038,1216,1965,1991]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="93" phylum="Platyhelminthes" rank="species" species="subterranea">
<emphasis id="B957EADEA76EFF8D0228F827FB6508FF" box="[1038,1216,1965,1991]" italics="true" pageId="23" pageNumber="93">K. subterranea</emphasis>
</taxonomicName>
, while they are very smooth in
<taxonomicName id="4C234D4FA761FF82075EFE80FE5E0E1B" authorityName="Reisinger" authorityYear="1924" box="[376,507,266,291]" family="Typhloplanidae" genus="Krumbachia" kingdom="Animalia" order="Rhabdocoela" pageId="24" pageNumber="94" phylum="Platyhelminthes" rank="species" species="styriaca">
<emphasis id="B957EADEA761FF82075EFE80FE5E0E1B" box="[376,507,266,291]" italics="true" pageId="24" pageNumber="94">K. styriaca</emphasis>
</taxonomicName>
. These branches are not that pronounced in our specimens, and also do not seem strongly developed in the specimens studied by
<bibRefCitation id="EFB24B3DA761FF820513FEA6FC540E7E" author="Schwank P." box="[821,1009,300,326]" pageId="24" pageNumber="94" pagination="101 - 131" refId="ref16081" refString="Schwank P. 1981. Seltene und wenig bekannte Strudelwurmer (Turbellaria) aus Quellen und Bergbachen Mitteleuropas. Beitrage zur Naturkunde in Osthessen 17: 101 - 131." type="journal article" year="1981">Schwank (1981</bibRefCitation>
: fig. 5).
</paragraph>
</subSubSection>
</treatment>
</document>
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