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<document id="E2F8FE84340C7FBDD1EABA95C8DEB9B7" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFB5FF8CF0D891C4FB6FB8E9" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus wittmeri Brancucci 1983" docType="treatment" docVersion="9" lastPageNumber="325" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="325" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="148D98D22F2598AE6947A9C7E4BC08A3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="078A2D8CB89DF1E33D8BFD3A46B7E6EF">
<mods:title id="88242DE1C8C7773E44517FCBB472E14A">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="15141517BA7066D9BC20C1E434BC7FFF" type="personal">
<mods:role id="4D7E305592FCB7FED9230065C8CC3959">
<mods:roleTerm id="46AA087A998E4041BA3644AC0D48925B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B50C99499B02CE05254F90B16B8FC89">Hájek, Jiří</mods:namePart>
<mods:affiliation id="BC811CCD6E6E41DFCC639A1CE1584ED3">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 – Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="4A60EAF27AD05376D1E302B3F87EAEF0" type="email">jiri_hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="013C66EF852999D006FF0D5C8E0E5FA7" type="personal">
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<mods:roleTerm id="B1571AF441699C2BF63FA0EEA341355B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9DBB3D5505E723D568C982979E4215CE">Brancucci, Michel</mods:namePart>
<mods:affiliation id="15B189D62421F72317B87F05440D5F31">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5E32489C5100D55394C02F3D2E3658C2">text</mods:typeOfResource>
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<mods:title id="8A169B2E09D31E01494C60FF369AD2F8">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="FB44E0040DA8C9FD106347E82D7DC59E">
<mods:date id="58F40821E1409817B87150C1B422F989">2015</mods:date>
<mods:detail id="ABA7BDF14C7D606299156BE65625ED9D" type="pubDate">
<mods:number id="C4D724A38FD74A8C24A0207BBC0A0068">2015-08-06</mods:number>
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<mods:detail id="A29B5F7FF9944E34B2623A1AE142B6B0" type="volume">
<mods:number id="6706C363327489869788A4F51EAF17B8">63</mods:number>
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<mods:start id="4E84DB25E296B56C3A737BC17D64AF22">309</mods:start>
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<mods:classification id="1C9C44C3E477462BF580749CFD1B6C41">journal article</mods:classification>
<mods:identifier id="24555CD8AC5EF9CF980F60FC46C22501" type="CLB-Dataset">55750</mods:identifier>
<mods:identifier id="9C903F86BED976F8984552623B16D4C5" type="DOI">10.5281/zenodo.5385198</mods:identifier>
<mods:identifier id="8A858AE317A7F5A58B15957A9A568141" type="GBIF-Dataset">c7f95b94-53f8-420d-97c5-4e27d7c211bf</mods:identifier>
<mods:identifier id="E771F8CD35A99777ACD9F0940C980141" type="ISSN">2345-7600</mods:identifier>
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<treatment id="038A87C4FFB5FF8CF0D891C4FB6FB8E9" ID-DOI="http://doi.org/10.5281/zenodo.5466909" ID-GBIF-Taxon="194955221" ID-Zenodo-Dep="5466909" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9" httpUri="http://treatment.plazi.org/id/038A87C4FFB5FF8CF0D891C4FB6FB8E9" lastPageNumber="325" pageId="16" pageNumber="325">
<subSubSection id="C3396559FFB5FF8CF0D891C4FD4BB889" box="[242,670,957,981]" pageId="16" pageNumber="325" type="nomenclature">
<paragraph id="8B9C36D2FFB5FF8CF0D891C4FD4BB889" blockId="16.[242,670,957,1013]" box="[242,670,957,981]" pageId="16" pageNumber="325">
<heading id="D0D481BEFFB5FF8CF0D891C4FD4BB889" bold="true" box="[242,670,957,981]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="325" reason="2">
<taxonomicName id="4C234D51FFB5FF8CF0D891C4FD4BB889" ID-CoL="3RMCZ" authority="Brancucci, 1983" authorityName="Brancucci" authorityYear="1983" box="[242,670,957,981]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="325" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="B957EAC0FFB5FF8CF0D891C4FD4BB889" bold="true" box="[242,670,957,981]" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF0D891C4FE09B889" bold="true" box="[242,476,957,981]" italics="true" pageId="16" pageNumber="325">Laccophilus wittmeri</emphasis>
<bibRefCitation id="EFB24B23FFB5FF8CF1CD91C4FD4BB889" author="Brancucci M" box="[487,670,957,981]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="16" pageNumber="325" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFB5FF8CF15491A4FDC4B8A9" box="[382,529,989,1013]" pageId="16" pageNumber="325" type="description">
<paragraph id="8B9C36D2FFB5FF8CF15491A4FDC4B8A9" blockId="16.[242,670,957,1013]" box="[382,529,989,1013]" pageId="16" pageNumber="325">
(
<figureCitation id="13182A57FFB5FF8CF1AF91A4FE34B8A9" box="[389,481,989,1013]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="16" pageNumber="325">Figs. 17</figureCitation>
,
<figureCitation id="13182A57FFB5FF8CF1C491A4FDDFB8A9" box="[494,522,989,1013]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="16" pageNumber="325">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFB5FF8CF0A29665FDEFBF13" pageId="16" pageNumber="325" type="reference_group">
<paragraph id="8B9C36D2FFB5FF8CF0A29665FDEFBF13" blockId="16.[136,777,1052,1103]" pageId="16" pageNumber="325">
<treatmentCitationGroup id="AB3311FCFFB5FF8CF0A29665FDEFBF13" pageId="16" pageNumber="325">
<treatmentCitation id="0A8210C3FFB5FF8CF0A29665FDFBBF6E" author="Brancucci M" box="[136,558,1052,1074]" page="260" pageId="16" pageNumber="325" year="1983">
<taxonomicName id="4C234D51FFB5FF8CF0A29665FDFBBF6E" ID-CoL="3RMCZ" authority="Brancucci, 1983: 260" authorityName="Brancucci" authorityPageNumber="260" authorityYear="1983" box="[136,558,1052,1074]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="325" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="B957EAC0FFB5FF8CF0A29665FE80BF6E" box="[136,341,1052,1074]" italics="true" pageId="16" pageNumber="325">Laccophilus wittmeri</emphasis>
<bibRefCitation id="EFB24B23FFB5FF8CF1779665FDFBBF6E" author="Brancucci M" box="[349,558,1052,1074]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" page="260" pageId="16" pageNumber="325" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983: 260</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="0A8210C3FFB5FF8CF0849640FE82BF12" author="Hajek J" box="[174,343,1081,1103]" page="120" pageId="16" pageNumber="325" year="2003">
<bibRefCitation id="EFB24B23FFB5FF8CF0849640FE82BF12" author="Hajek J" box="[174,343,1081,1103]" editor="Jach MA &amp; Ji L" firstAuthor="Hajek" journalOrPublisher="Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien." page="120" pageId="16" pageNumber="325" pagination="115 - 123" refId="ref15355" refString="Hajek J (2003) Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera). In: Jach MA &amp; Ji L (eds.) Water Beetles of China, Volume III. Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien. Pp. 115 - 123." title="Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera)" type="book chapter" volumeTitle="Water Beetles of China, Volume III" year="2003">Hájek, 2003: 120</bibRefCitation>
</treatmentCitation>
(in part, new records).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFB5FF8CF0A2960EFA78BAE9" pageId="16" pageNumber="325" type="materials_examined">
<paragraph id="8B9C36D2FFB5FF8CF0A2960EFF23BFF2" blockId="16.[136,774,1143,1199]" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF0A2960EFEFCBFD3" bold="true" box="[136,297,1143,1167]" pageId="16" pageNumber="325">
<typeStatus id="54988870FFB5FF8CF0A2960EFF16BFD3" box="[136,195,1143,1167]" pageId="16" pageNumber="325">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3347642FFB5FF8CF119960EFE51BFD3" box="[307,388,1143,1167]" name="Vietnam" pageId="16" pageNumber="325">Tonkin</collectingCountry>
,
<collectingRegion id="49E7F830FFB5FF8CF1B9960EFE2ABFD3" box="[403,511,1143,1167]" country="Vietnam" name="Hoa Binh" pageId="16" pageNumber="325">Hoa Binh</collectingRegion>
[=
<collectingCountry id="F3347642FFB5FF8CF206960EFD59BFD3" box="[556,652,1143,1167]" name="Vietnam" pageId="16" pageNumber="325">Vietnam</collectingCountry>
,
<collectingRegion id="49E7F830FFB5FF8CF2B1960EFF32BFF3" country="Vietnam" name="Hoa Binh" pageId="16" pageNumber="325">Hoa Binh Province</collectingRegion>
].
</paragraph>
<paragraph id="8B9C36D2FFB5FF8CF0A296AEFA78BAE9" blockId="16.[135,776,1239,2095]" lastBlockId="16.[835,1475,188,438]" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF0A296AEFEB0BFB3" bold="true" box="[136,357,1239,1263]" pageId="16" pageNumber="325">Material examined.</emphasis>
<specimenCount id="9D25FD5BFFB5FF8CF14096AEFE29BFB3" box="[362,508,1239,1263]" pageId="16" pageNumber="325" type="generic">62 specimens</specimenCount>
.
<materialsCitation id="3B4B3C8FFFB5FF8CF22E96AEFD43BFB3" ID-GBIF-Occurrence="3760488385" box="[516,662,1239,1263]" country="Myanmar" location="MYANMAR" pageId="16" pageNumber="325" specimenCount="62">
<emphasis id="B957EAC0FFB5FF8CF22E96AEFD43BFB3" bold="true" box="[516,662,1239,1263]" pageId="16" pageNumber="325">
<collectingCountry id="F3347642FFB5FF8CF22E96AEFD43BFB3" box="[516,662,1239,1263]" name="Myanmar" pageId="16" pageNumber="325">MYANMAR</collectingCountry>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B4B3C8FFFB5FF8CF28996AEFE87BE13" ID-GBIF-Occurrence="3760488323" collectingDate="1998-10-24" collectionCode="NHMW, NMPC" collectorName="H. Schillhammer" country="Myanmar" elevation="720" latitude="20.7063" location="Kalaw - Thazi" longLatPrecision="1" longitude="96.50363" pageId="16" pageNumber="325" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Shan">
<specimenCount id="9D25FD5BFFB5FF8CF28996AEFD21BFB3" box="[675,756,1239,1263]" pageId="16" pageNumber="325" type="male">2 males</specimenCount>
<specimenCount id="9D25FD5BFFB5FF8CF2D396AEFF08BE53" pageId="16" pageNumber="325" type="female">2 females</specimenCount>
,
<collectingRegion id="49E7F830FFB5FF8CF0CC968EFE86BE52" box="[230,339,1271,1295]" country="Myanmar" name="Shan" pageId="16" pageNumber="325">Shan state</collectingRegion>
, road
<location id="8EFC6009FFB5FF8CF1A5968EFDFEBE53" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF1A5968EFDFEBE53" box="[399,555,1271,1295]" country="Myanmar" latitude="20.7063" longLatPrecision="1" longitude="96.50363" name="Kalaw - Thazi" pageId="16" pageNumber="325" stateProvince="Shan">Kalaw – Thazi</location>
, km 23,
<geoCoordinate id="EE175015FFB5FF8CF2AB968CFCD2BE52" box="[641,775,1269,1294]" degrees="20" direction="north" minutes="42.378" orientation="latitude" pageId="16" pageNumber="325" precision="1" value="20.7063">20°42.378′N</geoCoordinate>
<geoCoordinate id="EE175015FFB5FF8CF0A2976CFEDBBE72" box="[136,270,1301,1326]" degrees="96" direction="east" minutes="30.218" orientation="longitude" pageId="16" pageNumber="325" precision="1" value="96.50363">96°30.218′E</geoCoordinate>
,
<quantity id="4CDB9B37FFB5FF8CF13D976EFE8EBE72" box="[279,347,1303,1327]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="16" pageNumber="325" unit="m" value="720.0">
<elevation id="000ED1E1FFB5FF8CF13D976EFE8EBE72" box="[279,347,1303,1327]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="16" pageNumber="325" unit="m" value="720.0">720 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF14F976EFDC3BE73" box="[357,534,1303,1327]" pageId="16" pageNumber="325" value="1998-10-24">
<collectingDate id="EFD9E9FAFFB5FF8CF14F976EFDC3BE73" box="[357,534,1303,1327]" pageId="16" pageNumber="325" value="1998-10-24">24 October 1998</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF279976EFCD2BE73" box="[595,775,1303,1327]" pageId="16" pageNumber="325">H. Schillhammer</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF0A5974EFF3EBE13" box="[143,235,1335,1359]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="325" type="Museum">NHMW</collectionCode>
,
<collectionCode id="ED32AE17FFB5FF8CF0D0974EFE9EBE13" box="[250,331,1335,1359]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="325">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFB5FF8CF14A974EFE5FBED3" ID-GBIF-Occurrence="3760488302" collectingDate="1987-03-05" collectionCode="LHCM" collectorName="P. Schwendinger" country="Thailand" elevation="950" location="Doi Suthep" pageId="16" pageNumber="325" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
<emphasis id="B957EAC0FFB5FF8CF14A974EFE24BE13" bold="true" box="[352,497,1335,1359]" pageId="16" pageNumber="325">
<collectingCountry id="F3347642FFB5FF8CF14A974EFE24BE13" box="[352,497,1335,1359]" name="Thailand" pageId="16" pageNumber="325">THAILAND</collectingCountry>
</emphasis>
:
<emphasis id="B957EAC0FFB5FF8CF229974EFCD5BE13" bold="true" box="[515,768,1335,1359]" pageId="16" pageNumber="325">
<collectingRegion id="49E7F830FFB5FF8CF229974EFCD5BE13" box="[515,768,1335,1359]" country="Thailand" name="Chiang Mai" pageId="16" pageNumber="325">Chiang Mai Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFB5FF8CF0A2972EFF27BE33" box="[136,242,1367,1391]" pageId="16" pageNumber="325" type="female">1 female</specimenCount>
,
<location id="8EFC6009FFB5FF8CF128972EFE5FBE33" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF128972EFE5FBE33" box="[258,394,1367,1391]" country="Thailand" name="Doi Suthep" pageId="16" pageNumber="325" stateProvince="Chiang Mai">Doi Suthep</location>
,
<quantity id="4CDB9B37FFB5FF8CF1B0972EFE3DBE32" box="[410,488,1367,1391]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="16" pageNumber="325" unit="m" value="950.0">
<elevation id="000ED1E1FFB5FF8CF1B0972EFE3DBE32" box="[410,488,1367,1391]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="16" pageNumber="325" unit="m" value="950.0">950 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF1D2972EFD74BE33" box="[504,673,1367,1391]" pageId="16" pageNumber="325" value="1987-03-05">
<collectingDate id="EFD9E9FAFFB5FF8CF1D2972EFD74BE33" box="[504,673,1367,1391]" pageId="16" pageNumber="325" value="1987-03-05">5 March 1987</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF2DA972EFEF7BED3" pageId="16" pageNumber="325">P. Schwendinger</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF105970EFE57BED3" box="[303,386,1399,1423]" pageId="16" pageNumber="325">LHCM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFB5FF8CF1BF970EFE0FBEF3" ID-GBIF-Occurrence="3760488324" collectingDate="1995-11-05" collectionCode="NBCL" collectorName="N. Nieser" country="Thailand" location="Doi Suthep" pageId="16" pageNumber="325" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D25FD5BFFB5FF8CF1BF970EFE26BED3" box="[405,499,1399,1423]" pageId="16" pageNumber="325" type="female">1 female</specimenCount>
,
<location id="8EFC6009FFB5FF8CF1D4970EFDACBED3" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF1D4970EFDACBED3" box="[510,633,1399,1423]" country="Thailand" name="Doi Suthep" pageId="16" pageNumber="325">Doi Suthep</location>
,
<date id="FF9D1012FFB5FF8CF2A9970EFF15BEF3" pageId="16" pageNumber="325" value="1995-11-05">
<collectingDate id="EFD9E9FAFFB5FF8CF2A9970EFF15BEF3" pageId="16" pageNumber="325" value="1995-11-05">5 November 1995</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF12297EEFEA7BEF3" box="[264,370,1431,1455]" pageId="16" pageNumber="325">N. Nieser</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF1AE97EEFE07BEF3" box="[388,466,1431,1455]" pageId="16" pageNumber="325">NBCL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFB5FF8CF1C297EEFD38BE93" ID-GBIF-Occurrence="3760488352" collectingDate="1995-05-21" collectingDateMax="1995-06-04" collectingDateMin="1995-05-21" collectionCode="JSCL" collectorName="M. Snizek" country="Thailand" location="Chiang Dao" pageId="16" pageNumber="325" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D25FD5BFFB5FF8CF1C297EEFDEBBEF3" box="[488,574,1431,1455]" pageId="16" pageNumber="325" type="male">2 males</specimenCount>
<specimenCount id="9D25FD5BFFB5FF8CF26297EEFD7EBEF3" box="[584,683,1431,1455]" pageId="16" pageNumber="325" type="female">1 female</specimenCount>
,
<location id="8EFC6009FFB5FF8CF29397EEFF61BE92" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF29397EEFF61BE92" country="Thailand" name="Chiang Dao" pageId="16" pageNumber="325">Chiang Dao</location>
env.,
<date id="FF9D1012FFB5FF8CF0D397CEFE34BE93" box="[249,481,1463,1487]" pageId="16" pageNumber="325" value="1995-05-21" valueMax="1995-06-04" valueMin="1995-05-21">
<collectingDate id="EFD9E9FAFFB5FF8CF0D397CEFE34BE93" box="[249,481,1463,1487]" pageId="16" pageNumber="325" value="1995-05-21" valueMax="1995-06-04" valueMin="1995-05-21">21 May–4 June 1995</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF20F97CEFD40BE93" box="[549,661,1463,1487]" pageId="16" pageNumber="325">M. Snížek</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF28E97CEFD30BE93" box="[676,741,1463,1487]" pageId="16" pageNumber="325">JSCL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFB5FF8CF2D097CEFE72BD53" ID-GBIF-Occurrence="3760488429" collectingDate="2000-08-02" collectionCode="NMPC" collectorName="L. Kratzerova" country="Thailand" location="Chiang Mai" pageId="16" pageNumber="325" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
<specimenCount id="9D25FD5BFFB5FF8CF2D097CEFF06BEB3" pageId="16" pageNumber="325" type="female">1 female</specimenCount>
,
<quantity id="4CDB9B37FFB5FF8CF0F497AEFEF4BEB3" box="[222,289,1495,1519]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="325" unit="km" value="50.0">50 km</quantity>
N
<collectingRegion id="49E7F830FFB5FF8CF16897AEFE11BEB3" box="[322,452,1495,1519]" country="Thailand" name="Chiang Mai" pageId="16" pageNumber="325">Chiang Mai</collectingRegion>
,
<date id="FF9D1012FFB5FF8CF1E497AEFDBBBEB3" box="[462,622,1495,1519]" pageId="16" pageNumber="325" value="2000-08-02">
<collectingDate id="EFD9E9FAFFB5FF8CF1E497AEFDBBBEB3" box="[462,622,1495,1519]" pageId="16" pageNumber="325" value="2000-08-02">2 August 2000</collectingDate>
</date>
, coll. KROLL &amp;
<collectorName id="26D65304FFB5FF8CF08C978EFEE8BD53" box="[166,317,1527,1551]" pageId="16" pageNumber="325">L. Kratzerová</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF164978EFE75BD53" box="[334,416,1527,1551]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="325">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFB5FF8CF19E978EFD82BD13" ID-GBIF-Occurrence="3760488439" collectingDate="1995-12-25" collectionCode="PMCB" collectorName="P. Mazzoldi" country="Thailand" elevation="1250" location="Huai Khamunoi" municipality="Phu Hin Rongkla National Park" pageId="16" pageNumber="325" specimenCount="8" stateProvince="Loei">
<emphasis id="B957EAC0FFB5FF8CF19E978EFD80BD53" bold="true" box="[436,597,1527,1551]" pageId="16" pageNumber="325">
<collectingRegion id="49E7F830FFB5FF8CF19E978EFD80BD53" box="[436,597,1527,1551]" country="Thailand" name="Loei" pageId="16" pageNumber="325">Loei Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFB5FF8CF24F978EFD4BBD52" box="[613,670,1527,1551]" pageId="16" pageNumber="325" type="generic">8 ex.</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFB5FF8CF286978EFEA2BD73" pageId="16" pageNumber="325">Phu Hin Rongkla National Park</collectingMunicipality>
,
<location id="8EFC6009FFB5FF8CF1AA946EFDF3BD73" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF1AA946EFDF3BD73" box="[384,550,1559,1583]" country="Thailand" municipality="Phu Hin Rongkla National Park" name="Huai Khamunoi" pageId="16" pageNumber="325" stateProvince="Loei">Huai Khamunoi</location>
waterfall,
<quantity id="4CDB9B37FFB5FF8CF2B9946EFD36BD72" box="[659,739,1559,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="16" pageNumber="325" unit="m" value="1250.0">
<elevation id="000ED1E1FFB5FF8CF2B9946EFD36BD72" box="[659,739,1559,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="16" pageNumber="325" unit="m" value="1250.0">1250 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF2C6946EFEE6BD13" pageId="16" pageNumber="325" value="1995-12-25">
<collectingDate id="EFD9E9FAFFB5FF8CF2C6946EFEE6BD13" pageId="16" pageNumber="325" value="1995-12-25">25 December 1995</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF15B944EFE27BD13" box="[369,498,1591,1615]" pageId="16" pageNumber="325">P. Mazzoldi</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF1D5944EFD9ABD13" box="[511,591,1591,1615]" pageId="16" pageNumber="325">PMCB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFB5FF8CF24A944EFEA0BDD3" ID-GBIF-Occurrence="3760488321" collectingDate="1992-05-08" collectingDateMax="1992-05-17" collectingDateMin="1992-05-08" collectionCode="NHMB" collectorName="S. Bily" country="Myanmar" elevation="1600" location="Ban Huei Po" municipality="Ban Huei Po" pageId="16" pageNumber="325" specimenCount="1" specimenCount-female="1" stateProvince="Mae Hong Son Province">
<emphasis id="B957EAC0FFB5FF8CF24A944EFF39BD33" bold="true" pageId="16" pageNumber="325">
<collectingRegion id="49E7F830FFB5FF8CF24A944EFF39BD33" country="Thailand" name="Mae Hong Son" pageId="16" pageNumber="325">Mae Hong Son Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFB5FF8CF0D1942EFE8FBD33" box="[251,346,1623,1647]" pageId="16" pageNumber="325" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFB5FF8CF14E942EFE3ABD32" box="[356,495,1623,1647]" country="Thailand" name="Ban Huei Po" pageId="16" pageNumber="325" stateProvince="Mae Hong Son">Ban Huei Po</collectingMunicipality>
,
<quantity id="4CDB9B37FFB5FF8CF1D0942EFD99BD32" box="[506,588,1623,1647]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="325" unit="m" value="1600.0">
<elevation id="000ED1E1FFB5FF8CF1D0942EFD99BD32" box="[506,588,1623,1647]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="325" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF27D942EFCD6BD33" box="[599,771,1623,1647]" pageId="16" pageNumber="325" value="1992-05-08" valueMax="1992-05-17" valueMin="1992-05-08">
<collectingDate id="EFD9E9FAFFB5FF8CF27D942EFCD6BD33" box="[599,771,1623,1647]" pageId="16" pageNumber="325" value="1992-05-08" valueMax="1992-05-17" valueMin="1992-05-08">8–17 May 1992</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF097940EFED3BDD3" box="[189,262,1655,1679]" pageId="16" pageNumber="325">S. Bílý</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF13E940EFEBFBDD3" box="[276,362,1655,1679]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="325">NHMB</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4B3C8FFFB5FF8CF156940EFDDFBD93" ID-GBIF-Occurrence="3760488329" collectingDate="1999-12-29" collectionCode="NMPC, PMCB" collectorName="P. Mazzoldi" country="Thailand" elevation="1000" location="Huai Boing" municipality="Road Mae Chem" pageId="16" pageNumber="325" specimenCount="17" stateProvince="Mae Hong Son">
<specimenCount id="9D25FD5BFFB5FF8CF156940EFE6BBDD2" box="[380,446,1655,1679]" pageId="16" pageNumber="325" type="generic">17 ex.</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFB5FF8CF1E3940EFDA8BDD3" box="[457,637,1655,1679]" pageId="16" pageNumber="325">Road Mae Chem</collectingMunicipality>
–
<collectingRegion id="49E7F830FFB5FF8CF2BC940EFF61BDF3" country="Thailand" name="Mae Hong Son" pageId="16" pageNumber="325">Mae Hong Son</collectingRegion>
,
<quantity id="4CDB9B37FFB5FF8CF0E894EEFF2FBDF3" box="[194,250,1687,1711]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="325" unit="km" value="4.0">4 km</quantity>
after
<location id="8EFC6009FFB5FF8CF11494EEFE6ABDF3" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF11494EEFE6ABDF3" box="[318,447,1687,1711]" country="Thailand" municipality="Road Mae Chem" name="Huai Boing" pageId="16" pageNumber="325" stateProvince="Mae Hong Son">Huai Boing</location>
,
<quantity id="4CDB9B37FFB5FF8CF1E694EEFDF7BDF2" box="[460,546,1687,1711]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="325" unit="m" value="1000.0">
<elevation id="000ED1E1FFB5FF8CF1E694EEFDF7BDF2" box="[460,546,1687,1711]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="325" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF21A94EEFCD6BDF3" box="[560,771,1687,1711]" pageId="16" pageNumber="325" value="1999-12-29">
<collectingDate id="EFD9E9FAFFB5FF8CF21A94EEFCD6BDF3" box="[560,771,1687,1711]" pageId="16" pageNumber="325" value="1999-12-29">29 December 1999</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF0EB94CEFE90BD93" box="[193,325,1719,1743]" pageId="16" pageNumber="325">P. Mazzoldi</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF17C94CEFE73BD93" box="[342,422,1719,1743]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="325">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFB5FF8CF19E94CEFDD6BD93" box="[436,515,1719,1743]" pageId="16" pageNumber="325">PMCB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFB5FF8CF23D94CEFCD2BC73" ID-GBIF-Occurrence="3760488371" collectedFrom="forest stream" collectingDate="2010-07-08" collectingDateMax="2010-07-11" collectingDateMin="2010-07-08" collectionCode="NHMB" collectorName="M. Brancucci &amp; M. Geiser" country="Laos" elevation="300" latitude="18.386168" location="Tad Paloy campsite" longLatPrecision="13" longitude="104.160835" municipality="Nam Kading" pageId="16" pageNumber="325" specimenCount="25" specimenCount-female="9" specimenCount-male="16" stateProvince="Bolikhamsay Province">
<emphasis id="B957EAC0FFB5FF8CF23D94CEFF38BDB3" bold="true" pageId="16" pageNumber="325">
<collectingCountry id="F3347642FFB5FF8CF23D94CEFDB5BD93" box="[535,608,1719,1743]" name="Laos" pageId="16" pageNumber="325">LAOS</collectingCountry>
:
<collectingRegion id="49E7F830FFB5FF8CF25994CEFF38BDB3" inferredBySuffix="true" pageId="16" pageNumber="325">Bolikhamsay Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFB5FF8CF0D494AEFEB1BDB3" box="[254,356,1751,1775]" pageId="16" pageNumber="325" type="male">16 males</specimenCount>
,
<specimenCount id="9D25FD5BFFB5FF8CF15894AEFE0ABDB3" box="[370,479,1751,1775]" pageId="16" pageNumber="325" type="female">9 females</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFB5FF8CF1C694AEFDACBDB3" box="[492,633,1751,1775]" pageId="16" pageNumber="325">Nam Kading</collectingMunicipality>
NBCA,
<location id="8EFC6009FFB5FF8CF2F794AEFEFEBC53" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF2F794AEFEFEBC53" country="Laos" latitude="18.386168" longLatPrecision="13" longitude="104.160835" municipality="Nam Kading" name="Tad Paloy campsite" pageId="16" pageNumber="325" stateProvince="Bolikhamsay Province">Tad Paloy campsite</location>
,
<collectedFrom id="383F752FFFB5FF8CF11E948EFE15BC52" box="[308,448,1783,1807]" pageId="16" pageNumber="325">forest stream</collectedFrom>
,
<geoCoordinate id="EE175015FFB5FF8CF1E2948CFD90BC52" box="[456,581,1781,1806]" degrees="18" direction="north" minutes="23.17" orientation="latitude" pageId="16" pageNumber="325" precision="9" value="18.386168">18°23.17′N</geoCoordinate>
,
<geoCoordinate id="EE175015FFB5FF8CF264948CFD00BC52" box="[590,725,1781,1806]" degrees="104" direction="east" minutes="09.65" orientation="longitude" pageId="16" pageNumber="325" precision="9" value="104.160835">104°09.65′E</geoCoordinate>
,
<quantity id="4CDB9B37FFB5FF8CF2F4948EFF4ABC72" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="325" unit="m" value="300.0">
<elevation id="000ED1E1FFB5FF8CF2F4948EFF4ABC72" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="325" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF082956EFE9FBC73" box="[168,330,1815,1839]" pageId="16" pageNumber="325" value="2010-07-08" valueMax="2010-07-11" valueMin="2010-07-08">
<collectingDate id="EFD9E9FAFFB5FF8CF082956EFE9FBC73" box="[168,330,1815,1839]" pageId="16" pageNumber="325" value="2010-07-08" valueMax="2010-07-11" valueMin="2010-07-08">8–11 July 2010</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF1AC956EFDC1BC73" box="[390,532,1815,1839]" pageId="16" pageNumber="325">M. Brancucci</collectorName>
&amp;
<collectorName id="26D65304FFB5FF8CF218956EFD4FBC73" box="[562,666,1815,1839]" pageId="16" pageNumber="325">M. Geiser</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF28C956EFD2EBC73" box="[678,763,1815,1839]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="325">NHMB</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4B3C8FFFB5FF8CF0A2954EFDB1BCD3" ID-GBIF-Occurrence="3760488396" collectingDate="2011-05-24" collectingDateMax="2011-05-28" collectingDateMin="2011-05-24" collectionCode="NHMB, NMPC" collectorName="M. Geiser &amp; D. Hauck" elevation="325" latitude="18.350000" location="Tad Paloy campsite" longitude="104.150000" municipality="Nam Kading" pageId="16" pageNumber="325" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="9D25FD5BFFB5FF8CF0A2954EFF09BC13" box="[136,220,1847,1871]" pageId="16" pageNumber="325" type="male">3 males</specimenCount>
<specimenCount id="9D25FD5BFFB5FF8CF0CE954EFE90BC13" box="[228,325,1847,1871]" pageId="16" pageNumber="325" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFB5FF8CF178954EFE08BC13" box="[338,477,1847,1871]" pageId="16" pageNumber="325">Nam Kading</collectingMunicipality>
NPA,
<location id="8EFC6009FFB5FF8CF200954EFCD6BC13" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF200954EFCD6BC13" box="[554,771,1847,1871]" longitude="104.15" municipality="Nam Kading" name="Tad Paloy campsite" pageId="16" pageNumber="325">Tad Paloy campsite</location>
, 18°21–23′N, 104°09′E,
<quantity id="4CDB9B37FFB5FF8CF1A3952EFDD0BC32" box="[393,517,1879,1903]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="16" pageNumber="325" unit="m" value="325.0" valueMax="400.0" valueMin="250.0">
<elevation id="000ED1E1FFB5FF8CF1A3952EFDD0BC32" box="[393,517,1879,1903]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="16" pageNumber="325" unit="m" value="325.0" valueMax="400.0" valueMin="250.0">250–400 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF23B952EFD19BC33" box="[529,716,1879,1903]" pageId="16" pageNumber="325" value="2011-05-24" valueMax="2011-05-28" valueMin="2011-05-24">
<collectingDate id="EFD9E9FAFFB5FF8CF23B952EFD19BC33" box="[529,716,1879,1903]" pageId="16" pageNumber="325" value="2011-05-24" valueMax="2011-05-28" valueMin="2011-05-24">24–28 May 2011</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF0A2950EFF2DBCD3" box="[136,248,1911,1935]" pageId="16" pageNumber="325">M. Geiser</collectorName>
&amp;
<collectorName id="26D65304FFB5FF8CF109950EFE5ABCD3" box="[291,399,1911,1935]" pageId="16" pageNumber="325">D. Hauck</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF188950EFE2DBCD3" box="[418,504,1911,1935]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="325">NHMB</collectionCode>
,
<collectionCode id="ED32AE17FFB5FF8CF22C950EFD8DBCD3" box="[518,600,1911,1935]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="325">NMPC</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4B3C8FFFB5FF8CF244950EFD7EB353" ID-GBIF-Occurrence="3760488311" collectingDate="2012-05-18" collectingDateMax="2012-05-21" collectingDateMin="2012-05-18" collectionCode="NHMB" collectorName="M. Brancucci &amp; M. Geiser &amp; K Phanthavong &amp; S. Xayalath" elevation="675" latitude="17.950000" location="Ban Navang" longitude="105.216667" municipality="Nakai-Nam Theun" pageId="16" pageNumber="325" specimenCount="1" specimenCount-male="1" stateProvince="Khammouan Province">
<emphasis id="B957EAC0FFB5FF8CF244950EFF3BBCF3" bold="true" pageId="16" pageNumber="325">
<collectingRegion id="49E7F830FFB5FF8CF244950EFF3BBCF3" country="Laos" name="Khammouan" pageId="16" pageNumber="325">Khammouan Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFB5FF8CF12A95EEFE9ABCF3" box="[256,335,1943,1967]" pageId="16" pageNumber="325" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFB5FF8CF17495EEFDFEBCF3" box="[350,555,1943,1967]" pageId="16" pageNumber="325">Nakai-Nam Theun</collectingMunicipality>
NPA,
<location id="8EFC6009FFB5FF8CF25795EEFCD2BCF3" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF25795EEFCD2BCF3" box="[637,775,1943,1967]" country="Laos" municipality="Nakai-Nam Theun" name="Ban Navang" pageId="16" pageNumber="325" stateProvince="Khammouan Province">Ban Navang</location>
env., 17°57–59′N, 105°13–16′E,
<quantity id="4CDB9B37FFB5FF8CF1D095CEFDACBC92" box="[506,633,1975,1999]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="16" pageNumber="325" unit="m" value="675.0" valueMax="750.0" valueMin="600.0">
<elevation id="000ED1E1FFB5FF8CF1D095CEFDACBC92" box="[506,633,1975,1999]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="16" pageNumber="325" unit="m" value="675.0" valueMax="750.0" valueMin="600.0">600–750 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF2A295CEFF15BCB3" pageId="16" pageNumber="325" value="2012-05-18" valueMax="2012-05-21" valueMin="2012-05-18">
<collectingDate id="EFD9E9FAFFB5FF8CF2A295CEFF15BCB3" pageId="16" pageNumber="325" value="2012-05-18" valueMax="2012-05-21" valueMin="2012-05-18">18–21 May 2012</collectingDate>
</date>
, NHMB Basel, Laos 2012 Expedition,
<collectorName id="26D65304FFB5FF8CF24795AEFCD6BCB3" box="[621,771,2007,2031]" pageId="16" pageNumber="325">M. Brancucci</collectorName>
,
<collectorName id="26D65304FFB5FF8CF0A2958EFF21B353" box="[136,244,2039,2063]" pageId="16" pageNumber="325">M. Geiser</collectorName>
,
<collectorName id="26D65304FFB5FF8CF0D7958EFE74B353" box="[253,417,2039,2063]" pageId="16" pageNumber="325">K Phanthavong</collectorName>
&amp;
<collectorName id="26D65304FFB5FF8CF1EB958EFDE9B353" box="[449,572,2039,2063]" pageId="16" pageNumber="325">S. Xayalath</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF263958EFD4AB353" box="[585,671,2039,2063]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="325">NHMB</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4B3C8FFFB5FF8CF285958EFB94BA49" ID-GBIF-Occurrence="3760488351" collectingDate="2011-05-14" collectingDateMax="2011-05-20" collectingDateMin="2011-05-14" collectionCode="NHMB" collectorName="D. Hauck &amp; M. Geiser" elevation="1075" latitude="21.150000" location="Ban Oudomsinh-B" longitude="101.216667" pageId="16" pageNumber="325" specimenCount="2" stateProvince="Louang Namtha Province">
<emphasis id="B957EAC0FFB5FF8CF285958EFE81B373" bold="true" pageId="16" pageNumber="325">
<collectingRegion id="49E7F830FFB5FF8CF285958EFE81B373" country="Laos" name="Louangnamtha" pageId="16" pageNumber="325">Louang Namtha Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFB5FF8CF14C9A6EFE75B372" box="[358,416,2071,2095]" pageId="16" pageNumber="325" type="generic">2 ex.</specimenCount>
,
<locationDeviation id="9A2E2FF0FFB5FF8CF1849A6EFD18B373" box="[430,717,2071,2095]" pageId="16" pageNumber="325">
ca.
<quantity id="4CDB9B37FFB5FF8CF1FD9A6EFDCBB373" box="[471,542,2071,2095]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="325" unit="km" value="10.0">10 km</quantity>
E Muang Sing
</locationDeviation>
,
<location id="8EFC6009FFB5FF8CF2F69A6EFC34BB89" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF2F69A6EFC34BB89" box="[732,993,189,2094]" name="Ban Oudomsinh-B" pageId="16" pageNumber="325" stateProvince="Louang Namtha Province">Ban Oudomsinh-B</location>
.
<location id="8EFC6009FFB5FF8CF3C592C7FBA1BB89" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF3C592C7FBA1BB89" box="[1007,1140,190,213]" name="Nam Det-B" pageId="16" pageNumber="325" stateProvince="Louang Namtha Province">Nam Det-B</location>
.
<collectingRegion id="49E7F830FFB5FF8CF4A992C7FB3BBB89" box="[1155,1262,189,213]" inferredBySuffix="true" pageId="16" pageNumber="325">Nam Mai</collectingRegion>
env., 21°09–10′N, 101°13–15′E,
<quantity id="4CDB9B37FFB5FF8CF3F692A4FBB4BBA9" box="[988,1121,221,245]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.4" metricValueMin="0.75" pageId="16" pageNumber="325" unit="m" value="1075.0" valueMax="1400.0" valueMin="750.0">
<elevation id="000ED1E1FFB5FF8CF3F692A4FBB4BBA9" box="[988,1121,221,245]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.4" metricValueMin="0.75" pageId="16" pageNumber="325" unit="m" value="1075.0" valueMax="1400.0" valueMin="750.0">750–1400 m</elevation>
</quantity>
<date id="FF9D1012FFB5FF8CF44D92A4FACBBBA9" box="[1127,1310,221,246]" pageId="16" pageNumber="325" value="2011-05-14" valueMax="2011-05-20" valueMin="2011-05-14">
<collectingDate id="EFD9E9FAFFB5FF8CF44D92A4FACBBBA9" box="[1127,1310,221,246]" pageId="16" pageNumber="325" value="2011-05-14" valueMax="2011-05-20" valueMin="2011-05-14">14–20 May 2011</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF57792A7FA16BBA9" box="[1373,1475,221,245]" pageId="16" pageNumber="325">D. Hauck</collectorName>
&amp;
<collectorName id="26D65304FFB5FF8CF34B9287FC1ABA49" box="[865,975,253,277]" pageId="16" pageNumber="325">M. Geiser</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF3CA9284FBE3BA49" box="[992,1078,253,277]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="325">NHMB</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4B3C8FFFB5FF8CF4609284FBA0BA29" ID-GBIF-Occurrence="3760488423" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="NHMB" collectorName="M. Brancucci" elevation="1400" latitude="19.616667" location="Phou Sane Mt." longitude="103.333333" municipality="Ban Na Lam" pageId="16" pageNumber="325" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Xieng Khouang Prov.">
<collectingRegion id="49E7F830FFB5FF8CF4609284FA93BA49" box="[1098,1350,253,277]" country="Laos" name="Xiangkhouang" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF4609284FAEABA49" bold="true" box="[1098,1343,253,277]" pageId="16" pageNumber="325">Xieng Khouang Prov</emphasis>
.
</collectingRegion>
:
<specimenCount id="9D25FD5BFFB5FF8CF57D9284FA7EBA49" box="[1367,1451,253,277]" pageId="16" pageNumber="325" type="male">4 males</specimenCount>
<specimenCount id="9D25FD5BFFB5FF8CF59F9284FC4CBA69" pageId="16" pageNumber="325" type="female">2 females</specimenCount>
,
<locationDeviation id="9A2E2FF0FFB5FF8CF38C9364FB44BA69" box="[934,1169,285,309]" location="Phonsavan" pageId="16" pageNumber="325">
<quantity id="4CDB9B37FFB5FF8CF38C9364FC36BA69" box="[934,995,285,309]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="325" unit="km" value="30.0">30km</quantity>
NE Phonsavan
</locationDeviation>
:
<collectingMunicipality id="6BF8ACA8FFB5FF8CF4B79367FAF9BA69" box="[1181,1324,286,309]" pageId="16" pageNumber="325">Ban Na Lam</collectingMunicipality>
→
<location id="8EFC6009FFB5FF8CF5649364FCB9BA09" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF5649364FCB9BA09" country="Laos" longitude="103.333336" municipality="Ban Na Lam" name="Phou Sane Mt." pageId="16" pageNumber="325" stateProvince="Xieng Khouang Prov.">Phou Sane Mt.</location>
, 19°37–8′N 103°20′E,
<quantity id="4CDB9B37FFB5FF8CF4499344FB2CBA09" box="[1123,1273,317,341]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="16" pageNumber="325" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">
<elevation id="000ED1E1FFB5FF8CF4499344FB2CBA09" box="[1123,1273,317,341]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="16" pageNumber="325" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300–1500 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF52E9344FA6BBA09" box="[1284,1470,317,342]" pageId="16" pageNumber="325" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<collectingDate id="EFD9E9FAFFB5FF8CF52E9344FA6BBA09" box="[1284,1470,317,342]" pageId="16" pageNumber="325" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">10–30 May 2009</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF3509324FBD1BA29" box="[890,1028,349,373]" pageId="16" pageNumber="325">M.Brancucci</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF43E9324FBBFBA29" box="[1044,1130,349,373]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="325">NHMB</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4B3C8FFFB5FF8CF4579324FB2DBAC9" ID-GBIF-Occurrence="3760488405" collectingDate="1995-05-04" collectionCode="EUMJ" collectorName="M. Sato" country="Vietnam" location="Deo Pha Din" pageId="16" pageNumber="325" specimenCount="1" specimenCount-male="1">
<emphasis id="B957EAC0FFB5FF8CF4579324FAD3BA29" bold="true" box="[1149,1286,349,373]" pageId="16" pageNumber="325">
<collectingCountry id="F3347642FFB5FF8CF4579324FB2BBA29" box="[1149,1278,349,373]" name="Vietnam" pageId="16" pageNumber="325">VIETNAM</collectingCountry>
:
</emphasis>
<specimenCount id="9D25FD5BFFB5FF8CF5249324FA8CBA29" box="[1294,1369,349,373]" pageId="16" pageNumber="325" type="male">1 male</specimenCount>
,
<location id="8EFC6009FFB5FF8CF54C9327FCBBBAC9" LSID="urn:lsid:plazi:treatment:038A87C4FFB5FF8CF0D891C4FB6FB8E9:8EFC6009FFB5FF8CF54C9327FCBBBAC9" country="Vietnam" name="Deo Pha Din" pageId="16" pageNumber="325">Deo Pha Din</location>
,
<date id="FF9D1012FFB5FF8CF3539304FC2ABAC9" box="[889,1023,381,406]" pageId="16" pageNumber="325" value="1995-05-04">
<collectingDate id="EFD9E9FAFFB5FF8CF3539304FC2ABAC9" box="[889,1023,381,406]" pageId="16" pageNumber="325" value="1995-05-04">4 May 1995</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF46A9307FB40BAC9" box="[1088,1173,381,405]" pageId="16" pageNumber="325">M. Sâto</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF48F9304FB24BAC9" box="[1189,1265,381,405]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="16" pageNumber="325">EUMJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFB5FF8CF5299304FA7FBAE9" ID-GBIF-Occurrence="3760488430" collectingDate="1995-05-18" collectionCode="EUMJ, NHMB" collectorName="M. Sato" country="Vietnam" location="Lai Chau" pageId="16" pageNumber="325" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Lai Chau">
<specimenCount id="9D25FD5BFFB5FF8CF5299304FA99BAC9" box="[1283,1356,381,405]" pageId="16" pageNumber="325" type="male">1 male</specimenCount>
<specimenCount id="9D25FD5BFFB5FF8CF57E9304FA6BBAC9" box="[1364,1470,381,405]" pageId="16" pageNumber="325" type="female">3 females</specimenCount>
,
<collectingRegion id="49E7F830FFB5FF8CF36993E4FC7EBAE9" box="[835,939,413,437]" country="Vietnam" name="Lai Chau" pageId="16" pageNumber="325">Lai Chau</collectingRegion>
,
<date id="FF9D1012FFB5FF8CF39D93E4FB99BAE9" box="[951,1100,413,438]" pageId="16" pageNumber="325" value="1995-05-18">
<collectingDate id="EFD9E9FAFFB5FF8CF39D93E4FB99BAE9" box="[951,1100,413,438]" pageId="16" pageNumber="325" value="1995-05-18">18 May 1995</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF4BA93E7FB33BAE9" box="[1168,1254,413,437]" pageId="16" pageNumber="325">M. Satô</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF4DC93E4FA95BAE9" box="[1270,1344,413,437]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="16" pageNumber="325">EUMJ</collectionCode>
,
<collectionCode id="ED32AE17FFB5FF8CF56793E4FA77BAE9" box="[1357,1442,413,437]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="325">NHMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFB5FF8CF36993A4FB05B9E9" pageId="16" pageNumber="325" type="diagnosis">
<paragraph id="8B9C36D2FFB5FF8CF36993A4FB05B9E9" blockId="16.[835,1475,477,693]" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF36993A4FC6DBAA9" bold="true" box="[835,952,477,501]" pageId="16" pageNumber="325">Diagnosis.</emphasis>
Large broadly oval species (body length
<quantity id="4CDB9B37FFB5FF8CF55893A4FCA4B949" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="16" pageNumber="325" unit="mm" value="4.8" valueMax="5.1" valueMin="4.5">4.5–5.1 mm</quantity>
). Reticulation of elytra composed of shallowly impressed very large meshes. Ground colour of elytra dark, brownblackish; mediolateral spot present (
<figureCitation id="13182A57FFB5FF8CF4F09044FAFBB909" box="[1242,1326,573,597]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="16" pageNumber="325">Fig. 17</figureCitation>
). Apical part of median lobe relatively shorter, with dorsal side lightly curved, and short and hooked apex; in lateral view with dorsal side almost straight (
<figureCitation id="13182A57FFB5FF8CF45B90E4FB15B9E9" box="[1137,1216,669,693]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="16" pageNumber="325">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFB5FF8CF36990A4FB6FB8E9" pageId="16" pageNumber="325" type="distribution">
<paragraph id="8B9C36D2FFB5FF8CF36990A4FB6FB8E9" blockId="16.[835,1475,733,949]" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF36990A4FC03B9A9" bold="true" box="[835,982,733,757]" pageId="16" pageNumber="325">Distribution.</emphasis>
<bibRefCitation id="EFB24B23FFB5FF8CF3F490A4FB4CB9A9" author="Brancucci M" box="[990,1177,733,757]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="16" pageNumber="325" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci (1983)</bibRefCitation>
described the species from northern
<collectingCountry id="F3347642FFB5FF8CF3899087FC02B849" box="[931,983,766,789]" name="Laos" pageId="16" pageNumber="325">Laos</collectingCountry>
and
<collectingCountry id="F3347642FFB5FF8CF4229084FBB2B849" box="[1032,1127,765,789]" name="Vietnam" pageId="16" pageNumber="325">Vietnam</collectingCountry>
. Later,
<bibRefCitation id="EFB24B23FFB5FF8CF49E9084FA87B849" author="Nilsson AN" box="[1204,1362,765,789]" editor="Jach MA &amp; Ji L" firstAuthor="Nilsson" journalOrPublisher="Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien." pageId="16" pageNumber="325" pagination="35 - 96" refId="ref15576" refString="Nilsson AN (1995) Noteridae and Dytiscidae: Annotated check list of the Noteridae and Dytiscidae of China (Coleoptera). In: Jach MA &amp; Ji L (eds.) Water Beetles of China, Volume I. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien. Pp. 35 - 96." title="Noteridae and Dytiscidae: Annotated check list of the Noteridae and Dytiscidae of China (Coleoptera)" type="book chapter" volumeTitle="Water Beetles of China, Volume I" year="1995">Nilsson (1995)</bibRefCitation>
and
<bibRefCitation id="EFB24B23FFB5FF8CF5A99084FC5EB869" author="Hajek J" editor="Jach MA &amp; Ji L" firstAuthor="Hajek" journalOrPublisher="Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien." pageId="16" pageNumber="325" pagination="115 - 123" refId="ref15355" refString="Hajek J (2003) Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera). In: Jach MA &amp; Ji L (eds.) Water Beetles of China, Volume III. Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien. Pp. 115 - 123." title="Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera)" type="book chapter" volumeTitle="Water Beetles of China, Volume III" year="2003">Hájek (2003)</bibRefCitation>
published the species from southern
<collectingCountry id="F3347642FFB5FF8CF53A9164FA85B869" box="[1296,1360,797,821]" name="China" pageId="16" pageNumber="325">China</collectingCountry>
(
<collectingRegion id="49E7F830FFB5FF8CF5779164FA62B869" box="[1373,1463,797,821]" country="China" name="Yunnan" pageId="16" pageNumber="325">Yünnan</collectingRegion>
). The present records from
<collectingCountry id="F3347642FFB5FF8CF44A9147FB1DB80A" box="[1120,1224,830,854]" name="Myanmar" pageId="16" pageNumber="325">Myanmar</collectingCountry>
and northern
<collectingCountry id="F3347642FFB5FF8CF5499144FA17B809" box="[1379,1474,829,853]" name="Thailand" pageId="16" pageNumber="325">Thailand</collectingCountry>
extend the distributional area of
<taxonomicName id="4C234D51FFB5FF8CF4E79124FA16B829" authorityName="Brancucci" authorityYear="1983" box="[1229,1475,861,885]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="325" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="B957EAC0FFB5FF8CF4E79124FA16B829" box="[1229,1475,861,885]" italics="true" pageId="16" pageNumber="325">Laccophilus wittmeri</emphasis>
</taxonomicName>
in the border area between the Palaearctic and Oriental zoogeographical regions (
<figureCitation id="13182A57FFB5FF8CF47691E4FB7BB8E9" box="[1116,1198,925,949]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 40–41. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="16" pageNumber="325">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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